262 lines
30 KiB
XML
262 lines
30 KiB
XML
<document id="342664E467B91E3EC83D3330C662EF7C" ID-DOI="10.1111/j.1096-3642.2008.00415.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5446700" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630461491835" checkinUser="felipe" docAuthor="Wu, Shu-Ping, Hwang, Chung-Chi & Lin, Yao-Sung" docDate="2008" docId="E45287EFFFE6430DFF46FE0B8650578E" docLanguage="en" docName="j.1096-3642.2008.00415.x.pdf" docOrigin="Zoological Journal of the Linnean Society 154 (3)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00415.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Satsuma phoenicis Wu & Hwang & Lin 2008, SP. NOV." docType="treatment" docVersion="4" lastPageNumber="458" masterDocId="186BFF97FFF54318FF92FFF482155506" masterDocTitle="Systematic revision of the arboreal snail Satsuma albida species complex (Mollusca: Camaenidae) with descriptions of 14 new species from Taiwan" masterLastPageNumber="493" masterPageNumber="437" pageNumber="456" updateTime="1699279363994" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="25C792EE444F04E29DA57507DBF97046">Systematic revision of the arboreal snail Satsuma albida species complex (Mollusca: Camaenidae) with descriptions of 14 new species from Taiwan</mods:title>
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<mods:namePart id="27260F00424AAD6A22A036A5F8E0B30D">Wu, Shu-Ping</mods:namePart>
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<mods:namePart id="57B3DE7077AD7ADEFB5E865E50AA19A0">Hwang, Chung-Chi</mods:namePart>
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<mods:roleTerm id="9883ECF814F18D689E09CA5EBE395631">Author</mods:roleTerm>
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<mods:namePart id="A8F9D4E0B468DBD7EA7471EB01CB5ADE">Lin, Yao-Sung</mods:namePart>
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<mods:title id="19DF1B54646EB82C3E3DA52BC6D2E107">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="E59DB0A7EF9FEE9A270285BB383C2CF7">
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<mods:date id="0FDBB8890A221A6BD1097F4983EA4C8C">2008</mods:date>
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<mods:number id="BAEC4C54E2C6FB9B973EEDD6289114FD">2008-10-31</mods:number>
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<mods:number id="5430E6F1B2E1E674ED8B9125A723419C">154</mods:number>
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<treatment id="E45287EFFFE6430DFF46FE0B8650578E" ID-DOI="http://doi.org/10.5281/zenodo.5492604" ID-Zenodo-Dep="5492604" LSID="urn:lsid:plazi:treatment:E45287EFFFE6430DFF46FE0B8650578E" httpUri="http://treatment.plazi.org/id/E45287EFFFE6430DFF46FE0B8650578E" lastPageId="21" lastPageNumber="458" pageId="19" pageNumber="456">
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<subSubSection id="24E16572FFE6430BFF46FE0B80CF5713" box="[212,730,511,533]" pageId="19" pageNumber="456" type="nomenclature">
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<paragraph id="6C4436F9FFE6430BFF46FE0B80CF5713" blockId="19.[212,730,511,533]" box="[212,730,511,533]" pageId="19" pageNumber="456">
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<heading id="370C8195FFE6430BFF46FE0B80CF5713" box="[212,730,511,533]" centered="true" fontSize="9" level="2" pageId="19" pageNumber="456" reason="2">
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<taxonomicName id="ABFB4D7AFFE6430BFF46FE0B83DE5712" ID-CoL="79QBJ" authority="Wu & Hwang & Lin, 2008" authorityName="Wu & Hwang & Lin" authorityYear="2008" box="[212,459,511,532]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="456" phylum="Mollusca" rank="species" species="phoenicis" status="sp. nov.">SATSUMA PHOENICIS</taxonomicName>
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||
<emphasis id="5E8FEAEBFFE6430BFE47FDF7802F5712" bold="true" box="[469,570,511,532]" pageId="19" pageNumber="456">
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||
