154 lines
26 KiB
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154 lines
26 KiB
XML
<document id="7EEA365ED55DC2DD15A8C64CE664E7E6" ID-CLB-Dataset="6754" ID-DOI="10.5281/zenodo.4648451" ID-GBIF-Dataset="70db1eca-74c8-4901-823a-79cfd01d993f" ID-ISSN="1536-9307" ID-Zenodo-Dep="4648451" ID-ZooBank="BF89F25E-D5F8-4519-A0E4-6DF2D12CCFE8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617135433681" checkinUser="carolina" docAuthor="Kovařík, František & Lowe, Graeme" docDate="2020" docId="E46A87C8FFE5FFDDFC7D20A5FDEAF833" docLanguage="en" docName="Euscorpius.309.1-19.pdf" docOrigin="Euscorpius 309" docStyle="DocumentStyle:5E96F6379CE20B0E8E4F9E7DF8E1C0DF.2:Euscorpius.2020-.journal_article" docStyleId="5E96F6379CE20B0E8E4F9E7DF8E1C0DF" docStyleName="Euscorpius.2020-.journal_article" docStyleVersion="2" docTitle="Leiurus Ehrenberg 1828" docType="treatment" docVersion="11" lastPageNumber="1" masterDocId="1853FFB0FFE7FFD9FFC12425FFDAFFC4" masterDocTitle="Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XXIV. Leiurus (Buthidae), with description of Leiurus gubanensis sp. n." masterLastPageNumber="19" masterPageNumber="1" pageNumber="1" updateTime="1698935465433" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="57E90F15807133DB37CF5116A4AD655F">Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XXIV. Leiurus (Buthidae), with description of Leiurus gubanensis sp. n.</mods:title>
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<mods:namePart id="52E1D0D1145697A143172824D055660A">Kovařík, František</mods:namePart>
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<mods:namePart id="EC325EF44CF34CC3B9DB9B03AB030C86">Lowe, Graeme</mods:namePart>
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<mods:title id="1966F1798D726F2FD14F92FD6BD892C1">Euscorpius</mods:title>
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<mods:date id="FAF19F06680361726DE135A3EC09A57E">2020</mods:date>
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<treatment id="E46A87C8FFE5FFDDFC7D20A5FDEAF833" ID-DOI="http://doi.org/10.5281/zenodo.4770595" ID-GBIF-Taxon="182216078" ID-Zenodo-Dep="4770595" LSID="urn:lsid:plazi:treatment:E46A87C8FFE5FFDDFC7D20A5FDEAF833" httpUri="http://treatment.plazi.org/id/E46A87C8FFE5FFDDFC7D20A5FDEAF833" lastPageId="4" lastPageNumber="1" pageId="2" pageNumber="1">
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<subSubSection id="24D96555FFE5FFDBFC7D20A5FAEDFB5E" box="[956,1335,1152,1178]" pageId="2" pageNumber="1" type="nomenclature">
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<paragraph id="6C7C36DEFFE5FFDBFC7D20A5FAEDFB5E" blockId="2.[932,1359,1117,1210]" box="[956,1335,1152,1178]" pageId="2" pageNumber="1">
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<heading id="373481B2FFE5FFDBFC7D20A5FAEDFB5E" box="[956,1335,1152,1178]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="1" reason="2">
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Genus
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<taxonomicName id="ABC34D5DFFE5FFDBFBCF20A5FAEDFB5E" ID-CoL="5C2F" authority="Ehrenberg, 1828" authorityName="Ehrenberg" authorityYear="1828" box="[1038,1335,1152,1178]" class="Arachnida" family="Buthidae" genus="Leiurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis id="5EB7EACCFFE5FFDBFBCF20A5FBB7FB5E" bold="true" box="[1038,1133,1152,1178]" italics="true" pageId="2" pageNumber="1">Leiurus</emphasis>
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Ehrenberg, 1828
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="24D96555FFE5FFDBFC3D2087FC40FAFE" pageId="2" pageNumber="1" type="description">
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<paragraph id="6C7C36DEFFE5FFDBFC3D2087FB2CFB7E" blockId="2.[932,1359,1117,1210]" box="[1020,1270,1186,1210]" pageId="2" pageNumber="1">
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(
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<figureCitation id="F4F82A5BFFE5FFDBFBC52087FB49FB7E" box="[1028,1171,1186,1210]" captionStart-0="Figures 1–4" captionStart-1="Figures 5–10" captionStart-10="Figures 86–87" captionStart-11="Figures 88–89" captionStart-12="Figures 90–91" captionStart-2="Figures 11–20" captionStart-3="Figures 21–32" captionStart-4="Figures 33–56" captionStart-5="Figures 57–69" captionStart-6="Figures 70–79" captionStart-7="Figures 80–81" captionStart-8="Figures 82–83" captionStart-9="Figures 84–85" captionStartId-0="3.[116,193,1981,2003]" captionStartId-1="5.[115,192,1981,2003]" captionStartId-10="16.[116,193,2015,2037]" captionStartId-11="17.[118,195,1986,2008]" captionStartId-12="18.[113,190,847,869]" captionStartId-2="6.[113,190,1956,1978]" captionStartId-3="7.[116,193,1964,1986]" captionStartId-4="8.[115,192,1897,1919]" captionStartId-5="9.[113,190,1931,1953]" captionStartId-6="11.[115,190,1958,1980]" captionStartId-7="12.[116,191,1989,2011]" captionStartId-8="13.[114,189,1989,2011]" captionStartId-9="15.[113,190,2013,2035]" captionTargetBox-0="[126,1465,261,1954]" captionTargetBox-1="[116,1465,244,1956]" captionTargetBox-10="[115,1472,232,1997]" captionTargetBox-11="[113,1474,232,1969]" captionTargetBox-12="[131,1466,247,795]" captionTargetBox-2="[119,1470,258,1931]" captionTargetBox-3="[113,1474,234,1945]" captionTargetBox-4="[115,1468,236,1874]" captionTargetBox-5="[122,1471,244,1886]" captionTargetBox-6="[115,1476,236,1944]" captionTargetBox-7="[113,1474,234,1982]" captionTargetBox-8="[113,1474,234,1851]" captionTargetBox-9="[114,1475,234,1995]" captionTargetId-0="figure-50@3.[113,1474,234,1958]" captionTargetId-1="figure-63@5.[116,1477,234,1960]" captionTargetId-10="figure-37@16.[115,1474,229,1997]" captionTargetId-11="figure-45@17.[113,1474,229,1969]" captionTargetId-12="figure-131@18.[113,1474,234,812]" captionTargetId-2="figure-94@6.[113,1474,234,1936]" captionTargetId-3="figure-77@7.[113,1474,234,1957]" captionTargetId-4="figure-128@8.[115,1476,236,1875]" captionTargetId-5="figure-90@9.[113,1474,234,1895]" captionTargetId-6="figure-88@11.[115,1476,233,1948]" captionTargetId-7="figure-15@12.[113,1474,234,1982]" captionTargetId-8="figure-9@13.[113,1474,234,1851]" captionTargetId-9="figure-30@15.[114,1475,234,1998]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-10="16" captionTargetPageId-11="17" captionTargetPageId-12="18" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionTargetPageId-6="11" captionTargetPageId-7="12" captionTargetPageId-8="13" captionTargetPageId-9="15" captionText-0="Figures 1–4: Leiurus gubanensis sp. n., 18SF.Figures 1–2. Male holotype in dorsal (1) andventral (2) views.Figures 3–4. Female paratopotype in dorsal (3) and ventral (4) views. Scale bar: 10 mm." captionText-1="Figures 5–10: Leiurus gubanensis sp. n., male subadult paratype, 11SG. Figures 5–6. Dorsal (5) and ventral (6) views. Figures 7–10. Telson and metasoma V lateral (7), and metasoma and telson lateral (8), dorsal (9), and ventral (10) views. Scale bars: 10 mm (5–6, 8–10)." captionText-10="Figures 86–87: Leiurus gubanensis sp. n., 11SG, male subadult paratype (86) in vivo habitus and its locality (87)." captionText-11="Figures 88–89: Figure 88. Distribution of Leiurus gubanensis sp. n., L. quinquestriatus and L. somalicus in the region. Figure 89. Type locality (18SF) of L. gubanensis sp. n." captionText-12="Figures 90–91: Scaling of relative size of median ocular tubercle with relative size of median eyes in the genus Leiurus. Ordinates of scatter plots are the ratio median ocular tubercle width/ carapace L, and abscissas are the ratio median eye diameter/ carapace L, in males (90) and females (91). Plotted symbols represent measurements from a single sample of each species, extracted or estimated from published images and data, or from our own material. Lines are least squares regression fits, and the corresponding correlation coeeficients R, and P-values are indicated. In both sexes, L. gubanensis sp. n. is well separated from other species in having the largest eyes in the genus." captionText-2="Figures 11–20: Leiurus gubanensis sp. n., 18SF. Figures 12–13, 15–17. Male holotype, metasoma V and telson ventral (12) and lateral (13), and metasoma and telson lateral (15), dorsal (16), and ventral (17) views. Figures 11, 14, 18–20. Female paratopotype, metasoma V and telson ventral (11) and lateral (14), and metasoma and telson lateral (18), dorsal (19), and ventral (20) views. Scale bars: 10 mm (15–17, 18–20)." captionText-3="Figures 21–32: Leiurus gubanensis sp. n., 18SF. Figures 21, 23, 25–28. Male holotype, carapace and tergites I–III (21), coxosternal area and sternites (23), left legs I–IV, retrolateral aspect (25–28 respectively). Figures 22, 24, 29–32. Female paratopotype, carapace and tergites I–III (22), coxosternal area and sternites (24), left legs I–IV, retrolateral aspect (29–32 respectively)." captionText-4="Figures 33–56: Leiurus gubanensis sp. n., 11SG. Figures 33–45, 54. Male subadult paratype, carapace and tergites I–IV (33), coxosternal area and sternites (34), left legs I–IV, retrolateral aspect (35–38 respectively), chela dorsal (39), external (40) and ventral (41) views, patella dorsal (42), external (43) and ventral (44) views, trochanter and femur dorsal (45), and movable finger dentition (54). Figures 46–53, 55–56. Female adult paratype, chela dorsal (46), external (47) and ventral (48) views, patella dorsal (49), external (50) and ventral (51) views, trochanter and femur dorsal (52) and ventral (53) views, and movable (55) and fixed (56) finger dentition." captionText-5="Figures 57–69: Leiurus gubanensis sp. n., 18SF. Figures 57–58. Female paratopotype, chela dorsal (57) and external (58) views. Figures 59– 69. Male holotype, chela dorsal (59), external (60) and ventral (61) views, patella dorsal (62), external (63) and ventral (64) views, trochanter and femur internal (65), dorsaloexternal (66) and ventral (67) views, and right chelicera in dorsal (68) and ventral (69) views. Trichobothrial pattern is indicated by white circles (59–63, 65–66)." captionText-6="Figures 70–79: Leiurus gubanensis sp. n., comparison of key characters of male subadult paratype from 11SG (70, 72, 74, 76, 78) and female paratopotype from 18SF (71, 73, 75, 77, 79). Figures 70–73. Tergites I–III under white light (70–71) and UV fluorescence (72–73). Figures 74– 75. Sternite III under UV fluorescence.Figures 76–79. Metasoma V and telson ventral under UV fluorescence (76–77) and white light (78–79)." captionText-7="Figures 80–81: Leiurus gubanensis sp. n., 18SF, male holotype, carapace and tergites (80) and coxosternal area and sternites (81) under UV fluorescence." captionText-8="Figures 82–83: Leiurus gubanensis sp. n., 18SF, female paratopotype, carapace and tergites (82) and coxosternal area and sternites (83) under UV fluorescence." captionText-9="Figures 84–85: Leiurus gubanensis sp. n., 18SF, male holotype (84) and female paratopotype (85) in vivo habitus." figureDoi-0="http://doi.org/10.5281/zenodo.4648453" figureDoi-1="http://doi.org/10.5281/zenodo.4648455" figureDoi-10="http://doi.org/10.5281/zenodo.4648474" figureDoi-11="http://doi.org/10.5281/zenodo.4648476" figureDoi-12="http://doi.org/10.5281/zenodo.4648478" figureDoi-2="http://doi.org/10.5281/zenodo.4648457" figureDoi-3="http://doi.org/10.5281/zenodo.4648459" figureDoi-4="http://doi.org/10.5281/zenodo.4648461" figureDoi-5="http://doi.org/10.5281/zenodo.4648464" figureDoi-6="http://doi.org/10.5281/zenodo.4648466" figureDoi-7="http://doi.org/10.5281/zenodo.4648468" figureDoi-8="http://doi.org/10.5281/zenodo.4648470" figureDoi-9="http://doi.org/10.5281/zenodo.4648472" httpUri-0="https://zenodo.org/record/4648453/files/figure.png" httpUri-1="https://zenodo.org/record/4648455/files/figure.png" httpUri-10="https://zenodo.org/record/4648474/files/figure.png" httpUri-11="https://zenodo.org/record/4648476/files/figure.png" httpUri-12="https://zenodo.org/record/4648478/files/figure.png" httpUri-2="https://zenodo.org/record/4648457/files/figure.png" httpUri-3="https://zenodo.org/record/4648459/files/figure.png" httpUri-4="https://zenodo.org/record/4648461/files/figure.png" httpUri-5="https://zenodo.org/record/4648464/files/figure.png" httpUri-6="https://zenodo.org/record/4648466/files/figure.png" httpUri-7="https://zenodo.org/record/4648468/files/figure.png" httpUri-8="https://zenodo.org/record/4648470/files/figure.png" httpUri-9="https://zenodo.org/record/4648472/files/figure.png" pageId="2" pageNumber="1">Figures 1–91</figureCitation>
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,
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<tableCitation id="21410365FFE5FFDBFB5F2087FB36FB7E" box="[1182,1260,1186,1210]" captionStart="Table 1" captionStartId="4.[113,169,1065,1086]" captionTargetId="graphics-533@4.[118,1473,228,1057]" captionTargetPageId="4" captionText="Table 1. Comparative measurements of adults of Leiurus gubanensis sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." httpUri="http://table.plazi.org/id/38BC6656FFE3FFDDFFB0200CFEA0FB98" pageId="2" pageNumber="1" tableUuid="38BC6656FFE3FFDDFFB0200CFEA0FB98">Table 1</tableCitation>
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)
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</paragraph>
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<paragraph id="6C7C36DEFFE5FFDBFCF120C7FC40FAFE" blockId="2.[816,1474,1250,1434]" pageId="2" pageNumber="1">
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<taxonomicName id="ABC34D5DFFE5FFDBFCF120C7FB40FB3E" authority="Ehrenberg" authorityName="Ehrenberg" box="[816,1178,1250,1274]" class="Arachnida" family="Buthidae" genus="Androctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="subGenus" subGenus="Leiurus">
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<emphasis id="5EB7EACCFFE5FFDBFCF120C7FC6DFB3E" box="[816,951,1250,1274]" italics="true" pageId="2" pageNumber="1">Androctonus</emphasis>
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(
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<emphasis id="5EB7EACCFFE5FFDBFC0920C6FBC3FB3E" box="[968,1049,1251,1274]" italics="true" pageId="2" pageNumber="1">Leiurus</emphasis>
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) Ehrenberg
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</taxonomicName>
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in Hemprich & Ehrenberg, 1828, pl. I, fig. 5; Hemprich & Ehrenberg, 1829: 353 (in part).
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</paragraph>
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</subSubSection>
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<subSubSection id="24D96555FFE5FFDBFCF12166FBB6FA5E" pageId="2" pageNumber="1" type="reference_group">
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<paragraph id="6C7C36DEFFE5FFDBFCF12166FBB6FA5E" blockId="2.[816,1474,1250,1434]" pageId="2" pageNumber="1">
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<treatmentCitationGroup id="4CD311F0FFE5FFDBFCF12166FBB6FA5E" pageId="2" pageNumber="1">
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<taxonomicName id="ABC34D5DFFE5FFDBFCF12166FBA9FA9E" ID-CoL="5C2F" authority=": Vachon, 1949: 83 - 88" authorityName=": Vachon" authorityPageNumber="83 - 88" authorityYear="1949" box="[816,1139,1346,1370]" class="Arachnida" family="Buthidae" genus="Leiurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis id="5EB7EACCFFE5FFDBFCF12166FC5BFA9E" box="[816,897,1347,1370]" italics="true" pageId="2" pageNumber="1">Leiurus</emphasis>
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: Vachon, 1949: 83–88
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</taxonomicName>
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(1952: 203–208); Fet & Lowe, 2000: 155–157;
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<treatmentCitation id="ED6210CFFFE5FFDBFBD92146FAE2FABE" ID-DOI="http://doi.org/10.5281/zenodo.7117227" ID-Zenodo-Dep="7117227" author="LOWE" box="[1048,1336,1378,1402]" httpUri="http://treatment.plazi.org/id/038C87BF20561948FC778E96DE5DFC2E" page="1" pageId="2" pageNumber="1" year="2014">
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<bibRefCitation id="08524B2FFFE5FFDBFBD92146FAE2FABE" author="LOWE" box="[1048,1336,1378,1402]" pageId="2" pageNumber="1" pagination="1 - 129" refId="ref7557" refString="LOWE, G., E. A. YAGMUR & F. KOVARIK. 2014. A revision of the genus Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) with description of four new species from the Arabian Peninsula. Euscorpius, 191: 1 - 129." type="journal article" year="2014">Lowe et al., 2014: 1–129</bibRefCitation>
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</treatmentCitation>
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||
, figs. 1–100 (complete reference list).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="24D96555FFE5FFDBFCF121E6FC73FA3E" pageId="2" pageNumber="1" type="materials_examined">
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<paragraph id="6C7C36DEFFE5FFDBFCF121E6FC73FA3E" blockId="2.[816,1474,1474,1530]" pageId="2" pageNumber="1">
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<materialsCitation id="DCAB3C83FFE5FFDBFCF121E6FC7CFA3E" ID-GBIF-Occurrence="3113736304" collectionCode="SPECIES" pageId="2" pageNumber="1" specimenCount="1" typeStatus="holotype">
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<typeStatus id="B378887CFFE5FFDBFCF121E6FCBCFA1E" box="[816,870,1475,1498]" pageId="2" pageNumber="1">TYPE</typeStatus>
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<collectionCode id="0AD2AE1BFFE5FFDBFCB121E7FC1BFA1E" box="[880,961,1474,1498]" pageId="2" pageNumber="1">SPECIES</collectionCode>
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.
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<taxonomicName id="ABC34D5DFFE5FFDBFC0C21E7FC7CFA3E" authority="Ehrenberg, 1828" authorityName="Ehrenberg" authorityYear="1828" class="Arachnida" family="Buthidae" genus="Androctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="quinquestriatus" subGenus="Leiurus">
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<emphasis id="5EB7EACCFFE5FFDBFC0C21E7FB8EFA1E" box="[973,1108,1474,1498]" italics="true" pageId="2" pageNumber="1">Androctonus</emphasis>
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(
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<emphasis id="5EB7EACCFFE5FFDBFBA721E6FB6DFA1E" box="[1126,1207,1475,1498]" italics="true" pageId="2" pageNumber="1">Leiurus</emphasis>
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||
)
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<emphasis id="5EB7EACCFFE5FFDBFB0B21E6FAAAFA1E" box="[1226,1392,1475,1498]" italics="true" pageId="2" pageNumber="1">quinquestriatus</emphasis>
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Ehrenberg, 1828
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</taxonomicName>
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</materialsCitation>
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.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="24D96555FFE5FFDDFCF12207FDEAF833" lastPageId="4" lastPageNumber="3" pageId="2" pageNumber="1" type="diagnosis">
|
||
<paragraph id="6C7C36DEFFE5FFDDFCF12207FDEAF833" blockId="2.[816,1475,1570,2043]" lastBlockId="4.[113,772,1151,2039]" lastPageId="4" lastPageNumber="3" pageId="2" pageNumber="1">
|
||
DIAGNOSIS. Medium to large sized buthid scorpions, total length
|
||
<quantity id="AB3B9B3BFFE5FFDBFCF12267FC6AF99E" box="[816,944,1602,1626]" metricMagnitude="-2" metricUnit="m" metricValue="8.25" metricValueMax="11.5" metricValueMin="5.0" pageId="2" pageNumber="1" unit="mm" value="82.5" valueMax="115.0" valueMin="50.0">50–115 mm</quantity>
|
||
. The carapace is almost flat, weakly or moderately trapezoidal, with anterior margin straight or slightly concave, and 5 lateral eyes in ‘
|
||
<typeStatus id="B378887CFFE5FFDBFBD122A7FB98F95E" box="[1040,1090,1666,1690]" pageId="2" pageNumber="1">type</typeStatus>
|
||
5’ pattern (
|
||
<bibRefCitation id="08524B2FFFE5FFDBFB7A22A7FA6DF95E" author="LORIA & PRENDINI" box="[1211,1463,1666,1690]" pageId="2" pageNumber="1" pagination="112913" refId="ref7308" refString="LORIA, S. F. & L. PRENDINI. 2014. Homology of the lateral eyes of Scorpiones: a six-ocellus model. PLoS ONE 9 (12): e 112913. doi: 10.1371 / journal. pone. 0112913." type="journal article" year="2014">Loria & Prendini, 2014</bibRefCitation>
|
||
). Both carapace and tergites bear granulated carinae. The carapace bears distinct anterior, superciliary, central median, central lateral, posterior median and posterior lateral carinae. The central lateral and posterior median carinae are fused into a lyre configuration. Tergites I–II, VII bear 5 carinae, and III– VI bear 3 carinae. Tergites lack macrosetae. The metasoma is elongate, metasoma I–III bear 10 carinae, with median lateral carinae complete on I, and reduced on II–III. Metasoma IV bears 8 carinae, and metasoma V bears 7 carinae of which the dorsolateral carinae are weak, and ventrolateral carinae are strong with serrate or lobate dentition. The telson has a relatively bulbous vesicle lacking a subaculear spine or tubercle. Pectines bear fulcra. Pectinal tooth count ranges are
|
||
<specimenCount id="7AC5FD57FFE3FFDDFDF62085FD91FB73" box="[567,587,1184,1207]" pageId="4" pageNumber="3" type="male">♂</specimenCount>
|
||
28–43,
|
||
<specimenCount id="7AC5FD57FFE3FFDDFD612085FD6EFB73" box="[672,692,1184,1207]" pageId="4" pageNumber="3" type="female">♀</specimenCount>
|
||
25–38. The hemispermatophore is flagelliform, the capsule having 3 sperm hemiduct lobes well separated from the flagellum, and a strong hook-like basal lobe. The chelicerae exhibit characteristic buthid dentition (
|
||
<bibRefCitation id="08524B2FFFE3FFDDFEF0213AFE11FAF3" author="VACHON" box="[305,459,1311,1335]" pageId="4" pageNumber="3" pagination="161 - 166" refId="ref7760" refString="VACHON, M. 1963. De l'utilite, en systematique d'une nomenclature des dents des cheliceres chez les scorpions. Bulletin du Museum National d'Histoire Naturelle Paris, 35 (2): 161 - 166." type="journal article" year="1963">Vachon, 1963</bibRefCitation>
|
||
), with two denticles on the ventral aspect of the fixed finger. The pedipalp chela is slender with long fingers, the surface smooth with carinae reduced or obsolete, and the dentate margins of the fingers are armed with linear, non-imbricated subrows of primary denticles, flanked by internal and external accessory denticles. The movable finger bears two enlarged subdistal internal denticles. The pedipalps are orthobothriotaxic, pattern
|
||
<typeStatus id="B378887CFFE3FFDDFE742225FE38F9DC" box="[437,482,1536,1560]" pageId="4" pageNumber="3">type</typeStatus>
|
||
Ab (
|
||
<bibRefCitation id="08524B2FFFE3FFDDFDDE21DAFD6BF9D3" author="VACHON" box="[543,689,1535,1559]" pageId="4" pageNumber="3" pagination="857 - 958" refId="ref7798" refString="VACHON, M. 1974. Etudes des caracteres utilises pour classer les familles et les genres des scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, 3 e serie, 140 (Zoologie, 104): 857 - 958." type="journal article" year="1974">Vachon 1974</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="08524B2FFFE3FFDDFD7E21DAFD2DF9D3" author="VACHON" box="[703,759,1535,1559]" pageId="4" pageNumber="3" pagination="1597 - 1599" refId="ref7860" refString="VACHON, M. 1975. Sur l'utilisation de la trichobothriotaxie du bras des pedipalpes des scorpions (Arachnides) dans le classement des genres de la famille des Buthidae Simon. Comptes Rendus de l'Academie des Sciences, Paris, D, 281: 1597 - 1599." type="journal article" year="1975">1975</bibRefCitation>
|
||
), the femur with trichobothrium
|
||
<emphasis id="5EB7EACCFFE3FFDDFE79223AFE1FF9F3" box="[440,453,1567,1591]" italics="true" pageId="4" pageNumber="3">d</emphasis>
|
||
<subScript id="F047349BFFE3FFDDFE04220AFE17F9F9" attach="left" box="[453,461,1583,1597]" fontSize="6" pageId="4" pageNumber="3">2</subScript>
|
||
on its dorsal surface, and the patella with
|
||
<emphasis id="5EB7EACCFFE3FFDDFF38221AFEDCF993" box="[249,262,1599,1623]" italics="true" pageId="4" pageNumber="3">d</emphasis>
|
||
<subScript id="F047349BFFE3FFDDFEC7226AFED4F999" attach="left" box="[262,270,1615,1629]" fontSize="6" pageId="4" pageNumber="3">3</subScript>
|
||
internal to the dorsomedian carina. The chela manus has
|
||
<emphasis id="5EB7EACCFFE3FFDDFF2B227AFEDDF9B3" box="[234,263,1631,1655]" italics="true" pageId="4" pageNumber="3">Eb</emphasis>
|
||
<subScript id="F047349BFFE3FFDDFEC9224AFECAF9B9" attach="right" box="[264,272,1647,1661]" fontSize="6" pageId="4" pageNumber="3">1</subScript>
|
||
-
|
||
<emphasis id="5EB7EACCFFE3FFDDFED9227AFEEFF9B3" box="[280,309,1631,1655]" italics="true" pageId="4" pageNumber="3">Eb</emphasis>
|
||
<subScript id="F047349BFFE3FFDDFEF4224AFEE7F9B9" attach="left" box="[309,317,1647,1661]" fontSize="6" pageId="4" pageNumber="3">2</subScript>
|
||
angled proximally, and
|
||
<emphasis id="5EB7EACCFFE3FFDDFD82227AFDBAF9B3" box="[579,608,1631,1655]" italics="true" pageId="4" pageNumber="3">Eb</emphasis>
|
||
<subScript id="F047349BFFE3FFDDFDA1224AFDB2F9B9" attach="left" box="[608,616,1647,1661]" fontSize="6" pageId="4" pageNumber="3">1</subScript>
|
||
-
|
||
<emphasis id="5EB7EACCFFE3FFDDFDB0227AFD54F9B3" box="[625,654,1631,1655]" italics="true" pageId="4" pageNumber="3">Eb</emphasis>
|
||
<subScript id="F047349BFFE3FFDDFD4F224AFD4CF9B9" attach="left" box="[654,662,1647,1661]" fontSize="6" pageId="4" pageNumber="3">2</subScript>
|
||
-
|
||
<emphasis id="5EB7EACCFFE3FFDDFD5F227AFD61F9B3" box="[670,699,1631,1655]" italics="true" pageId="4" pageNumber="3">Eb</emphasis>
|
||
<subScript id="F047349BFFE3FFDDFD7A224AFD19F9B9" attach="left" box="[699,707,1647,1661]" fontSize="6" pageId="4" pageNumber="3">3</subScript>
|
||
acute angle opening in the distal direction (
|
||
<emphasis id="5EB7EACCFFE3FFDDFDCC225AFDC3F953" bold="true" box="[525,537,1663,1687]" italics="true" pageId="4" pageNumber="3">δ</emphasis>
|
||
-configuration). Tibial spurs are present on legs III–IV, and basitarsi I–III bear regular series of macrosetae on retrosuperior, retroinferior and inferior margins. On the ventral surfaces of telotarsi are paired rows of macrosetae, and prolateral and retrolateral tarsal spurs are present on all legs.
|
||
<emphasis id="5EB7EACCFFE3FFDDFE82233AFDD5F8F3" box="[323,527,1823,1847]" italics="true" pageId="4" pageNumber="3">Sexual dimorphism</emphasis>
|
||
: compared to females, males have a narrower mesosoma, more robust carination on tergites and sternites III–V, more slender pedipalps and metasoma, longer pectines with larger teeth, and weaker dentition or granulation on ventromedian carinae of metasoma II–III. Males are similar to females in lacking undulate dentate margins at the base of the pedipalp fingers.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |