303 lines
38 KiB
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303 lines
38 KiB
XML
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<mods:title id="867342A950D752226B73B1D2A4E42A92">Systematics and Phylogenetics of Indo-Pacific Luciolinae Fireflies (Coleoptera: Lampyridae) and the Description of new Genera</mods:title>
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<mods:namePart id="5DB4B160F35BC66D5EC2D97433991D11">Ballantyne, Lesley A.</mods:namePart>
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<mods:nameIdentifier id="8778FA7C55F35411DFCBA7453EDDBC9A" type="email">lballantyne@csu.edu.au</mods:nameIdentifier>
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<mods:namePart id="F8B12C5FA71724469596082FC3A6F14C">Lambkin, Christine L.</mods:namePart>
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<mods:title id="C0922481958AEB7903E89E9CAE10834B">Zootaxa</mods:title>
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<mods:date id="A6AEE8C7E5CA105AE8CF3E11B64E811F">2013</mods:date>
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<mods:number id="92AE1437D5B2B66993589171BB6E171D">2013-05-22</mods:number>
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<paragraph id="6C7943005E6CC523FF6FBF84FE53EDDA" blockId="110.[151,389,151,210]" box="[151,389,151,177]" pageId="110" pageNumber="111">
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<heading id="3731F46C5E6CC523FF6FBF84FE53EDDA" bold="true" box="[151,389,151,177]" fontSize="11" level="2" pageId="110" pageNumber="111" reason="4">
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<taxonomicName id="ABC638835E6CC523FF6FBF84FE53EDDA" ID-CoL="765M" authority="Olivier" authorityName="Olivier" box="[151,389,151,177]" class="Insecta" family="Lampyridae" genus="Pyrophanes" kingdom="Animalia" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="genus">
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<emphasis id="5EB29F125E6CC523FF6FBF84FE53EDDA" bold="true" box="[151,389,151,177]" pageId="110" pageNumber="111">
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<emphasis id="5EB29F125E6CC523FF6FBF84FEF0EDDA" bold="true" box="[151,294,151,177]" italics="true" pageId="110" pageNumber="111">Pyrophanes</emphasis>
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Olivier
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="24DC108B5E6CC523FF6FBFA9FE94EDB9" box="[151,322,186,210]" pageId="110" pageNumber="111" type="description">
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<paragraph id="6C7943005E6CC523FF6FBFA9FE94EDB9" blockId="110.[151,389,151,210]" box="[151,322,186,210]" pageId="110" pageNumber="111">
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[
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<figureCitation id="F4FD5F855E6CC523FF65BFA9FEEAEDB9" box="[157,316,186,210]" captionStart="FIGURES 221–227" captionStartId="109.[151,263,1767,1790]" captionTargetBox="[151,1436,193,1742]" captionTargetId="figure-116@109.[151,1436,193,1746]" captionTargetPageId="109" captionText="FIGURES 221–227. Pyrophanes spp. 221, 222 P. appendiculata (slides, USNM). 223, 224 Luciola venusta type male (MNHN). 225, 226 P. quadrimaculata type female (MNHN). 227 Luciola semilimbata type female (MNHN). 221 male, 222 female abdomens; 223, 225 dorsal; 224 ventral; 226 type labels. Figure legends: 1 anterior projection of aedeagal sheath sternite; 2 basal piece aedeagus; 3 lateral area anterior prolongation of T8; 4 posterior area of aedeagal sheath in region of paraprocts; 5 ventrolateral surface of T8 in spined area; 6 posterolateral area of T8; 7 anterior plates bursa; 8 posterior plates bursa; 9 median oviduct plate." figureDoi="http://doi.org/10.5281/zenodo.5265424" httpUri="https://zenodo.org/record/5265424/files/figure.png" pageId="110" pageNumber="111">Figs 221–227</figureCitation>
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]
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</paragraph>
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</subSubSection>
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<subSubSection id="24DC108B5E6CC523FF6FBFEFFE8FEC24" pageId="110" pageNumber="111" type="reference_group">
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<paragraph id="6C7943005E6CC523FF6FBFEFFE8FEC24" blockId="110.[151,1436,251,335]" pageId="110" pageNumber="111">
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<treatmentCitation id="ED6765115E6CC523FF6FBFEFFE1DEC7A" author="Olivier, E." box="[151,459,251,274]" page="368" pageId="110" pageNumber="111" year="1885">
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<taxonomicName id="ABC638835E6CC523FF6FBFEFFE1DEC7A" ID-CoL="765M" authority="Olivier, 1885: 368" authorityName="Olivier" authorityPageNumber="368" authorityYear="1885" box="[151,459,251,274]" class="Insecta" family="Lampyridae" genus="Pyrophanes" kingdom="Animalia" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="genus">
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<emphasis id="5EB29F125E6CC523FF6FBFEFFED9EC79" box="[151,271,252,274]" italics="true" pageId="110" pageNumber="111">Pyrophanes</emphasis>
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<bibRefCitation id="08573EF15E6CC523FEEDBFE8FE1DEC7A" author="Olivier, E." box="[277,459,251,273]" pageId="110" pageNumber="111" pagination="333 - 374" refId="ref128538" refString="Olivier, E. (1885) Catalogue des Lampyrides faisant partie des collections du Musee Civique de Genes. Annali del Museo Civico di Storia Naturale di Genova, (2 a) 2, 333 - 374." type="journal article" year="1885">Olivier, 1885:368</bibRefCitation>
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</taxonomicName>
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; in Baer, 1886:132; 1902:72; 1907:56; 1911b:102.
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<treatmentCitation id="ED6765115E6CC523FC28BFEFFB7AEC7A" author="McDermott, F. A." box="[976,1196,251,273]" page="46" pageId="110" pageNumber="111" year="1964">
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<bibRefCitation id="08573EF15E6CC523FC28BFEFFB7AEC7A" author="McDermott, F. A." box="[976,1196,251,273]" pageId="110" pageNumber="111" pagination="1 - 72" refId="ref127702" refString="McDermott, F. A. (1964) The taxonomy of the Lampyridae. Transactions of the American Entomological Society, 90, 1 - 72." type="journal article" year="1964">McDermott, 1964:46</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="ED6765115E6CC523FB41BFE8FACDEC7A" author="McDermott, F. A." box="[1209,1307,251,273]" page="116" pageId="110" pageNumber="111" year="1966">
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<bibRefCitation id="08573EF15E6CC523FB41BFE8FACDEC7A" author="McDermott, F. A." box="[1209,1307,251,273]" pageId="110" pageNumber="111" pagination="98 - 118" refId="ref127730" refString="McDermott, F. A. (1966) Lampyridae Pars. 9. Coleopterorum Catalogus (Junk-Schenkling) (Ed. Sec), 1 - 149 (Luciolini, 98 - 118)." type="book chapter" year="1966">1966:116</bibRefCitation>
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</treatmentCitation>
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.
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<treatmentCitation id="ED6765115E6CC523FAD0BFE8FEFEEC5B" author="Ballantyne, L. A." page="106" pageId="110" pageNumber="111" year="1968">
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<bibRefCitation id="08573EF15E6CC523FAD0BFE8FEFEEC5B" author="Ballantyne, L. A." pageId="110" pageNumber="111" pagination="103 - 139" refId="ref122148" refString="Ballantyne, L. A. (1968) Revisional Studies of Australian and Indomalayan Luciolini (Coleoptera, Lampyridae, Luciolinae). University of Queensland Papers, Department of Entomology, University of Queensland Press, II, no 6, 103 - 139." type="journal article" year="1968">Ballantyne, 1968:106</bibRefCitation>
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</treatmentCitation>
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, 107;
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||
<treatmentCitation id="ED6765115E6CC523FE9EBE09FDDEEC5B" author="Ballantyne, L. A." box="[358,520,282,304]" page="173" pageId="110" pageNumber="111" year="1987">
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<bibRefCitation id="08573EF15E6CC523FE9EBE09FDDEEC5B" author="Ballantyne, L. A." box="[358,520,282,304]" pageId="110" pageNumber="111" pagination="171 - 188" refId="ref122235" refString="Ballantyne, L. A. (1987 b) Lucioline Morphology, Taxonomy and Behaviour: A Reappraisal. (Coleoptera, Lampyridae). Transactions of the American Entomological Society, 113, 171 - 188." type="journal article" year="1987">1987b:173–176</bibRefCitation>
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</treatmentCitation>
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. Ballantyne in
|
||
<treatmentCitation id="ED6765115E6CC523FD58BE09FC84EC5B" author="Calder, A. A." box="[672,850,282,304]" page="180" pageId="110" pageNumber="111" year="1998">
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<bibRefCitation id="08573EF15E6CC523FD58BE09FC84EC5B" author="Calder, A. A." box="[672,850,282,304]" pageId="110" pageNumber="111" refId="ref123793" refString="Calder, A. A. (1998) Coleoptera: Elateroidea In Wells, A. (ed.) Zoological Catalogue of Australia, 29.6 (CSIRO Publishing Melbourne)." type="book" year="1998">Calder, 1998:180</bibRefCitation>
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</treatmentCitation>
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||
.
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||
<treatmentCitation id="ED6765115E6CC523FCA3BE09FB3EEC5B" author="Ballantyne, L. A. & Lambkin, C." box="[859,1256,282,304]" page="2009" pageId="110" pageNumber="111" year="2000">
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<bibRefCitation id="08573EF15E6CC523FCA3BE09FB7FEC5B" author="Ballantyne, L. A. & Lambkin, C." box="[859,1193,282,304]" pageId="110" pageNumber="111" pagination="15 - 93" refId="ref122603" refString="Ballantyne, L. A. & Lambkin, C. (2000) Lampyridae of Australia (Coleoptera: Lampyridae: Luciolinae: Luciolini). Memoirs of the Queensland Museum, 46 (1), 15 - 93." type="journal article" year="2000">Ballantyne & Lambkin, 2000:70</bibRefCitation>
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||
: 2009
|
||
</treatmentCitation>
|
||
:
|
||
<figureCitation id="F4FD5F855E6CC523FB09BE09FAB2EC5B" box="[1265,1380,282,304]" captionStart="FIGURES 92–98" captionStartId="60.[151,264,1948,1971]" captionTargetBox="[151,1436,475,1930]" captionTargetId="figure-174@60.[151,1436,475,1936]" captionTargetPageId="60" captionText="FIGURES 92–98. Colophotia plagiata bred female (USNM). 92, 93 dorsal (92) and ventral body; 94, 95 dorsal (94) and ventral abdomen (soaked in cold KOH), anterior apodeme of V8 arrowed in 95; 96–98 bursa ventral (96), and lateral (97, 98). Figure legends: 2 posterior plate; 3 anterior plate." figureDoi="http://doi.org/10.5281/zenodo.5266139" httpUri="https://zenodo.org/record/5266139/files/figure.png" pageId="110" pageNumber="111">Figs 94, 95</figureCitation>
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,
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||
<figureCitation id="F4FD5F855E6CC523FA88BE09FF39EC24" captionStart="FIGURES 104–108" captionStartId="62.[151,263,1642,1665]" captionTargetBox="[151,1435,189,1622]" captionTargetId="figure-167@62.[151,1436,183,1626]" captionTargetPageId="62" captionText="FIGURES 104–108. Colophotia females. 104–106 Colophotia praeusta female (USNM). 107, 108 C. plagiata male (USNM). 104, 105 dorsal (104) and ventral; 106 bursa and V8 ventral (2 posterior plate; 3 anterior plate); 107 abdomen LO segments ventral; 108 abdomen cleared in cold KOH from beneath. Figure legends: T7 tergite 7 anterior prolongations right side; T8 tergite 8." figureDoi="http://doi.org/10.5281/zenodo.5265386" httpUri="https://zenodo.org/record/5265386/files/figure.png" pageId="110" pageNumber="111">106, 107</figureCitation>
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, 503, 509.
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||
</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="24DC108B5E6CC523FF6FBE66FC13ECE0" box="[151,965,372,395]" pageId="110" pageNumber="111" type="type_taxon">
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<paragraph id="6C7943005E6CC523FF6FBE66FC13ECE0" blockId="110.[151,965,372,395]" box="[151,965,372,395]" pageId="110" pageNumber="111">
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<emphasis id="5EB29F125E6CC523FF6FBE66FEFCECE0" bold="true" box="[151,298,372,395]" pageId="110" pageNumber="111">
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<typeStatus id="B37DFDA25E6CC523FF6FBE66FF1AECE1" box="[151,204,373,394]" pageId="110" pageNumber="111">Type</typeStatus>
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Species:
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</emphasis>
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||
<taxonomicName id="ABC638835E6CC523FEC8BE66FDE9ECE0" ID-CoL="4QV6L" authority="Olivier" authorityName="Olivier" box="[304,575,373,395]" class="Insecta" family="Lampyridae" genus="Pyrophanes" kingdom="Animalia" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="similis">
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<emphasis id="5EB29F125E6CC523FEC8BE66FE26ECE0" box="[304,496,373,395]" italics="true" pageId="110" pageNumber="111">Pyrophanes similis</emphasis>
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||
Olivier
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||
</taxonomicName>
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||
, designated by
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<bibRefCitation id="08573EF15E6CC523FD24BE65FC17ECE0" author="McDermott, F. A." box="[732,961,373,395]" pageId="110" pageNumber="111" pagination="98 - 118" refId="ref127730" refString="McDermott, F. A. (1966) Lampyridae Pars. 9. Coleopterorum Catalogus (Junk-Schenkling) (Ed. Sec), 1 - 149 (Luciolini, 98 - 118)." type="book chapter" year="1966">McDermott, 1966:116</bibRefCitation>
|
||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="24DC108B5E6CC523FF6FBEA3FDF2E929" pageId="110" pageNumber="111" type="diagnosis">
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||
<paragraph id="6C7943005E6CC523FF6FBEA3FC46EE5A" blockId="110.[151,1436,432,818]" pageId="110" pageNumber="111">
|
||
<emphasis id="5EB29F125E6CC523FF6FBEA3FEC7ECA2" bold="true" box="[151,273,432,457]" pageId="110" pageNumber="111">Diagnosis.</emphasis>
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||
<taxonomicName id="ABC638835E6CC523FEE0BEA2FE4BECA2" box="[280,413,433,457]" class="Insecta" family="Lampyridae" genus="Pyrophanes" kingdom="Animalia" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="genus">
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||
<emphasis id="5EB29F125E6CC523FEE0BEA2FE4BECA2" box="[280,413,433,457]" italics="true" pageId="110" pageNumber="111">Pyrophanes</emphasis>
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||
</taxonomicName>
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||
belongs to a group of 7
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||
<taxonomicName id="ABC638835E6CC523FD54BEA2FCF5ECA2" authorityName="sensu McDermott" authorityYear="1966" box="[684,803,433,457]" class="Insecta" family="Lampyridae" genus="Luciolinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="110" pageNumber="126" phylum="Arthropoda" rank="genus">Luciolinae</taxonomicName>
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||
genera with males characterized by: an elongate slender aedeagus with LL concealed behind the ML when viewed from beneath, pronotal width less than width across the elytral humeri, parallel-sided elytra, aedeagal sheath elongate slender, widest across the middle, with posterior half of sternite not emarginate on either side, and tapering evenly towards a narrow entire apex. It is distinguished from all other genera by a combination of MFC and paraprocts on the aedeagal sheath; without deflexed elytral apices; femora 3 are swollen and curved in four of the five species; the posterior margin of V7 is trilobed with incurving lobes and small pointed projections between the PLP and MPP; the underside of T8 has depressions at the sides housing very short spines and hairs. Females are macropterous and the bursa in certain species has wide paired plates. Larvae lack laterally explanate margins in
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<taxonomicName id="ABC638835E6CC523FD29BDC2FC7EEF82" box="[721,936,721,745]" class="Insecta" family="Lampyridae" genus="Pyrophanes" kingdom="Animalia" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="similis">
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||
<emphasis id="5EB29F125E6CC523FD29BDC2FC7EEF82" box="[721,936,721,745]" italics="true" pageId="110" pageNumber="111">Pyrophanes similis</emphasis>
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||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="08573EF15E6CC523FC41BDC2FBEAEF82" author="Blair, K. G." box="[953,1084,721,745]" pageId="110" pageNumber="111" pagination="43 - 45" refId="ref123002" refString="Blair, K. G. (1927) An aquatic Lampyrid larva from S. Celebes. Transactions of the Entomological Society of London, Part 1. 75, 43 - 45." type="journal article" year="1927">Blair, 1927</bibRefCitation>
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||
; Bertrand, 1972). This larva is not however reliably associated and an accurate identification of larval
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||
<typeStatus id="B37DFDA25E6CC523FC34BDE5FC2AEE65" box="[972,1020,758,782]" pageId="110" pageNumber="111">type</typeStatus>
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for this genus is not available. This genus needs more extensive revision than is currently possible here.
|
||
</paragraph>
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||
<paragraph id="6C7943005E6CC523FF6FBC64FE93EEFA" blockId="110.[151,325,887,913]" box="[151,325,887,913]" pageId="110" pageNumber="111">
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||
<emphasis id="5EB29F125E6CC523FF6FBC64FE93EEFA" bold="true" box="[151,325,887,913]" pageId="110" pageNumber="111">List of species</emphasis>
|
||
</paragraph>
|
||
<paragraph id="6C7943005E6CC523FF6FBCABFE62EEA7" blockId="110.[151,548,951,1090]" box="[151,436,951,973]" pageId="110" pageNumber="111">
|
||
<emphasis id="5EB29F125E6CC523FF6FBCABFF49EEA6" box="[151,159,952,973]" italics="true" pageId="110" pageNumber="111">-</emphasis>
|
||
<taxonomicName id="ABC638835E6CC523FF19BCABFE62EEA7" authority="Olivier" authorityName="Olivier" box="[225,436,951,973]" genus="Pyr" pageId="110" pageNumber="111" rank="species" species="appendiculata">
|
||
<emphasis id="5EB29F125E6CC523FF19BCABFEBCEEA6" box="[225,362,952,973]" italics="true" pageId="110" pageNumber="111">appendiculata</emphasis>
|
||
Olivier
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="6C7943005E6CC523FF6FBCC6FEAFEE81" blockId="110.[151,548,951,1090]" box="[151,377,981,1002]" pageId="110" pageNumber="111">
|
||
<emphasis id="5EB29F125E6CC523FF6FBCC6FF49EE81" box="[151,159,981,1002]" italics="true" pageId="110" pageNumber="111">-</emphasis>
|
||
<taxonomicName id="ABC638835E6CC523FF19BCC6FEAFEE81" authority="Olivier" authorityName="Olivier" box="[225,377,981,1002]" genus="Pyro" pageId="110" pageNumber="111" rank="species" species="beccarii">
|
||
<emphasis id="5EB29F125E6CC523FF19BCC6FEE6EE81" box="[225,304,981,1002]" italics="true" pageId="110" pageNumber="111">beccarii</emphasis>
|
||
Olivier
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="6C7943005E6CC523FF6FBCE1FE12E96C" blockId="110.[151,548,951,1090]" box="[151,452,1010,1031]" pageId="110" pageNumber="111">
|
||
<emphasis id="5EB29F125E6CC523FF6FBCE1FF49E96C" box="[151,159,1010,1031]" italics="true" pageId="110" pageNumber="111">-</emphasis>
|
||
<taxonomicName id="ABC638835E6CC523FF19BCE1FE12E96C" authority="Olivier" authorityName="Olivier" box="[225,452,1010,1031]" genus="Pyro" pageId="110" pageNumber="111" rank="species" species="quadrimaculata">
|
||
<emphasis id="5EB29F125E6CC523FF19BCE1FEACE96C" box="[225,378,1010,1031]" italics="true" pageId="110" pageNumber="111">quadrimaculata</emphasis>
|
||
Olivier
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="6C7943005E6CC523FF6FBB03FEBFE94F" blockId="110.[151,548,951,1090]" box="[151,361,1039,1061]" pageId="110" pageNumber="111">
|
||
<emphasis id="5EB29F125E6CC523FF6FBB03FF49E94E" box="[151,159,1040,1061]" italics="true" pageId="110" pageNumber="111">-</emphasis>
|
||
<taxonomicName id="ABC638835E6CC523FF19BB03FEBFE94F" authority="Olivier" authorityName="Olivier" box="[225,361,1039,1061]" class="Insecta" family="Lampyridae" genus="Pyrophanes" kingdom="Animalia" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="5EB29F125E6CC523FF19BB03FEF6E94E" box="[225,288,1040,1061]" italics="true" pageId="110" pageNumber="111">similis</emphasis>
|
||
Olivier
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="6C7943005E6CC523FF6FBB3EFDF2E929" blockId="110.[151,548,951,1090]" box="[151,548,1069,1090]" pageId="110" pageNumber="111">
|
||
<emphasis id="5EB29F125E6CC523FF6FBB3EFF49E929" box="[151,159,1069,1090]" italics="true" pageId="110" pageNumber="111">-</emphasis>
|
||
<taxonomicName id="ABC638835E6CC523FF19BB3EFE66E929" authority="(Olivier)" baseAuthorityName="Olivier" box="[225,432,1069,1090]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="semilimbata" status="comb. nov.">
|
||
<emphasis id="5EB29F125E6CC523FF19BB3EFE80E929" box="[225,342,1069,1090]" italics="true" pageId="110" pageNumber="111">semilimbata</emphasis>
|
||
(Olivier)
|
||
</taxonomicName>
|
||
<emphasis id="5EB29F125E6CC523FE4EBB3EFDF2E929" bold="true" box="[438,548,1069,1090]" pageId="110" pageNumber="111">
|
||
<taxonomicNameLabel id="458122695E6CC523FE4EBB3EFDF2E929" box="[438,548,1069,1090]" pageId="110" pageNumber="111" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="24DC108B5E6CC523FF6FBB96FA4AE8CC" pageId="110" pageNumber="111" type="key">
|
||
<paragraph id="6C7943005E6CC523FF6FBB96FD23E9F4" blockId="110.[151,757,1157,1183]" box="[151,757,1157,1183]" pageId="110" pageNumber="111">
|
||
<emphasis id="5EB29F125E6CC523FF6FBB96FD23E9F4" bold="true" box="[151,757,1157,1183]" pageId="110" pageNumber="111">
|
||
Key to species of
|
||
<taxonomicName id="ABC638835E6CC523FE95BB96FD8AE9F4" ID-CoL="765M" authority="Olivier" authorityName="Olivier" box="[365,604,1157,1183]" class="Insecta" family="Lampyridae" genus="Pyrophanes" kingdom="Animalia" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5EB29F125E6CC523FE95BB96FE2AE9F4" bold="true" box="[365,508,1157,1183]" italics="true" pageId="110" pageNumber="111">Pyrophanes</emphasis>
|
||
Olivier
|
||
</taxonomicName>
|
||
using males
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="6C7943005E6CC523FF6FBBD6FA4AE99C" blockId="110.[151,1436,1221,1447]" pageId="110" pageNumber="111">
|
||
1. Hind femora not swollen or curved (
|
||
<figureCitation id="F4FD5F855E6CC523FDC6BBD6FD15E9B1" box="[574,707,1221,1242]" captionStart="FIGURES 221–227" captionStartId="109.[151,263,1767,1790]" captionTargetBox="[151,1436,193,1742]" captionTargetId="figure-116@109.[151,1436,193,1746]" captionTargetPageId="109" captionText="FIGURES 221–227. Pyrophanes spp. 221, 222 P. appendiculata (slides, USNM). 223, 224 Luciola venusta type male (MNHN). 225, 226 P. quadrimaculata type female (MNHN). 227 Luciola semilimbata type female (MNHN). 221 male, 222 female abdomens; 223, 225 dorsal; 224 ventral; 226 type labels. Figure legends: 1 anterior projection of aedeagal sheath sternite; 2 basal piece aedeagus; 3 lateral area anterior prolongation of T8; 4 posterior area of aedeagal sheath in region of paraprocts; 5 ventrolateral surface of T8 in spined area; 6 posterolateral area of T8; 7 anterior plates bursa; 8 posterior plates bursa; 9 median oviduct plate." figureDoi="http://doi.org/10.5281/zenodo.5265424" httpUri="https://zenodo.org/record/5265424/files/figure.png" pageId="110" pageNumber="111">Figs 223, 224</figureCitation>
|
||
)................................
