308 lines
36 KiB
XML
308 lines
36 KiB
XML
<document id="CFB4A5F7F959E58FE3270347ADFB2EA0" ID="10.11646/zootaxa.4262.1.1SLASH10959" ID-CLB-Dataset="33105" ID-DOI="10.5281/zenodo.571683" ID-GBIF-Dataset="5ab1c651-825a-43bc-af00-11b77a792de3" ID-ISSN="1175-5326" ID-Zenodo-Dep="571683" ID-ZooBank="894795FB-ED38-4132-92B9-6AC8C3E41566" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1493972067874" checkinUser="plazi" docAuthor="Maya-Morales, Julieta & Jiménez, María Luisa" docDate="2017" docId="E45787C5FFA7FFDCE1A163CBFD4CFA0F" docLanguage="en" docName="zootaxa.4262.1.1SLASH10959.pdf" docOrigin="Zootaxa 4262 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Novalena clara Maya-Morales & Jiménez, 2017, new species" docType="treatment" docVersion="7" lastPageNumber="40" masterDocId="186EFFBDFF81FFFBE136627DFF86FFE9" masterDocTitle="Revision of the funnel-web spider genus Novalena (Araneae: Agelenidae)" masterLastPageNumber="88" masterPageNumber="1" pageNumber="39" updateTime="1698439297005" updateUser="ExternalLinkService">
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<mods:mods id="EBB5993ECBEA840AACED6C4B8AFD2A47" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B36179CD69D014287A887DF59249D7ED">
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<mods:title id="12B067BA7A6FEC6DAD39C5187CF5A07B">Revision of the funnel-web spider genus Novalena (Araneae: Agelenidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="88BB907C0FC0538760BECBAF42726154" type="personal">
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<mods:roleTerm id="C6DE75A09494BF6DA33D2F556468BD39">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="62424E99945FACC375B29C697B1C64D1">Maya-Morales, Julieta</mods:namePart>
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</mods:name>
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<mods:name id="FD4049FBB2AE975325C5407B7176D521" type="personal">
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<mods:role id="BF11E57CBC651E4D87238CA40661EC0D">
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<mods:roleTerm id="E97814C4EFDC0EC4E30DD7F6244C14C7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FBD42AB45FE8AED4F526606130E57AD1">Jiménez, María Luisa</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="CE095BB72A7B258CFAB97A5CBFEC2646">text</mods:typeOfResource>
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<mods:titleInfo id="533016D6EA3324B16597A57B307F4C5A">
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<mods:title id="AF9B15232B5C515D4A1B3D04E50F5283">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="7ABC54DCA5639D516915A5C3470C07AB">
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<mods:date id="6A0EC70520083F8F0C7742CC13A5436C">2017</mods:date>
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<mods:detail id="6FE325656C2806442306D89E9113F43C" type="volume">
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<mods:number id="AD3F0A256BFC57AE5FFAB3D04AD291DC">4262</mods:number>
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</mods:detail>
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<mods:detail id="3C1C9DCDAA0B559EC6A380CA8D0C0BDB" type="issue">
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<mods:number id="0163DFE5CE51C9D9526F938C24C6A61F">1</mods:number>
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<mods:end id="DF31B871F3009FA851D0162056BA72FE">88</mods:end>
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</mods:part>
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<mods:classification id="4EC0D302D89BA5923467A1A17AB42A37">journal article</mods:classification>
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<mods:identifier id="763BA7AF3E5A319895BF19829725AC91" type="CLB-Dataset">33105</mods:identifier>
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<mods:identifier id="9CAB3802ECF3FF4875F75C209BB0F652" type="DOI">10.5281/zenodo.571683</mods:identifier>
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<mods:identifier id="21836D74333B3E33DEFD34F1F2247DD3" type="GBIF-Dataset">5ab1c651-825a-43bc-af00-11b77a792de3</mods:identifier>
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<mods:identifier id="BD004785B92D9031A8FFD3284DD34D98" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="2038E9585DEA91F9F16028D5A7A43A80" type="Zenodo-Dep">571683</mods:identifier>
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<mods:identifier id="56840B9626C748E2CEFCF9EB8B0A247F" type="ZooBank">894795FB-ED38-4132-92B9-6AC8C3E41566</mods:identifier>
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<treatment id="E45787C5FFA7FFDCE1A163CBFD4CFA0F" ID-DOI="http://doi.org/10.5281/zenodo.6031574" ID-GBIF-Taxon="128848554" ID-Zenodo-Dep="6031574" LSID="urn:lsid:plazi:treatment:E45787C5FFA7FFDCE1A163CBFD4CFA0F" httpUri="http://treatment.plazi.org/id/E45787C5FFA7FFDCE1A163CBFD4CFA0F" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
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<subSubSection id="24E46558FFA7FFDDE1A163CBFE2DFE18" pageId="38" pageNumber="39" type="nomenclature">
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<paragraph id="6C4136D3FFA7FFDDE1A163CBFE61FE39" blockId="38.[151,487,438,498]" box="[151,487,438,464]" pageId="38" pageNumber="39">
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<heading id="370981BFFFA7FFDDE1A163CBFE61FE39" bold="true" box="[151,487,438,464]" fontSize="11" level="1" pageId="38" pageNumber="39" reason="1">
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<emphasis id="5E8AEAC1FFA7FFDDE1A163CBFE61FE39" bold="true" box="[151,487,438,464]" pageId="38" pageNumber="39">
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||
<taxonomicName id="ABFE4D50FFA7FFDDE1A163CBFED6FE39" ID-CoL="47ZY9" box="[151,336,438,464]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="clara" status="sp. nov.">
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<emphasis id="5E8AEAC1FFA7FFDDE1A163CBFED6FE39" bold="true" box="[151,336,438,464]" italics="true" pageId="38" pageNumber="39">Novalena clara</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="45B957BAFFA7FFDDE06163CBFE61FE39" box="[343,487,438,464]" pageId="38" pageNumber="39" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="6C4136D3FFA7FFDDE1A163A4FE2DFE18" blockId="38.[151,487,438,498]" box="[151,427,473,498]" pageId="38" pageNumber="39">
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<figureCitation id="F4C52A56FFA7FFDDE1A163A4FF7BFE1B" box="[151,253,473,498]" captionStart="FIGURE 1" captionStartId="6.[151,250,1888,1910]" captionTargetBox="[186,1400,348,1856]" captionTargetId="figure@6.[176,1411,339,1866]" captionTargetPageId="6" captionText="FIGURE 1. Novalena spp., habitus: a, e, N. approximata (Gertsch & Ivie 1936), female; b, f, N. alvarezi sp. nov., female; c, g, N. chamberlini sp. nov., male; d, h, N. clara sp. nov., female; i, m, N. dentata sp. nov., male; j, n, N. irazu sp. nov., female; k, o, N. iviei sp. nov., male; l, p, N. leonensis sp. nov., female; q, u, N. mexiquensis sp. nov., male; r, v, N. poncei sp. nov., female; s, w, N. puebla sp. nov., female; t, x, N. rothi sp. nov., female. a – d, i – l, q – t, dorsal view; e – h, m – p, v – x, ventral view. Scale bars: a, d, j, q, s, t = 4 mm; b, i, l, r = 3 mm; c, k = 2 mm." httpUri="https://zenodo.org/record/571684/files/figure.png" pageId="38" pageNumber="39">Figures 1</figureCitation>
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d, h, 19; Map 4
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</paragraph>
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</subSubSection>
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<subSubSection id="24E46558FFA7FFDDE1A1605CFCCAFDB4" pageId="38" pageNumber="39" type="materials_examined">
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<paragraph id="6C4136D3FFA7FFDDE1A1605CFCCAFDB4" blockId="38.[151,1437,545,2010]" pageId="38" pageNumber="39">
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<materialsCitation id="DC963C8EFFA7FFDDE1A1605CFCCEFDB4" ID-GBIF-Occurrence="1500194654" collectingDate="1969-12-27" collectionCode="CAS" collectorName="V. Roth" country="Mexico" county="Arroyo" latitude="29.333334" location="Sierra" longLatPrecision="1227" longitude="-106.833336" municipality="Nido" pageId="38" pageNumber="39" specimenCount="2" stateProvince="Chihuahua" typeStatus="holotype">
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<emphasis id="5E8AEAC1FFA7FFDDE1A1605CFF56FDD3" bold="true" box="[151,208,545,570]" pageId="38" pageNumber="39">
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<typeStatus id="B3458871FFA7FFDDE1A1605CFF56FDD3" box="[151,208,545,570]" pageId="38" pageNumber="39">Type</typeStatus>
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</emphasis>
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.
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<emphasis id="5E8AEAC1FFA7FFDDE1DC605CFE24FDD3" bold="true" box="[234,418,545,570]" pageId="38" pageNumber="39">
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<typeStatus id="B3458871FFA7FFDDE1DC605CFED1FDD3" box="[234,343,545,570]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
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male
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</emphasis>
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:
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<collectingCountry id="14E97643FFA7FFDDE08A605CFDA9FDD0" box="[444,559,545,569]" name="Mexico" pageId="38" pageNumber="39">MEXICO</collectingCountry>
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:
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<emphasis id="5E8AEAC1FFA7FFDDE372605CFD46FDD3" box="[580,704,545,570]" italics="true" pageId="38" pageNumber="39">
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||
<collectingRegion id="AE3AF831FFA7FFDDE372605CFD46FDD3" box="[580,704,545,570]" country="Mexico" name="Chihuahua" pageId="38" pageNumber="39">Chihuahua</collectingRegion>
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</emphasis>
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:
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<location id="69216008FFA7FFDDE3EC605CFC9BFDD3" LSID="urn:lsid:plazi:treatment:E45787C5FFA7FFDCE1A163CBFD4CFA0F:69216008FFA7FFDDE3EC605CFC9BFDD3" box="[730,797,545,570]" country="Mexico" county="Arroyo" latitude="29.333334" longLatPrecision="1227" longitude="-106.833336" municipality="Nido" name="Sierra" pageId="38" pageNumber="39" stateProvince="Chihuahua">Sierra</location>
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del
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<collectingMunicipality id="8C25ACA9FFA7FFDDE255605CFC18FDD3" box="[867,926,545,570]" pageId="38" pageNumber="39">Nido</collectingMunicipality>
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,
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<collectingCounty id="85204E5FFFA7FFDDE285605CFB82FDD0" box="[947,1028,545,569]" pageId="38" pageNumber="39">Arroyo</collectingCounty>
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del
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<taxonomicName id="ABFE4D50FFA7FFDDE57C605CFB1FFDD3" box="[1098,1177,545,570]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="85" phylum="Arthropoda" rank="species" species="alamo" status="sp. nov.">Álamo</taxonomicName>
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,
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<geoCoordinate id="09CA5014FFA7FFDDE599605CFA89FDD3" box="[1199,1295,545,570]" direction="north" orientation="latitude" pageId="38" pageNumber="39" precision="925" value="29.333334">29°20′N</geoCoordinate>
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,
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<geoCoordinate id="09CA5014FFA7FFDDE410605CFA11FDD3" box="[1318,1431,545,570]" direction="west" orientation="longitude" pageId="38" pageNumber="39" precision="925" value="-106.833336">106°50′W</geoCoordinate>
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,
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<date id="18401013FFA7FFDDE1A16039FEA5FDB4" box="[151,291,580,605]" pageId="38" pageNumber="39" value="1969-12-27">
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<collectingDate id="0804E9FBFFA7FFDDE1A16039FEA5FDB4" box="[151,291,580,605]" pageId="38" pageNumber="39" value="1969-12-27">27.XII.1969</collectingDate>
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</date>
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,
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<collectorName id="C10B5305FFA7FFDDE018603BFE04FDB4" box="[302,386,580,605]" pageId="38" pageNumber="39">V. Roth</collectorName>
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(CASENT 9048865), deposited at
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<collectionCode id="0AEFAE16FFA7FFDDE2386038FCCEFDB4" box="[782,840,581,605]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="38" pageNumber="39">CAS</collectionCode>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="24E46558FFA7FFDDE1F16014FEB4FD4C" pageId="38" pageNumber="39" type="etymology">
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<paragraph id="6C4136D3FFA7FFDDE1F16014FEB4FD4C" blockId="38.[151,1437,545,2010]" pageId="38" pageNumber="39">
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<emphasis id="5E8AEAC1FFA7FFDDE1F16014FEC0FD6B" bold="true" box="[199,326,617,642]" pageId="38" pageNumber="39">Etymology</emphasis>
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. The specific name is a noun in apposition referring to the color tone of the
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<typeStatus id="B3458871FFA7FFDDE5AC6014FB7AFD6B" box="[1178,1276,617,642]" pageId="38" pageNumber="39" type="holotype">holotype</typeStatus>
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<emphasis id="5E8AEAC1FFA7FFDDE4356014FAD6FD6B" box="[1283,1360,617,642]" italics="true" pageId="38" pageNumber="39">‘clara’</emphasis>
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which means ‘
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<emphasis id="5E8AEAC1FFA7FFDDE1D960F1FEADFD4C" box="[239,299,652,677]" italics="true" pageId="38" pageNumber="39">light’</emphasis>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="24E46558FFA7FFDDE1F160CCFB39FC94" pageId="38" pageNumber="39" type="diagnosis">
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<paragraph id="6C4136D3FFA7FFDDE1F160CCFB39FC94" blockId="38.[151,1437,545,2010]" pageId="38" pageNumber="39">
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<emphasis id="5E8AEAC1FFA7FFDDE1F160CCFEBCFD23" bold="true" box="[199,314,689,714]" pageId="38" pageNumber="39">Diagnosis</emphasis>
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. The males of this species differ from
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<taxonomicName id="ABFE4D50FFA7FFDDE23F60CFFC3DFD20" box="[777,955,690,713]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="approximata">
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<emphasis id="5E8AEAC1FFA7FFDDE23F60CFFC3DFD20" box="[777,955,690,713]" italics="true" pageId="38" pageNumber="39">N. approximata</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="ABFE4D50FFA7FFDDE2CF60CFFB19FD23" box="[1017,1183,689,714]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="bosencheve">
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<emphasis id="5E8AEAC1FFA7FFDDE2CF60CFFB19FD23" box="[1017,1183,689,714]" italics="true" pageId="38" pageNumber="39">N. bosencheve</emphasis>
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</taxonomicName>
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in having the tegular median process with one projection (
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<figureCitation id="F4C52A56FFA7FFDDE30860A9FD16FD04" box="[574,656,724,749]" captionStart="FIGURE 19" captionStartId="39.[151,250,1194,1216]" captionTargetBox="[152,1426,450,1169]" captionTargetId="figure@39.[151,1436,445,1172]" captionTargetPageId="39" captionText="FIGURE 19. Novalena clara sp. nov.: a, male pedipalp, ventral view; b, same, retrolateral view; c, epigynum, ventral view; d, internal genitalia, dorsal view; e, epigynum, posterior view. Abbreviations: A, atrium; C, conductor; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process. Scale bars: a = 0.5 mm; e = 0.25 mm." httpUri="https://zenodo.org/record/571703/files/figure.png" pageId="38" pageNumber="39">Fig. 19</figureCitation>
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a) and the RTA with a denticle on ventral projection (
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<figureCitation id="F4C52A56FFA7FFDDE5C160A9FACCFD04" box="[1271,1354,724,749]" captionStart="FIGURE 19" captionStartId="39.[151,250,1194,1216]" captionTargetBox="[152,1426,450,1169]" captionTargetId="figure@39.[151,1436,445,1172]" captionTargetPageId="39" captionText="FIGURE 19. Novalena clara sp. nov.: a, male pedipalp, ventral view; b, same, retrolateral view; c, epigynum, ventral view; d, internal genitalia, dorsal view; e, epigynum, posterior view. Abbreviations: A, atrium; C, conductor; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process. Scale bars: a = 0.5 mm; e = 0.25 mm." httpUri="https://zenodo.org/record/571703/files/figure.png" pageId="38" pageNumber="39">Fig. 19</figureCitation>
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b). The females differ from
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<taxonomicName id="ABFE4D50FFA7FFDDE0B76087FDB5FCF8" box="[385,563,762,785]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="approximata">
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<emphasis id="5E8AEAC1FFA7FFDDE0B76087FDB5FCF8" box="[385,563,762,785]" italics="true" pageId="38" pageNumber="39">N. approximata</emphasis>
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</taxonomicName>
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in having the spurs more conspicuous (
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<figureCitation id="F4C52A56FFA7FFDDE53F6084FBDBFCF8" box="[1033,1117,761,786]" captionStart="FIGURE 19" captionStartId="39.[151,250,1194,1216]" captionTargetBox="[152,1426,450,1169]" captionTargetId="figure@39.[151,1436,445,1172]" captionTargetPageId="39" captionText="FIGURE 19. Novalena clara sp. nov.: a, male pedipalp, ventral view; b, same, retrolateral view; c, epigynum, ventral view; d, internal genitalia, dorsal view; e, epigynum, posterior view. Abbreviations: A, atrium; C, conductor; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process. Scale bars: a = 0.5 mm; e = 0.25 mm." httpUri="https://zenodo.org/record/571703/files/figure.png" pageId="38" pageNumber="39">Fig. 19</figureCitation>
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c), from
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<taxonomicName id="ABFE4D50FFA7FFDDE5F36087FAC3FCFB" box="[1221,1349,761,786]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="franckei">
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<emphasis id="5E8AEAC1FFA7FFDDE5F36087FAC3FCFB" box="[1221,1349,761,786]" italics="true" pageId="38" pageNumber="39">N. franckei</emphasis>
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||
</taxonomicName>
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and
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<taxonomicName id="ABFE4D50FFA7FFDDE4B46087FF7FFCDC" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="volcanes">
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<emphasis id="5E8AEAC1FFA7FFDDE4B46087FF7FFCDC" italics="true" pageId="38" pageNumber="39">N. volcanes</emphasis>
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</taxonomicName>
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in having the primary spermathecae with posterior part wider than anterior part (
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<figureCitation id="F4C52A56FFA7FFDDE5F76161FA91FCDC" box="[1217,1303,796,821]" captionStart="FIGURE 19" captionStartId="39.[151,250,1194,1216]" captionTargetBox="[152,1426,450,1169]" captionTargetId="figure@39.[151,1436,445,1172]" captionTargetPageId="39" captionText="FIGURE 19. Novalena clara sp. nov.: a, male pedipalp, ventral view; b, same, retrolateral view; c, epigynum, ventral view; d, internal genitalia, dorsal view; e, epigynum, posterior view. Abbreviations: A, atrium; C, conductor; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process. Scale bars: a = 0.5 mm; e = 0.25 mm." httpUri="https://zenodo.org/record/571703/files/figure.png" pageId="38" pageNumber="39">Fig. 19</figureCitation>
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d), from
|
||
<taxonomicName id="ABFE4D50FFA7FFDDE4B46163FF78FCB0" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="paricutin">
|
||
<emphasis id="5E8AEAC1FFA7FFDDE4B46163FF78FCB0" italics="true" pageId="38" pageNumber="39">N. paricutin</emphasis>
|
||
</taxonomicName>
|
||
in having the spurs directed mesally, from
|
||
<taxonomicName id="ABFE4D50FFA7FFDDE3DE613FFCF1FCB3" box="[744,887,833,858]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="saltoensis">
|
||
<emphasis id="5E8AEAC1FFA7FFDDE3DE613FFCF1FCB3" box="[744,887,833,858]" italics="true" pageId="38" pageNumber="39">N. saltoensis</emphasis>
|
||
</taxonomicName>
|
||
in having the spurs separated from the posterior margin by their basal width, and from
|
||
<taxonomicName id="ABFE4D50FFA7FFDDE373611BFD1BFC94" box="[581,669,868,893]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="creel">
|
||
<emphasis id="5E8AEAC1FFA7FFDDE373611BFD1BFC94" box="[581,669,868,893]" italics="true" pageId="38" pageNumber="39">N. creel</emphasis>
|
||
</taxonomicName>
|
||
in having the spurs in lateral position (
|
||
<figureCitation id="F4C52A56FFA7FFDDE5646119FB22FC94" box="[1106,1188,868,893]" captionStart="FIGURE 19" captionStartId="39.[151,250,1194,1216]" captionTargetBox="[152,1426,450,1169]" captionTargetId="figure@39.[151,1436,445,1172]" captionTargetPageId="39" captionText="FIGURE 19. Novalena clara sp. nov.: a, male pedipalp, ventral view; b, same, retrolateral view; c, epigynum, ventral view; d, internal genitalia, dorsal view; e, epigynum, posterior view. Abbreviations: A, atrium; C, conductor; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process. Scale bars: a = 0.5 mm; e = 0.25 mm." httpUri="https://zenodo.org/record/571703/files/figure.png" pageId="38" pageNumber="39">Fig. 19</figureCitation>
|
||
c).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="24E46558FFA7FFDCE1F161F4FCFBFA2B" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39" type="description">
|
||
<paragraph id="6C4136D3FFA7FFDDE1F161F4FB47F9C3" blockId="38.[151,1437,545,2010]" pageId="38" pageNumber="39">
|
||
<emphasis id="5E8AEAC1FFA7FFDDE1F161F4FED7FC4B" bold="true" box="[199,337,905,930]" pageId="38" pageNumber="39">Description</emphasis>
|
||
.
|
||
<emphasis id="5E8AEAC1FFA7FFDDE05661F4FE1BFC4B" bold="true" box="[352,413,905,930]" pageId="38" pageNumber="39">Male</emphasis>
|
||
(
|
||
<typeStatus id="B3458871FFA7FFDDE09A61F4FD93FC4B" box="[428,533,905,930]" pageId="38" pageNumber="39" type="holotype">holotype</typeStatus>
|
||
): Carapace yellow. Chelicerae yellow, condyles yellow. Endites yellow. Labium light brown with white tip. Sternum brown with a central yellow band with arrowhead shaped spot. Legs yellow with black spots on coxa and trochanter, three black rings on femur, two on patella, and two on tibia. Opisthosoma light brown with an anterior reddish spot and one yellow dorsal band. PLS with basal segment brown. Total length 10.13. Carapace length 4.5, width 3.13, cephalic region width 1.88, ocular region width 1. Eye diameter: AME 0.13, ALE 0.19, PME and PLE 0.17. Distance between eyes: AME-AME 0.15, AME-ALE 0.12, AME-PME 0.1, ALE-PLE 0.1, ALE-ALE 0.5, PME-PME 0.19, PME-PLE 0.19. Clypeus height 0.52. Chelicerae with four retromarginal teeth; basal segment length 1.33, fang length 0.76. Labium as long as wide (0.61/0.64). Endites slightly convergent (distance at their base compared at their tips 0.64/0.42). Sternum longer than wide (2.24/1.9). Opisthosoma longer than wide (5.88/3.38). ALS separated by less their basal diameter (0.19/0.48), PLS with distal segment shorter than basal segment (0.43/0.71). Leg lengths: I—femur 3.5/ patella-tibia 4.5/ metatarsus 3.38/ tarsus 2.38; II—3.75/ 4.5/ 3.5/ 2.13; III—3.75/ 4.38/ 3.5/ 1.88; IV—4.38/ 5.13/ 5/ 2.38. Spination: femur dorsal
|
||
<date id="18401013FFA7FFDDE441676BFF3FFAB8" pageId="38" pageNumber="39">I— 2-1</date>
|
||
-2/
|
||
<date id="18401013FFA7FFDDE1D26744FEBEFAB8" box="[228,312,1337,1361]" pageId="38" pageNumber="39">II—2-1</date>
|
||
-3/
|
||
<date id="18401013FFA7FFDDE0696744FE3BFAB8" box="[351,445,1337,1361]" pageId="38" pageNumber="39">III—2-1</date>
|
||
-3/
|
||
<date id="18401013FFA7FFDDE0D56744FDC0FAB8" box="[483,582,1337,1361]" pageId="38" pageNumber="39">IV—2-1</date>
|
||
-2; patella I—dorsal 1-1/ prolateral 1/ retrolateral 0;
|
||
<date id="18401013FFA7FFDDE5A16744FB6EFAB8" box="[1175,1256,1337,1361]" pageId="38" pageNumber="39">II—1-1</date>
|
||
/ 1/ 0;
|
||
<date id="18401013FFA7FFDDE4016744FA14FAB8" box="[1335,1426,1337,1361]" pageId="38" pageNumber="39">III—1-1</date>
|
||
/ 1/ 1;
|
||
<date id="18401013FFA7FFDDE1E46720FEA8FA9C" box="[210,302,1373,1397]" pageId="38" pageNumber="39">IV—1-1</date>
|
||
/ 1/ 1; tibia I—dorsal 1-1-0/ ventral 2-2-2/ prolateral 1-1-0/ retrolateral 0;
|
||
<date id="18401013FFA7FFDDE5456720FB42FA9C" box="[1139,1220,1373,1397]" pageId="38" pageNumber="39">II—1-1</date>
|
||
-0/ 2-2-2/ 1-1-0/ 0;
|
||
<date id="18401013FFA7FFDDE1A167FCFF74FA70" box="[151,242,1409,1433]" pageId="38" pageNumber="39">III—1-1</date>
|
||
-0/ 2-2-2/ 1-1-0/ 1-1-0;
|
||
<date id="18401013FFA7FFDDE0C867FCFDDDFA70" box="[510,603,1409,1433]" pageId="38" pageNumber="39">IV—1-1</date>
|
||
-0/ 2-2-2/ 1-1-0/ 1-1-0; metatarsus I—0/ 2-2-2/ 0-1-1/ 0; II—0-0-2/ 2-2-2/ 1-1-1/ 0-0-1;
|
||
<date id="18401013FFA7FFDDE01867D8FE0AFA54" box="[302,396,1445,1469]" pageId="38" pageNumber="39">III—2-1</date>
|
||
-2/ 2-2-2/ 0-1-1/ 1-0-1;
|
||
<date id="18401013FFA7FFDDE3A067D8FD72FA54" box="[662,756,1445,1469]" pageId="38" pageNumber="39">IV—2-1</date>
|
||
-2/ 2-1-2-2/ 0-1-1/ 0-1-1. Number of trichobothria on tarsus:
|
||
<date id="18401013FFA7FFDDE1A167B4FF4AFA08" box="[151,204,1481,1505]" pageId="38" pageNumber="39">I—8</date>
|
||
,
|
||
<date id="18401013FFA7FFDDE1EC67B7FE9CFA08" box="[218,282,1482,1505]" pageId="38" pageNumber="39">II—7</date>
|
||
,
|
||
<date id="18401013FFA7FFDDE01167B4FEE9FA0B" box="[295,367,1481,1506]" pageId="38" pageNumber="39">III—5</date>
|
||
,
|
||
<date id="18401013FFA7FFDDE04B67B7FE4BFA08" box="[381,461,1482,1505]" pageId="38" pageNumber="39">IV—7.</date>
|
||
Number of dorsal spines on pedipalp: femur 3, tibia 2. Cymbium length 1.76, width 1.12. Embolus straight, conductor distally notched; tegular median process with two broad and overlapping tips; dorsal projection of RTA longer than ventral projection, which presents a denticle (
|
||
<figureCitation id="F4C52A56FFA7FFDDE50E646CFB0FF9C0" box="[1080,1161,1553,1578]" captionStart="FIGURE 19" captionStartId="39.[151,250,1194,1216]" captionTargetBox="[152,1426,450,1169]" captionTargetId="figure@39.[151,1436,445,1172]" captionTargetPageId="39" captionText="FIGURE 19. Novalena clara sp. nov.: a, male pedipalp, ventral view; b, same, retrolateral view; c, epigynum, ventral view; d, internal genitalia, dorsal view; e, epigynum, posterior view. Abbreviations: A, atrium; C, conductor; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process. Scale bars: a = 0.5 mm; e = 0.25 mm." httpUri="https://zenodo.org/record/571703/files/figure.png" pageId="38" pageNumber="39">Fig. 19</figureCitation>
|
||
a, b).
|
||
</paragraph>
|
||
<paragraph id="6C4136D3FFA7FFDCE1F16449FDF7FE61" blockId="38.[151,1437,545,2010]" lastBlockId="39.[151,1436,151,392]" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
|
||
<emphasis id="5E8AEAC1FFA7FFDDE1F16449FE98F9A4" bold="true" box="[199,286,1588,1613]" pageId="38" pageNumber="39">Female</emphasis>
|
||
(
|
||
<typeStatus id="B3458871FFA7FFDDE01A6448FE16F9A4" box="[300,400,1589,1613]" pageId="38" pageNumber="39" type="paratype">paratype</typeStatus>
|
||
) (CASENT 9048874): Carapace yellow, white feathery scales on cephalic region. Chelicerae reddish, condyles orange. Endites light brown. Labium brown with white tip. Sternum brown with a median yellow band with arrowhead shaped spot directed posteriorly. Legs yellow, tibia to tarsus reddish. Three black rings on femur and two on patella. Opisthosoma yellow with several lateral and ventral black spots and yellow foliage. Spinnerets yellow. Total length 11.63. Carapace length 5.63, width 3.75, cephalic region width 2.38, ocular region width 1.25. Eye diameter: AME, PME and PLE 0.21, ALE 0.25. Distance between eyes: AME-AME 0.12, AME- ALE 0.13, AME-PME 0.25, ALE-PLE 0.1, ALE-ALE 0.58, PME-PME 0.15, PME-PLE 0.23. Clypeus height 0.38. Chelicerae with four retromarginal teeth; basal segment length 1.29, fang length 0.95. Labium wider than long (0.95/0.76). Endites slightly convergent (distance at their base compared at their tips 0.95/0.48). Sternum longer than wide (2.63/2.25). Opisthosoma longer than wide (6.38/3.75). ALS separated by half their basal diameter (0.3/ 0.61), PLS with distal segment as long as their basal segment (0.64/0.64). Leg lengths: I—femur 4/ patella-tibia 4.5/ metatarsus 3.63/ tarsus 1.88;
|
||
<date id="18401013FFA7FFDDE32665BCFDCAF830" box="[528,588,1985,2009]" pageId="38" pageNumber="39">II—4</date>
|
||
/ 4.38/ 2.5/ 1.88; III—3.75/ 4/ 5/ 1.88; IV—4.63/ 5.63/ 4.75/ 1.5. Spination: femur dorsal
|
||
<date id="18401013FFA6FFDCE01A62E5FEF3FF59" box="[300,373,152,176]" pageId="39" pageNumber="40">I—1-2</date>
|
||
-2/
|
||
<date id="18401013FFA6FFDCE0AA62E5FE69FF59" box="[412,495,152,176]" pageId="39" pageNumber="40">II—1-2</date>
|
||
-2/
|
||
<date id="18401013FFA6FFDCE32062E5FDF2FF59" box="[534,628,152,176]" pageId="39" pageNumber="40">III—1-3</date>
|
||
-3/
|
||
<date id="18401013FFA6FFDCE3AF62E5FD7BFF59" box="[665,765,152,176]" pageId="39" pageNumber="40">IV—1-2</date>
|
||
-3; patella I—dorsal 1-1/ prolateral 1/ retrolateral 0;
|
||
<date id="18401013FFA6FFDCE47662E5FA14FF59" box="[1344,1426,152,176]" pageId="39" pageNumber="40">II—1-1</date>
|
||
/ 1/ 0;
|
||
<date id="18401013FFA6FFDCE1E662C1FEADFF3D" box="[208,299,188,212]" pageId="39" pageNumber="40">III—1-1</date>
|
||
/ 1/ 1;
|
||
<date id="18401013FFA6FFDCE04362C1FE57FF3D" box="[373,465,188,212]" pageId="39" pageNumber="40">IV—1-1</date>
|
||
/ 1/ 1; tibia I—dorsal 1-1-0/ ventral 2-2-2/ prolateral 1-1-0/ retrolateral 0; II—0-1-0/ 2- 2-2/ 1-1-0/ 0;
|
||
<date id="18401013FFA6FFDCE002629DFE10FF11" box="[308,406,224,248]" pageId="39" pageNumber="40">III—1-2</date>
|
||
-1-2/ 2-2-2/ 0/ 0;
|
||
<date id="18401013FFA6FFDCE363629DFD3EFF11" box="[597,696,224,248]" pageId="39" pageNumber="40">IV—1-2</date>
|
||
-1-2/ 2-2-2/ 0/ 0; metatarsus I—0/ 2-2-2/ 0-1-1/ 0; II—0/ 2-2-2/ 0- 1-1/ 0;
|
||
<date id="18401013FFA6FFDCE1DA6379FECDFEF5" box="[236,331,260,284]" pageId="39" pageNumber="40">III—2-2</date>
|
||
-2/ 2-2-2/ 0-1-1/ 0-1-1;
|
||
<date id="18401013FFA6FFDCE36C6379FD31FEF5" box="[602,695,260,284]" pageId="39" pageNumber="40">IV—1-1</date>
|
||
-2/ 1-1-1-1-1-2/ 1-1-1/ 0-0-1. Number of trichobothria on tarsus:
|
||
<date id="18401013FFA6FFDCE1A16355FF4AFEA9" box="[151,204,296,320]" pageId="39" pageNumber="40">I—8</date>
|
||
,
|
||
<date id="18401013FFA6FFDCE1EE6355FE91FED6" box="[216,279,296,319]" pageId="39" pageNumber="40">II—7</date>
|
||
,
|
||
<date id="18401013FFA6FFDCE0146355FEECFEA9" box="[290,362,296,320]" pageId="39" pageNumber="40">III—9</date>
|
||
,
|
||
<date id="18401013FFA6FFDCE043635AFE42FEA9" box="[373,452,295,320]" pageId="39" pageNumber="40">IV—6.</date>
|
||
Dorsal spines on pedipalp femur: 3. Prolateral spines on tibia: 1-2. Epigynal plate wider than long (1.58/0.91); atrium superficial; spurs lateral (
|
||
<figureCitation id="F4C52A56FFA6FFDCE2276331FCE3FE8D" box="[785,869,332,357]" captionStart="FIGURE 19" captionStartId="39.[151,250,1194,1216]" captionTargetBox="[152,1426,450,1169]" captionTargetId="figure@39.[151,1436,445,1172]" captionTargetPageId="39" captionText="FIGURE 19. Novalena clara sp. nov.: a, male pedipalp, ventral view; b, same, retrolateral view; c, epigynum, ventral view; d, internal genitalia, dorsal view; e, epigynum, posterior view. Abbreviations: A, atrium; C, conductor; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process. Scale bars: a = 0.5 mm; e = 0.25 mm." httpUri="https://zenodo.org/record/571703/files/figure.png" pageId="39" pageNumber="40">Fig. 19</figureCitation>
|
||
c, e). Internal genitalia: copulatory ducts straight, primary spermathecae adjacent (
|
||
<figureCitation id="F4C52A56FFA6FFDCE3346312FDD2FE61" box="[514,596,367,392]" captionStart="FIGURE 19" captionStartId="39.[151,250,1194,1216]" captionTargetBox="[152,1426,450,1169]" captionTargetId="figure@39.[151,1436,445,1172]" captionTargetPageId="39" captionText="FIGURE 19. Novalena clara sp. nov.: a, male pedipalp, ventral view; b, same, retrolateral view; c, epigynum, ventral view; d, internal genitalia, dorsal view; e, epigynum, posterior view. Abbreviations: A, atrium; C, conductor; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process. Scale bars: a = 0.5 mm; e = 0.25 mm." httpUri="https://zenodo.org/record/571703/files/figure.png" pageId="39" pageNumber="40">Fig. 19</figureCitation>
|
||
d).
|
||
</paragraph>
|
||
<caption id="3881665BFFA6FFDCE1A166D7FD92FAD3" httpUri="https://zenodo.org/record/571703/files/figure.png" pageId="39" pageNumber="40" targetBox="[152,1426,450,1169]" targetPageId="39">
|
||
<paragraph id="6C4136D3FFA6FFDCE1A166D7FD92FAD3" blockId="39.[151,1436,1194,1338]" pageId="39" pageNumber="40">
|
||
<emphasis id="5E8AEAC1FFA6FFDCE1A166D7FE9FFB56" bold="true" box="[151,281,1194,1216]" pageId="39" pageNumber="40">FIGURE 19</emphasis>
|
||
.
|
||
<taxonomicName id="ABFE4D50FFA6FFDCE01366D7FE44FB29" box="[293,450,1194,1216]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="clara" status="sp. nov.">
|
||
<emphasis id="5E8AEAC1FFA6FFDCE01366D7FE44FB29" box="[293,450,1194,1216]" italics="true" pageId="39" pageNumber="40">Novalena clara</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="5E8AEAC1FFA6FFDCE0FF66D7FD9FFB56" bold="true" box="[457,537,1194,1215]" pageId="39" pageNumber="40">
|
||
<taxonomicNameLabel id="45B957BAFFA6FFDCE0FF66D7FD9FFB56" box="[457,537,1194,1215]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: a, male pedipalp, ventral view; b, same, retrolateral view; c, epigynum, ventral view; d, internal genitalia, dorsal view; e, epigynum, posterior view. Abbreviations: A, atrium; C, conductor; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S, spurs; S1, primary spermathecae; S2, secondary spermathecae; T, tegulum; TMP, tegular median process. Scale bars: a = 0.5 mm; e = 0.25 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="6C4136D3FFA6FFDCE1F1671CFCFBFA2B" blockId="39.[151,1436,1377,1510]" pageId="39" pageNumber="40">
|
||
<materialsCitation id="DC963C8EFFA6FFDCE1F1671CFCC7FA77" ID-GBIF-Occurrence="1500194658" country="Mexico" location="MEXICO" pageId="39" pageNumber="40" specimenCount="1" typeStatus="paratype">
|
||
<emphasis id="5E8AEAC1FFA6FFDCE1F1671CFEB3FA93" bold="true" box="[199,309,1377,1402]" pageId="39" pageNumber="40">Variation</emphasis>
|
||
. Males (n=2): body length 10.13–10.75; carapace length 4.5–5.58; patella-tibia I length 4.5–5.
|
||
<emphasis id="5E8AEAC1FFA6FFDCE1F167F8FDB5FA77" bold="true" box="[199,563,1413,1438]" pageId="39" pageNumber="40">Additional material examined</emphasis>
|
||
.
|
||
<emphasis id="5E8AEAC1FFA6FFDCE37267F8FD3AFA77" bold="true" box="[580,700,1413,1438]" pageId="39" pageNumber="40">
|
||
<typeStatus id="B3458871FFA6FFDCE37267F8FD3AFA77" box="[580,700,1413,1438]" pageId="39" pageNumber="40" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<collectingCountry id="14E97643FFA6FFDCE3F867FBFCC7FA77" box="[718,833,1414,1438]" name="Mexico" pageId="39" pageNumber="40">MEXICO</collectingCountry>
|
||
</materialsCitation>
|
||
:
|
||
<materialsCitation id="DC963C8EFFA6FFDCE27867F8FE80FA2B" ID-GBIF-Occurrence="1500194659" collectingDate="1969-12-27" collectionCode="CAS" collectorName="V. Roth" country="Mexico" county="Arroyo" latitude="29.333334" location="Sierra" longLatPrecision="1227" longitude="-106.833336" municipality="Nido" pageId="39" pageNumber="40" specimenCount="1" specimenCount-male="1" stateProvince="Chihuahua" typeStatus="paratype">
|
||
<emphasis id="5E8AEAC1FFA6FFDCE27867F8FC4CFA77" box="[846,970,1413,1438]" italics="true" pageId="39" pageNumber="40">
|
||
<collectingRegion id="AE3AF831FFA6FFDCE27867F8FC4CFA77" box="[846,970,1413,1438]" country="Mexico" name="Chihuahua" pageId="39" pageNumber="40">Chihuahua</collectingRegion>
|
||
</emphasis>
|
||
: same data as holotype,
|
||
<specimenCount id="7AF8FD5AFFA6FFDCE5D867FBFA9AFA77" box="[1262,1308,1414,1438]" pageId="39" pageNumber="40" type="male">1 ♂</specimenCount>
|
||
(CASENT 9048864)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="DC963C8EFFA6FFDCE02767D7FCFBFA2B" ID-GBIF-Occurrence="1500194677" box="[273,893,1449,1474]" collectingDate="1969-10-14" collectionCode="CAS" collectorName="V. Roth" country="Mexico" county="Arroyo" latitude="29.333334" location="Sierra" longLatPrecision="1227" longitude="-106.833336" municipality="Nido" pageId="39" pageNumber="40" specimenCount="1" specimenCount-female="1" stateProvince="Chihuahua" typeStatus="paratype">
|
||
same data except
|
||
<date id="18401013FFA6FFDCE0E167D4FDC9FA2B" box="[471,591,1449,1474]" pageId="39" pageNumber="40" value="1969-10-14">
|
||
<collectingDate id="0804E9FBFFA6FFDCE0E167D4FDC9FA2B" box="[471,591,1449,1474]" pageId="39" pageNumber="40" value="1969-10-14">14.X.1969</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="7AF8FD5AFFA6FFDCE36D67D7FD03FA28" box="[603,645,1450,1474]" pageId="39" pageNumber="40" type="female">1 ♀</specimenCount>
|
||
(CASENT 9048874).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="24E46558FFA6FFDCE1F167B0FD4CFA0F" box="[199,714,1485,1510]" pageId="39" pageNumber="40" type="distribution">
|
||
<paragraph id="6C4136D3FFA6FFDCE1F167B0FD4CFA0F" blockId="39.[151,1436,1377,1510]" box="[199,714,1485,1510]" pageId="39" pageNumber="40">
|
||
<emphasis id="5E8AEAC1FFA6FFDCE1F167B0FEDFFA0F" bold="true" box="[199,345,1485,1510]" pageId="39" pageNumber="40">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="14E97643FFA6FFDCE05167B3FE5CFA0F" box="[359,474,1486,1510]" name="Mexico" pageId="39" pageNumber="40">
|
||
<collectingRegion id="AE3AF831FFA6FFDCE05167B3FE5CFA0F" box="[359,474,1486,1510]" country="Mexico" name="Mexico" pageId="39" pageNumber="40">MEXICO</collectingRegion>
|
||
</collectingCountry>
|
||
:
|
||
<emphasis id="5E8AEAC1FFA6FFDCE0D267B0FDE7FA0F" box="[484,609,1485,1510]" italics="true" pageId="39" pageNumber="40">
|
||
<collectingRegion id="AE3AF831FFA6FFDCE0D267B0FDE7FA0F" box="[484,609,1485,1510]" country="Mexico" name="Chihuahua" pageId="39" pageNumber="40">Chihuahua</collectingRegion>
|
||
</emphasis>
|
||
(Map 4).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |