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<document id="ECDB0054F18DCF8AF8A536B227286D7F" ID-DOI="10.5281/zenodo.191352" ID-GBIF-Dataset="3f63d847-9e21-40f9-a5fd-14fc0809a9b4" ID-ISSN="1175-5326" ID-Zenodo-Dep="191352" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461127920264" checkinUser="plazi" docAuthor="Pesic, Vladimir, Gerecke, Reinhard &amp; Smit, Harry" docDate="2009" docId="E45C367308357252B5B5F2AE0622ED77" docLanguage="en" docName="zt02290p064.pdf" docOrigin="Zootaxa 2290" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Iranothyas alhajarica Pesic, Gerecke &amp; Smit, 2009, spec. nov." docType="treatment" docVersion="9" lastPageNumber="64" masterDocId="18654E0B08367257B522F3630578EE6B" masterDocTitle="A redefinition of Iranothyas Bader, 1984 with the description of a new species from Oman" masterLastPageNumber="64" masterPageNumber="59" pageNumber="62" updateTime="1698597096250" updateUser="plazi">
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<mods:title id="FDD25EE295D294CB2D42E7B51FA0D96C">A redefinition of Iranothyas Bader, 1984 with the description of a new species from Oman</mods:title>
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<mods:namePart id="4BB10169113267B44FEC3BFD36A2C8CF">Pesic, Vladimir</mods:namePart>
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<mods:namePart id="5C3C27A0D5608D03401CB34B01B307AE">Gerecke, Reinhard</mods:namePart>
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<mods:namePart id="EEDE28723198ACBE5FDB7D3E98C665E2">Smit, Harry</mods:namePart>
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<mods:date id="2E0D80BE41E385C6464C4397D2F6A85A">2009</mods:date>
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<treatment id="E45C367308357252B5B5F2AE0622ED77" ID-DOI="http://doi.org/10.5281/zenodo.6216279" ID-GBIF-Taxon="119626606" ID-Zenodo-Dep="6216279" LSID="urn:lsid:plazi:treatment:E45C367308357252B5B5F2AE0622ED77" httpUri="http://treatment.plazi.org/id/E45C367308357252B5B5F2AE0622ED77" lastPageId="5" lastPageNumber="64" pageId="3" pageNumber="62">
<subSubSection id="24EFD4EE08357254B5B5F2AE045CEC61" pageId="3" pageNumber="62" type="nomenclature">
<paragraph id="6C4A876508357254B5B5F2AE0765EF8C" blockId="3.[151,541,461,522]" box="[151,541,461,487]" pageId="3" pageNumber="62">
<heading id="3702300908357254B5B5F2AE0765EF8C" bold="true" box="[151,541,461,487]" fontSize="11" level="1" pageId="3" pageNumber="62" reason="1">
<emphasis id="5E815B7708357254B5B5F2AE0765EF8C" bold="true" box="[151,541,461,487]" pageId="3" pageNumber="62">
<taxonomicName id="ABF5FCE608357254B5B5F2AE04E6EF8C" box="[151,414,461,487]" class="Arachnida" family="Hydryphantidae" genus="Iranothyas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="alhajarica" status="sp. nov.">
<emphasis id="5E815B7708357254B5B5F2AE04E6EF8C" bold="true" box="[151,414,461,487]" italics="true" pageId="3" pageNumber="62">Iranothyas alhajarica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45B2E60C08357254B487F2AE0765EF8C" box="[421,541,461,487]" pageId="3" pageNumber="62" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6C4A876508357254B5B5F293045CEC61" blockId="3.[151,541,461,522]" box="[151,292,496,522]" pageId="3" pageNumber="62">
(
<figureCitation id="F4CE9BE008357254B5BDF2930464EC61" box="[159,284,496,522]" captionStart="FIGURE 2 A E" captionStartId="4.[151,255,1890,1914]" captionTargetBox="[215,1379,270,1863]" captionTargetId="figure@4.[207,1379,258,1864]" captionTargetPageId="4" captionText="FIGURE 2 A E: Iranothyas alhajarica sp. nov.: A = female frontal shield; B = male palp; C = female palp (P- 1 lacking); D = male genital field; E = male IV-L. Scale bars = 100 µm (Figs. A, C E), 50 µm (Figs. B)." httpUri="https://zenodo.org/record/191354/files/figure.png" pageId="3" pageNumber="62">Fig. 2AE</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="24EFD4EE08357254B5B5F15E073CECA0" pageId="3" pageNumber="62" type="materials_examined">
<paragraph id="6C4A876508357254B5B5F15E073CECA0" blockId="3.[151,1437,572,2030]" pageId="3" pageNumber="62">
<emphasis id="5E815B7708357254B5B5F15E045BEC3C" bold="true" box="[151,291,573,599]" pageId="3" pageNumber="62">
<typeStatus id="B34E39C708357254B5B5F15E05ADEC3C" box="[151,213,573,599]" pageId="3" pageNumber="62">Type</typeStatus>
series
</emphasis>
.
<typeStatus id="B34E39C708357254B411F15E04D9EC3C" box="[307,417,573,599]" pageId="3" pageNumber="62" type="holotype">Holotype</typeStatus>
3, Zoological Museum of the University of Amsterdam,
<collectingCountry id="14E2C7F508357254B170F15E01E5EC3C" box="[1106,1181,573,599]" name="Oman" pageId="3" pageNumber="62">Oman</collectingCountry>
, Al Hajar mountains, spring Al Khremh, crossing road to
<collectingCountry id="14E2C7F508357254B760F107063DEC15" box="[578,837,612,638]" name="United Arab Emirates" pageId="3" pageNumber="62">United Arab Emirates</collectingCountry>
border,
<geoCoordinate id="09C1E1A208357254B68AF1070143EC15" box="[936,1083,612,638]" direction="north" orientation="latitude" pageId="3" pageNumber="62" precision="1" value="24.79035">24 47.421 N</geoCoordinate>
<geoCoordinate id="09C1E1A208357254B166F10701AEEC15" box="[1092,1238,612,638]" direction="east" orientation="longitude" pageId="3" pageNumber="62" precision="1" value="55.941715">55 56.503 E</geoCoordinate>
,
<date id="184BA1A508357254B1C1F107001CEC15" box="[1251,1380,612,638]" pageId="3" pageNumber="62" value="2008-11-15">15.xi.2008</date>
, leg. Smit, slide-mounted.
<typeStatus id="B34E39C708357254B4B6F1E90772ECCF" box="[404,522,650,676]" pageId="3" pageNumber="62" type="paratype">Paratypes</typeStatus>
: 1 3, 2 Ƥ, 11 deutonymphs, same date as
<typeStatus id="B34E39C708357254B6DBF1E9011CECCF" box="[1017,1124,650,676]" pageId="3" pageNumber="62" type="holotype">holotype</typeStatus>
. 1 Ƥ, 1 deutonymph slidemounted, 1 Ƥ,
<quantity id="AB0D2A8008357254B465F1D204FAECA0" box="[327,386,689,715]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="62" unit="in" value="10.0">10 in</quantity>
Koenike's fluid.
</paragraph>
</subSubSection>
<subSubSection id="24EFD4EE08357254B5E4F1B40580ED0D" pageId="3" pageNumber="62" type="diagnosis">
<paragraph id="6C4A876508357254B5E4F1B40580ED0D" blockId="3.[151,1437,572,2030]" pageId="3" pageNumber="62">
<emphasis id="5E815B7708357254B5E4F1B40447EC9A" bold="true" box="[198,319,727,753]" pageId="3" pageNumber="62">Diagnosis</emphasis>
. Both sexes: Acetabula circular in shape, Ac-2 located halfway between Ac-1 and Ac-3; Ac-3 diameter 3040. Sexual dimorphism in shape of IV-L (males: stouter, terminal segment subrectangular in shape, claws robust; females: more slender, terminal segment distally widened, claws more slender and less curved).
</paragraph>
</subSubSection>
<subSubSection id="24EFD4EE08357252B5E4F0110493EF18" lastPageId="5" lastPageNumber="64" pageId="3" pageNumber="62" type="description">
<paragraph id="6C4A876508357254B5E4F01100E4EBBB" blockId="3.[151,1437,572,2030]" pageId="3" pageNumber="62">
<emphasis id="5E815B7708357254B5E4F011042FEDE7" bold="true" box="[198,343,882,908]" pageId="3" pageNumber="62">Description</emphasis>
. Both sexes: Papillae covering membranous integument flat in anterior part of idiosoma, elevated and directed posteriorly in posterior part. Sclerotized parts with fine round pores (dorsal plates) or larger pores surrounded by a ring of fine dots (coxae, appendages). Frontal shield with a rounded posteromedial extension; Dc-2 not fused to frontal shield; Dc-3 fused medially to a small roundish platelet, Dc-5 fused to a large transverse dorsocaudal plate, Dc-2, Dc-4 and Dl-2-4 rather large, roundish. Coxae with numerous long setae, a little stronger and particularly densely arranged near tips of Cx-I-IV, finer and scattered along medial margin of Cx-I and suture line Cx-I/II. Genital flaps with longish pores, anteriorly narrow, posteriorly enlarged and including Ac-3; a line of densely-arranged long and fine setae extending from the anterior edge to the area lateral to Ac-3, stronger and larger setae arranged in a group on a mediocaudal extension, anteromedially to Ac-3. Ac-1 and Ac-2 small and circular, in the membranous area flanking the gonopore. Excretory pore surrounded by a sclerite ring; except for a pair of small sclerite platelets lying posterior to excretory pore, ventrocaudal idiosoma membranous. Gnathosoma elevated, with welldeveloped rostrum. Chelicera slender, with relatively short claw (basal segment/claw ratio 2.2). Palp stout, all segments strongly thickened, P-4 shortened and extremely narrowed from the base to the tip; P-1 bearing one dorsal seta; P-2, 45 stronger setae, dorsally and medially and 13 finer setae laterally; P-3, 2 dorsodistal setae, one longer, medially, one shorter laterally; P-4, 3 fine distal setae, in lateral, dorsal and medial position.
</paragraph>
<paragraph id="6C4A876508357254B5E4F6BE04BFE938" blockId="3.[151,1437,572,2030]" pageId="3" pageNumber="62">
Male: Idiosoma L/W 720/550; frontal shield L/W 290/280; W fused Dc-5 205; genital flap L 200, gonopore area with a pair of tongue-shaped extensions facing each other and densely covered by heavy denticulation (
<figureCitation id="F4CE9BE008357254B466F54904DDE82F" box="[324,421,1578,1604]" captionStart="FIGURE 2 A E" captionStartId="4.[151,255,1890,1914]" captionTargetBox="[215,1379,270,1863]" captionTargetId="figure@4.[207,1379,258,1864]" captionTargetPageId="4" captionText="FIGURE 2 A E: Iranothyas alhajarica sp. nov.: A = female frontal shield; B = male palp; C = female palp (P- 1 lacking); D = male genital field; E = male IV-L. Scale bars = 100 µm (Figs. A, C E), 50 µm (Figs. B)." httpUri="https://zenodo.org/record/191354/files/figure.png" pageId="3" pageNumber="62">Fig. 2D</figureCitation>
); in the area between these extensions, tip of ejaculatory complex located between a pair of very finely denticulated membranous humps; anterior to this area a pair of weekly sclerotized, crossed pointed extensions; pregenital sclerite knob-shaped, postgenital sclerite rod-shaped; posteromedial extensions of genital flaps strongly protruding, with densely-arranged strong setae; diameter Ac-1-3, 18-15-30; gnathosoma ventral L 190; chelicera basal segment L/H 205/52, claw L 95; palp (
<figureCitation id="F4CE9BE008357254B173F5A601D6E8B4" box="[1105,1198,1733,1759]" captionStart="FIGURE 2 A E" captionStartId="4.[151,255,1890,1914]" captionTargetBox="[215,1379,270,1863]" captionTargetId="figure@4.[207,1379,258,1864]" captionTargetPageId="4" captionText="FIGURE 2 A E: Iranothyas alhajarica sp. nov.: A = female frontal shield; B = male palp; C = female palp (P- 1 lacking); D = male genital field; E = male IV-L. Scale bars = 100 µm (Figs. A, C E), 50 µm (Figs. B)." httpUri="https://zenodo.org/record/191354/files/figure.png" pageId="3" pageNumber="62">Fig. 2B</figureCitation>
): L/H P-1, 24/65; P- 2, 90/75; P-3, 40/67; P-4, 102/52; P-5, 38/12; IV-L very stout, terminal segments subrectangular in shape, with very robust claws; L/H (in brackets, ratio) IV-L-3, 108/70 (1.5), IV-L-4, 145/61 (2.4); IV-L-5, 114/54 (2.1); IV-L-6, 67/45 (1.5).
</paragraph>
<paragraph id="6C4A876508357254B5E7F40300E4E985" blockId="3.[151,1437,572,2030]" pageId="3" pageNumber="62">
Female: Idiosoma L/W 850/600; frontal shield L/W 350/350; W fused Dc-5, 280; genital flap L 170; diameter Ac-
<date id="184BA1A508357254B41AF4E504DFE9CB" box="[312,423,1926,1952]" pageId="3" pageNumber="62" value="1920-01-18" valueMax="1920-03-18" valueMin="1920-01-18">1-3 20-18</date>
-38; gnathosoma ventral L 270; chelicera basal segment L/H 250/62, claw L 115; palp (
<figureCitation id="F4CE9BE008357254B5BDF4CE0586E9AC" box="[159,254,1965,1991]" captionStart="FIGURE 2 A E" captionStartId="4.[151,255,1890,1914]" captionTargetBox="[215,1379,270,1863]" captionTargetId="figure@4.[207,1379,258,1864]" captionTargetPageId="4" captionText="FIGURE 2 A E: Iranothyas alhajarica sp. nov.: A = female frontal shield; B = male palp; C = female palp (P- 1 lacking); D = male genital field; E = male IV-L. Scale bars = 100 µm (Figs. A, C E), 50 µm (Figs. B)." httpUri="https://zenodo.org/record/191354/files/figure.png" pageId="3" pageNumber="62">Fig. 2C</figureCitation>
): L/H P-1, 25/65; P-2, 112/95; P-3, 42/67; P-4, 120/50; P-5, 42/15; IV-L much more slender than in males in setation and shape of segments and claws similar to those of male
<taxonomicName id="ABF5FCE608357254B121F4B701D4E986" box="[1027,1196,2004,2029]" class="Arachnida" family="Hydryphantidae" genus="Balaneothyas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="marismortui">
<emphasis id="5E815B7708357254B121F4B701D4E986" box="[1027,1196,2004,2029]" italics="true" pageId="3" pageNumber="62">I. marismortui</emphasis>
</taxonomicName>
(
<bibRefCitation id="0864FA9408357254B198F4B700E4E985" author="Gerecke" box="[1210,1436,2004,2030]" pageId="3" pageNumber="64" refString="Gerecke, R. (1999) Further studies on hydryphantoid water mites in the W Palaearctic region (Acari, Actinedida). Archiv fur Hydrobiologie, Supplement, 121, 119 - 158." type="journal article" year="1999">Gerecke 1999, Fig.</bibRefCitation>
</paragraph>
<paragraph id="6C4A876508327253B5B5F3FB05AFEEB3" blockId="4.[151,1436,152,216]" pageId="4" pageNumber="63">3F), L/H (in brackets, ratio) IV-L-3, 100/58 (1.7), IV-L-4, 166/54 (3.1); IV-L-5, 139/41 (3.4); IV-L-6, 90/41 (2.2).</paragraph>
<caption id="388AD7ED08327253B5B5F401010DE9F0" httpUri="https://zenodo.org/record/191354/files/figure.png" pageId="4" pageNumber="63" targetBox="[215,1379,270,1863]" targetPageId="4">
<paragraph id="6C4A876508327253B5B5F401010DE9F0" blockId="4.[151,1436,1890,1947]" pageId="4" pageNumber="63">
<emphasis id="5E815B7708327253B5B5F401043DE911" bold="true" box="[151,325,1890,1914]" pageId="4" pageNumber="63">FIGURE 2AE</emphasis>
:
<taxonomicName id="ABF5FCE608327253B470F4010742E912" box="[338,570,1890,1913]" class="Arachnida" family="Hydryphantidae" genus="Iranothyas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="alhajarica" status="sp. nov.">
<emphasis id="5E815B7708327253B470F4010742E912" box="[338,570,1890,1913]" italics="true" pageId="4" pageNumber="63">Iranothyas alhajarica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45B2E60C08327253B762F40107EEE912" box="[576,662,1890,1913]" pageId="4" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
: A = female frontal shield; B = male palp; C = female palp (P-1 lacking); D = male genital field; E = male IV-L. Scale bars = 100 µm (Figs. A, CE), 50 µm (Figs. B).
</paragraph>
</caption>
<paragraph id="6C4A876508337252B5E7F3FB0493EF18" blockId="5.[151,1437,152,796]" pageId="5" pageNumber="64">Deutonymph: As described for adults, but dorsalia very small and Dc-5 unfused. Idiosoma L/W 580/500; frontal shield L/W 130/161, posterior margin irregularly convex, without a medial extension; provisory genital organ with two pairs of round Ac (diameter Ac-1, 9, Ac-2, 7) and three paired setae, L/W 40/71; gnathosoma ventral L 212; chelicera basal segment L/H 220/29, claw L 50; L/H P-1, 12/35; P-2, 51/41; P-3, 25/32; P-4, 63/25; P-5, 17/8; L/H IV-L-3, 58/29, IV-L-4, 99/29; IV-L-5, 9/22; IV-L-6, 58/22; proportions of IV-L segments as in females.</paragraph>
</subSubSection>
<subSubSection id="24EFD4EE08337252B5E7F21C0471EFAB" pageId="5" pageNumber="64" type="etymology">
<paragraph id="6C4A876508337252B5E7F21C0471EFAB" blockId="5.[151,1437,152,796]" pageId="5" pageNumber="64">
<emphasis id="5E815B7708337252B5E7F21C0436EFF2" bold="true" box="[197,334,383,409]" pageId="5" pageNumber="64">Etymology</emphasis>
. Named after the Al-Hajar mountain range in northern
<collectingCountry id="14E2C7F508337252B6D5F2E3013CEFF1" box="[1015,1092,384,410]" name="Oman" pageId="5" pageNumber="64">Oman</collectingCountry>
, where the new species was collected.
</paragraph>
</subSubSection>
<subSubSection id="24EFD4EE08337252B5E4F2AE073AEC9D" pageId="5" pageNumber="64" type="discussion">
<paragraph id="6C4A876508337252B5E4F2AE073AEC9D" blockId="5.[151,1437,152,796]" pageId="5" pageNumber="64">
<emphasis id="5E815B7708337252B5E4F2AE0432EF8C" bold="true" box="[198,330,461,487]" pageId="5" pageNumber="64">Discussion</emphasis>
. Both sexes of
<taxonomicName id="ABF5FCE608337252B72FF2AE0669EF8D" box="[525,785,461,486]" class="Arachnida" family="Hydryphantidae" genus="Iranothyas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="alhajarica">
<emphasis id="5E815B7708337252B72FF2AE0669EF8D" box="[525,785,461,486]" italics="true" pageId="5" pageNumber="64">Iranothyas alhajarica</emphasis>
</taxonomicName>
differ from the two other species in the shape of the frontal shield (with undulating outline, having a posteromedial extension), and from
<taxonomicName id="ABF5FCE608337252B198F2970035EC66" box="[1210,1357,500,525]" class="Arachnida" family="Hydryphantidae" genus="Iranothyas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="circularis">
<emphasis id="5E815B7708337252B198F2970035EC66" box="[1210,1357,500,525]" italics="true" pageId="5" pageNumber="64">I. circularis</emphasis>
</taxonomicName>
in the position of Ac-2 (in agreement with
<taxonomicName id="ABF5FCE608337252B76BF179078BEC58" box="[585,755,538,563]" class="Arachnida" family="Hydryphantidae" genus="Balaneothyas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="marismortui">
<emphasis id="5E815B7708337252B76BF179078BEC58" box="[585,755,538,563]" italics="true" pageId="5" pageNumber="64">I. marismortui</emphasis>
</taxonomicName>
, halfway between Ac-1 and Ac-3, not approached to Ac- 3). Males of
<taxonomicName id="ABF5FCE608337252B40DF12204A1EC31" box="[303,473,577,602]" class="Arachnida" family="Hydryphantidae" genus="Balaneothyas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="marismortui">
<emphasis id="5E815B7708337252B40DF12204A1EC31" box="[303,473,577,602]" italics="true" pageId="5" pageNumber="64">I. marismortui</emphasis>
</taxonomicName>
differ from
<taxonomicName id="ABF5FCE608337252B749F1220787EC31" box="[619,767,577,602]" class="Arachnida" family="Hydryphantidae" genus="Iranothyas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="alhajarica">
<emphasis id="5E815B7708337252B749F1220787EC31" box="[619,767,577,602]" italics="true" pageId="5" pageNumber="64">I. alhajarica</emphasis>
</taxonomicName>
in a series of further characters that cannot be discussed for
<taxonomicName id="ABF5FCE608337252B5E6F10B0421ECEA" box="[196,345,616,641]" class="Arachnida" family="Hydryphantidae" genus="Iranothyas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="circularis">
<emphasis id="5E815B7708337252B5E6F10B0421ECEA" box="[196,345,616,641]" italics="true" pageId="5" pageNumber="64">I. circularis</emphasis>
</taxonomicName>
, this species being known in the female sex only: (1) Dc-2 fused to frontal shield; (2) posteroventral area bearing two pairs of small platelets; (3) IV-L with weakly developed sexual dimorphism and segments consequently much more slender (L/H ratio IV-L-3, 2.0; IV-L-4, 3.1; IV-L-5, 3.8; IV-L-6, 1.8), claws less strongly curved and finer.
</paragraph>
</subSubSection>
<subSubSection id="24EFD4EE08337252B5E4F0610622ED77" box="[198,858,770,796]" pageId="5" pageNumber="64" type="distribution">
<paragraph id="6C4A876508337252B5E4F0610622ED77" blockId="5.[151,1437,152,796]" box="[198,858,770,796]" pageId="5" pageNumber="64">
<emphasis id="5E815B7708337252B5E4F0610427ED77" bold="true" box="[198,351,770,796]" pageId="5" pageNumber="64">Distribution</emphasis>
.
<collectingCountry id="14E2C7F508337252B44CF06104C1ED77" box="[366,441,770,796]" name="Oman" pageId="5" pageNumber="64">Oman</collectingCountry>
, only known from the
<typeStatus id="B34E39C708337252B7E0F061078CED77" box="[706,756,770,796]" pageId="5" pageNumber="64">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>