|
||
<taxonomicName id="ABC638835E6CC523FBA1BBD6FAFEE9B1" ID-CoL="3WCVV" authority="(Olivier)" baseAuthorityName="Olivier" box="[1113,1320,1221,1242]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="semilimbata" status="comb. nov.">
|
||
<emphasis id="5EB29F125E6CC523FBA1BBD6FB18E9B1" box="[1113,1230,1221,1242]" italics="true" pageId="110" pageNumber="111">semilimbata</emphasis>
|
||
(Olivier)
|
||
</taxonomicName>
|
||
<emphasis id="5EB29F125E6CC523FAD6BBD6FA4AE9B1" bold="true" box="[1326,1436,1221,1242]" pageId="110" pageNumber="111">
|
||
<taxonomicNameLabel id="458122695E6CC523FAD6BBD6FA4AE9B1" box="[1326,1436,1221,1242]" pageId="110" pageNumber="111" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Hind femora swollen and curved......................................................................... 2
|
||
</paragraph>
|
||
<paragraph id="6C7943005E6CC523FF6FBBECFA4AE859" blockId="110.[151,1436,1221,1447]" pageId="110" pageNumber="111">
|
||
2. Elytra medium to dark brown, often with lateral and sutural margins semitransparent and pale (
|
||
<figureCitation id="F4FD5F855E6CC523FB88BBECFB23E87F" box="[1136,1269,1279,1300]" captionStart="FIGURES 221–227" captionStartId="109.[151,263,1767,1790]" captionTargetBox="[151,1436,193,1742]" captionTargetId="figure-116@109.[151,1436,193,1746]" captionTargetPageId="109" captionText="FIGURES 221–227. Pyrophanes spp. 221, 222 P. appendiculata (slides, USNM). 223, 224 Luciola venusta type male (MNHN). 225, 226 P. quadrimaculata type female (MNHN). 227 Luciola semilimbata type female (MNHN). 221 male, 222 female abdomens; 223, 225 dorsal; 224 ventral; 226 type labels. Figure legends: 1 anterior projection of aedeagal sheath sternite; 2 basal piece aedeagus; 3 lateral area anterior prolongation of T8; 4 posterior area of aedeagal sheath in region of paraprocts; 5 ventrolateral surface of T8 in spined area; 6 posterolateral area of T8; 7 anterior plates bursa; 8 posterior plates bursa; 9 median oviduct plate." figureDoi="http://doi.org/10.5281/zenodo.5265424" httpUri="https://zenodo.org/record/5265424/files/figure.png" pageId="110" pageNumber="111">Figs 223, 224</figureCitation>
|
||
)............ 3 Elytra dingy or pale, clear yellow, always with apex dark brown to black; base of elytra may be brown (
|
||
<figureCitation id="F4FD5F855E6CC523FB3ABA0EFAC2E859" box="[1218,1300,1309,1330]" captionStart="FIGURES 221–227" captionStartId="109.[151,263,1767,1790]" captionTargetBox="[151,1436,193,1742]" captionTargetId="figure-116@109.[151,1436,193,1746]" captionTargetPageId="109" captionText="FIGURES 221–227. Pyrophanes spp. 221, 222 P. appendiculata (slides, USNM). 223, 224 Luciola venusta type male (MNHN). 225, 226 P. quadrimaculata type female (MNHN). 227 Luciola semilimbata type female (MNHN). 221 male, 222 female abdomens; 223, 225 dorsal; 224 ventral; 226 type labels. Figure legends: 1 anterior projection of aedeagal sheath sternite; 2 basal piece aedeagus; 3 lateral area anterior prolongation of T8; 4 posterior area of aedeagal sheath in region of paraprocts; 5 ventrolateral surface of T8 in spined area; 6 posterolateral area of T8; 7 anterior plates bursa; 8 posterior plates bursa; 9 median oviduct plate." figureDoi="http://doi.org/10.5281/zenodo.5265424" httpUri="https://zenodo.org/record/5265424/files/figure.png" pageId="110" pageNumber="111">Fig. 225</figureCitation>
|
||
)......... 4
|
||
</paragraph>
|
||
<paragraph id="6C7943005E6CC523FF6FBA29FA4AE807" blockId="110.[151,1436,1221,1447]" pageId="110" pageNumber="111">
|
||
3 Elytra medium to dark brown, lateral margin, and often also sutural margin, yellow................
|
||
<taxonomicName id="ABC638835E6CC523FB31BA29FECAE807" authority="Olivier Elytra" authorityName="Olivier Elytra" genus="Pyr" pageId="110" pageNumber="111" rank="species" species="appendiculata">
|
||
<emphasis id="5EB29F125E6CC523FB31BA29FA84E824" box="[1225,1362,1338,1359]" italics="true" pageId="110" pageNumber="111">appendiculata</emphasis>
|
||
Olivier Elytra
|
||
</taxonomicName>
|
||
medium to dark brown, without any paler margins..........................................
|
||
<taxonomicName id="ABC638835E6CC523FAFBBA4BFA4AE807" authority="Olivier" authorityName="Olivier" box="[1283,1436,1367,1389]" genus="Pyro" pageId="110" pageNumber="111" rank="species" species="beccarii">
|
||
<emphasis id="5EB29F125E6CC523FAFBBA4BFA84E806" box="[1283,1362,1368,1389]" italics="true" pageId="110" pageNumber="111">beccarii</emphasis>
|
||
Olivier
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="6C7943005E6CC523FF6FBA66FA4AE8CC" blockId="110.[151,1436,1221,1447]" pageId="110" pageNumber="111">
|
||
4 Elytra yellow with apical and basal dark markings.........................................
|
||
<taxonomicName id="ABC638835E6CC523FB41BA66FECAE8CC" authority="Olivier Elytra" authorityName="Olivier Elytra" genus="Pyro" pageId="110" pageNumber="111" rank="species" species="quadrimaculata">
|
||
<emphasis id="5EB29F125E6CC523FB41BA66FA84E8E1" box="[1209,1362,1397,1418]" italics="true" pageId="110" pageNumber="111">quadrimaculata</emphasis>
|
||
Olivier Elytra
|
||
</taxonomicName>
|
||
yellow with only apices dark brown......................................................
|
||
<taxonomicName id="ABC638835E6CC523FAEBBA81FA4AE8CC" ID-CoL="4QV6L" authority="Olivier" authorityName="Olivier" box="[1299,1436,1426,1447]" class="Insecta" family="Lampyridae" genus="Pyrophanes" kingdom="Animalia" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="5EB29F125E6CC523FAEBBA81FA84E8CC" box="[1299,1362,1426,1447]" italics="true" pageId="110" pageNumber="111">similis</emphasis>
|
||
Olivier
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="24DC108B5E6CC523FF3FBADEFE99EB8A" pageId="110" pageNumber="111" type="discussion">
|
||
<paragraph id="6C7943005E6CC523FF3FBADEFE99EB8A" blockId="110.[151,1440,1485,1761]" pageId="110" pageNumber="111">
|
||
<emphasis id="5EB29F125E6CC523FF3FBADEFEE2E88E" bold="true" box="[199,308,1485,1509]" pageId="110" pageNumber="111">Remarks</emphasis>
|
||
.
|
||
<bibRefCitation id="08573EF15E6CC523FEBDBADEFE3CE88E" author="Olivier, E." box="[325,490,1485,1509]" pageId="110" pageNumber="111" pagination="333 - 374" refId="ref128538" refString="Olivier, E. (1885) Catalogue des Lampyrides faisant partie des collections du Musee Civique de Genes. Annali del Museo Civico di Storia Naturale di Genova, (2 a) 2, 333 - 374." type="journal article" year="1885">Olivier (1885)</bibRefCitation>
|
||
considered
|
||
<taxonomicName id="ABC638835E6CC523FD81BADEFC50E88E" authority="Motsch." authorityName="Motsch." box="[633,902,1485,1509]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="indica">
|
||
<emphasis id="5EB29F125E6CC523FD81BADEFCF6E88E" box="[633,800,1485,1509]" italics="true" pageId="110" pageNumber="111">Luciola indica</emphasis>
|
||
Motsch.
|
||
</taxonomicName>
|
||
approached
|
||
<emphasis id="5EB29F125E6CC523FBE4BADEFB7EE88E" box="[1052,1192,1485,1509]" italics="true" pageId="110" pageNumber="111">
|
||
<taxonomicName id="ABC638835E6CC523FBE4BADEFB72E88E" box="[1052,1188,1485,1509]" class="Insecta" family="Lampyridae" genus="Pyrophanes" kingdom="Animalia" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="genus">Pyrophanes</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and (1902) formally incorporated it. This species does not belong in
|
||
<taxonomicName id="ABC638835E6CC523FD22BAE2FC88EB62" box="[730,862,1521,1545]" class="Insecta" family="Lampyridae" genus="Pyrophanes" kingdom="Animalia" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5EB29F125E6CC523FD22BAE2FC88EB62" box="[730,862,1521,1545]" italics="true" pageId="110" pageNumber="111">Pyrophanes</emphasis>
|
||
</taxonomicName>
|
||
and is being addressed (Boontop, Lambkin and Ballantyne in prep.). Certain diagrams of the terminal abdomen of
|
||
<taxonomicName id="ABC638835E6CC523FC6DB906FBCCEB46" box="[917,1050,1557,1581]" class="Insecta" family="Lampyridae" genus="Pyrophanes" kingdom="Animalia" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5EB29F125E6CC523FC6DB906FBCCEB46" box="[917,1050,1557,1581]" italics="true" pageId="110" pageNumber="111">Pyrophanes</emphasis>
|
||
</taxonomicName>
|
||
are inaccurate e.g.
|
||
<bibRefCitation id="08573EF15E6CC523FB03B906FA41EB46" author="Olivier, E." box="[1275,1431,1557,1581]" pageId="110" pageNumber="111" pagination="333 - 374" refId="ref128538" refString="Olivier, E. (1885) Catalogue des Lampyrides faisant partie des collections du Musee Civique de Genes. Annali del Museo Civico di Storia Naturale di Genova, (2 a) 2, 333 - 374." type="journal article" year="1885">Olivier (1885</bibRefCitation>
|
||
, Plate V,
|
||
<figureCitation id="F4FD5F855E6CC523FEFBB92AFE9AEB3A" box="[259,332,1593,1617]" captionStart="FIGURES 9–17" captionStartId="39.[151,264,1950,1973]" captionTargetBox="[151,1436,265,1919]" captionTargetId="figure-85@39.[151,1436,265,1935]" captionTargetPageId="39" captionText="FIGURES 9–17. Australoluciola anthracina type male, Cyclops (MNHN). 9, 10 dorsal (9) and ventral; 11 V6, 7 ventral; 12 pronotum dorsal; 13 type labels; 14 aedeagal sheath ventral; 15-17 aedeagus, dorsal (15), ventral (16) and lateral. These figures share scale lines: 15–17." figureDoi="http://doi.org/10.5281/zenodo.5265360" httpUri="https://zenodo.org/record/5265360/files/figure.png" pageId="110" pageNumber="111">Fig.12</figureCitation>
|
||
) and
|
||
<bibRefCitation id="08573EF15E6CC523FE61B92AFDE1EB3A" author="Olivier, E." box="[409,567,1593,1617]" pageId="110" pageNumber="111" pagination="1 - 74" refId="ref128760" refString="Olivier, E. (1907) Coleoptera, Lampyridae. Genera Insectorum, 53, 1 - 74 [Luciolinae, 48 - 56]." type="journal article" year="1907">Olivier (1907</bibRefCitation>
|
||
Plate 3
|
||
<figureCitation id="F4FD5F855E6CC523FD5FB92AFD29EB3A" box="[679,767,1593,1617]" captionStart="FIGURES 9–17" captionStartId="39.[151,264,1950,1973]" captionTargetBox="[151,1436,265,1919]" captionTargetId="figure-85@39.[151,1436,265,1935]" captionTargetPageId="39" captionText="FIGURES 9–17. Australoluciola anthracina type male, Cyclops (MNHN). 9, 10 dorsal (9) and ventral; 11 V6, 7 ventral; 12 pronotum dorsal; 13 type labels; 14 aedeagal sheath ventral; 15-17 aedeagus, dorsal (15), ventral (16) and lateral. These figures share scale lines: 15–17." figureDoi="http://doi.org/10.5281/zenodo.5265360" httpUri="https://zenodo.org/record/5265360/files/figure.png" pageId="110" pageNumber="111">Fig. 10</figureCitation>
|
||
) are inaccurate representations of the terminal ventrite;
|
||
<bibRefCitation id="08573EF15E6CC523FF6FB94EFE57EB1E" author="McDermott, F. A." box="[151,385,1629,1653]" pageId="110" pageNumber="111" pagination="1 - 72" refId="ref127702" refString="McDermott, F. A. (1964) The taxonomy of the Lampyridae. Transactions of the American Entomological Society, 90, 1 - 72." type="journal article" year="1964">McDermott’s (1964)</bibRefCitation>
|
||
brief redescription of
|
||
<taxonomicName id="ABC638835E6CC523FD72B94EFCD8EB1E" box="[650,782,1629,1653]" class="Insecta" family="Lampyridae" genus="Pyrophanes" kingdom="Animalia" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5EB29F125E6CC523FD72B94EFCD8EB1E" box="[650,782,1629,1653]" italics="true" pageId="110" pageNumber="111">Pyrophanes</emphasis>
|
||
</taxonomicName>
|
||
was probably influenced by the atypical abdomen of
|
||
<taxonomicName id="ABC638835E6CC523FA7FB94DFE5AEBF2" authority="McLean" authorityName="McLean" class="Insecta" family="Lampyridae" genus="Photinus" kingdom="Animalia" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="macdermotti">
|
||
<emphasis id="5EB29F125E6CC523FA7FB94DFEF2EBF2" italics="true" pageId="110" pageNumber="111">P. macdermotti</emphasis>
|
||
McLean
|
||
</taxonomicName>
|
||
which he is known to have examined at this time and he misinterpreted various aspects. Ballantyne in
|
||
<bibRefCitation id="08573EF15E6CC523FEC2B9B6FE0CEBD6" author="Calder, A. A." box="[314,474,1701,1725]" pageId="110" pageNumber="111" refId="ref123793" refString="Calder, A. A. (1998) Coleoptera: Elateroidea In Wells, A. (ed.) Zoological Catalogue of Australia, 29.6 (CSIRO Publishing Melbourne)." type="book" year="1998">Calder (1998)</bibRefCitation>
|
||
synonymised
|
||
<taxonomicName id="ABC638835E6CC523FD7AB9B6FC45EBD6" authority="Lea" authorityName="Lea" authorityYear="1921" box="[642,915,1701,1725]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="complicata">
|
||
<emphasis id="5EB29F125E6CC523FD7AB9B6FC88EBD6" box="[642,862,1701,1725]" italics="true" pageId="110" pageNumber="111">Luciola complicata</emphasis>
|
||
Lea
|
||
</taxonomicName>
|
||
from far north
|
||
<collectingRegion id="AE028DE25E6CC523FBB5B9B6FB07EBD6" box="[1101,1233,1701,1725]" country="Australia" name="Queensland" pageId="110" pageNumber="111">Queensland</collectingRegion>
|
||
with
|
||
<taxonomicName id="ABC638835E6CC523FAEFB9B6FE99EB8A" authority="Olivier." authorityName="Olivier." class="Insecta" family="Lampyridae" genus="Pyrophanes" kingdom="Animalia" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="beccarii">
|
||
<emphasis id="5EB29F125E6CC523FAEFB9B6FF25EB8A" italics="true" pageId="110" pageNumber="111">Pyrophanes beccarii</emphasis>
|
||
Olivier.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |