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<document id="2B0484007817011FF633228CD5723634" ID="10.11646/zootaxa.4262.1.1SLASH10959" ID-CLB-Dataset="33105" ID-DOI="10.5281/zenodo.571683" ID-GBIF-Dataset="5ab1c651-825a-43bc-af00-11b77a792de3" ID-ISSN="1175-5326" ID-Zenodo-Dep="571683" ID-ZooBank="894795FB-ED38-4132-92B9-6AC8C3E41566" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1493972067874" checkinUser="plazi" docAuthor="Maya-Morales, Julieta &amp; Jiménez, María Luisa" docDate="2017" docId="E45787C5FF85FFFDE1A1646DFC55FEF5" docLanguage="en" docName="zootaxa.4262.1.1SLASH10959.pdf" docOrigin="Zootaxa 4262 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Novalena Chamberlin &amp; Ivie 1942" docType="treatment" docVersion="7" lastPageNumber="7" masterDocId="186EFFBDFF81FFFBE136627DFF86FFE9" masterDocTitle="Revision of the funnel-web spider genus Novalena (Araneae: Agelenidae)" masterLastPageNumber="88" masterPageNumber="1" pageNumber="5" updateTime="1698439297005" updateUser="ExternalLinkService">
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<mods:titleInfo id="E8E84FD9C66711056C4350E8B00B71FC">
<mods:title id="BA8875E0F828183D8D99BBCE3AB1B985">Revision of the funnel-web spider genus Novalena (Araneae: Agelenidae)</mods:title>
</mods:titleInfo>
<mods:name id="E64A274DFEF90279B5F337BE53EC3229" type="personal">
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<mods:namePart id="E5B26EDD144FDC31D97A58D8930EB0A4">Maya-Morales, Julieta</mods:namePart>
</mods:name>
<mods:name id="8B607FE9874981890B646C334AFB6FB6" type="personal">
<mods:role id="483F387ACE853453CC0176634E50C7D2">
<mods:roleTerm id="A316F6967FED9CA7B51D73B84E1524F7">Author</mods:roleTerm>
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<mods:namePart id="EA94723E1969514CF79CB608F639B203">Jiménez, María Luisa</mods:namePart>
</mods:name>
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<mods:title id="BDE09B71D8C7645442929BA47756F835">Zootaxa</mods:title>
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<mods:date id="EF30C4561B47C74F66E384A6590F3865">2017</mods:date>
<mods:detail id="B16C81A351E45C811BBB0C8A43DD7102" type="volume">
<mods:number id="0C618BAD08B44036712F6660F58B5D6C">4262</mods:number>
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<mods:classification id="4F5E8903FA9A6524E3DC285E8214621C">journal article</mods:classification>
<mods:identifier id="34C6FE1A28E6AF634B59C3C68DD90456" type="CLB-Dataset">33105</mods:identifier>
<mods:identifier id="4FC1B2CC6773C127F1CE93A5E1A9BD99" type="DOI">10.5281/zenodo.571683</mods:identifier>
<mods:identifier id="5D719C0BCAB2FC086F8103B8BF158236" type="GBIF-Dataset">5ab1c651-825a-43bc-af00-11b77a792de3</mods:identifier>
<mods:identifier id="30A557127863B1BD28D66814B1BA5BCF" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="118AA62467F9C13B77FF29A25B6F54F8" type="Zenodo-Dep">571683</mods:identifier>
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<treatment id="E45787C5FF85FFFDE1A1646DFC55FEF5" ID-DOI="http://doi.org/10.5281/zenodo.6031524" ID-GBIF-Taxon="128848526" ID-Zenodo-Dep="6031524" LSID="urn:lsid:plazi:treatment:E45787C5FF85FFFDE1A1646DFC55FEF5" httpUri="http://treatment.plazi.org/id/E45787C5FF85FFFDE1A1646DFC55FEF5" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
<subSubSection id="24E46558FF85FFFFE1A1646DFD1FF9C3" box="[151,665,1551,1578]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="6C4136D3FF85FFFFE1A1646DFD1FF9C3" blockId="4.[151,665,1551,1578]" box="[151,665,1551,1578]" pageId="4" pageNumber="5">
<heading id="370981BFFF85FFFFE1A1646DFD1FF9C3" bold="true" box="[151,665,1551,1578]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<emphasis id="5E8AEAC1FF85FFFFE1A1646DFD1FF9C3" bold="true" box="[151,665,1551,1578]" pageId="4" pageNumber="5">
Genus
<taxonomicName id="ABFE4D50FF85FFFFE1D86472FD1FF9C3" ID-CoL="65NL" ID-ENA="313399" authority="Chamberlin &amp; Ivie 1942" authorityName="Chamberlin &amp; Ivie" authorityYear="1942" box="[238,665,1551,1578]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="5E8AEAC1FF85FFFFE1D86472FEE4F9C0" bold="true" box="[238,354,1551,1577]" italics="true" pageId="4" pageNumber="5">Novalena</emphasis>
<bibRefCitation id="086F4B22FF85FFFFE05F646DFD1FF9C3" author="Chamberlin" box="[361,665,1552,1578]" pageId="4" pageNumber="5" refString="Chamberlin, R. V. &amp; Ivie, W. (1942) Agelenidae of the genera Hololena, Novalena, Rualena, and Melpomene. Annals of the Entomological Society of America, 35, 203 - 241." type="journal article" year="1942">Chamberlin &amp; Ivie 1942</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="24E46558FF85FFFFE1A1642BFA1BF83F" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="6C4136D3FF85FFFFE1A1642BFF59F97A" blockId="4.[151,1437,1622,2007]" pageId="4" pageNumber="5">
<emphasis id="5E8AEAC1FF85FFFFE1A1642BFEA9F986" bold="true" box="[151,303,1622,1647]" pageId="4" pageNumber="5">Type species</emphasis>
.
<taxonomicName id="ABFE4D50FF85FFFFE077642BFC1EF986" authority="Chamberlin &amp; Gertsch 1930" authorityName="Chamberlin &amp; Gertsch" authorityYear="1930" box="[321,920,1622,1647]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="5E8AEAC1FF85FFFFE077642BFDB7F986" box="[321,561,1622,1647]" italics="true" pageId="4" pageNumber="5">Novalena intermedia</emphasis>
(Chamberlin &amp; Gertsch 1930)
</taxonomicName>
by original designation (
<bibRefCitation id="086F4B22FF85FFFFE589642BFF55F97A" author="Chamberlin" pageId="4" pageNumber="5" refString="Chamberlin, R. V. &amp; Ivie, W. (1942) Agelenidae of the genera Hololena, Novalena, Rualena, and Melpomene. Annals of the Entomological Society of America, 35, 203 - 241." type="journal article" year="1942">Chamberlin &amp; Ivie 1942</bibRefCitation>
).
</paragraph>
<paragraph id="6C4136D3FF85FFFFE1F164E3FA1BF83F" blockId="4.[151,1437,1622,2007]" pageId="4" pageNumber="5">
<emphasis id="5E8AEAC1FF85FFFFE1F164E3FED9F95E" bold="true" box="[199,351,1694,1719]" pageId="4" pageNumber="5">Composition</emphasis>
. Fifty-three species:
<taxonomicName id="ABFE4D50FF85FFFFE36D64E2FB94F95E" authority="Gertsch &amp; Davis 1940" authorityName="Gertsch &amp; Davis" authorityYear="1940" box="[603,1042,1694,1719]" class="Arachnida" family="Agelenidae" genus="Agelena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="annamae">
<emphasis id="5E8AEAC1FF85FFFFE36D64E2FD60F95F" box="[603,742,1695,1718]" italics="true" pageId="4" pageNumber="5">N. annamae</emphasis>
(Gertsch &amp; Davis 1940)
</taxonomicName>
,
<taxonomicName id="ABFE4D50FF85FFFFE51564E2FF5CF932" authority="Gertsch &amp; Ivie 1936" authorityName="Gertsch &amp; Ivie" authorityYear="1936" class="Arachnida" family="Agelenidae" genus="Agelena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="approximata">
<emphasis id="5E8AEAC1FF85FFFFE51564E2FB50F95F" box="[1059,1238,1695,1718]" italics="true" pageId="4" pageNumber="5">N. approximata</emphasis>
(Gertsch &amp; Ivie 1936)
</taxonomicName>
,
<taxonomicName id="ABFE4D50FF85FFFFE1D064B9FD5BF932" authority="F.O. Pickard-Cambridge 1902" authorityName="F.O. Pickard-Cambridge" authorityYear="1902" box="[230,733,1730,1755]" class="Arachnida" family="Agelenidae" genus="Agelenopsis" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="attenuata">
<emphasis id="5E8AEAC1FF85FFFFE1D064B9FEF7F932" box="[230,369,1732,1755]" italics="true" pageId="4" pageNumber="5">N. attenuata</emphasis>
(F.O. Pickard-Cambridge 1902)
</taxonomicName>
,
<taxonomicName id="ABFE4D50FF85FFFFE3DE64B9FB8EF932" authority="Kraus 1955" authorityName="Kraus" authorityYear="1955" box="[744,1032,1730,1755]" class="Arachnida" family="Agelenidae" genus="Agelenopsis" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bipartita">
<emphasis id="5E8AEAC1FF85FFFFE3DE64B9FCEDF932" box="[744,875,1730,1755]" italics="true" pageId="4" pageNumber="5">N. bipartita</emphasis>
(Kraus 1955)
</taxonomicName>
,
<taxonomicName id="ABFE4D50FF85FFFFE52264B9FAAFF932" authority="Roth 1967" authorityName="Roth" authorityYear="1967" box="[1044,1321,1730,1755]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bipunctata" status="stat. nov.">
<emphasis id="5E8AEAC1FF85FFFFE52264B9FB2BF932" box="[1044,1197,1730,1755]" italics="true" pageId="4" pageNumber="5">N. bipunctata</emphasis>
<bibRefCitation id="086F4B22FF85FFFFE58264BFFAAFF932" author="Roth" box="[1204,1321,1730,1755]" pageId="4" pageNumber="5" refString="Roth, V. D. (1967) A review of the South American spiders of the family Agelenidae (Arachnida, Araneae). Bulletin of the American Museum of Natural History, 134, 297 - 346." type="journal article" year="1967">Roth 1967</bibRefCitation>
</taxonomicName>
<emphasis id="5E8AEAC1FF85FFFFE40664BFFA13F932" bold="true" box="[1328,1429,1730,1755]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="45B957BAFF85FFFFE40664BFFA13F932" box="[1328,1429,1730,1755]" pageId="4" pageNumber="5">stat. rev.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF85FFFFE1A1649AFDC5F916" authority="Chamberlin &amp; Ivie 1942" authorityName="Chamberlin &amp; Ivie" authorityYear="1942" box="[151,579,1766,1791]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="calavera">
<emphasis id="5E8AEAC1FF85FFFFE1A1649AFE9AF916" box="[151,284,1766,1791]" italics="true" pageId="4" pageNumber="5">N. calavera</emphasis>
<bibRefCitation id="086F4B22FF85FFFFE010649BFDC5F916" author="Chamberlin" box="[294,579,1766,1791]" pageId="4" pageNumber="5" refString="Chamberlin, R. V. &amp; Ivie, W. (1942) Agelenidae of the genera Hololena, Novalena, Rualena, and Melpomene. Annals of the Entomological Society of America, 35, 203 - 241." type="journal article" year="1942">Chamberlin &amp; Ivie 1942</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="ABFE4D50FF85FFFFE367649AFBBCF916" authority="F.O. Pickard-Cambridge 1902" authorityName="F.O. Pickard-Cambridge" authorityYear="1902" box="[593,1082,1766,1791]" class="Arachnida" family="Agelenidae" genus="Agelenopsis" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="costata">
<emphasis id="5E8AEAC1FF85FFFFE367649AFD43F917" box="[593,709,1767,1790]" italics="true" pageId="4" pageNumber="5">N. costata</emphasis>
(F.O. Pickard-Cambridge 1902)
</taxonomicName>
,
<taxonomicName id="ABFE4D50FF85FFFFE57E649AFEBFF8CA" authority="Chamberlin &amp; Gertsch 1930" authorityName="Chamberlin &amp; Gertsch" authorityYear="1930" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="5E8AEAC1FF85FFFFE57E649AFB63F916" box="[1096,1253,1766,1791]" italics="true" pageId="4" pageNumber="5">N. intermedia</emphasis>
(Chamberlin &amp; Gertsch 1930)
</taxonomicName>
,
<taxonomicName id="ABFE4D50FF85FFFFE0706571FDE4F8CA" authority="Kraus 1955" authorityName="Kraus" authorityYear="1955" box="[326,610,1802,1827]" class="Arachnida" family="Agelenidae" genus="Agelenopsis" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="laticava">
<emphasis id="5E8AEAC1FF85FFFFE0706571FE44F8CA" box="[326,450,1802,1827]" italics="true" pageId="4" pageNumber="5">N. laticava</emphasis>
(Kraus 1955)
</taxonomicName>
,
<taxonomicName id="ABFE4D50FF85FFFFE3586571FCF2F8CA" authority="Gertsch 1933" authorityName="Gertsch" authorityYear="1933" box="[622,884,1802,1827]" class="Arachnida" family="Agelenidae" genus="Agelena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="lutzi">
<emphasis id="5E8AEAC1FF85FFFFE3586571FD46F8CA" box="[622,704,1802,1827]" italics="true" pageId="4" pageNumber="5">N. lutzi</emphasis>
(Gertsch 1933)
</taxonomicName>
,
<taxonomicName id="ABFE4D50FF85FFFFE2B76571FB1DF8CA" authority="Banks 1898" authorityName="Banks" authorityYear="1898" box="[897,1179,1802,1827]" class="Arachnida" family="Agelenidae" genus="Agelena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="orizaba">
<emphasis id="5E8AEAC1FF85FFFFE2B76571FC7FF8CA" box="[897,1017,1802,1827]" italics="true" pageId="4" pageNumber="5">N. orizaba</emphasis>
(Banks 1898)
</taxonomicName>
,
<taxonomicName id="ABFE4D50FF85FFFFE59E6571FAB1F8CA" box="[1192,1335,1802,1827]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="shlomitae">
<emphasis id="5E8AEAC1FF85FFFFE59E6571FAB1F8CA" box="[1192,1335,1802,1827]" italics="true" pageId="4" pageNumber="5">N. shlomitae</emphasis>
</taxonomicName>
(García- Villafuerte 2009),
<taxonomicName id="ABFE4D50FF85FFFFE0526552FCC3F8AE" authority="F.O. Pickard-Cambridge 1902" authorityName="F.O. Pickard-Cambridge" authorityYear="1902" box="[356,837,1838,1863]" class="Arachnida" family="Agelenidae" genus="Rualena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="5E8AEAC1FF85FFFFE0526552FE5FF8AE" box="[356,473,1838,1863]" italics="true" pageId="4" pageNumber="5">N. simplex</emphasis>
(F.O. Pickard-Cambridge 1902)
</taxonomicName>
,
<taxonomicName id="ABFE4D50FF85FFFFE2666552FC3CF8AF" box="[848,954,1839,1862]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ajusco" status="sp. nov.">
<emphasis id="5E8AEAC1FF85FFFFE2666552FC3CF8AF" box="[848,954,1839,1862]" italics="true" pageId="4" pageNumber="5">N. ajusco</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF85FFFFE2F66553FB9FF8AE" bold="true" box="[960,1049,1838,1863]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="45B957BAFF85FFFFE2F66553FB9FF8AE" box="[960,1049,1838,1863]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF85FFFFE5116552FB0BF8AE" box="[1063,1165,1838,1863]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="alamo" status="sp. nov.">
<emphasis id="5E8AEAC1FF85FFFFE5116552FB0BF8AE" box="[1063,1165,1838,1863]" italics="true" pageId="4" pageNumber="5">N. alamo</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF85FFFFE5A26553FB6BF8AE" bold="true" box="[1172,1261,1838,1863]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="45B957BAFF85FFFFE5A26553FB6BF8AE" box="[1172,1261,1838,1863]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF85FFFFE5CC6552FAF5F8AE" box="[1274,1395,1838,1863]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="alvarezi" status="sp. nov.">
<emphasis id="5E8AEAC1FF85FFFFE5CC6552FAF5F8AE" box="[1274,1395,1838,1863]" italics="true" pageId="4" pageNumber="5">N. alvarezi</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF85FFFFE44C6553FF4EF882" bold="true" pageId="4" pageNumber="5">
<taxonomicNameLabel id="45B957BAFF85FFFFE44C6553FF4EF882" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF85FFFFE1E36529FEC9F882" box="[213,335,1874,1899]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="atzimbo" status="sp. nov.">
<emphasis id="5E8AEAC1FF85FFFFE1E36529FEC9F882" box="[213,335,1874,1899]" italics="true" pageId="4" pageNumber="5">N. atzimbo</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF85FFFFE063652FFE28F882" bold="true" box="[341,430,1874,1899]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="45B957BAFF85FFFFE063652FFE28F882" box="[341,430,1874,1899]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF85FFFFE08A6529FDDBF882" box="[444,605,1874,1899]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bosencheve" status="sp. nov.">
<emphasis id="5E8AEAC1FF85FFFFE08A6529FDDBF882" box="[444,605,1874,1899]" italics="true" pageId="4" pageNumber="5">N. bosencheve</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF85FFFFE352652FFD3BF882" bold="true" box="[612,701,1874,1899]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="45B957BAFF85FFFFE352652FFD3BF882" box="[612,701,1874,1899]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF85FFFFE3FD6529FCF4F882" box="[715,882,1874,1899]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="chamberlini" status="sp. nov.">
<emphasis id="5E8AEAC1FF85FFFFE3FD6529FCF4F882" box="[715,882,1874,1899]" italics="true" pageId="4" pageNumber="5">N. chamberlini</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF85FFFFE24F652FFC54F882" bold="true" box="[889,978,1874,1899]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="45B957BAFF85FFFFE24F652FFC54F882" box="[889,978,1874,1899]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF85FFFFE2D66529FBFBF882" box="[992,1149,1874,1899]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cieneguilla" status="sp. nov.">
<emphasis id="5E8AEAC1FF85FFFFE2D66529FBFBF882" box="[992,1149,1874,1899]" italics="true" pageId="4" pageNumber="5">N. cieneguilla</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF85FFFFE5B5652FFB5AF882" bold="true" box="[1155,1244,1874,1899]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="45B957BAFF85FFFFE5B5652FFB5AF882" box="[1155,1244,1874,1899]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF85FFFFE5DC6529FAF2F882" box="[1258,1396,1874,1899]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cintalapa" status="sp. nov.">
<emphasis id="5E8AEAC1FF85FFFFE5DC6529FAF2F882" box="[1258,1396,1874,1899]" italics="true" pageId="4" pageNumber="5">N. cintalapa</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF85FFFFE44C652FFF4EF866" bold="true" pageId="4" pageNumber="5">
<taxonomicNameLabel id="45B957BAFF85FFFFE44C652FFF4EF866" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF85FFFFE1EE650AFEB3F866" box="[216,309,1910,1935]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="clara" status="sp. nov.">
<emphasis id="5E8AEAC1FF85FFFFE1EE650AFEB3F866" box="[216,309,1910,1935]" italics="true" pageId="4" pageNumber="5">N. clara</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF85FFFFE008650BFE1FF866" bold="true" box="[318,409,1910,1935]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="45B957BAFF85FFFFE008650BFE1FF866" box="[318,409,1910,1935]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF85FFFFE09F650AFDCAF866" box="[425,588,1910,1935]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="comaltepec" status="sp. nov.">
<emphasis id="5E8AEAC1FF85FFFFE09F650AFDCAF866" box="[425,588,1910,1935]" italics="true" pageId="4" pageNumber="5">N. comaltepec</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF85FFFFE362650BFD29F866" bold="true" box="[596,687,1910,1935]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="45B957BAFF85FFFFE362650BFD29F866" box="[596,687,1910,1935]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF85FFFFE389650AFC9FF866" box="[703,793,1910,1935]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="creel" status="sp. nov.">
<emphasis id="5E8AEAC1FF85FFFFE389650AFC9FF866" box="[703,793,1910,1935]" italics="true" pageId="4" pageNumber="5">N. creel</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF85FFFFE217650BFCFBF866" bold="true" box="[801,893,1910,1935]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="45B957BAFF85FFFFE217650BFCFBF866" box="[801,893,1910,1935]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF85FFFFE2BA650AFB85F866" box="[908,1027,1910,1935]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dentata" status="sp. nov.">
<emphasis id="5E8AEAC1FF85FFFFE2BA650AFB85F866" box="[908,1027,1910,1935]" italics="true" pageId="4" pageNumber="5">N. dentata</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF85FFFFE53A650BFBE1F866" bold="true" box="[1036,1127,1910,1935]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="45B957BAFF85FFFFE53A650BFBE1F866" box="[1036,1127,1910,1935]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF85FFFFE541650AFA88F866" box="[1143,1294,1910,1935]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="divisadero" status="sp. nov.">
<emphasis id="5E8AEAC1FF85FFFFE541650AFA88F866" box="[1143,1294,1910,1935]" italics="true" pageId="4" pageNumber="5">N. divisadero</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF85FFFFE421650BFAF4F866" bold="true" box="[1303,1394,1910,1935]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="45B957BAFF85FFFFE421650BFAF4F866" box="[1303,1394,1910,1935]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF85FFFFE4B4650AFF70F85A" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="durango" status="sp. nov.">
<emphasis id="5E8AEAC1FF85FFFFE4B4650AFF70F85A" italics="true" pageId="4" pageNumber="5">N. durango</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF85FFFFE1CA65E7FED3F85A" bold="true" box="[252,341,1946,1971]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="45B957BAFF85FFFFE1CA65E7FED3F85A" box="[252,341,1946,1971]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF85FFFFE05565E1FE66F85A" box="[355,480,1946,1971]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="franckei" status="sp. nov.">
<emphasis id="5E8AEAC1FF85FFFFE05565E1FE66F85A" box="[355,480,1946,1971]" italics="true" pageId="4" pageNumber="5">N. franckei</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF85FFFFE0D065E7FDB9F85A" bold="true" box="[486,575,1946,1971]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="45B957BAFF85FFFFE0D065E7FDB9F85A" box="[486,575,1946,1971]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF85FFFFE37B65E1FD42F85A" box="[589,708,1948,1971]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="garnica" status="sp. nov.">
<emphasis id="5E8AEAC1FF85FFFFE37B65E1FD42F85A" box="[589,708,1948,1971]" italics="true" pageId="4" pageNumber="5">N. garnica</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF85FFFFE3FD65E7FCA2F85A" bold="true" box="[715,804,1946,1971]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="45B957BAFF85FFFFE3FD65E7FCA2F85A" box="[715,804,1946,1971]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF85FFFFE20465E1FC29F85A" box="[818,943,1946,1971]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="gibarrai" status="sp. nov.">
<emphasis id="5E8AEAC1FF85FFFFE20465E1FC29F85A" box="[818,943,1946,1971]" italics="true" pageId="4" pageNumber="5">N. gibarrai</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF85FFFFE28065E7FB89F85A" bold="true" box="[950,1039,1946,1971]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="45B957BAFF85FFFFE28065E7FB89F85A" box="[950,1039,1946,1971]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF85FFFFE52B65E1FBF1F85A" box="[1053,1143,1948,1971]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="irazu" status="sp. nov.">
<emphasis id="5E8AEAC1FF85FFFFE52B65E1FBF1F85A" box="[1053,1143,1948,1971]" italics="true" pageId="4" pageNumber="5">N. irazu</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF85FFFFE54B65E7FB50F85A" bold="true" box="[1149,1238,1946,1971]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="45B957BAFF85FFFFE54B65E7FB50F85A" box="[1149,1238,1946,1971]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF85FFFFE5D265E1FAB3F85A" box="[1252,1333,1948,1971]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="iviei" status="sp. nov.">
<emphasis id="5E8AEAC1FF85FFFFE5D265E1FAB3F85A" box="[1252,1333,1948,1971]" italics="true" pageId="4" pageNumber="5">N. iviei</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF85FFFFE40A65E7FA13F85A" bold="true" box="[1340,1429,1946,1971]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="45B957BAFF85FFFFE40A65E7FA13F85A" box="[1340,1429,1946,1971]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF85FFFFE1A165C2FF7CF83E" box="[151,250,1982,2007]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ixtlan" status="sp. nov.">
<emphasis id="5E8AEAC1FF85FFFFE1A165C2FF7CF83E" box="[151,250,1982,2007]" italics="true" pageId="4" pageNumber="5">N. ixtlan</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF85FFFFE03265C3FEE6F83E" bold="true" box="[260,352,1982,2007]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="45B957BAFF85FFFFE03265C3FEE6F83E" box="[260,352,1982,2007]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF85FFFFE04765C2FE7FF83E" box="[369,505,1982,2007]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="jiquilpan" status="sp. nov.">
<emphasis id="5E8AEAC1FF85FFFFE04765C2FE7FF83E" box="[369,505,1982,2007]" italics="true" pageId="4" pageNumber="5">N. jiquilpan</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF85FFFFE33265C3FDE6F83E" bold="true" box="[516,608,1982,2007]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="45B957BAFF85FFFFE33265C3FDE6F83E" box="[516,608,1982,2007]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF85FFFFE34765C2FD7AF83E" box="[625,764,1982,2007]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="leonensis" status="sp. nov.">
<emphasis id="5E8AEAC1FF85FFFFE34765C2FD7AF83E" box="[625,764,1982,2007]" italics="true" pageId="4" pageNumber="5">N. leonensis</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF85FFFFE23065C3FCE5F83E" bold="true" box="[774,867,1982,2007]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="45B957BAFF85FFFFE23065C3FCE5F83E" box="[774,867,1982,2007]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF85FFFFE24265C2FBA6F83F" box="[884,1056,1983,2006]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="mexiquensis" status="sp. nov.">
<emphasis id="5E8AEAC1FF85FFFFE24265C2FBA6F83F" box="[884,1056,1983,2006]" italics="true" pageId="4" pageNumber="5">N. mexiquensis</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF85FFFFE51C65C3FB00F83E" bold="true" box="[1066,1158,1982,2007]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="45B957BAFF85FFFFE51C65C3FB00F83E" box="[1066,1158,1982,2007]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF85FFFFE5A165C2FA8DF83F" box="[1175,1291,1983,2006]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="oaxaca" status="sp. nov.">
<emphasis id="5E8AEAC1FF85FFFFE5A165C2FA8DF83F" box="[1175,1291,1983,2006]" italics="true" pageId="4" pageNumber="5">N. oaxaca</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF85FFFFE42365C3FAF7F83E" bold="true" box="[1301,1393,1982,2007]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="45B957BAFF85FFFFE42365C3FAF7F83E" box="[1301,1393,1982,2007]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="5E8AEAC1FF85FFFFE4B465C2FA1BF83F" box="[1410,1437,1983,2006]" italics="true" pageId="4" pageNumber="5">N.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="24E46558FF84FFFEE1A162E4FB0EFF11" pageId="5" pageNumber="6" type="description">
<paragraph id="6C4136D3FF84FFFEE1A162E4FB0EFF11" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="6">
<taxonomicName id="ABFE4D50FF84FFFEE1A162E4FF78FF59" box="[151,254,153,176]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="paricutin" status="sp. nov.">
<emphasis id="5E8AEAC1FF84FFFEE1A162E4FF78FF59" box="[151,254,153,176]" italics="true" pageId="5" pageNumber="6">paricutin</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF84FFFEE03362EAFED9FF59" bold="true" box="[261,351,151,176]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="45B957BAFF84FFFEE03362EAFED9FF59" box="[261,351,151,176]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF84FFFEE05B62E4FE53FF59" box="[365,469,153,176]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="perote" status="sp. nov.">
<emphasis id="5E8AEAC1FF84FFFEE05B62E4FE53FF59" box="[365,469,153,176]" italics="true" pageId="5" pageNumber="6">N. perote</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF84FFFEE0EA62EAFDB0FF59" bold="true" box="[476,566,151,176]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="45B957BAFF84FFFEE0EA62EAFDB0FF59" box="[476,566,151,176]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF84FFFEE37362E4FD26FF59" box="[581,672,151,176]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="plata" status="sp. nov.">
<emphasis id="5E8AEAC1FF84FFFEE37362E4FD26FF59" box="[581,672,151,176]" italics="true" pageId="5" pageNumber="6">N. plata</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF84FFFEE39162EAFC87FF59" bold="true" box="[679,769,151,176]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="45B957BAFF84FFFEE39162EAFC87FF59" box="[679,769,151,176]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF84FFFEE22662E4FCFAFF59" box="[784,892,153,176]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="poncei" status="sp. nov.">
<emphasis id="5E8AEAC1FF84FFFEE22662E4FCFAFF59" box="[784,892,153,176]" italics="true" pageId="5" pageNumber="6">N. poncei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45B957BAFF84FFFEE2B562EAFC5BFF59" box="[899,989,151,176]" pageId="5" pageNumber="6" rank="species">
<emphasis id="5E8AEAC1FF84FFFEE2B562EAFC5BFF59" bold="true" box="[899,989,151,176]" pageId="5" pageNumber="6">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="ABFE4D50FF84FFFEE2DA62E4FBD9FF59" box="[1004,1119,153,176]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="popoca" status="sp. nov.">
<emphasis id="5E8AEAC1FF84FFFEE2DA62E4FBD9FF59" box="[1004,1119,153,176]" italics="true" pageId="5" pageNumber="6">N. popoca</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF84FFFEE55162EAFB47FF59" bold="true" box="[1127,1217,151,176]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="45B957BAFF84FFFEE55162EAFB47FF59" box="[1127,1217,151,176]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF84FFFEE5F962E4FAB2FF59" box="[1231,1332,153,176]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="prieta" status="sp. nov.">
<emphasis id="5E8AEAC1FF84FFFEE5F962E4FAB2FF59" box="[1231,1332,153,176]" italics="true" pageId="5" pageNumber="6">N. prieta</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF84FFFEE40D62EAFA13FF59" bold="true" box="[1339,1429,151,176]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="45B957BAFF84FFFEE40D62EAFA13FF59" box="[1339,1429,151,176]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF84FFFEE1A162C0FE83FF3C" box="[151,261,188,213]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="puebla" status="sp. nov.">
<emphasis id="5E8AEAC1FF84FFFEE1A162C0FE83FF3C" box="[151,261,188,213]" italics="true" pageId="5" pageNumber="6">N. puebla</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF84FFFEE03862C1FEEFFF3C" bold="true" box="[270,361,188,213]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="45B957BAFF84FFFEE03862C1FEEFFF3C" box="[270,361,188,213]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF84FFFEE04E62C0FE5DFF3D" box="[376,475,189,212]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="punta" status="sp. nov.">
<emphasis id="5E8AEAC1FF84FFFEE04E62C0FE5DFF3D" box="[376,475,189,212]" italics="true" pageId="5" pageNumber="6">N. punta</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF84FFFEE0D562C1FDB8FF3C" bold="true" box="[483,574,188,213]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="45B957BAFF84FFFEE0D562C1FDB8FF3C" box="[483,574,188,213]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF84FFFEE37B62C0FD22FF3C" box="[589,676,188,213]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="rothi" status="sp. nov.">
<emphasis id="5E8AEAC1FF84FFFEE37B62C0FD22FF3C" box="[589,676,188,213]" italics="true" pageId="5" pageNumber="6">N. rothi</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF84FFFEE39B62C1FC8EFF3C" bold="true" box="[685,776,188,213]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="45B957BAFF84FFFEE39B62C1FC8EFF3C" box="[685,776,188,213]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF84FFFEE22E62C0FC2EFF3C" box="[792,936,188,213]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="saltoensis" status="sp. nov.">
<emphasis id="5E8AEAC1FF84FFFEE22E62C0FC2EFF3C" box="[792,936,188,213]" italics="true" pageId="5" pageNumber="6">N. saltoensis</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF84FFFEE28762C1FB8AFF3C" bold="true" box="[945,1036,188,213]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="45B957BAFF84FFFEE28762C1FB8AFF3C" box="[945,1036,188,213]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF84FFFEE52A62C0FB17FF3C" box="[1052,1169,188,213]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sinaloa" status="sp. nov.">
<emphasis id="5E8AEAC1FF84FFFEE52A62C0FB17FF3C" box="[1052,1169,188,213]" italics="true" pageId="5" pageNumber="6">N. sinaloa</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF84FFFEE5AF62C1FB72FF3C" bold="true" box="[1177,1268,188,213]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="45B957BAFF84FFFEE5AF62C1FB72FF3C" box="[1177,1268,188,213]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF84FFFEE43262C0FAF4FF3D" box="[1284,1394,189,212]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="tacana" status="sp. nov.">
<emphasis id="5E8AEAC1FF84FFFEE43262C0FAF4FF3D" box="[1284,1394,189,212]" italics="true" pageId="5" pageNumber="6">N. tacana</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF84FFFEE44C62C1FF4EFF11" bold="true" pageId="5" pageNumber="6">
<taxonomicNameLabel id="45B957BAFF84FFFEE44C62C1FF4EFF11" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF84FFFEE1E0629CFEC4FF11" box="[214,322,224,248]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="triunfo" status="sp. nov.">
<emphasis id="5E8AEAC1FF84FFFEE1E0629CFEC4FF11" box="[214,322,224,248]" italics="true" pageId="5" pageNumber="6">N. triunfo</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF84FFFEE07C62A2FE25FF11" bold="true" box="[330,419,223,248]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="45B957BAFF84FFFEE07C62A2FE25FF11" box="[330,419,223,248]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF84FFFEE087629CFDA7FF11" box="[433,545,223,248]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="valdezi" status="sp. nov.">
<emphasis id="5E8AEAC1FF84FFFEE087629CFDA7FF11" box="[433,545,223,248]" italics="true" pageId="5" pageNumber="6">N. valdezi</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF84FFFEE31E62A2FD07FF11" bold="true" box="[552,641,223,248]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="45B957BAFF84FFFEE31E62A2FD07FF11" box="[552,641,223,248]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ABFE4D50FF84FFFEE3A6629CFC81FF11" box="[656,775,225,248]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="victoria" status="sp. nov.">
<emphasis id="5E8AEAC1FF84FFFEE3A6629CFC81FF11" box="[656,775,225,248]" italics="true" pageId="5" pageNumber="6">N. victoria</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF84FFFEE23862A2FCEEFF11" bold="true" box="[782,872,223,248]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="45B957BAFF84FFFEE23862A2FCEEFF11" box="[782,872,223,248]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="ABFE4D50FF84FFFEE293629CFBAEFF11" box="[933,1064,223,248]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="volcanes" status="sp. nov.">
<emphasis id="5E8AEAC1FF84FFFEE293629CFBAEFF11" box="[933,1064,223,248]" italics="true" pageId="5" pageNumber="6">N. volcanes</emphasis>
</taxonomicName>
<emphasis id="5E8AEAC1FF84FFFEE51962A2FB0EFF11" bold="true" box="[1071,1160,223,248]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="45B957BAFF84FFFEE51962A2FB0EFF11" box="[1071,1160,223,248]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="24E46558FF84FFFEE1F16379FB02FD41" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="6C4136D3FF84FFFEE1F16379FB02FD41" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="6">
<emphasis id="5E8AEAC1FF84FFFEE1F16379FEBCFEF4" bold="true" box="[199,314,260,285]" pageId="5" pageNumber="6">Diagnosis</emphasis>
. The genus is diagnosed by the following characteristics in combination: the male pedipalp with a short and simple curved embolus (
<figureCitation id="F4C52A56FF84FFFEE321635AFDE2FEA9" box="[535,612,295,320]" captionStart="FIGURE 3" captionStartId="12.[151,250,1065,1087]" captionTargetBox="[211,1375,196,1039]" captionTargetId="figure@12.[208,1379,193,1044]" captionTargetPageId="12" captionText="FIGURE 3. Novalena approximata (Gertsch &amp; Ivie 1936), male pedipalp (SEM): a, ventral view; b, prolateral view; c, d, retrolateral view. Abbreviations: Cd, dorsal projection of conductor; Cv, ventral projection of conductor; E, embolus; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; St, subtegulum; T, tegulum; TMP, tegular median process." httpUri="https://zenodo.org/record/571686/files/figure.png" pageId="5" pageNumber="6">Figs. 3</figureCitation>
b, 13a, 18a, 32a, 43d); conductor with two projections (
<figureCitation id="F4C52A56FF84FFFEE5F9635AFA9AFEA9" box="[1231,1308,295,320]" captionStart="FIGURE 3" captionStartId="12.[151,250,1065,1087]" captionTargetBox="[211,1375,196,1039]" captionTargetId="figure@12.[208,1379,193,1044]" captionTargetPageId="12" captionText="FIGURE 3. Novalena approximata (Gertsch &amp; Ivie 1936), male pedipalp (SEM): a, ventral view; b, prolateral view; c, d, retrolateral view. Abbreviations: Cd, dorsal projection of conductor; Cv, ventral projection of conductor; E, embolus; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; St, subtegulum; T, tegulum; TMP, tegular median process." httpUri="https://zenodo.org/record/571686/files/figure.png" pageId="5" pageNumber="6">Figs. 3</figureCitation>
c, 10a, 14c, 27a, 41f); tegular median process present (
<figureCitation id="F4C52A56FF84FFFEE34A6331FD5CFE8D" box="[636,730,332,357]" captionStart="FIGURE 10" captionStartId="22.[151,250,1507,1529]" captionTargetBox="[209,1373,205,1475]" captionTargetId="figure@22.[196,1390,193,1486]" captionTargetPageId="22" captionText="FIGURE 10. Novalena intermedia (Chamberlin &amp; Gertsch 1930) (SEM): a, c, e, male pedipalp; b, d, f, internal female genitalia. a, c, ventral view; b, d, dorsal view; e, prolateral view, arrow to prolateral protuberance; f, anterior view. Abbreviations: CD, copulatory ducts; Cd, dorsal projection of conductor; Cv, ventral projection of conductor; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process." httpUri="https://zenodo.org/record/571693/files/figure.png" pageId="5" pageNumber="6">Figs. 10</figureCitation>
a, 13a, 26a, 34d, 38a); RTA with distal and dorsal projections, ventral projection usually present (
<figureCitation id="F4C52A56FF84FFFEE30B6312FD14FE61" box="[573,658,367,392]" captionStart="FIGURE 3" captionStartId="12.[151,250,1065,1087]" captionTargetBox="[211,1375,196,1039]" captionTargetId="figure@12.[208,1379,193,1044]" captionTargetPageId="12" captionText="FIGURE 3. Novalena approximata (Gertsch &amp; Ivie 1936), male pedipalp (SEM): a, ventral view; b, prolateral view; c, d, retrolateral view. Abbreviations: Cd, dorsal projection of conductor; Cv, ventral projection of conductor; E, embolus; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; St, subtegulum; T, tegulum; TMP, tegular median process." httpUri="https://zenodo.org/record/571686/files/figure.png" pageId="5" pageNumber="6">Figs. 3</figureCitation>
d, 11b,
<quantity id="AB069B36FF84FFFEE3C06312FCACFE61" box="[758,810,367,392]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="6" unit="m" value="16.0">16m</quantity>
, 30e, 33e) (absent in
<taxonomicName id="ABFE4D50FF84FFFEE574630CFB51FE61" box="[1090,1239,367,392]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="shlomitae">
<emphasis id="5E8AEAC1FF84FFFEE574630CFB51FE61" box="[1090,1239,367,392]" italics="true" pageId="5" pageNumber="6">N. shlomitae</emphasis>
</taxonomicName>
,
<taxonomicName id="ABFE4D50FF84FFFEE5D8630CFAEDFE61" box="[1262,1387,367,392]" class="Arachnida" family="Agelenidae" genus="Rualena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="5E8AEAC1FF84FFFEE5D8630CFAEDFE61" box="[1262,1387,367,392]" italics="true" pageId="5" pageNumber="6">N. simplex</emphasis>
</taxonomicName>
,
<taxonomicName id="ABFE4D50FF84FFFEE4B4630CFE98FE44" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="chamberlini">
<emphasis id="5E8AEAC1FF84FFFEE4B4630CFE98FE44" italics="true" pageId="5" pageNumber="6">N. chamberlini</emphasis>
</taxonomicName>
,
<taxonomicName id="ABFE4D50FF84FFFEE01B63E8FE3FFE44" box="[301,441,404,429]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="cintalapa">
<emphasis id="5E8AEAC1FF84FFFEE01B63E8FE3FFE44" box="[301,441,404,429]" italics="true" pageId="5" pageNumber="6">N. cintalapa</emphasis>
</taxonomicName>
,
<taxonomicName id="ABFE4D50FF84FFFEE0F163E8FDC0FE44" box="[455,582,404,429]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="gibarrai">
<emphasis id="5E8AEAC1FF84FFFEE0F163E8FDC0FE44" box="[455,582,404,429]" italics="true" pageId="5" pageNumber="6">N. gibarrai</emphasis>
</taxonomicName>
,
<taxonomicName id="ABFE4D50FF84FFFEE36363E8FD4EFE45" box="[597,712,405,428]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="oaxaca">
<emphasis id="5E8AEAC1FF84FFFEE36363E8FD4EFE45" box="[597,712,405,428]" italics="true" pageId="5" pageNumber="6">N. oaxaca</emphasis>
</taxonomicName>
, and
<taxonomicName id="ABFE4D50FF84FFFEE23163E8FCF3FE45" box="[775,885,405,428]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="tacana">
<emphasis id="5E8AEAC1FF84FFFEE23163E8FCF3FE45" box="[775,885,405,428]" italics="true" pageId="5" pageNumber="6">N. tacana</emphasis>
</taxonomicName>
,
<figureCitation id="F4C52A56FF84FFFEE2B263E9FC59FE44" box="[900,991,404,429]" captionStart="FIGURE 15" captionStartId="29.[151,250,1265,1287]" captionTargetBox="[161,1431,449,1235]" captionTargetId="figure@29.[151,1435,445,1243]" captionTargetPageId="29" captionText="FIGURE 15. Novalena simplex (F. O. Pickard-Cambridge 1902): a, male pedipalp, ventral view; b, same, retrolateral view; c, epigynum, ventral view; d, internal genitalia, dorsal view; e, epigynum, posterior view. Abbreviations: A, atrium; C, conductor; CD, copulatory ducts; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process. Scale bars: a = 0.5 mm; e = 0.25 mm." httpUri="https://zenodo.org/record/571699/files/figure.png" pageId="5" pageNumber="6">Figs. 15</figureCitation>
b, 18b, 32b); atrium posteriorly located on the epigynal plate, wider than long (
<figureCitation id="F4C52A56FF84FFFEE36563CAFD26FE39" box="[595,672,439,464]" captionStart="FIGURE 5" captionStartId="14.[151,250,1484,1506]" captionTargetBox="[161,1424,865,1453]" captionTargetId="figure@14.[150,1437,860,1463]" captionTargetPageId="14" captionText="FIGURE 5. Novalena spp.: a – c, Novalena attenuata (F. O. Pickard-Cambridge 1902); d – f, Novalena bipunctata Roth 1967 stat. rev. a, d, epigynum, ventral view; b, e, internal genitalia, dorsal view; c, f, epigynum, posterior view. Abbreviations: A, atrium; CD, copulatory ducts; CO, copulatory openings; FD, fertilization ducts; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae. Dashed line indicates distance between anterior and posterior margins. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/571688/files/figure.png" pageId="5" pageNumber="6">Figs. 5</figureCitation>
a, 13c, 19c, 24d, 40a), and usually lacking a division (if divided [
<taxonomicName id="ABFE4D50FF84FFFEE4B463C4FF7FFE1C" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="calavera">
<emphasis id="5E8AEAC1FF84FFFEE4B463C4FF7FFE1C" italics="true" pageId="5" pageNumber="6">N. calavera</emphasis>
</taxonomicName>
and
<taxonomicName id="ABFE4D50FF84FFFEE01963A0FEF9FE1C" box="[303,383,476,501]" class="Arachnida" family="Agelenidae" genus="Agelena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="lutzi">
<emphasis id="5E8AEAC1FF84FFFEE01963A0FEF9FE1C" box="[303,383,476,501]" italics="true" pageId="5" pageNumber="6">N. lutzi</emphasis>
</taxonomicName>
] then copulatory openings visible in posterior view,
<figureCitation id="F4C52A56FF84FFFEE2FD63A1FB9EFE1D" box="[971,1048,476,501]" captionStart="FIGURE 7" captionStartId="17.[151,250,1332,1354]" captionTargetBox="[160,1423,201,1300]" captionTargetId="figure@17.[150,1436,193,1310]" captionTargetPageId="17" captionText="FIGURE 7. Novalena spp. genitalia: a – e, Novalena calavera Chamberlin &amp; Ivie 1942; f – h, Novalena costata (F. O. Pickard- Cambridge 1902). a, male pedipalp, ventral view; b, same, retrolateral view; c, f, epigynum, ventral view; d, g, internal genitalia, dorsal view; e, h, epigynum, posterior view. Abbreviations: A, atrium; C, conductor; CD, copulatory ducts; CO, copulatory openings; E, embolus; FD, fertilization ducts; H, hoods; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process. Scale bars: a = 1 mm; e, h = 0.5 mm." httpUri="https://zenodo.org/record/571690/files/figure.png" pageId="5" pageNumber="6">Figs. 7</figureCitation>
e, 12e); atrial spurs usually present (
<figureCitation id="F4C52A56FF84FFFEE1A96382FF6AFDF1" box="[159,236,511,536]" captionStart="FIGURE 5" captionStartId="14.[151,250,1484,1506]" captionTargetBox="[161,1424,865,1453]" captionTargetId="figure@14.[150,1437,860,1463]" captionTargetPageId="14" captionText="FIGURE 5. Novalena spp.: a – c, Novalena attenuata (F. O. Pickard-Cambridge 1902); d – f, Novalena bipunctata Roth 1967 stat. rev. a, d, epigynum, ventral view; b, e, internal genitalia, dorsal view; c, f, epigynum, posterior view. Abbreviations: A, atrium; CD, copulatory ducts; CO, copulatory openings; FD, fertilization ducts; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae. Dashed line indicates distance between anterior and posterior margins. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/571688/files/figure.png" pageId="5" pageNumber="6">Figs. 5</figureCitation>
d, 15c, 24a, 33a, 42c) (absent in
<taxonomicName id="ABFE4D50FF84FFFEE363607CFD51FDF1" box="[597,727,511,536]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="calavera">
<emphasis id="5E8AEAC1FF84FFFEE363607CFD51FDF1" box="[597,727,511,536]" italics="true" pageId="5" pageNumber="6">N. calavera</emphasis>
</taxonomicName>
,
<taxonomicName id="ABFE4D50FF84FFFEE3D3607CFCB3FDF1" box="[741,821,511,536]" class="Arachnida" family="Agelenidae" genus="Agelena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="lutzi">
<emphasis id="5E8AEAC1FF84FFFEE3D3607CFCB3FDF1" box="[741,821,511,536]" italics="true" pageId="5" pageNumber="6">N. lutzi</emphasis>
</taxonomicName>
, and
<taxonomicName id="ABFE4D50FF84FFFEE244607CFC6FFDF1" box="[882,1001,513,536]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="garnica">
<emphasis id="5E8AEAC1FF84FFFEE244607CFC6FFDF1" box="[882,1001,513,536]" italics="true" pageId="5" pageNumber="6">N. garnica</emphasis>
</taxonomicName>
,
<figureCitation id="F4C52A56FF84FFFEE2C16382FBC2FDFE" box="[1015,1092,511,536]" captionStart="FIGURE 7" captionStartId="17.[151,250,1332,1354]" captionTargetBox="[160,1423,201,1300]" captionTargetId="figure@17.[150,1436,193,1310]" captionTargetPageId="17" captionText="FIGURE 7. Novalena spp. genitalia: a – e, Novalena calavera Chamberlin &amp; Ivie 1942; f – h, Novalena costata (F. O. Pickard- Cambridge 1902). a, male pedipalp, ventral view; b, same, retrolateral view; c, f, epigynum, ventral view; d, g, internal genitalia, dorsal view; e, h, epigynum, posterior view. Abbreviations: A, atrium; C, conductor; CD, copulatory ducts; CO, copulatory openings; E, embolus; FD, fertilization ducts; H, hoods; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process. Scale bars: a = 1 mm; e, h = 0.5 mm." httpUri="https://zenodo.org/record/571690/files/figure.png" pageId="5" pageNumber="6">Figs. 7</figureCitation>
c, 12c, 26c); short and strongly sclerotized copulatory ducts, straight or curved (
<figureCitation id="F4C52A56FF84FFFEE3996059FD7AFDD4" box="[687,764,548,573]" captionStart="FIGURE 5" captionStartId="14.[151,250,1484,1506]" captionTargetBox="[161,1424,865,1453]" captionTargetId="figure@14.[150,1437,860,1463]" captionTargetPageId="14" captionText="FIGURE 5. Novalena spp.: a – c, Novalena attenuata (F. O. Pickard-Cambridge 1902); d – f, Novalena bipunctata Roth 1967 stat. rev. a, d, epigynum, ventral view; b, e, internal genitalia, dorsal view; c, f, epigynum, posterior view. Abbreviations: A, atrium; CD, copulatory ducts; CO, copulatory openings; FD, fertilization ducts; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae. Dashed line indicates distance between anterior and posterior margins. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/571688/files/figure.png" pageId="5" pageNumber="6">Figs. 5</figureCitation>
b, 14d, 20b, 36a, 38d), connected ectally to longer than wide primary spermathecae (
<figureCitation id="F4C52A56FF84FFFEE09A603AFD86FD89" box="[428,512,583,608]" captionStart="FIGURE 5" captionStartId="14.[151,250,1484,1506]" captionTargetBox="[161,1424,865,1453]" captionTargetId="figure@14.[150,1437,860,1463]" captionTargetPageId="14" captionText="FIGURE 5. Novalena spp.: a – c, Novalena attenuata (F. O. Pickard-Cambridge 1902); d – f, Novalena bipunctata Roth 1967 stat. rev. a, d, epigynum, ventral view; b, e, internal genitalia, dorsal view; c, f, epigynum, posterior view. Abbreviations: A, atrium; CD, copulatory ducts; CO, copulatory openings; FD, fertilization ducts; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae. Dashed line indicates distance between anterior and posterior margins. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/571688/files/figure.png" pageId="5" pageNumber="6">Figs. 5</figureCitation>
e, 10b, 15d, 25d, 31d); secondary spermathecae blind-ended receptacles with primary pores and each connected to the union of copulatory duct and primary spermathecae by short diverticulum (
<figureCitation id="F4C52A56FF84FFFEE1A960F2FF7CFD41" box="[159,250,655,680]" captionStart="FIGURE 10" captionStartId="22.[151,250,1507,1529]" captionTargetBox="[209,1373,205,1475]" captionTargetId="figure@22.[196,1390,193,1486]" captionTargetPageId="22" captionText="FIGURE 10. Novalena intermedia (Chamberlin &amp; Gertsch 1930) (SEM): a, c, e, male pedipalp; b, d, f, internal female genitalia. a, c, ventral view; b, d, dorsal view; e, prolateral view, arrow to prolateral protuberance; f, anterior view. Abbreviations: CD, copulatory ducts; Cd, dorsal projection of conductor; Cv, ventral projection of conductor; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process." httpUri="https://zenodo.org/record/571693/files/figure.png" pageId="5" pageNumber="6">Figs. 10</figureCitation>
f, 14f,
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, 36b, 44e); and short fertilization ducts (
<figureCitation id="F4C52A56FF84FFFEE20E60F2FC00FD41" box="[824,902,655,680]" captionStart="FIGURE 2" captionStartId="10.[151,250,1518,1540]" captionTargetBox="[156,1432,687,1497]" captionTargetId="figure@10.[152,1438,681,1505]" captionTargetPageId="10" captionText="FIGURE 2. Novalena annamae (Gertsch &amp; Davis 1940): a, male pedipalp, ventral view; b, same, retrolateral view; c, epigynum, ventral view; d, internal genitalia, dorsal view; e, epigynum, posterior view. Abbreviations: A, atrium; C, conductor; CD, copulatory ducts; CO, copulatory openings; E, embolus; F, fulcrum; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process. Scale bars: a = 0.5 mm; e = 0.25 mm." httpUri="https://zenodo.org/record/571685/files/figure.png" pageId="5" pageNumber="6">Figs. 2</figureCitation>
d, 13d, 22d, 31d, 39d).
</paragraph>
</subSubSection>
<subSubSection id="24E46558FF84FFFEE1F160C9FD74FB49" pageId="5" pageNumber="6" type="reference_group">
<paragraph id="6C4136D3FF84FFFEE1F160C9FD74FB49" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="6">
<taxonomicName id="ABFE4D50FF84FFFEE1F160C9FEB5FD24" ID-CoL="65NL" box="[199,307,692,717]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="5E8AEAC1FF84FFFEE1F160C9FEB5FD24" box="[199,307,692,717]" italics="true" pageId="5" pageNumber="6">Novalena</emphasis>
</taxonomicName>
differs from
<taxonomicName id="ABFE4D50FF84FFFEE0EB60C8FCCFFD25" ID-CoL="633YC" authority="Bolzern et al. 2013" authorityName="Bolzern et al." authorityYear="2013" box="[477,841,692,717]" class="Arachnida" family="Agelenidae" genus="Eratigena" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="5E8AEAC1FF84FFFEE0EB60C8FDCBFD25" box="[477,589,693,716]" italics="true" pageId="5" pageNumber="6">Eratigena</emphasis>
Bolzern
<emphasis id="5E8AEAC1FF84FFFEE3F460C8FD7CFD24" box="[706,762,692,717]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
. 2013
</taxonomicName>
and
<taxonomicName id="ABFE4D50FF84FFFEE2B860C8FB35FD25" ID-CoL="645LS" authority="Latreille 1804" authorityName="Latreille" authorityYear="1804" box="[910,1203,692,717]" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="5E8AEAC1FF84FFFEE2B860C8FC79FD25" box="[910,1023,693,716]" italics="true" pageId="5" pageNumber="6">Tegenaria</emphasis>
Latreille
<quantity id="AB069B36FF84FFFEE54D60C9FB35FD25" box="[1147,1203,692,716]" metricMagnitude="1" metricUnit="m" metricValue="4.58216" pageId="5" pageNumber="6" unit="in" value="1804.0">1804</quantity>
</taxonomicName>
in having strongly procurved eye rows in frontal view; from
<taxonomicName id="ABFE4D50FF84FFFEE3AA60AAFC43FD19" ID-CoL="QNM" authority="Giebel 1869" authorityName="Giebel" authorityYear="1869" box="[668,965,727,752]" class="Arachnida" family="Agelenidae" genus="Agelenopsis" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="5E8AEAC1FF84FFFEE3AA60AAFCA5FD19" box="[668,803,727,752]" italics="true" pageId="5" pageNumber="6">Agelenopsis</emphasis>
Giebel 1869
</taxonomicName>
,
<taxonomicName id="ABFE4D50FF84FFFEE2E060A4FA11FD19" ID-CoL="387T" authority="Chamberlin &amp; Ivie 1941" authorityName="Chamberlin &amp; Ivie" authorityYear="1941" box="[982,1431,727,752]" class="Arachnida" family="Agelenidae" genus="Barronopsis" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="5E8AEAC1FF84FFFEE2E060A4FBD9FD19" box="[982,1119,729,752]" italics="true" pageId="5" pageNumber="6">Barronopsis</emphasis>
<bibRefCitation id="086F4B22FF84FFFEE55860AAFA11FD19" author="Chamberlin" box="[1134,1431,727,752]" pageId="5" pageNumber="6" refString="Chamberlin, R. V. &amp; Ivie, W. (1941) North American Agelenidae of the genera Agelenopsis, Calilena, Ritalena, and Tortolena. Annals of the Entomological Society of America, 34, 585 - 628." type="journal article" year="1941">Chamberlin &amp; Ivie 1941</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="ABFE4D50FF84FFFEE1A16081FE9CFCFC" ID-CoL="5NWP" box="[151,282,764,789]" class="Arachnida" family="Agelenidae" genus="Melpomene" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="5E8AEAC1FF84FFFEE1A16081FE9CFCFC" box="[151,282,764,789]" italics="true" pageId="5" pageNumber="6">Melpomene</emphasis>
</taxonomicName>
, and
<taxonomicName id="ABFE4D50FF84FFFEE06D6081FD6FFCFD" ID-CoL="7XWX" authority="Chamberlin &amp; Ivie 1941" authorityName="Chamberlin &amp; Ivie" authorityYear="1941" box="[347,745,764,789]" class="Arachnida" family="Agelenidae" genus="Tortolena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="5E8AEAC1FF84FFFEE06D6081FE42FCFC" box="[347,452,764,789]" italics="true" pageId="5" pageNumber="6">Tortolena</emphasis>
<bibRefCitation id="086F4B22FF84FFFEE0E66081FD6FFCFD" author="Chamberlin" box="[464,745,764,789]" pageId="5" pageNumber="6" refString="Chamberlin, R. V. &amp; Ivie, W. (1941) North American Agelenidae of the genera Agelenopsis, Calilena, Ritalena, and Tortolena. Annals of the Entomological Society of America, 34, 585 - 628." type="journal article" year="1941">
Chamberlin &amp; Ivie
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</bibRefCitation>
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in lacking both a strongly modified embolus and projecting tegular lateral process in males (= tegular apophysis, Stocks 2009: fig. 39), and lacking long, membranous and plicate copulatory ducts in females (as in Stocks 2009: figs. 52, 61); from
<taxonomicName id="ABFE4D50FF84FFFEE2C16139FAFBFCB5" ID-CoL="3F9H" authority="Chamberlin &amp; Ivie 1941" authorityName="Chamberlin &amp; Ivie" authorityYear="1941" box="[1015,1405,836,861]" class="Arachnida" family="Agelenidae" genus="Calilena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="5E8AEAC1FF84FFFEE2C16139FBDEFCB4" box="[1015,1112,836,861]" italics="true" pageId="5" pageNumber="6">Calilena</emphasis>
<bibRefCitation id="086F4B22FF84FFFEE5546139FAFBFCB5" author="Chamberlin" box="[1122,1405,836,861]" pageId="5" pageNumber="6" refString="Chamberlin, R. V. &amp; Ivie, W. (1941) North American Agelenidae of the genera Agelenopsis, Calilena, Ritalena, and Tortolena. Annals of the Entomological Society of America, 34, 585 - 628." type="journal article" year="1941">
Chamberlin &amp; Ivie
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</bibRefCitation>
</taxonomicName>
in lacking a membranous fulcrum (as in
<bibRefCitation id="086F4B22FF84FFFEE375611AFCDAFC69" author="Chamberlin" box="[579,860,871,896]" pageId="5" pageNumber="6" refString="Chamberlin, R. V. &amp; Ivie, W. (1941) North American Agelenidae of the genera Agelenopsis, Calilena, Ritalena, and Tortolena. Annals of the Entomological Society of America, 34, 585 - 628." type="journal article" year="1941">Chamberlin &amp; Ivie 1941</bibRefCitation>
: figs. 68, 70, 74) (only present in
<taxonomicName id="ABFE4D50FF84FFFEE5ED6114FAE5FC69" box="[1243,1379,873,896]" class="Arachnida" family="Agelenidae" genus="Agelena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="annamae">
<emphasis id="5E8AEAC1FF84FFFEE5ED6114FAE5FC69" box="[1243,1379,873,896]" italics="true" pageId="5" pageNumber="6">N. annamae</emphasis>
</taxonomicName>
) and an epigynal scape (as in
<bibRefCitation id="086F4B22FF84FFFEE08561F1FD56FC4D" author="Chamberlin" box="[435,720,908,933]" pageId="5" pageNumber="6" refString="Chamberlin, R. V. &amp; Ivie, W. (1941) North American Agelenidae of the genera Agelenopsis, Calilena, Ritalena, and Tortolena. Annals of the Entomological Society of America, 34, 585 - 628." type="journal article" year="1941">Chamberlin &amp; Ivie 1941</bibRefCitation>
: figs. 48–60); from
<taxonomicName id="ABFE4D50FF84FFFEE28361F1FAFBFC4D" ID-CoL="4XZC" authority="Chamberlin &amp; Gertsch 1929" authorityName="Chamberlin &amp; Gertsch" authorityYear="1929" box="[949,1405,908,933]" class="Arachnida" family="Agelenidae" genus="Hololena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="5E8AEAC1FF84FFFEE28361F1FB9BFC4C" box="[949,1053,908,933]" italics="true" pageId="5" pageNumber="6">Hololena</emphasis>
(Chamberlin &amp; Gertsch 1929)
</taxonomicName>
in lacking a fulcrum supporting the embolus (as in
<bibRefCitation id="086F4B22FF84FFFEE38E61D2FC56FC21" author="Chamberlin" box="[696,976,943,968]" pageId="5" pageNumber="6" refString="Chamberlin, R. V. &amp; Ivie, W. (1942) Agelenidae of the genera Hololena, Novalena, Rualena, and Melpomene. Annals of the Entomological Society of America, 35, 203 - 241." type="journal article" year="1942">Chamberlin &amp; Ivie 1942</bibRefCitation>
: figs. 6, 11, 12) and an anteriorly located epigynal atrium (as in
<bibRefCitation id="086F4B22FF84FFFEE0A461A9FD2EFC05" author="Chamberlin" box="[402,680,980,1005]" pageId="5" pageNumber="6" refString="Chamberlin, R. V. &amp; Ivie, W. (1942) Agelenidae of the genera Hololena, Novalena, Rualena, and Melpomene. Annals of the Entomological Society of America, 35, 203 - 241." type="journal article" year="1942">Chamberlin &amp; Ivie 1942</bibRefCitation>
: fig. 10); from
<taxonomicName id="ABFE4D50FF84FFFEE26661A9FAA0FC05" ID-CoL="643WZ" authority="Maya-Morales &amp; Jimenez 2013" authorityName="Maya-Morales &amp; Jimenez" authorityYear="2013" box="[848,1318,980,1005]" class="Arachnida" family="Agelenidae" genus="Rothilena" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="5E8AEAC1FF84FFFEE26661A9FC3BFC04" box="[848,957,980,1005]" italics="true" pageId="5" pageNumber="6">Rothilena</emphasis>
<bibRefCitation id="086F4B22FF84FFFEE2F261A9FAA0FC05" author="Maya-Morales" box="[964,1318,980,1005]" pageId="5" pageNumber="6" refString="Maya-Morales, J. &amp; Jimenez, M. L. (2013) Rothilena (Araneae: Agelenidae), a new genus of funnel-web spiders endemic to the Baja California Peninsula, Mexico. Zootaxa, 3718 (5), 441 - 466." type="journal article" year="2013">
Maya-Morales &amp; Jiménez
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</bibRefCitation>
</taxonomicName>
in lacking three projections on conductor (as in
<bibRefCitation id="086F4B22FF84FFFEE30C618AFC24FBF9" author="Maya-Morales" box="[570,930,1015,1040]" pageId="5" pageNumber="6" refString="Maya-Morales, J. &amp; Jimenez, M. L. (2013) Rothilena (Araneae: Agelenidae), a new genus of funnel-web spiders endemic to the Baja California Peninsula, Mexico. Zootaxa, 3718 (5), 441 - 466." type="journal article" year="2013">Maya-Morales &amp; Jiménez 2013</bibRefCitation>
: fig. 18) and spherical primary spermathecae (as in
<bibRefCitation id="086F4B22FF84FFFEE1D56661FDD3FBDD" author="Maya-Morales" box="[227,597,1052,1077]" pageId="5" pageNumber="6" refString="Maya-Morales, J. &amp; Jimenez, M. L. (2013) Rothilena (Araneae: Agelenidae), a new genus of funnel-web spiders endemic to the Baja California Peninsula, Mexico. Zootaxa, 3718 (5), 441 - 466." type="journal article" year="2013">Maya-Morales &amp; Jiménez 2013</bibRefCitation>
: fig. 29); from
<taxonomicName id="ABFE4D50FF84FFFEE23A6661FCEFFBDC" ID-CoL="6449H" box="[780,873,1052,1077]" class="Arachnida" family="Agelenidae" genus="Rualena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="5E8AEAC1FF84FFFEE23A6661FCEFFBDC" box="[780,873,1052,1077]" italics="true" pageId="5" pageNumber="6">Rualena</emphasis>
</taxonomicName>
in lacking a membranous fulcrum (as in Maya-
<bibRefCitation id="086F4B22FF84FFFEE1A16642FE3CFBB1" author="Morales" box="[151,442,1087,1112]" pageId="5" pageNumber="6" refString="Maya-Morales, J. &amp; Jimenez, M. L. (2016) Taxonomic revision of the spider genus Rualena Chamberlin &amp; Ivie 1942 and description of Hoffmannilena, a new genus from Mexico (Araneae: Agelenidae). Zootaxa, 4084 (1), 1 - 49. https: // doi. org / 10.11646 / zootaxa. 4084.1.1" type="journal article" year="2016">Morales &amp; Jiménez 2016</bibRefCitation>
: fig. 43) and an atrium as long as wide (as in
<bibRefCitation id="086F4B22FF84FFFEE2FE6642FAB2FBB1" author="Maya-Morales" box="[968,1332,1087,1112]" pageId="5" pageNumber="6" refString="Maya-Morales, J. &amp; Jimenez, M. L. (2016) Taxonomic revision of the spider genus Rualena Chamberlin &amp; Ivie 1942 and description of Hoffmannilena, a new genus from Mexico (Araneae: Agelenidae). Zootaxa, 4084 (1), 1 - 49. https: // doi. org / 10.11646 / zootaxa. 4084.1.1" type="journal article" year="2016">Maya-Morales &amp; Jiménez 2016</bibRefCitation>
: fig. 30); and from
<taxonomicName id="ABFE4D50FF84FFFEE0356619FE2DFB94" ID-CoL="4XRZ" box="[259,427,1124,1149]" class="Arachnida" family="Agelenidae" genus="Hoffmannilena" higherTaxonomySource="CoL" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="5E8AEAC1FF84FFFEE0356619FE2DFB94" box="[259,427,1124,1149]" italics="true" pageId="5" pageNumber="6">Hoffmannilena</emphasis>
</taxonomicName>
in lacking a basal projection on RTA (as in
<figureCitation id="F4C52A56FF84FFFEE2A76619FC64FB94" box="[913,994,1124,1149]" captionStart="FIGURE 45" captionStartId="81.[151,250,1896,1918]" captionTargetBox="[199,1375,1092,1848]" captionTargetId="figure@81.[186,1399,1074,1864]" captionTargetPageId="81" captionText="FIGURE 45. Hoffmannilena nova (O. Pickard-Cambridge 1896) comb. nov., male pedipalp. a, prolateral view; b, ventral view; c, retrolateral view. Abbreviations: C, conductor; E, embolus; MA, median apophysis; RTAb, basal projection of RTA; RTAd, distal projection of RTA; T, tegulum; TMP, tegular median process. Scale bar: 0.25 mm." httpUri="https://zenodo.org/record/571729/files/figure.png" pageId="5" pageNumber="6">Fig. 45</figureCitation>
b, c) and a strongly sclerotized epigynal plate (as in
<bibRefCitation id="086F4B22FF84FFFEE02F66FAFD07FB49" author="Maya-Morales" box="[281,641,1159,1184]" pageId="5" pageNumber="6" refString="Maya-Morales, J. &amp; Jimenez, M. L. (2016) Taxonomic revision of the spider genus Rualena Chamberlin &amp; Ivie 1942 and description of Hoffmannilena, a new genus from Mexico (Araneae: Agelenidae). Zootaxa, 4084 (1), 1 - 49. https: // doi. org / 10.11646 / zootaxa. 4084.1.1" type="journal article" year="2016">Maya-Morales &amp; Jiménez 2016</bibRefCitation>
: fig. 116).
</paragraph>
</subSubSection>
<subSubSection id="24E46558FF84FFFDE1F166D1FDC7FF3C" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
<paragraph id="6C4136D3FF84FFFEE1F166D1FCD7F87C" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="6">
<emphasis id="5E8AEAC1FF84FFFEE1F166D1FED7FB2C" bold="true" box="[199,337,1196,1221]" pageId="5" pageNumber="6">Description</emphasis>
. Medium-sized spiders,
<quantity id="AB069B36FF84FFFEE34566D1FD64FB2D" box="[627,738,1196,1220]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="14.0" metricValueMin="4.0" pageId="5" pageNumber="6" unit="mm" value="9.0" valueMax="14.0" valueMin="4.0">4–14 mm</quantity>
total length. Eight eyes in two rows strongly procurved in frontal view (
<figureCitation id="F4C52A56FF84FFFEE01966B2FEF7FB01" box="[303,369,1231,1256]" captionStart="FIGURE 8" captionStartId="20.[151,250,1504,1526]" captionTargetBox="[211,1372,203,1468]" captionTargetId="figure@20.[200,1387,193,1483]" captionTargetPageId="20" captionText="FIGURE 8. Novalena intermedia (Chamberlin &amp; Gertsch 1930), female (SEM): a, carapace; b, eyes; c, cheliceral fang; d, serrula; e, tarsus II; f, tarsal organ. a, d, f, dorsal view; b, frontal view; c, apical view; e, lateral view." httpUri="https://zenodo.org/record/571691/files/figure.png" pageId="5" pageNumber="6">Fig. 8</figureCitation>
b). Carapace with two symmetrical longitudinal dark bands intensified by feathery scales and a black band around the border of thoracic region (
<figureCitation id="F4C52A56FF84FFFEE3F26689FC80FAE5" box="[708,774,1268,1293]" captionStart="FIGURE 1" captionStartId="6.[151,250,1888,1910]" captionTargetBox="[186,1400,348,1856]" captionTargetId="figure@6.[176,1411,339,1866]" captionTargetPageId="6" captionText="FIGURE 1. Novalena spp., habitus: a, e, N. approximata (Gertsch &amp; Ivie 1936), female; b, f, N. alvarezi sp. nov., female; c, g, N. chamberlini sp. nov., male; d, h, N. clara sp. nov., female; i, m, N. dentata sp. nov., male; j, n, N. irazu sp. nov., female; k, o, N. iviei sp. nov., male; l, p, N. leonensis sp. nov., female; q, u, N. mexiquensis sp. nov., male; r, v, N. poncei sp. nov., female; s, w, N. puebla sp. nov., female; t, x, N. rothi sp. nov., female. a – d, i – l, q – t, dorsal view; e – h, m – p, v – x, ventral view. Scale bars: a, d, j, q, s, t = 4 mm; b, i, l, r = 3 mm; c, k = 2 mm." httpUri="https://zenodo.org/record/571684/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
c, i, r). Chelicerae with three promarginal teeth and two to four retromarginal teeth (
<figureCitation id="F4C52A56FF84FFFEE0F5676AFD8AFAD9" box="[451,524,1303,1328]" captionStart="FIGURE 8" captionStartId="20.[151,250,1504,1526]" captionTargetBox="[211,1372,203,1468]" captionTargetId="figure@20.[200,1387,193,1483]" captionTargetPageId="20" captionText="FIGURE 8. Novalena intermedia (Chamberlin &amp; Gertsch 1930), female (SEM): a, carapace; b, eyes; c, cheliceral fang; d, serrula; e, tarsus II; f, tarsal organ. a, d, f, dorsal view; b, frontal view; c, apical view; e, lateral view." httpUri="https://zenodo.org/record/571691/files/figure.png" pageId="5" pageNumber="6">Fig. 8</figureCitation>
c). Sternum longer than wide (
<figureCitation id="F4C52A56FF84FFFEE248676AFC42FAD9" box="[894,964,1303,1328]" captionStart="FIGURE 1" captionStartId="6.[151,250,1888,1910]" captionTargetBox="[186,1400,348,1856]" captionTargetId="figure@6.[176,1411,339,1866]" captionTargetPageId="6" captionText="FIGURE 1. Novalena spp., habitus: a, e, N. approximata (Gertsch &amp; Ivie 1936), female; b, f, N. alvarezi sp. nov., female; c, g, N. chamberlini sp. nov., male; d, h, N. clara sp. nov., female; i, m, N. dentata sp. nov., male; j, n, N. irazu sp. nov., female; k, o, N. iviei sp. nov., male; l, p, N. leonensis sp. nov., female; q, u, N. mexiquensis sp. nov., male; r, v, N. poncei sp. nov., female; s, w, N. puebla sp. nov., female; t, x, N. rothi sp. nov., female. a – d, i – l, q – t, dorsal view; e – h, m – p, v – x, ventral view. Scale bars: a, d, j, q, s, t = 4 mm; b, i, l, r = 3 mm; c, k = 2 mm." httpUri="https://zenodo.org/record/571684/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
f). Feathery scales present on carapace, opisthosoma, pedipalps, and legs (
<figureCitation id="F4C52A56FF84FFFEE3066741FDFFFABD" box="[560,633,1340,1365]" captionStart="FIGURE 8" captionStartId="20.[151,250,1504,1526]" captionTargetBox="[211,1372,203,1468]" captionTargetId="figure@20.[200,1387,193,1483]" captionTargetPageId="20" captionText="FIGURE 8. Novalena intermedia (Chamberlin &amp; Gertsch 1930), female (SEM): a, carapace; b, eyes; c, cheliceral fang; d, serrula; e, tarsus II; f, tarsal organ. a, d, f, dorsal view; b, frontal view; c, apical view; e, lateral view." httpUri="https://zenodo.org/record/571691/files/figure.png" pageId="5" pageNumber="6">Fig. 8</figureCitation>
a, b). Opisthosoma oval with dorsal foliate pattern and/or posterior chevrons (
<figureCitation id="F4C52A56FF84FFFEE03D6722FECFFA91" box="[267,329,1375,1400]" captionStart="FIGURE 1" captionStartId="6.[151,250,1888,1910]" captionTargetBox="[186,1400,348,1856]" captionTargetId="figure@6.[176,1411,339,1866]" captionTargetPageId="6" captionText="FIGURE 1. Novalena spp., habitus: a, e, N. approximata (Gertsch &amp; Ivie 1936), female; b, f, N. alvarezi sp. nov., female; c, g, N. chamberlini sp. nov., male; d, h, N. clara sp. nov., female; i, m, N. dentata sp. nov., male; j, n, N. irazu sp. nov., female; k, o, N. iviei sp. nov., male; l, p, N. leonensis sp. nov., female; q, u, N. mexiquensis sp. nov., male; r, v, N. poncei sp. nov., female; s, w, N. puebla sp. nov., female; t, x, N. rothi sp. nov., female. a – d, i – l, q – t, dorsal view; e – h, m – p, v – x, ventral view. Scale bars: a, d, j, q, s, t = 4 mm; b, i, l, r = 3 mm; c, k = 2 mm." httpUri="https://zenodo.org/record/571684/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
b, k, s). Colulus divided with few hairs on each side (
<figureCitation id="F4C52A56FF84FFFEE2AF6722FC5BFA91" box="[921,989,1375,1400]" captionStart="FIGURE 9" captionStartId="21.[151,250,1205,1227]" captionTargetBox="[211,1375,339,1180]" captionTargetId="figure@21.[208,1379,337,1184]" captionTargetPageId="21" captionText="FIGURE 9. Novalena intermedia (Chamberlin &amp; Gertsch 1930), female (SEM): a, spinnerets; b, anal tubercle; c, colulus and tracheal spiracle; d, ALS, details. Abbreviations: Cl, colulus; ALS, anterior lateral spinnerets; AT, anal tubercle; MAS, major ampullate glans spigots; PGS, piriform gland spigots; PLS, posterior lateral spinnerets; PMS, posterior median spinnerets; TH, thick hairs; TS, tracheal spiracle." httpUri="https://zenodo.org/record/571692/files/figure.png" pageId="5" pageNumber="6">Fig. 9</figureCitation>
c). PLS longest with distal segment 0.5– 1.25 times basal (
<figureCitation id="F4C52A56FF84FFFEE06967F9FE2DFA74" box="[351,427,1412,1437]" captionStart="FIGURE 1" captionStartId="6.[151,250,1888,1910]" captionTargetBox="[186,1400,348,1856]" captionTargetId="figure@6.[176,1411,339,1866]" captionTargetPageId="6" captionText="FIGURE 1. Novalena spp., habitus: a, e, N. approximata (Gertsch &amp; Ivie 1936), female; b, f, N. alvarezi sp. nov., female; c, g, N. chamberlini sp. nov., male; d, h, N. clara sp. nov., female; i, m, N. dentata sp. nov., male; j, n, N. irazu sp. nov., female; k, o, N. iviei sp. nov., male; l, p, N. leonensis sp. nov., female; q, u, N. mexiquensis sp. nov., male; r, v, N. poncei sp. nov., female; s, w, N. puebla sp. nov., female; t, x, N. rothi sp. nov., female. a – d, i – l, q – t, dorsal view; e – h, m – p, v – x, ventral view. Scale bars: a, d, j, q, s, t = 4 mm; b, i, l, r = 3 mm; c, k = 2 mm." httpUri="https://zenodo.org/record/571684/files/figure.png" pageId="5" pageNumber="6">Figs. 1</figureCitation>
f, 9a). Legs formula usually 4123. Legs usually annulated (
<figureCitation id="F4C52A56FF84FFFEE57967F9FB09FA74" box="[1103,1167,1412,1437]" captionStart="FIGURE 1" captionStartId="6.[151,250,1888,1910]" captionTargetBox="[186,1400,348,1856]" captionTargetId="figure@6.[176,1411,339,1866]" captionTargetPageId="6" captionText="FIGURE 1. Novalena spp., habitus: a, e, N. approximata (Gertsch &amp; Ivie 1936), female; b, f, N. alvarezi sp. nov., female; c, g, N. chamberlini sp. nov., male; d, h, N. clara sp. nov., female; i, m, N. dentata sp. nov., male; j, n, N. irazu sp. nov., female; k, o, N. iviei sp. nov., male; l, p, N. leonensis sp. nov., female; q, u, N. mexiquensis sp. nov., male; r, v, N. poncei sp. nov., female; s, w, N. puebla sp. nov., female; t, x, N. rothi sp. nov., female. a – d, i – l, q – t, dorsal view; e – h, m – p, v – x, ventral view. Scale bars: a, d, j, q, s, t = 4 mm; b, i, l, r = 3 mm; c, k = 2 mm." httpUri="https://zenodo.org/record/571684/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
b–d, k). Patella I and II with two dorsal spines and one prolateral spine, patella III and VI with two dorsal spines, one prolateral spine, and one retrolateral spine. Leg tarsi with five to eight trichobothria. Capsulate tarsal organ distad of trichobothrial row (
<figureCitation id="F4C52A56FF84FFFEE1A96792FF65F9E1" box="[159,227,1519,1544]" captionStart="FIGURE 8" captionStartId="20.[151,250,1504,1526]" captionTargetBox="[211,1372,203,1468]" captionTargetId="figure@20.[200,1387,193,1483]" captionTargetPageId="20" captionText="FIGURE 8. Novalena intermedia (Chamberlin &amp; Gertsch 1930), female (SEM): a, carapace; b, eyes; c, cheliceral fang; d, serrula; e, tarsus II; f, tarsal organ. a, d, f, dorsal view; b, frontal view; c, apical view; e, lateral view." httpUri="https://zenodo.org/record/571691/files/figure.png" pageId="5" pageNumber="6">Fig. 8</figureCitation>
f). Pedipalp femur with two to four dorsal spines. Female pedipalp tibia usually with one + two prolateral spines. Male pedipalp tibia with two to four spines and with prolateroventral protuberance and retrolateroventral groove (
<figureCitation id="F4C52A56FF84FFFEE1C2644AFEC0F9B9" box="[244,326,1591,1616]" captionStart="FIGURE 10" captionStartId="22.[151,250,1507,1529]" captionTargetBox="[209,1373,205,1475]" captionTargetId="figure@22.[196,1390,193,1486]" captionTargetPageId="22" captionText="FIGURE 10. Novalena intermedia (Chamberlin &amp; Gertsch 1930) (SEM): a, c, e, male pedipalp; b, d, f, internal female genitalia. a, c, ventral view; b, d, dorsal view; e, prolateral view, arrow to prolateral protuberance; f, anterior view. Abbreviations: CD, copulatory ducts; Cd, dorsal projection of conductor; Cv, ventral projection of conductor; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process." httpUri="https://zenodo.org/record/571693/files/figure.png" pageId="5" pageNumber="6">Fig. 10</figureCitation>
e). RTA of male pedipalp with distal and dorsal projections; ventral projection usually present (
<figureCitation id="F4C52A56FF84FFFEE459644AFF33F99C" captionStart="FIGURE 21" captionStartId="43.[151,250,1716,1738]" captionTargetBox="[209,1376,845,1690]" captionTargetId="figure@43.[205,1381,841,1694]" captionTargetPageId="43" captionText="FIGURE 21. Novalena dentata sp. nov., male pedipalp (SEM): a, c, ventral view; b, prolateral view; d, retrolateral view. Abbreviations: C, conductor; E, embolus; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; T, tegulum; TMP, tegular median process." httpUri="https://zenodo.org/record/571705/files/figure.png" pageId="5" pageNumber="6">Fig. 21</figureCitation>
d). Embolus a short and simple curve (
<figureCitation id="F4C52A56FF84FFFEE3BC6421FD64F99D" box="[650,738,1628,1653]" captionStart="FIGURE 14" captionStartId="28.[151,250,1512,1534]" captionTargetBox="[208,1375,206,1477]" captionTargetId="figure@28.[196,1390,195,1490]" captionTargetPageId="28" captionText="FIGURE 14. Novalena simplex (F. O. Pickard-Cambridge 1902) (SEM): a, c, e, male pedipalp; b, d, f, internal female genitalia. a, ventral view; b, dorsal view; c, e, retrolateral view; d, f, anterior view. Abbreviations: CD, copulatory ducts; Cd, dorsal projection of conductor; Cv, ventral projection of conductor; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process." httpUri="https://zenodo.org/record/571698/files/figure.png" pageId="5" pageNumber="6">Fig. 14</figureCitation>
a). Conductor with two short projections (
<figureCitation id="F4C52A56FF84FFFEE5E06421FA98F99D" box="[1238,1310,1628,1653]" captionStart="FIGURE 3" captionStartId="12.[151,250,1065,1087]" captionTargetBox="[211,1375,196,1039]" captionTargetId="figure@12.[208,1379,193,1044]" captionTargetPageId="12" captionText="FIGURE 3. Novalena approximata (Gertsch &amp; Ivie 1936), male pedipalp (SEM): a, ventral view; b, prolateral view; c, d, retrolateral view. Abbreviations: Cd, dorsal projection of conductor; Cv, ventral projection of conductor; E, embolus; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; St, subtegulum; T, tegulum; TMP, tegular median process." httpUri="https://zenodo.org/record/571686/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
c). Median apophysis spoon-shaped (
<figureCitation id="F4C52A56FF84FFFEE0886402FD94F971" box="[446,530,1663,1688]" captionStart="FIGURE 10" captionStartId="22.[151,250,1507,1529]" captionTargetBox="[209,1373,205,1475]" captionTargetId="figure@22.[196,1390,193,1486]" captionTargetPageId="22" captionText="FIGURE 10. Novalena intermedia (Chamberlin &amp; Gertsch 1930) (SEM): a, c, e, male pedipalp; b, d, f, internal female genitalia. a, c, ventral view; b, d, dorsal view; e, prolateral view, arrow to prolateral protuberance; f, anterior view. Abbreviations: CD, copulatory ducts; Cd, dorsal projection of conductor; Cv, ventral projection of conductor; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process." httpUri="https://zenodo.org/record/571693/files/figure.png" pageId="5" pageNumber="6">Fig. 10</figureCitation>
a) (only reduced in
<taxonomicName id="ABFE4D50FF84FFFEE3CC64FCFCE5F971" box="[762,867,1663,1688]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="alamo">
<emphasis id="5E8AEAC1FF84FFFEE3CC64FCFCE5F971" box="[762,867,1663,1688]" italics="true" pageId="5" pageNumber="6">N. alamo</emphasis>
</taxonomicName>
,
<figureCitation id="F4C52A56FF84FFFEE2436402FC4CF971" box="[885,970,1663,1688]" captionStart="FIGURE 16" captionStartId="31.[151,250,1804,1826]" captionTargetBox="[245,1339,202,1769]" captionTargetId="figure@31.[233,1350,193,1779]" captionTargetPageId="31" captionText="FIGURE 16. Novalena spp. genitalia: a – c, Novalena ajusco sp. nov.; d, e, Novalena alamo sp. nov.; f – h, Novalena alvarezi sp. nov.; i – k, Novalena atzimbo sp. nov.; l, m, Novalena bosencheve sp. nov. a, f, i, epigynum, ventral view; b, g, j, internal genitalia, dorsal view; c, h, k, epigynum, posterior view; d, l, male pedipalp, ventral view; e, m, same retrolateral view. Abbreviations: A, atrium; C, conductor; CD, copulatory ducts; CO, copulatory openings; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process. Dashed line indicates distance between anterior and posterior margins. Scale bars: c, h, k = 0.25 mm; d, l = 0.5 mm." httpUri="https://zenodo.org/record/571700/files/figure.png" pageId="5" pageNumber="6">Fig. 16</figureCitation>
d). Tegular median process present (
<figureCitation id="F4C52A56FF84FFFEE4596402FF33F954" captionStart="FIGURE 21" captionStartId="43.[151,250,1716,1738]" captionTargetBox="[209,1376,845,1690]" captionTargetId="figure@43.[205,1381,841,1694]" captionTargetPageId="43" captionText="FIGURE 21. Novalena dentata sp. nov., male pedipalp (SEM): a, c, ventral view; b, prolateral view; d, retrolateral view. Abbreviations: C, conductor; E, embolus; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; T, tegulum; TMP, tegular median process." httpUri="https://zenodo.org/record/571705/files/figure.png" pageId="5" pageNumber="6">Fig. 21</figureCitation>
b). Fulcrum absent (only present in
<taxonomicName id="ABFE4D50FF84FFFEE36364D8FD59F955" box="[597,735,1701,1724]" class="Arachnida" family="Agelenidae" genus="Agelena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="annamae">
<emphasis id="5E8AEAC1FF84FFFEE36364D8FD59F955" box="[597,735,1701,1724]" italics="true" pageId="5" pageNumber="6">N. annamae</emphasis>
</taxonomicName>
,
<figureCitation id="F4C52A56FF84FFFEE3C764D9FCB1F955" box="[753,823,1700,1725]" captionStart="FIGURE 2" captionStartId="10.[151,250,1518,1540]" captionTargetBox="[156,1432,687,1497]" captionTargetId="figure@10.[152,1438,681,1505]" captionTargetPageId="10" captionText="FIGURE 2. Novalena annamae (Gertsch &amp; Davis 1940): a, male pedipalp, ventral view; b, same, retrolateral view; c, epigynum, ventral view; d, internal genitalia, dorsal view; e, epigynum, posterior view. Abbreviations: A, atrium; C, conductor; CD, copulatory ducts; CO, copulatory openings; E, embolus; F, fulcrum; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process. Scale bars: a = 0.5 mm; e = 0.25 mm." httpUri="https://zenodo.org/record/571685/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
a). Female epigynal plate wider than long (
<figureCitation id="F4C52A56FF84FFFEE41A64D9FA04F954" box="[1324,1410,1700,1725]" captionStart="FIGURE 25" captionStartId="47.[151,250,1346,1368]" captionTargetBox="[162,1430,593,1320]" captionTargetId="figure@47.[152,1436,587,1324]" captionTargetPageId="47" captionText="FIGURE 25. Novalena franckei sp. nov.: a, male pedipalp, ventral view; b, same, retrolateral view; c, epigynum, ventral view; d, internal genitalia, dorsal view; e, epigynum, posterior view. Abbreviations: A, atrium; C, conductor; CD, copulatory ducts; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process. Scale bars: a = 0.5 mm; e = 0.25 mm." httpUri="https://zenodo.org/record/571709/files/figure.png" pageId="5" pageNumber="6">Fig. 25</figureCitation>
c). Atrium on posterior part of plate, usually wider than long (
<figureCitation id="F4C52A56FF84FFFEE21464BAFCF5F909" box="[802,883,1735,1760]" captionStart="FIGURE 30" captionStartId="55.[151,250,1651,1673]" captionTargetBox="[158,1425,602,1616]" captionTargetId="figure@55.[151,1437,591,1628]" captionTargetPageId="55" captionText="FIGURE 30. Novalena spp. genitalia: a – c, Novalena jiquilpan sp. nov.; d – h, Novalena leonensis sp. nov. a, f, epigynum, ventral view; b, g, internal genitalia, dorsal view; c, h, epigynum, posterior view; d, male pedipalp, ventral view; e, same, retrolateral view. Abbreviations: A, atrium; C, conductor; CD, copulatory ducts; CO, copulatory openings; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process. Dashed line indicates distance between anterior and posterior margins. Scale bars: c, h = 0.25 mm; d = 0.5 mm." httpUri="https://zenodo.org/record/571713/files/figure.png" pageId="5" pageNumber="6">Fig. 30</figureCitation>
f). Atrial spurs usually present (
<figureCitation id="F4C52A56FF84FFFEE5E764BAFA95F909" box="[1233,1299,1735,1760]" captionStart="FIGURE 5" captionStartId="14.[151,250,1484,1506]" captionTargetBox="[161,1424,865,1453]" captionTargetId="figure@14.[150,1437,860,1463]" captionTargetPageId="14" captionText="FIGURE 5. Novalena spp.: a – c, Novalena attenuata (F. O. Pickard-Cambridge 1902); d – f, Novalena bipunctata Roth 1967 stat. rev. a, d, epigynum, ventral view; b, e, internal genitalia, dorsal view; c, f, epigynum, posterior view. Abbreviations: A, atrium; CD, copulatory ducts; CO, copulatory openings; FD, fertilization ducts; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae. Dashed line indicates distance between anterior and posterior margins. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/571688/files/figure.png" pageId="5" pageNumber="6">Fig. 5</figureCitation>
a) (absent in
<taxonomicName id="ABFE4D50FF84FFFEE1A16490FE9DF8EC" box="[151,283,1772,1797]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="calavera">
<emphasis id="5E8AEAC1FF84FFFEE1A16490FE9DF8EC" box="[151,283,1772,1797]" italics="true" pageId="5" pageNumber="6">N. calavera</emphasis>
</taxonomicName>
,
<taxonomicName id="ABFE4D50FF84FFFEE01D6490FEF8F8EC" box="[299,382,1772,1797]" class="Arachnida" family="Agelenidae" genus="Agelena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="lutzi">
<emphasis id="5E8AEAC1FF84FFFEE01D6490FEF8F8EC" box="[299,382,1772,1797]" italics="true" pageId="5" pageNumber="6">N. lutzi</emphasis>
</taxonomicName>
, and
<taxonomicName id="ABFE4D50FF84FFFEE0896490FDBFF8ED" box="[447,569,1773,1796]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="garnica">
<emphasis id="5E8AEAC1FF84FFFEE0896490FDBFF8ED" box="[447,569,1773,1796]" italics="true" pageId="5" pageNumber="6">N. garnica</emphasis>
</taxonomicName>
). Internal genitalia with copulatory ducts strongly sclerotized, straight (
<figureCitation id="F4C52A56FF84FFFEE4596491FF32F8C1" captionStart="FIGURE 23" captionStartId="45.[151,250,793,815]" captionTargetBox="[211,1375,347,762]" captionTargetId="figure@45.[202,1385,337,772]" captionTargetPageId="45" captionText="FIGURE 23. Novalena divisadero sp. nov. (SEM), internal female genitalia, dorsal view. Abbreviations: CD, copulatory ducts; FD, fertilization ducts; S 1, primary spermathecae; S 2, secondary spermathecae." httpUri="https://zenodo.org/record/571707/files/figure.png" pageId="5" pageNumber="6">Fig. 23</figureCitation>
a) or curved (
<figureCitation id="F4C52A56FF84FFFEE0646572FE21F8C1" box="[338,423,1807,1832]" captionStart="FIGURE 14" captionStartId="28.[151,250,1512,1534]" captionTargetBox="[208,1375,206,1477]" captionTargetId="figure@28.[196,1390,195,1490]" captionTargetPageId="28" captionText="FIGURE 14. Novalena simplex (F. O. Pickard-Cambridge 1902) (SEM): a, c, e, male pedipalp; b, d, f, internal female genitalia. a, ventral view; b, dorsal view; c, e, retrolateral view; d, f, anterior view. Abbreviations: CD, copulatory ducts; Cd, dorsal projection of conductor; Cv, ventral projection of conductor; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process." httpUri="https://zenodo.org/record/571698/files/figure.png" pageId="5" pageNumber="6">Fig. 14</figureCitation>
d). Spermathecae bipartite (
<figureCitation id="F4C52A56FF84FFFEE3D26572FCBDF8C1" box="[740,827,1807,1832]" captionStart="FIGURE 36" captionStartId="67.[151,250,1897,1919]" captionTargetBox="[209,1377,1456,1872]" captionTargetId="figure@67.[204,1383,1452,1876]" captionTargetPageId="67" captionText="FIGURE 36. Novalena puebla sp. nov. (SEM), internal female genitalia: a, ectal view; b, dorsal view. Abbreviations: CD, copulatory ducts; S 1, primary spermathecae; S 2, secondary spermathecae." httpUri="https://zenodo.org/record/571720/files/figure.png" pageId="5" pageNumber="6">Fig. 36</figureCitation>
); primary spermathecae longer than wide (
<figureCitation id="F4C52A56FF84FFFEE41C6572FA06F8C1" box="[1322,1408,1807,1832]" captionStart="FIGURE 32" captionStartId="59.[151,250,1446,1468]" captionTargetBox="[157,1430,631,1418]" captionTargetId="figure@59.[151,1436,623,1424]" captionTargetPageId="59" captionText="FIGURE 32. Novalena oaxaca sp. nov.: a, male pedipalp, ventral view; b, same, retrolateral view; c, epigynum, ventral view; d, internal genitalia, dorsal view; e, epigynum, posterior view. Abbreviations: A, atrium; C, conductor; CD, copulatory ducts; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process. Scale bars: a = 0.5 mm; e = 0.25 mm." httpUri="https://zenodo.org/record/571716/files/figure.png" pageId="5" pageNumber="6">Fig. 32</figureCitation>
d), secondary spermathecae blind-ended receptacles with primary pores (
<bibRefCitation id="086F4B22FF84FFFEE28A6549FBDCF8A5" author="Bennett" box="[956,1114,1844,1868]" pageId="5" pageNumber="6" refString="Bennett, R. G. (1992) The spermathecal pores of spiders with special reference to dictynoids and amaurobioids (Araneae, Araneomorphae, Araneoclada). Proceedings of the Entomological Society of Ontario, 123, 1 - 21." type="journal article" year="1992">Bennett 1992</bibRefCitation>
) and each connected to the union of copulatory duct and primary spermathecae by short diverticulum (
<figureCitation id="F4C52A56FF84FFFEE530652AFBDBF899" box="[1030,1117,1879,1904]" captionStart="FIGURE 14" captionStartId="28.[151,250,1512,1534]" captionTargetBox="[208,1375,206,1477]" captionTargetId="figure@28.[196,1390,195,1490]" captionTargetPageId="28" captionText="FIGURE 14. Novalena simplex (F. O. Pickard-Cambridge 1902) (SEM): a, c, e, male pedipalp; b, d, f, internal female genitalia. a, ventral view; b, dorsal view; c, e, retrolateral view; d, f, anterior view. Abbreviations: CD, copulatory ducts; Cd, dorsal projection of conductor; Cv, ventral projection of conductor; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process." httpUri="https://zenodo.org/record/571698/files/figure.png" pageId="5" pageNumber="6">Fig. 14</figureCitation>
b). Fertilization ducts short, originating from the posterior part of spermathecae (
<figureCitation id="F4C52A56FF84FFFEE3D46501FCB3F87C" box="[738,821,1916,1941]" captionStart="FIGURE 10" captionStartId="22.[151,250,1507,1529]" captionTargetBox="[209,1373,205,1475]" captionTargetId="figure@22.[196,1390,193,1486]" captionTargetPageId="22" captionText="FIGURE 10. Novalena intermedia (Chamberlin &amp; Gertsch 1930) (SEM): a, c, e, male pedipalp; b, d, f, internal female genitalia. a, c, ventral view; b, d, dorsal view; e, prolateral view, arrow to prolateral protuberance; f, anterior view. Abbreviations: CD, copulatory ducts; Cd, dorsal projection of conductor; Cv, ventral projection of conductor; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process." httpUri="https://zenodo.org/record/571693/files/figure.png" pageId="5" pageNumber="6">Fig. 10</figureCitation>
d).
</paragraph>
<paragraph id="6C4136D3FF84FFFEE1F165DDFA1AF835" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="6">
<emphasis id="5E8AEAC1FF84FFFEE1F165DDFEA5F850" bold="true" box="[199,291,1952,1977]" pageId="5" pageNumber="6">Habitat</emphasis>
. In North America,
<taxonomicName id="ABFE4D50FF84FFFEE33365DDFDF7F850" box="[517,625,1952,1977]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="5E8AEAC1FF84FFFEE33365DDFDF7F850" box="[517,625,1952,1977]" italics="true" pageId="5" pageNumber="6">Novalena</emphasis>
</taxonomicName>
spiders are usually found in webs under loose bark of standing trees and stumps in dense coniferous forest, and occasionally in webs on the ground in shaded areas (
<bibRefCitation id="086F4B22FF84FFFEE59265B9FA16F835" author="Roth" box="[1188,1424,1988,2013]" pageId="5" pageNumber="6" refString="Roth, V. D. &amp; Brame, P. L. (1972) Nearctic genera of the spider family Agelenidae (Arachnida, Araneida). American Museum Novitates, 2505, 1 - 52." type="journal article" year="1972">Roth &amp; Brame 1972</bibRefCitation>
).
</paragraph>
<paragraph id="6C4136D3FF87FFFDE1A162E5FDC7FF3C" blockId="6.[151,1436,151,285]" pageId="6" pageNumber="7">In tropical montane cloud forests of southeastern Mexico, they are found in webs within epiphytes and on understory vegetation and the ground.</paragraph>
</subSubSection>
<subSubSection id="24E46558FF87FFFDE1F162A2FC55FEF5" pageId="6" pageNumber="7" type="distribution">
<paragraph id="6C4136D3FF87FFFDE1F162A2FC55FEF5" blockId="6.[151,1436,151,285]" pageId="6" pageNumber="7">
<emphasis id="5E8AEAC1FF87FFFDE1F162A2FEDFFF11" bold="true" box="[199,345,223,248]" pageId="6" pageNumber="7">Distribution</emphasis>
.
<taxonomicName id="ABFE4D50FF87FFFDE04462A2FE58FF11" box="[370,478,223,248]" class="Arachnida" family="Agelenidae" genus="Novalena" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="5E8AEAC1FF87FFFDE04462A2FE58FF11" box="[370,478,223,248]" italics="true" pageId="6" pageNumber="7">Novalena</emphasis>
</taxonomicName>
is widely distributed in the western Nearctic and northern Neotropics from southwestern
<collectingCountry id="14E97643FF87FFFDE0046379FE00FEF4" box="[306,390,260,285]" name="Canada" pageId="6" pageNumber="7">Canada</collectingCountry>
to
<collectingCountry id="14E97643FF87FFFDE09F6379FDA5FEF4" box="[425,547,260,285]" name="Costa Rica" pageId="6" pageNumber="7">
<collectingRegion id="AE3AF831FF87FFFDE09F6379FE6FFEF5" box="[425,489,260,284]" country="New Zealand" name="West Coast" pageId="6" pageNumber="7">Costa</collectingRegion>
Rica
</collectingCountry>
and
<collectingCountry id="14E97643FF87FFFDE36C6379FCC2FEF4" box="[602,836,260,285]" name="Trinidad and Tobago" pageId="6" pageNumber="7">
Trinidad and
<collectingRegion id="AE3AF831FF87FFFDE3C66379FCC2FEF4" box="[752,836,260,285]" country="Trinidad and Tobago" name="Tobago" pageId="6" pageNumber="7">Tobago</collectingRegion>
</collectingCountry>
(Maps 1–8).
</paragraph>
</subSubSection>
</treatment>
</document>
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