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<document id="545FC570780F510C1B5C8128F51BCE3D" ID-CLB-Dataset="6775" ID-DOI="10.11646/zootaxa.4970.3.4" ID-GBIF-Dataset="3d4e2d68-0b43-468d-a921-75a749c3169c" ID-ISSN="1175-5326" ID-Zenodo-Dep="4766798" ID-ZooBank="CEFDA917-6594-4F5E-9956-917B72113CE5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621232145645" checkinUser="plazi" docAuthor="Cadena-Castañeda, Oscar J., Díaz, Carlos Julio Arango, López, Víctor Hugo Grande &amp; Cárdenas, Andrea Del Pilar Floréz" docDate="2021" docId="E47A8798FF90FF9B58E80686FB33D0FE" docLanguage="en" docName="zootaxa.4970.3.4.pdf" docOrigin="Zootaxa 4970 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anurogryllus (Urogryllus) edithsantosum Cadena-Castaneda 2021, n. sp." docType="treatment" docVersion="7" lastPageNumber="518" masterDocId="1843FFE0FF92FF98587F047DFFAFD75C" masterDocTitle="Studies on Neotropical crickets: New species and notes on the classification of Field Crickets genera Anurogryllus and Gryllus (Orthoptera: Gryllidae: Gryllinae)" masterLastPageNumber="532" masterPageNumber="515" pageNumber="517" updateTime="1698939908919" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7A7CCBFC67681D1101D151403A37B29A">Studies on Neotropical crickets: New species and notes on the classification of Field Crickets genera Anurogryllus and Gryllus (Orthoptera: Gryllidae: Gryllinae)</mods:title>
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<mods:namePart id="1AB0E0BDB6C819C5C89B9A06AEF3B229">Cadena-Castañeda, Oscar J.</mods:namePart>
<mods:affiliation id="65EFCFB1265E6A06F5196ADECFEB9A3C">Universidad Distrital Francisco José de Caldas. Grupo de Investigación en Artrópodos “ Kumangui ”, Bogotá, Colombia. Universidad INCCA de Colombia. Grupo en Ecología Evolutiva y Biogeografía Tropical ECOBIT.</mods:affiliation>
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<mods:namePart id="8710C6C2EC8F330F0FC6DE5C9282B7F0">Díaz, Carlos Julio Arango</mods:namePart>
<mods:affiliation id="08E7D6F941A0C197E796F1E282842EDE">Universidad Distrital Francisco José de Caldas. Grupo de Investigación en Artrópodos “ Kumangui ”, Bogotá, Colombia. Universidad INCCA de Colombia. Grupo en Ecología Evolutiva y Biogeografía Tropical ECOBIT.</mods:affiliation>
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<mods:namePart id="6F1B8E00888E19AD2714A63B6DEA0680">López, Víctor Hugo Grande</mods:namePart>
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<mods:affiliation id="3D832E0514B94E82503C7693B67A2D61">vhgrandel @ unincca. edu. co; https: // orcid. org / 0000 - 0003 - 0475 - 6975</mods:affiliation>
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<mods:namePart id="32F8983756F0182F1DDF85F728072AA8">Cárdenas, Andrea Del Pilar Floréz</mods:namePart>
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<mods:affiliation id="C68A4AE2DD8DCCD41184D99B83C8E263">biosilvestres @ yahoo. com; https: // orcid. org / 0000 - 0003 - 1218 - 0187</mods:affiliation>
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<mods:date id="33082D5B17EADFAE33474A6B2D75D89F">2021</mods:date>
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<taxonomicName id="ABD34D0DFF90FF9A58E80686FC2CD449" authority="Cadena-Castaneda" authorityName="Cadena-Castaneda" authorityYear="2021" box="[151,899,763,790]" class="Insecta" family="Haglotettigoniidae" genus="Anurogryllus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="species" species="edithsantosum" status="sp. nov." subGenus="Urogryllus">
<emphasis id="5EA7EA9CFF90FF9A58E80686FE97D44A" bold="true" box="[151,312,763,790]" italics="true" pageId="2" pageNumber="517">Anurogryllus</emphasis>
(
<emphasis id="5EA7EA9CFF90FF9A59350686FE63D44A" bold="true" box="[330,460,763,790]" italics="true" pageId="2" pageNumber="517">Urogryllus</emphasis>
)
<emphasis id="5EA7EA9CFF90FF9A59A20686FD22D44A" bold="true" box="[477,653,763,790]" italics="true" pageId="2" pageNumber="517">edithsantosum</emphasis>
Cadena-Castañeda
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(
<figureCitation id="F4E82A0BFF90FF9A58DF0762FF51D465" box="[160,254,799,825]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1462,1486]" captionStartId-1="4.[151,250,1927,1951]" captionStartId-2="5.[151,250,1943,1967]" captionTargetBox-0="[152,1436,181,1429]" captionTargetBox-1="[151,1436,182,1903]" captionTargetBox-2="[215,1373,181,1920]" captionTargetId-0="figure-21@3.[151,1436,181,1438]" captionTargetId-1="figure-34@4.[151,1436,181,1903]" captionTargetId-2="figure-21@5.[215,1373,181,1920]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Anurogryllus (Urogryllus) edithsantosum n. sp. male habitus A. Lateral and B. Dorsal view respectively." captionText-1="FIGURE 2. Anurogryllus (Urogryllus) edithsantosum n. sp. A. Frons and legs. B. Tegmina. C. Stridulation file." captionText-2="FIGURE 3. Anurogryllus (Urogryllus) edithsantosum n. sp. male genitalia A. Ventral. B. Axial. C. Dorsal and D. Lateral view respectively." figureDoi-0="http://doi.org/10.5281/zenodo.4766802" figureDoi-1="http://doi.org/10.5281/zenodo.4766808" figureDoi-2="http://doi.org/10.5281/zenodo.4766812" httpUri-0="https://zenodo.org/record/4766802/files/figure.png" httpUri-1="https://zenodo.org/record/4766808/files/figure.png" httpUri-2="https://zenodo.org/record/4766812/files/figure.png" pageId="2" pageNumber="517">Fig. 13</figureCitation>
)
</paragraph>
<paragraph id="6C6C368EFF90FF9A58E8073EFBB0D402" blockId="2.[151,1055,763,862]" box="[151,1055,835,862]" pageId="2" pageNumber="517">
http://lsid.speciesfile.org/urn:lsid:
<taxonomicName id="ABD34D0DFF90FF9A5A75073EFD25D402" authorityName="Olivier" authorityYear="1789" box="[522,650,835,862]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
.speciesfile.org:TaxonName:514177
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<subSubSection id="24C96505FF90FF9A58E807F6FAEFD496" pageId="2" pageNumber="517" type="etymology">
<paragraph id="6C6C368EFF90FF9A58E807F6FAEFD496" blockId="2.[151,1437,907,2014]" pageId="2" pageNumber="517">
<emphasis id="5EA7EA9CFF90FF9A58E807F6FEB1D4F9" bold="true" box="[151,286,907,933]" pageId="2" pageNumber="517">Etymology.</emphasis>
Dedicated to the memory of the social leader Edith del Consuelo Santos Jiménez, murdered in 2014, in the Chichimene village, Acacias,
<collectingRegion id="AE17F86CFF90FF9A5A7107CDFDE5D496" box="[526,586,944,970]" country="Colombia" name="Meta" pageId="2" pageNumber="517">Meta</collectingRegion>
, for her defense of the environment and the people of the oil sector.
</paragraph>
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<subSubSection id="24C96505FF90FF9A58B807AEFB64D34D" pageId="2" pageNumber="517" type="materials_examined">
<paragraph id="6C6C368EFF90FF9A58B807AEFB64D34D" blockId="2.[151,1437,907,2014]" pageId="2" pageNumber="517">
<emphasis id="5EA7EA9CFF90FF9A58B807AEFED7D4B1" bold="true" box="[199,376,979,1005]" pageId="2" pageNumber="517">Type material.</emphasis>
<materialsCitation id="DCBB3CD3FF90FF9A59FE07AEFCAFD34D" ID-GBIF-Occurrence="3113047303" collectingDate="2004-04-21" collectionCode="CAUD" collectorName="J. Molano" country="Colombia" county="Acacias" elevation="514" location="Finca La Bonita." municipality="Vrd. La Esmeralda" pageId="2" pageNumber="517" specimenCount="514" stateProvince="Meta" typeStatus="holotype">
<emphasis id="5EA7EA9CFF90FF9A59FE07AEFE5AD4B1" bold="true" box="[385,501,979,1005]" pageId="2" pageNumber="517">
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.
</emphasis>
Male.
<collectingCountry id="14C4761EFF90FF9A5A3607AEFD15D4B1" box="[585,698,979,1005]" name="Colombia" pageId="2" pageNumber="517">Colombia</collectingCountry>
,
<collectingRegion id="AE17F86CFF90FF9A5AB707A9FCABD4B2" box="[712,772,980,1006]" country="Colombia" name="Meta" pageId="2" pageNumber="517">Meta</collectingRegion>
,
<collectingCounty id="850D4E02FF90FF9A5B6E07AEFCC3D4B1" box="[785,876,979,1005]" pageId="2" pageNumber="517">Acacias</collectingCounty>
,
<collectingMunicipality id="8C08ACF4FF90FF9A5B0607AEFBFBD4B1" box="[889,1108,979,1006]" pageId="2" pageNumber="517">Vrd. La Esmeralda</collectingMunicipality>
,
<location id="690C6055FF90FF9A5C1D07AEFA8CD4B1" LSID="urn:lsid:plazi:treatment:E47A8798FF90FF9B58E80686FB33D0FE:690C6055FF90FF9A5C1D07AEFA8CD4B1" box="[1122,1315,979,1006]" country="Colombia" county="Acacias" municipality="Vrd. La Esmeralda" name="Finca La Bonita." pageId="2" pageNumber="517" stateProvince="Meta">Finca La Bonita.</location>
3.972 260, -73.726315.
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<quantity id="AB2B9B6BFF90FF9A5959078AFEDFD34E" box="[294,368,1015,1042]" metricMagnitude="2" metricUnit="m" metricValue="5.14" pageId="2" pageNumber="517" unit="m" value="514.0">
<elevation id="E7FED1BDFF90FF9A5959078AFEDFD34E" box="[294,368,1015,1042]" metricMagnitude="2" metricUnit="m" metricValue="5.14" pageId="2" pageNumber="517" unit="m" value="514.0">514 m</elevation>
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.
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<date id="186D104EFF90FF9A5905078AFDB5D34D" box="[378,538,1015,1041]" pageId="2" pageNumber="517" value="2004-04-21">
<collectingDate id="0829E9A6FF90FF9A5905078AFDB5D34D" box="[378,538,1015,1041]" pageId="2" pageNumber="517" value="2004-04-21">21 April 2004</collectingDate>
</date>
.
<collectorName id="C1265358FF90FF9A5A5B0785FD3BD34D" box="[548,660,1015,1042]" pageId="2" pageNumber="517">J. Molano</collectorName>
(
<collectionCode id="0AC2AE4BFF90FF9A5ADC078AFD56D34D" box="[675,761,1015,1041]" pageId="2" pageNumber="517">CAUD</collectionCode>
)
</materialsCitation>
.
<emphasis id="5EA7EA9CFF90FF9A5B74078AFCD7D34D" bold="true" box="[779,888,1015,1041]" pageId="2" pageNumber="517">
<typeStatus id="B368882CFF90FF9A5B74078AFCD7D34D" box="[779,888,1015,1041]" pageId="2" pageNumber="517" type="paratype">Paratype</typeStatus>
</emphasis>
. Male, same data as holotype.
</paragraph>
</subSubSection>
<subSubSection id="24C96505FF90FF9B58B80066FB33D0FE" lastPageId="3" lastPageNumber="518" pageId="2" pageNumber="517" type="description">
<paragraph id="6C6C368EFF90FF9A58B80066FBB8D0C9" blockId="2.[151,1437,907,2014]" pageId="2" pageNumber="517">
<emphasis id="5EA7EA9CFF90FF9A58B80066FE0CD369" bold="true" box="[199,419,1051,1077]" pageId="2" pageNumber="517">Description. Male.</emphasis>
Large size for the genus (
<figureCitation id="F4E82A0BFF90FF9A5ABB0066FC92D36A" box="[708,829,1051,1078]" captionStart="FIGURE 1" captionStartId="3.[151,250,1462,1486]" captionTargetBox="[152,1436,181,1429]" captionTargetId="figure-21@3.[151,1436,181,1438]" captionTargetPageId="3" captionText="FIGURE 1. Anurogryllus (Urogryllus) edithsantosum n. sp. male habitus A. Lateral and B. Dorsal view respectively." figureDoi="http://doi.org/10.5281/zenodo.4766802" httpUri="https://zenodo.org/record/4766802/files/figure.png" pageId="2" pageNumber="517">Fig. 1A, B</figureCitation>
).
<emphasis id="5EA7EA9CFF90FF9A5B2D0066FC75D369" bold="true" box="[850,986,1051,1077]" pageId="2" pageNumber="517">Coloration.</emphasis>
Head, thorax and abdomen dark brown; legs and ventral surface of body yellowish brown. Head with bands alternating shades of brown; eyes black with depigmented ommatidia in the supra-internal region; ocelli and mouthparts ocher, labial palps and jaws yellow; translucent tegminae with yellow venation.
<emphasis id="5EA7EA9CFF90FF9A5A0500FAFD6ED3FD" bold="true" box="[634,705,1159,1185]" pageId="2" pageNumber="517">Head.</emphasis>
Rounded and as wide as the pronotal disc; frons between antennal cavities 1.5 times as wide as scape; ocelli moderately large and located almost on one transverse line, median ocellus transverse and lateral ones round. Maxillary palp with five palpomeres, first and second shorter, third larger than fourth, fifth-largest, enlarged, apex truncated and grooved (
<figureCitation id="F4E82A0BFF90FF9A5B40008EFC33D251" box="[831,924,1267,1293]" captionStart="FIGURE 2" captionStartId="4.[151,250,1927,1951]" captionTargetBox="[151,1436,182,1903]" captionTargetId="figure-34@4.[151,1436,181,1903]" captionTargetPageId="4" captionText="FIGURE 2. Anurogryllus (Urogryllus) edithsantosum n. sp. A. Frons and legs. B. Tegmina. C. Stridulation file." figureDoi="http://doi.org/10.5281/zenodo.4766808" httpUri="https://zenodo.org/record/4766808/files/figure.png" pageId="2" pageNumber="517">Fig. 2A</figureCitation>
).
<emphasis id="5EA7EA9CFF90FF9A5BCD008EFBBDD251" bold="true" box="[946,1042,1267,1293]" pageId="2" pageNumber="517">Thorax.</emphasis>
Pronotum pubescent, with thicker bristles at the edges; anterior edge of the pronotal disc with concave central portion, straight posterior edge; midline crossing from the anterior to the posterior edge of the pronotal disc, and on each side, a mark in leaf form, no varying in color with the rest of the pronotum.
<emphasis id="5EA7EA9CFF90FF9A5A0B0122FD1CD225" bold="true" box="[628,691,1375,1401]" pageId="2" pageNumber="517">Legs.</emphasis>
Pubescents; fore and mid legs similar; fore tibia with one external and two internal apical spurs, outer tympanum large and ovoid, inner tympanum very small and rounded; hind tibia with seven pairs of dorsal spines; hind basitarsus with two rows of dorsal denticles, seven inner and eight outer, two apical spurs, the internal longer than the external.
<emphasis id="5EA7EA9CFF90FF9A5ABF01B6FCBCD2B9" bold="true" box="[704,787,1483,1509]" pageId="2" pageNumber="517">Wings.</emphasis>
Tegmina ovoid and covering only the first abdominal segment and the base of the second one (
<figureCitation id="F4E82A0BFF90FF9A5A4A0192FD23D155" box="[565,652,1519,1545]" captionStart="FIGURE 2" captionStartId="4.[151,250,1927,1951]" captionTargetBox="[151,1436,182,1903]" captionTargetId="figure-34@4.[151,1436,181,1903]" captionTargetPageId="4" captionText="FIGURE 2. Anurogryllus (Urogryllus) edithsantosum n. sp. A. Frons and legs. B. Tegmina. C. Stridulation file." figureDoi="http://doi.org/10.5281/zenodo.4766808" httpUri="https://zenodo.org/record/4766808/files/figure.png" pageId="2" pageNumber="517">Fig. 2B</figureCitation>
); stridulatory file with 101 teeth (
<figureCitation id="F4E82A0BFF90FF9A5C7F0192FBF7D155" box="[1024,1112,1519,1545]" captionStart="FIGURE 2" captionStartId="4.[151,250,1927,1951]" captionTargetBox="[151,1436,182,1903]" captionTargetId="figure-34@4.[151,1436,181,1903]" captionTargetPageId="4" captionText="FIGURE 2. Anurogryllus (Urogryllus) edithsantosum n. sp. A. Frons and legs. B. Tegmina. C. Stridulation file." figureDoi="http://doi.org/10.5281/zenodo.4766808" httpUri="https://zenodo.org/record/4766808/files/figure.png" pageId="2" pageNumber="517">Fig. 2C</figureCitation>
), basal area reduced, A3 vein very reduced almost covering the anterior edge of the pronotum when the tegmina is at rest, harp with two diagonal veins, the chordal area with veins Cu2 and 1A parallel and arcuate, veins 2A and 3A parallel to the tegmen edge, two small veins connecting the Cu2 vein to the mirror; apical area reduced and reticulate; mirror divided by a vein very close to the inner-lower edge (
<figureCitation id="F4E82A0BFF90FF9A59970202FDEFD1C5" box="[488,576,1663,1689]" captionStart="FIGURE 2" captionStartId="4.[151,250,1927,1951]" captionTargetBox="[151,1436,182,1903]" captionTargetId="figure-34@4.[151,1436,181,1903]" captionTargetPageId="4" captionText="FIGURE 2. Anurogryllus (Urogryllus) edithsantosum n. sp. A. Frons and legs. B. Tegmina. C. Stridulation file." figureDoi="http://doi.org/10.5281/zenodo.4766808" httpUri="https://zenodo.org/record/4766808/files/figure.png" pageId="2" pageNumber="517">Fig. 2B</figureCitation>
). Hind wings reduced.
<emphasis id="5EA7EA9CFF90FF9A5B3D0202FC11D1C5" bold="true" box="[834,958,1663,1689]" pageId="2" pageNumber="517">Abdomen.</emphasis>
Epiproctus with moderately elongated and rounded posterior edge; subgenital plate rectangular, longer than wide and with a rounded apex.
<emphasis id="5EA7EA9CFF90FF9A5CB702DEFA92D1E1" bold="true" box="[1224,1341,1699,1725]" pageId="2" pageNumber="517">Genitalia.</emphasis>
Pseudepiphallic median lophi elongated and lanceolated in ventral and dorsal view (
<figureCitation id="F4E82A0BFF90FF9A5B8A02BAFBC1D1BD" box="[1013,1134,1735,1761]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1920]" captionTargetId="figure-21@5.[215,1373,181,1920]" captionTargetPageId="5" captionText="FIGURE 3. Anurogryllus (Urogryllus) edithsantosum n. sp. male genitalia A. Ventral. B. Axial. C. Dorsal and D. Lateral view respectively." figureDoi="http://doi.org/10.5281/zenodo.4766812" httpUri="https://zenodo.org/record/4766812/files/figure.png" pageId="2" pageNumber="517">Fig. 3A, C</figureCitation>
), moderately flared, with a deep U-shaped notch at the apex, lateral edges with abundant hairs; pseudepiphallic parameres slender and shorter than the median lophi, apex truncated and ondulated (
<figureCitation id="F4E82A0BFF90FF9A5A920372FCC9D076" box="[749,870,1807,1834]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1920]" captionTargetId="figure-21@5.[215,1373,181,1920]" captionTargetPageId="5" captionText="FIGURE 3. Anurogryllus (Urogryllus) edithsantosum n. sp. male genitalia A. Ventral. B. Axial. C. Dorsal and D. Lateral view respectively." figureDoi="http://doi.org/10.5281/zenodo.4766812" httpUri="https://zenodo.org/record/4766812/files/figure.png" pageId="2" pageNumber="517">Fig. 3A, B</figureCitation>
); lateral lophi shorth and hook-shaped; ectophallic fold ribbon-shaped, narrow and sticking out between the pseudepiphallic median lophi and parameres, apex lanceolated (
<figureCitation id="F4E82A0BFF90FF9A58A1032AFE96D02D" box="[222,313,1879,1905]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1920]" captionTargetId="figure-21@5.[215,1373,181,1920]" captionTargetPageId="5" captionText="FIGURE 3. Anurogryllus (Urogryllus) edithsantosum n. sp. male genitalia A. Ventral. B. Axial. C. Dorsal and D. Lateral view respectively." figureDoi="http://doi.org/10.5281/zenodo.4766812" httpUri="https://zenodo.org/record/4766812/files/figure.png" pageId="2" pageNumber="517">Fig. 3A</figureCitation>
); arc narrow, as a plate up-curved dorsally (
<figureCitation id="F4E82A0BFF90FF9A5B54032AFC09D02E" box="[811,934,1879,1906]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1920]" captionTargetId="figure-21@5.[215,1373,181,1920]" captionTargetPageId="5" captionText="FIGURE 3. Anurogryllus (Urogryllus) edithsantosum n. sp. male genitalia A. Ventral. B. Axial. C. Dorsal and D. Lateral view respectively." figureDoi="http://doi.org/10.5281/zenodo.4766812" httpUri="https://zenodo.org/record/4766812/files/figure.png" pageId="2" pageNumber="517">Fig. 3C, D</figureCitation>
); endophallic cavity membranous and ovoid (
<figureCitation id="F4E82A0BFF90FF9A58DF0306FEB5D0CA" box="[160,282,1915,1942]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1920]" captionTargetId="figure-21@5.[215,1373,181,1920]" captionTargetPageId="5" captionText="FIGURE 3. Anurogryllus (Urogryllus) edithsantosum n. sp. male genitalia A. Ventral. B. Axial. C. Dorsal and D. Lateral view respectively." figureDoi="http://doi.org/10.5281/zenodo.4766812" httpUri="https://zenodo.org/record/4766812/files/figure.png" pageId="2" pageNumber="517">Fig. 3C, D</figureCitation>
); quadrangular rami connected in the posterior portion (
<figureCitation id="F4E82A0BFF90FF9A5BF20306FBA4D0C9" box="[909,1035,1915,1941]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1920]" captionTargetId="figure-21@5.[215,1373,181,1920]" captionTargetPageId="5" captionText="FIGURE 3. Anurogryllus (Urogryllus) edithsantosum n. sp. male genitalia A. Ventral. B. Axial. C. Dorsal and D. Lateral view respectively." figureDoi="http://doi.org/10.5281/zenodo.4766812" httpUri="https://zenodo.org/record/4766812/files/figure.png" pageId="2" pageNumber="517">Fig. 3AD</figureCitation>
).
</paragraph>
<paragraph id="6C6C368EFF90FF9A58B803E2FE0DD0E5" blockId="2.[151,1437,907,2014]" box="[199,418,1951,1977]" pageId="2" pageNumber="517">
<emphasis id="5EA7EA9CFF90FF9A58B803E2FE8AD0E5" bold="true" box="[199,293,1951,1977]" pageId="2" pageNumber="517">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="6C6C368EFF90FF9A58B803BEFBA8D081" blockId="2.[151,1437,907,2014]" box="[199,1031,1987,2014]" pageId="2" pageNumber="517">
<emphasis id="5EA7EA9CFF90FF9A58B803BEFE67D081" bold="true" box="[199,456,1987,2013]" pageId="2" pageNumber="517">Measurements (mm):</emphasis>
LB: 1112. Pr: 44.5. Teg.: 66.5. HF: 14. HT: 10.
</paragraph>
<caption id="38AC6606FF91FF9B58E801CBFAE0D292" ID-DOI="http://doi.org/10.5281/zenodo.4766802" ID-Zenodo-Dep="4766802" box="[151,1359,1461,1486]" httpUri="https://zenodo.org/record/4766802/files/figure.png" pageId="3" pageNumber="518" startId="3.[151,250,1462,1486]" targetBox="[152,1436,181,1429]" targetPageId="3">
<paragraph id="6C6C368EFF91FF9B58E801CBFAE0D292" blockId="3.[151,1359,1461,1486]" box="[151,1359,1461,1486]" pageId="3" pageNumber="518">
<emphasis id="5EA7EA9CFF91FF9B58E801CBFCA4D292" bold="true" box="[151,779,1461,1486]" pageId="3" pageNumber="518">
FIGURE 1.
<taxonomicName id="ABD34D0DFF91FF9B596601C8FD64D291" authority="Cadena-Castañeda &amp; Díaz &amp; López &amp; Cárdenas, 2021" authorityName="Cadena-Castañeda &amp; Díaz &amp; López &amp; Cárdenas" authorityYear="2021" box="[281,715,1461,1486]" class="Insecta" family="Haglotettigoniidae" genus="Anurogryllus" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="518" phylum="Arthropoda" rank="species" species="edithsantosum" status="sp. nov." subGenus="Urogryllus">
<emphasis id="5EA7EA9CFF91FF9B596601C8FE0BD291" bold="true" box="[281,420,1461,1485]" italics="true" pageId="3" pageNumber="518">Anurogryllus</emphasis>
(
<emphasis id="5EA7EA9CFF91FF9B59CB01C8FD8AD291" bold="true" box="[436,549,1461,1485]" italics="true" pageId="3" pageNumber="518">Urogryllus</emphasis>
)
<emphasis id="5EA7EA9CFF91FF9B5A4C01C8FD64D291" bold="true" box="[563,715,1461,1485]" italics="true" pageId="3" pageNumber="518">edithsantosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="459457E7FF91FF9B5AAE01CBFCA4D292" box="[721,779,1462,1486]" pageId="3" pageNumber="518" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
male habitus
<emphasis id="5EA7EA9CFF91FF9B5BE701CBFC1FD292" bold="true" box="[920,944,1462,1486]" pageId="3" pageNumber="518">A.</emphasis>
Lateral and
<emphasis id="5EA7EA9CFF91FF9B5C4F01CBFBE8D292" bold="true" box="[1072,1095,1462,1486]" pageId="3" pageNumber="518">B.</emphasis>
Dorsal view respectively.
</paragraph>
</caption>
<paragraph id="6C6C368EFF91FF9B58B80181FB33D0FE" blockId="3.[151,1437,1532,1955]" pageId="3" pageNumber="518">
<emphasis id="5EA7EA9CFF91FF9B58B80181FECDD14A" bold="true" box="[199,354,1532,1558]" pageId="3" pageNumber="518">Comparison.</emphasis>
This new species is more similar to Caribbean species such as
<emphasis id="5EA7EA9CFF91FF9B5C600181FB97D14A" box="[1055,1080,1532,1558]" italics="true" pageId="3" pageNumber="518">A.</emphasis>
(
<emphasis id="5EA7EA9CFF91FF9B5C380180FBF4D14A" box="[1095,1115,1533,1558]" italics="true" pageId="3" pageNumber="518">U</emphasis>
.)
<emphasis id="5EA7EA9CFF91FF9B5C0D0181FB7ED14A" box="[1138,1233,1532,1558]" italics="true" pageId="3" pageNumber="518">amolgos</emphasis>
<bibRefCitation id="08424B7FFF91FF9B5CA70181FE8FD166" author="Otte, D. &amp; Perez-Gelabert, D. E." pageId="3" pageNumber="518" refId="ref8268" refString="Otte, D. &amp; Perez-Gelabert, D. E. (2009) Caribbean crickets. The Orthopterists' Society, Philadelphia, Pennsylvania, iii + 792 pp." type="book" year="2009">Otte &amp; Perez-Gelabert, 2009</bibRefCitation>
,
<emphasis id="5EA7EA9CFF91FF9B5954025DFEEBD166" box="[299,324,1568,1594]" italics="true" pageId="3" pageNumber="518">A.</emphasis>
(
<emphasis id="5EA7EA9CFF91FF9B592B025CFEC7D166" box="[340,360,1569,1594]" italics="true" pageId="3" pageNumber="518">U</emphasis>
.)
<emphasis id="5EA7EA9CFF91FF9B59FF025CFE78D166" box="[384,471,1569,1594]" italics="true" pageId="3" pageNumber="518">gnomus</emphasis>
<bibRefCitation id="08424B7FFF91FF9B59A1025DFC8FD166" author="Otte, D. &amp; Perez-Gelabert, D. E." box="[478,800,1568,1594]" pageId="3" pageNumber="518" refId="ref8268" refString="Otte, D. &amp; Perez-Gelabert, D. E. (2009) Caribbean crickets. The Orthopterists' Society, Philadelphia, Pennsylvania, iii + 792 pp." type="book" year="2009">Otte &amp; Perez-Gelabert, 2009</bibRefCitation>
and
<emphasis id="5EA7EA9CFF91FF9B5B28025DFCC0D166" box="[855,879,1568,1594]" italics="true" pageId="3" pageNumber="518">A.</emphasis>
(
<emphasis id="5EA7EA9CFF91FF9B5B00025CFC3CD166" box="[895,915,1569,1594]" italics="true" pageId="3" pageNumber="518">U</emphasis>
.)
<emphasis id="5EA7EA9CFF91FF9B5BD4025CFC44D166" box="[939,1003,1569,1594]" italics="true" pageId="3" pageNumber="518">nigua</emphasis>
<bibRefCitation id="08424B7FFF91FF9B5B8D025DFA97D166" author="Otte, D. &amp; Perez-Gelabert, D. E." box="[1010,1336,1568,1594]" pageId="3" pageNumber="518" refId="ref8268" refString="Otte, D. &amp; Perez-Gelabert, D. E. (2009) Caribbean crickets. The Orthopterists' Society, Philadelphia, Pennsylvania, iii + 792 pp." type="book" year="2009">Otte &amp; Perez-Gelabert, 2009</bibRefCitation>
, than the continental species of the subgenus
<taxonomicName id="ABD34D0DFF91FF9B5A560239FD0CD102" box="[553,675,1604,1630]" class="Insecta" family="Gryllidae" genus="Urogryllus" higherTaxonomySource="CoL" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="518" phylum="Arthropoda" rank="genus">
<emphasis id="5EA7EA9CFF91FF9B5A560239FD0CD102" box="[553,675,1604,1630]" italics="true" pageId="3" pageNumber="518">Urogryllus</emphasis>
</taxonomicName>
. It is distinguished from
<emphasis id="5EA7EA9CFF91FF9B5BC30239FC7BD102" box="[956,980,1604,1630]" italics="true" pageId="3" pageNumber="518">A.</emphasis>
(
<emphasis id="5EA7EA9CFF91FF9B5B9B0238FC57D102" box="[996,1016,1605,1630]" italics="true" pageId="3" pageNumber="518">U</emphasis>
.)
<emphasis id="5EA7EA9CFF91FF9B5C6F0239FBC0D102" box="[1040,1135,1604,1630]" italics="true" pageId="3" pageNumber="518">amolgos</emphasis>
because the tegmina reach the base of the second abdominal tergite and the apex of the median lophi is lanceolate and without undulations on the lateral edges of the new species, in contrast,
<emphasis id="5EA7EA9CFF91FF9B5AC002F1FD78D1FA" box="[703,727,1676,1702]" italics="true" pageId="3" pageNumber="518">A.</emphasis>
(
<emphasis id="5EA7EA9CFF91FF9B5A9602F0FD52D1FA" box="[745,765,1677,1702]" italics="true" pageId="3" pageNumber="518">U</emphasis>
.)
<emphasis id="5EA7EA9CFF91FF9B5B6A02F1FCDBD1FA" box="[789,884,1676,1702]" italics="true" pageId="3" pageNumber="518">amolgos</emphasis>
the tegmina cover slightly more than half of the abdomen and the apex of the median lophi is narrow and wavy.
<emphasis id="5EA7EA9CFF91FF9B5B1702CDFC2FD196" box="[872,896,1712,1738]" italics="true" pageId="3" pageNumber="518">A.</emphasis>
<taxonomicName id="ABD34D0DFF91FF9B5BF802CDFBF0D196" authority="Cadena-Castañeda &amp; Díaz &amp; López &amp; Cárdenas, 2021" authorityName="Cadena-Castañeda &amp; Díaz &amp; López &amp; Cárdenas" authorityYear="2021" box="[903,1119,1712,1739]" class="Insecta" family="Haglotettigoniidae" genus="Anurogryllus" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="518" phylum="Arthropoda" rank="species" species="edithsantosum" status="sp. nov." subGenus="Urogryllus">
(
<emphasis id="5EA7EA9CFF91FF9B5BEE02CCFC0AD196" box="[913,933,1713,1738]" italics="true" pageId="3" pageNumber="518">U</emphasis>
.)
<emphasis id="5EA7EA9CFF91FF9B5BC202CDFBF0D196" box="[957,1119,1712,1738]" italics="true" pageId="3" pageNumber="518">edithsantosum</emphasis>
</taxonomicName>
<emphasis id="5EA7EA9CFF91FF9B5C1902CDFB0AD196" bold="true" box="[1126,1189,1712,1738]" pageId="3" pageNumber="518">
<taxonomicNameLabel id="459457E7FF91FF9B5C1902CDFB0AD196" box="[1126,1189,1712,1738]" pageId="3" pageNumber="518" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs in the incision of the median lophi, which is deeper and wider to
<emphasis id="5EA7EA9CFF91FF9B5AB302A9FD4BD1B2" box="[716,740,1748,1774]" italics="true" pageId="3" pageNumber="518">A.</emphasis>
(
<emphasis id="5EA7EA9CFF91FF9B5A8B02A8FCA7D1B2" box="[756,776,1749,1774]" italics="true" pageId="3" pageNumber="518">U</emphasis>
.)
<emphasis id="5EA7EA9CFF91FF9B5B5F02A8FCD8D1B2" box="[800,887,1749,1774]" italics="true" pageId="3" pageNumber="518">gnomus</emphasis>
. The new species has the ectophallic fold of medium size, but in
<emphasis id="5EA7EA9CFF91FF9B59210285FEC0D04E" box="[350,367,1784,1810]" italics="true" pageId="3" pageNumber="518">A</emphasis>
. (
<emphasis id="5EA7EA9CFF91FF9B59F70284FE33D04E" box="[392,412,1785,1810]" italics="true" pageId="3" pageNumber="518">U</emphasis>
.)
<emphasis id="5EA7EA9CFF91FF9B59CA0284FE5AD04E" box="[437,501,1785,1810]" italics="true" pageId="3" pageNumber="518">nigua</emphasis>
is elongated, in lateral view the median lophi and lateral lophi of pseudepiphallus looks altogether like a snout, and between these the ectophallic fold emerges like a tongue.
<emphasis id="5EA7EA9CFF91FF9B5CDA0361FB12D06A" box="[1189,1213,1820,1846]" italics="true" pageId="3" pageNumber="518">A.</emphasis>
<taxonomicName id="ABD34D0DFF91FF9B5CBA0361FA33D06A" authority="Cadena-Castañeda &amp; Díaz &amp; López &amp; Cárdenas, 2021" authorityName="Cadena-Castañeda &amp; Díaz &amp; López &amp; Cárdenas" authorityYear="2021" box="[1221,1436,1820,1847]" class="Insecta" family="Haglotettigoniidae" genus="Anurogryllus" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="518" phylum="Arthropoda" rank="species" species="edithsantosum" status="sp. nov." subGenus="Urogryllus">
(
<emphasis id="5EA7EA9CFF91FF9B5CB10360FB4DD06A" box="[1230,1250,1821,1846]" italics="true" pageId="3" pageNumber="518">U</emphasis>
.)
<emphasis id="5EA7EA9CFF91FF9B5C850361FA33D06A" box="[1274,1436,1820,1846]" italics="true" pageId="3" pageNumber="518">edithsantosum</emphasis>
</taxonomicName>
<emphasis id="5EA7EA9CFF91FF9B58E8033DFF78D006" bold="true" box="[151,215,1856,1882]" pageId="3" pageNumber="518">
<taxonomicNameLabel id="459457E7FF91FF9B58E8033DFF78D006" box="[151,215,1856,1882]" pageId="3" pageNumber="518" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs from the three mentioned species, in its larger size, brown color of the head, and the pseudepiphallic median lophi is covered with hairs (as the species of the subgenus
<emphasis id="5EA7EA9CFF91FF9B5BF90319FBB4D022" box="[902,1051,1892,1918]" italics="true" pageId="3" pageNumber="518">Pilosogryllus</emphasis>
), in contrast, the other species are of medium to small size, the head is black, and the pseudepiphallic median lophi is hairless.
</paragraph>
</subSubSection>
</treatment>
</document>