<taxonomicNameLabel id="45BC5790FFE6430BFE47FDF7802F5712" box="[469,570,511,532]" pageId="19" pageNumber="456" rank="species">SP. NOV.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="F4C02A7CFFE6430BFDDFFE0B80B25712" box="[589,679,511,532]" captionStart="Figure 13" captionStartId="19.[164,243,1853,1872]" captionTargetBox="[389,1219,1092,1824]" captionTargetId="figure-290@19.[387,1221,1090,1824]" captionTargetPageId="19" captionText="Figure 13. Satsuma phoenicis sp. nov. A–D, holotype (TMMT 0601, shell height = 19.4 mm). E, living specimen. Arrow indicates the pre-apertural constriction." figureDoi="http://doi.org/10.5281/zenodo.5446729" httpUri="https://zenodo.org/record/5446729/files/figure.png" pageId="19" pageNumber="456">FIGS 13</figureCitation>
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,
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<figureCitation id="F4C02A7CFFE6430BFD24FE0B80C75713" box="[694,722,511,533]" captionStart="Figure 14" captionStartId="20.[144,223,1834,1853]" captionTargetBox="[149,1418,197,1803]" captionTargetId="figure-0@20.[145,1421,195,1805]" captionTargetPageId="20" captionText="Figure 14. Reproductive system of Satsuma phoenicis sp. nov. (NMNS 005405-6). A, whole genitalia; B, interior of genitalia. Scale bar = 5 mm. See text for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5446731" httpUri="https://zenodo.org/record/5446731/files/figure.png" pageId="19" pageNumber="456">14</figureCitation>
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)
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="24E16572FFE6430BFF36FDD2800A575C" pageId="19" pageNumber="456" type="reference_group">
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||
<paragraph id="6C4436F9FFE6430BFF36FDD2800A575C" blockId="19.[164,779,550,602]" pageId="19" pageNumber="456">
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||
<treatmentCitationGroup id="4CEB11D7FFE6430BFF36FDD2800A575C" pageId="19" pageNumber="456">
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||
<taxonomicName id="ABFB4D7AFFE6430BFF36FDD28012573A" ID-CoL="7V2TQ" authority="(H. Adams)" baseAuthorityName="H. Adams" baseAuthorityYear="1870" box="[164,519,550,572]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="456" phylum="Mollusca" rank="species" species="albida">
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<emphasis id="5E8FEAEBFFE6430BFF36FDD28370573D" box="[164,357,550,571]" italics="true" pageId="19" pageNumber="456">Satsuma albida</emphasis>
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||
(H. Adams)
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||
</taxonomicName>
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||
,
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<treatmentCitation id="ED5A10E8FFE6430BFD8CFDD282FE575C" author="Chang KM & Chang HW & Hwang CC" page="25" pageId="19" pageNumber="456" year="1996">
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<bibRefCitation id="086A4B08FFE6430BFD8CFDD282FE575C" author="Chang KM & Chang HW & Hwang CC" pageId="19" pageNumber="456" pagination="25 - 30" refId="ref24521" refString="Chang KM, Chang HW, Hwang CC. 1996. Anatomy on Satsuma albida (H. Adams, 1870) from Taiwan (Pulmonata: Camaenidae). Bulletin of Malacology 20: 25 - 30." type="journal article" year="1996">
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Chang
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<emphasis id="5E8FEAEBFFE6430BFDEEFDD380A1573D" box="[636,692,550,572]" italics="true" pageId="19" pageNumber="456">et al.</emphasis>
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||
, 1996: 25–30
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||
</bibRefCitation>
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||
</treatmentCitation>
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||
, fig. 1 (shell). [non
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<taxonomicName id="ABFB4D7AFFE6430BFE5CFDB08002575F" ID-CoL="7V2TQ" baseAuthorityName="H. Adams" baseAuthorityYear="1870" box="[462,535,580,601]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="456" phylum="Mollusca" rank="species" species="albida">
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<emphasis id="5E8FEAEBFFE6430BFE5CFDB08002575F" box="[462,535,580,601]" italics="true" pageId="19" pageNumber="456">albida</emphasis>
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</taxonomicName>
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||
]
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="24E16572FFE6430DFF36FD6E8650578E" lastPageId="21" lastPageNumber="458" pageId="19" pageNumber="456" type="materials_examined">
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||
<paragraph id="6C4436F9FFE6430BFF36FD6E836957A9" blockId="19.[164,779,666,872]" box="[164,380,666,687]" pageId="19" pageNumber="456">
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||
<emphasis id="5E8FEAEBFFE6430BFF36FD6E836957A9" box="[164,380,666,687]" italics="true" pageId="19" pageNumber="456">Material examined</emphasis>
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||
</paragraph>
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||
<paragraph id="6C4436F9FFE6430BFF36FD4E83AA566E" blockId="19.[164,779,666,872]" pageId="19" pageNumber="456">
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<emphasis id="5E8FEAEBFFE6430BFF36FD4E834357C9" box="[164,342,698,719]" italics="true" pageId="19" pageNumber="456">Type specimen:</emphasis>
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||
<materialsCitation id="DC933CA4FFE6430BFEF5FD4D83AA566E" collectionCode="TMMT, NMNS, BMNH, ANSP, SMF" pageId="19" pageNumber="456" specimenCode="TMMT 0601, NMNS 005405-6, TMMT 0645, BMNH 20060772, ANSP 413687, SMF 329395" specimenCount="2" typeStatus="holotype">
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||
<typeStatus id="B340885BFFE6430BFEF5FD4D83C457C9" box="[359,465,697,719]" pageId="19" pageNumber="456" type="holotype">Holotype</typeStatus>
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||
:
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||
<specimenCode id="3C5D9E82FFE6430BFE76FD4E806E57C9" box="[484,635,697,719]" collectionCode="TMMT" pageId="19" pageNumber="456">TMMT 0601</specimenCode>
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||
(from type locality, dry shell, tissue in alcohol). Five
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<typeStatus id="B340885BFFE6430BFD01FD2C811257EB" box="[659,775,728,749]" pageId="19" pageNumber="456" type="paratype">paratypes</typeStatus>
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||
: all from type locality,
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||
<specimenCode id="3C5D9E82FFE6430BFE29FD028094560A" box="[443,641,758,780]" collectionCode="NMNS" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="19" pageNumber="456">NMNS 005405-6</specimenCode>
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||
(dry shell, tissue in alcohol, dissected);
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<specimenCode id="3C5D9E82FFE6430BFE6AFCE1809C562D" box="[504,649,789,811]" collectionCode="TMMT" pageId="19" pageNumber="456">TMMT 0645</specimenCode>
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(dry shell, tissue in alcohol);
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<specimenCode id="3C5D9E82FFE6430BFE10FCC08045564F" box="[386,592,819,841]" collectionCode="BMNH" pageId="19" pageNumber="456">BMNH 20060772</specimenCode>
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,
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<specimenCode id="3C5D9E82FFE6430BFDF2FCC08112564F" box="[608,775,819,841]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="19" pageNumber="456">ANSP 413687</specimenCode>
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,
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<specimenCode id="3C5D9E82FFE6430BFF36FCA68322566E" box="[164,311,850,872]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="19" pageNumber="456">SMF 329395</specimenCode>
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(dry shell).
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</materialsCitation>
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||
</paragraph>
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||
<paragraph id="6C4436F9FFE6430BFF36FC5D832056BB" blockId="19.[164,778,936,1019]" box="[164,309,936,958]" pageId="19" pageNumber="456">
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||
<emphasis id="5E8FEAEBFFE6430BFF36FC5D832056BB" box="[164,309,936,958]" italics="true" pageId="19" pageNumber="456">
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||
<typeStatus id="B340885BFFE6430BFF36FC5D82CD56B8" box="[164,216,937,958]" pageId="19" pageNumber="456">Type</typeStatus>
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locality
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</emphasis>
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</paragraph>
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<paragraph id="6C4436F9FFE6430BFF36FC33834A56FD" blockId="19.[164,778,936,1019]" pageId="19" pageNumber="456">
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<materialsCitation id="DC933CA4FFE6430BFF36FC33834F56FD" country="Taiwan" location="Nantou County" pageId="19" pageNumber="456" specimenCount="1" stateProvince="Nantou" typeStatus="holotype">
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Fenghuanggu, Lugu,
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<collectingRegion id="AE3FF81BFFE6430BFE0FFC33805A56DA" box="[413,591,967,988]" country="Taiwan" name="Nantou" pageId="19" pageNumber="456">Nantou County</collectingRegion>
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, central
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<collectingCountry id="14EC7669FFE6430BFD24FC33811F56DB" box="[694,778,967,989]" name="Taiwan" pageId="19" pageNumber="456">Taiwan</collectingCountry>
|
||
(
|
||
<figureCitation id="F4C02A7CFFE6430BFF3EFC1182F856FC" box="[172,237,997,1019]" captionStart="Figure 1" captionStartId="1.[164,243,1882,1901]" captionTargetBox="[387,1219,874,1853]" captionTargetId="graphics-312@1.[386,1220,874,1853]" captionTargetPageId="1" captionText="Figure 1. Sampling localities of all taxa in Taiwan. Bold line, Central Range; thin line, administrative boundary." figureDoi="http://doi.org/10.5281/zenodo.5446702" httpUri="https://zenodo.org/record/5446702/files/figure.png" pageId="19" pageNumber="456">Fig. 1</figureCitation>
|
||
,
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||
<tableCitation id="21790342FFE6430BFF6FFC11834556FD" box="[253,336,997,1019]" captionStart="Table 1" captionStartId="2.[144,208,200,219]" captionText="Table 1. List of the sampling localities showing altitude (metres) and WGS84 coordinate" pageId="19" pageNumber="456">Table 1</tableCitation>
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||
)
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||
</materialsCitation>
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||
.
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||
</paragraph>
|
||
<paragraph id="6C4436F9FFE6430BFCA8FF3281BF55DD" blockId="19.[826,1441,198,342]" box="[826,938,198,219]" pageId="19" pageNumber="456">
|
||
<emphasis id="5E8FEAEBFFE6430BFCA8FF3281BF55DD" box="[826,938,198,219]" italics="true" pageId="19" pageNumber="456">Diagnosis</emphasis>
|
||
</paragraph>
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||
<paragraph id="6C4436F9FFE6430BFCA8FF1081825450" blockId="19.[826,1441,198,342]" pageId="19" pageNumber="456">Shell and soft body white with grey pedal stripe; proximal penis present, swollen, grooved; principal pilaster present, bifurcate, not formed in single strong pilaster.</paragraph>
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||
<paragraph id="6C4436F9FFE6430BFCA8FE7D81A45498" blockId="19.[826,1441,393,476]" box="[826,945,393,414]" pageId="19" pageNumber="456">
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||
<emphasis id="5E8FEAEBFFE6430BFCA8FE7D81A45498" box="[826,945,393,414]" italics="true" pageId="19" pageNumber="456">Etymology</emphasis>
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||
</paragraph>
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||
<paragraph id="6C4436F9FFE6430BFCA8FE5C86BC54DA" blockId="19.[826,1441,393,476]" pageId="19" pageNumber="456">
|
||
L.
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||
<taxonomicName id="ABFB4D7AFFE6430BFCC9FE5381D054BA" authorityName="Wu & Hwang & Lin" authorityYear="2008" box="[859,965,423,444]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="456" phylum="Mollusca" rank="species" species="phoenicis">
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||
<emphasis id="5E8FEAEBFFE6430BFCC9FE5381D054BA" box="[859,965,423,444]" italics="true" pageId="19" pageNumber="456">phoenicis</emphasis>
|
||
</taxonomicName>
|
||
: phoenix, translation of the
|
||
<typeStatus id="B340885BFFE6430BFA81FE5C875154BB" box="[1299,1348,424,445]" pageId="19" pageNumber="456">type</typeStatus>
|
||
locality ‘phoenix valley’ (Fenghuanggu).
|
||
</paragraph>
|
||
<paragraph id="6C4436F9FFE6430BFCA8FDFB81AE5722" blockId="19.[826,1442,527,916]" box="[826,955,527,548]" pageId="19" pageNumber="456">
|
||
<emphasis id="5E8FEAEBFFE6430BFCA8FDFB81AE5722" box="[826,955,527,548]" italics="true" pageId="19" pageNumber="456">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="6C4436F9FFE6430BFCA8FDD986325692" blockId="19.[826,1442,527,916]" pageId="19" pageNumber="456">
|
||
<emphasis id="5E8FEAEBFFE6430BFCA8FDD981605744" box="[826,885,557,578]" italics="true" pageId="19" pageNumber="456">Shell</emphasis>
|
||
(
|
||
<figureCitation id="F4C02A7CFFE6430BFC15FDD981C35745" box="[903,982,557,579]" captionStart="Figure 13" captionStartId="19.[164,243,1853,1872]" captionTargetBox="[389,1219,1092,1824]" captionTargetId="figure-290@19.[387,1221,1090,1824]" captionTargetPageId="19" captionText="Figure 13. Satsuma phoenicis sp. nov. A–D, holotype (TMMT 0601, shell height = 19.4 mm). E, living specimen. Arrow indicates the pre-apertural constriction." figureDoi="http://doi.org/10.5281/zenodo.5446729" httpUri="https://zenodo.org/record/5446729/files/figure.png" pageId="19" pageNumber="456">Fig. 13</figureCitation>
|
||
): Dextral, conical, medium sized. Apex obtuse. Whorls expanded. Periphery bluntly angulated, extending to peristome. Base expanded. Preapertural constriction behind outer and inferior lips present. Surface glossy, with spiral striae. Shell colour milky white, covered with fine periostracum. Aperture diagonal, ovate-lunate. Peristome thin, expanded. Inferior lip reflected, curved downward. Superior columellar lip vertical, reflected. Umbilicus covered by columellar lip, crevice-like. Junction between oblique inferior columellar lip and inferior lip smoothly curved.
|
||
</paragraph>
|
||
<paragraph id="6C4436F9FFE6430BFCA8FC33870556FC" blockId="19.[826,1441,967,1019]" pageId="19" pageNumber="456">
|
||
<emphasis id="5E8FEAEBFFE6430BFCA8FC3381F256DA" box="[826,999,967,988]" italics="true" pageId="19" pageNumber="456">Band or stain:</emphasis>
|
||
Band or stain is not present in this species. Band formula = 0 0 0 0 0 0 0 0.
|
||
</paragraph>
|
||
<caption id="38846671FFE6430BFF36F8C9805B5268" ID-DOI="http://doi.org/10.5281/zenodo.5446729" ID-Zenodo-Dep="5446729" httpUri="https://zenodo.org/record/5446729/files/figure.png" pageId="19" pageNumber="456" startId="19.[164,243,1853,1872]" targetBox="[389,1219,1092,1824]" targetPageId="19">
|
||
<paragraph id="6C4436F9FFE6430BFF36F8C9805B5268" blockId="19.[164,1441,1852,1902]" pageId="19" pageNumber="456">
|
||
<emphasis id="5E8FEAEBFFE6430BFF36F8C9804B5256" bold="true" box="[164,606,1853,1872]" pageId="19" pageNumber="456">
|
||
Figure 13.
|
||
<taxonomicName id="ABFB4D7AFFE6430BFEB4F8C980145256" authority="Wu & Hwang & Lin, 2008" authorityName="Wu & Hwang & Lin" authorityYear="2008" box="[294,513,1853,1872]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="456" phylum="Mollusca" rank="species" species="phoenicis" status="sp. nov.">
|
||
<emphasis id="5E8FEAEBFFE6430BFEB4F8C980145256" bold="true" box="[294,513,1853,1872]" italics="true" pageId="19" pageNumber="456">Satsuma phoenicis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="45BC5790FFE6430BFD9AF8C9804B5256" box="[520,606,1853,1872]" pageId="19" pageNumber="456" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A–D, holotype (TMMT 0601, shell height = 19.4 mm). E, living specimen. Arrow indicates the pre-apertural constriction.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="38846671FFE1430CFF02F8DE80CB525D" ID-DOI="http://doi.org/10.5281/zenodo.5446731" ID-Zenodo-Dep="5446731" httpUri="https://zenodo.org/record/5446731/files/figure.png" pageId="20" pageNumber="457" startId="20.[144,223,1834,1853]" targetBox="[149,1418,197,1803]" targetPageId="20">
|
||
<paragraph id="6C4436F9FFE1430CFF02F8DE80CB525D" blockId="20.[144,1421,1834,1883]" pageId="20" pageNumber="457">
|
||
<emphasis id="5E8FEAEBFFE1430CFF02F8DE83105238" bold="true" box="[144,261,1834,1854]" pageId="20" pageNumber="457">Figure 14.</emphasis>
|
||
Reproductive system of
|
||
<emphasis id="5E8FEAEBFFE1430CFD86F8DF81435238" bold="true" box="[532,854,1834,1854]" pageId="20" pageNumber="457">
|
||
<taxonomicName id="ABFB4D7AFFE1430CFD86F8DF80E6523B" authority="Wu & Hwang & Lin, 2008" authorityName="Wu & Hwang & Lin" authorityYear="2008" box="[532,755,1834,1854]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="457" phylum="Mollusca" rank="species" species="phoenicis" status="sp. nov.">
|
||
<emphasis id="5E8FEAEBFFE1430CFD86F8DF80E6523B" bold="true" box="[532,755,1834,1854]" italics="true" pageId="20" pageNumber="457">Satsuma phoenicis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="45BC5790FFE1430CFD6FF8DF81435238" box="[765,854,1835,1854]" pageId="20" pageNumber="457" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(NMNS 005405-6). A, whole genitalia; B, interior of genitalia. Scale bar = 5 mm. See text for abbreviations.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="6C4436F9FFE0430DFF36FF32806A57C3" blockId="21.[164,779,198,709]" pageId="21" pageNumber="458">
|
||
<emphasis id="5E8FEAEBFFE0430DFF36FF32839E55DA" box="[164,395,198,220]" italics="true" pageId="21" pageNumber="458">Reproductive system</emphasis>
|
||
(
|
||
<figureCitation id="F4C02A7CFFE0430DFE0BFF3283F255DA" box="[409,487,198,220]" captionStart="Figure 14" captionStartId="20.[144,223,1834,1853]" captionTargetBox="[149,1418,197,1803]" captionTargetId="figure-0@20.[145,1421,195,1805]" captionTargetPageId="20" captionText="Figure 14. Reproductive system of Satsuma phoenicis sp. nov. (NMNS 005405-6). A, whole genitalia; B, interior of genitalia. Scale bar = 5 mm. See text for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5446731" httpUri="https://zenodo.org/record/5446731/files/figure.png" pageId="21" pageNumber="458">Fig. 14</figureCitation>
|
||
): Bursa stalk long, tapering with wiggly folds internally. Proximal vagina muscular, furrowed externally; middle vagina muscular, constrictive, with weak folds inside; distal vagina one-quarter length of vagina. Flagellum long, tapering smoothly towards tip. Penial caecum long, tapering with blunt tip; cecal pilasters two or three in number, surrounding epiphallic pore. Principal pilaster present, bifurcate, weak, not merged as single strong pilaster. Proximal penis widened, muscular, furrowed with a deep groove externally corresponding to strong, elevated, corrugated internal pilasters; middle penis short, suddenly constrictive from proximal penis, with weak and smooth pilasters inside; distal penis moderately slender, with one to two weak internal pilasters gradually weakened towards atrium. Three specimens were dissected.
|
||
</paragraph>
|
||
<paragraph id="6C4436F9FFE0430DFF36FD02833A560D" blockId="21.[164,779,758,841]" box="[164,303,758,779]" pageId="21" pageNumber="458">
|
||
<emphasis id="5E8FEAEBFFE0430DFF36FD02833A560D" box="[164,303,758,779]" italics="true" pageId="21" pageNumber="458">Distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="6C4436F9FFE0430DFF36FCE18313564F" blockId="21.[164,779,758,841]" pageId="21" pageNumber="458">
|
||
Lugu area,
|
||
<collectingRegion id="AE3FF81BFFE0430DFEA6FCE183FF562F" box="[308,490,788,810]" country="Taiwan" name="Nantou" pageId="21" pageNumber="458">Nantou County</collectingRegion>
|
||
, central
|
||
<collectingCountry id="14EC7669FFE0430DFDCBFCE080B8562C" box="[601,685,788,810]" name="Taiwan" pageId="21" pageNumber="458">Taiwan</collectingCountry>
|
||
(
|
||
<figureCitation id="F4C02A7CFFE0430DFD51FCE08111562F" box="[707,772,788,810]" captionStart="Figure 1" captionStartId="1.[164,243,1882,1901]" captionTargetBox="[387,1219,874,1853]" captionTargetId="graphics-312@1.[386,1220,874,1853]" captionTargetPageId="1" captionText="Figure 1. Sampling localities of all taxa in Taiwan. Bold line, Central Range; thin line, administrative boundary." figureDoi="http://doi.org/10.5281/zenodo.5446702" httpUri="https://zenodo.org/record/5446702/files/figure.png" pageId="21" pageNumber="458">Fig. 1</figureCitation>
|
||
,
|
||
<tableCitation id="21790342FFE0430DFF36FCC782E3564F" box="[164,246,819,841]" captionStart="Table 1" captionStartId="2.[144,208,200,219]" captionText="Table 1. List of the sampling localities showing altitude (metres) and WGS84 coordinate" pageId="21" pageNumber="458">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6C4436F9FFE0430DFF36FC8D831C5688" blockId="21.[164,779,889,1003]" box="[164,265,889,910]" pageId="21" pageNumber="458">
|
||
<emphasis id="5E8FEAEBFFE0430DFF36FC8D831C5688" box="[164,265,889,910]" italics="true" pageId="21" pageNumber="458">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="6C4436F9FFE0430DFF36FC6C80E956ED" blockId="21.[164,779,889,1003]" pageId="21" pageNumber="458">
|
||
Animals are arboreal, herbivores and nocturnal perching under leaves
|
||
<quantity id="AB039B1CFFE0430DFE28FC4383EA56CA" box="[442,511,951,972]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="21" pageNumber="458" unit="m" value="2.5" valueMax="3.0" valueMin="2.0">2–3 m</quantity>
|
||
above ground. Adults were found during summer and autumn (
|
||
<figureCitation id="F4C02A7CFFE0430DFD1FFC2180E556ED" box="[653,752,981,1003]" captionStart="Figure 13" captionStartId="19.[164,243,1853,1872]" captionTargetBox="[389,1219,1092,1824]" captionTargetId="figure-290@19.[387,1221,1090,1824]" captionTargetPageId="19" captionText="Figure 13. Satsuma phoenicis sp. nov. A–D, holotype (TMMT 0601, shell height = 19.4 mm). E, living specimen. Arrow indicates the pre-apertural constriction." figureDoi="http://doi.org/10.5281/zenodo.5446729" httpUri="https://zenodo.org/record/5446729/files/figure.png" pageId="21" pageNumber="458">Fig. 13E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6C4436F9FFE0430DFCC1FF328650578E" blockId="21.[826,1441,198,648]" pageId="21" pageNumber="458">
|
||
The swollen and grooved proximal penis and bifurcate principal pilaster is unique among species in west
|
||
<collectingCountry id="14EC7669FFE0430DFC37FEF781E8541F" box="[933,1021,259,281]" name="Taiwan" pageId="21" pageNumber="458">Taiwan</collectingCountry>
|
||
. The shell of the present species differs from that of
|
||
<taxonomicName id="ABFB4D7AFFE0430DFBBAFED686815431" baseAuthorityName="H. Adams" baseAuthorityYear="1870" box="[1064,1172,290,311]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="458" phylum="Mollusca" rank="species" species="albida">
|
||
<emphasis id="5E8FEAEBFFE0430DFBBAFED686815431" box="[1064,1172,290,311]" italics="true" pageId="21" pageNumber="458">S. albida</emphasis>
|
||
</taxonomicName>
|
||
in its larger size, and being thinner, translucent and more rigidness; differs from that of
|
||
<taxonomicName id="ABFB4D7AFFE0430DFBBFFEAB866C5472" authorityName="Wu & Hwang & Lin" authorityYear="2008" box="[1069,1145,351,372]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="458" phylum="Mollusca" rank="species" species="lini">
|
||
<emphasis id="5E8FEAEBFFE0430DFBBFFEAB866C5472" box="[1069,1145,351,372]" italics="true" pageId="21" pageNumber="458">S. lini</emphasis>
|
||
</taxonomicName>
|
||
in larger measurements (
|
||
<tableCitation id="21790342FFE0430DFCD3FE8981825492" box="[833,919,381,404]" captionStart="Table 2" captionStartId="6.[144,208,200,219]" captionText="Table 2. Satsumaalbida species complex showing measurements of holotype/lectotype (italic)" pageId="21" pageNumber="458">Table 2</tableCitation>
|
||
), more flattened base and inferior lip, more blunt periphery, more curved downward inferior lip and, hence, smooth junction between the inferior lip and the columellar lip. This species is morphologically similar to
|
||
<taxonomicName id="ABFB4D7AFFE0430DFB98FE0C864E570B" authority="Wu & Hwang & Lin, 2008" authorityName="Wu & Hwang & Lin" authorityYear="2008" box="[1034,1115,504,525]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="458" phylum="Mollusca" rank="species" species="lini" status="sp. nov.">
|
||
<emphasis id="5E8FEAEBFFE0430DFB98FE0C864E570B" box="[1034,1115,504,525]" italics="true" pageId="21" pageNumber="458">S. lini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="45BC5790FFE0430DFBFEFE0C86D8570B" box="[1132,1229,504,525]" pageId="21" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, but the genetic relationship was calculated to be closer to
|
||
<taxonomicName id="ABFB4D7AFFE0430DFAD4FDE3818F574D" authority="Wu & Hwang & Lin, 2008" authorityName="Wu & Hwang & Lin" authorityYear="2008" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="458" phylum="Mollusca" rank="species" species="careocaecum" status="sp. nov.">
|
||
<emphasis id="5E8FEAEBFFE0430DFAD4FDE3818F574D" italics="true" pageId="21" pageNumber="458">S. careocaecum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="45BC5790FFE0430DFC35FDC18617574C" box="[935,1026,565,586]" pageId="21" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
|
||
The photographed shell in
|
||
<bibRefCitation id="086A4B08FFE0430DFAC4FDC181DD576C" author="Chang KM & Chang HW & Hwang CC" pageId="21" pageNumber="458" pagination="25 - 30" refId="ref24521" refString="Chang KM, Chang HW, Hwang CC. 1996. Anatomy on Satsuma albida (H. Adams, 1870) from Taiwan (Pulmonata: Camaenidae). Bulletin of Malacology 20: 25 - 30." type="journal article" year="1996">
|
||
Chang
|
||
<emphasis id="5E8FEAEBFFE0430DFCA8FDA08167576F" box="[826,882,596,617]" italics="true" pageId="21" pageNumber="458">et al.</emphasis>
|
||
(1996
|
||
</bibRefCitation>
|
||
,
|
||
<figureCitation id="F4C02A7CFFE0430DFC72FDA0860A576C" box="[992,1055,596,618]" captionStart="Figure 1" captionStartId="1.[164,243,1882,1901]" captionTargetBox="[387,1219,874,1853]" captionTargetId="graphics-312@1.[386,1220,874,1853]" captionTargetPageId="1" captionText="Figure 1. Sampling localities of all taxa in Taiwan. Bold line, Central Range; thin line, administrative boundary." figureDoi="http://doi.org/10.5281/zenodo.5446702" httpUri="https://zenodo.org/record/5446702/files/figure.png" pageId="21" pageNumber="458">Fig. 1</figureCitation>
|
||
) belongs to this species (see remarks in
|
||
<taxonomicName id="ABFB4D7AFFE0430DFC5BFD8786235781" baseAuthorityName="H. Adams" baseAuthorityYear="1870" box="[969,1078,626,648]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="458" phylum="Mollusca" rank="species" species="albida">
|
||
<emphasis id="5E8FEAEBFFE0430DFC5BFD8786235781" box="[969,1078,626,648]" italics="true" pageId="21" pageNumber="458">S. albida</emphasis>
|
||
</taxonomicName>
|
||
.)
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |