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<document id="196D0726880EBE5C622582F50BD38C3A" ID-DOI="10.1111/j.1096-3642.2008.00415.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5446700" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630461491835" checkinUser="felipe" docAuthor="Wu, Shu-Ping, Hwang, Chung-Chi &amp; Lin, Yao-Sung" docDate="2008" docId="E45287EFFFDA4329FCE7FEBA86B65472" docLanguage="en" docName="j.1096-3642.2008.00415.x.pdf" docOrigin="Zoological Journal of the Linnean Society 154 (3)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00415.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Satsuma pilsbryi Wu &amp; Hwang &amp; Lin 2008, SP. NOV." docType="treatment" docVersion="4" lastPageNumber="486" masterDocId="186BFF97FFF54318FF92FFF482155506" masterDocTitle="Systematic revision of the arboreal snail Satsuma albida species complex (Mollusca: Camaenidae) with descriptions of 14 new species from Taiwan" masterLastPageNumber="493" masterPageNumber="437" pageNumber="484" updateTime="1699279363994" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="CDBE2E4CF4A17770E6A0B23264422611">Systematic revision of the arboreal snail Satsuma albida species complex (Mollusca: Camaenidae) with descriptions of 14 new species from Taiwan</mods:title>
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<mods:namePart id="1E60FEE17C8E82E35A6F3899F6B1B670">Wu, Shu-Ping</mods:namePart>
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<mods:namePart id="710ADE178D1B67CFE193B79D47C706EC">Hwang, Chung-Chi</mods:namePart>
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<mods:namePart id="7BFBE510FF160CF4D020E5516CBE2A5E">Lin, Yao-Sung</mods:namePart>
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<treatment id="E45287EFFFDA4329FCE7FEBA86B65472" ID-DOI="http://doi.org/10.5281/zenodo.5492614" ID-Zenodo-Dep="5492614" LSID="urn:lsid:plazi:treatment:E45287EFFFDA4329FCE7FEBA86B65472" httpUri="http://treatment.plazi.org/id/E45287EFFFDA4329FCE7FEBA86B65472" lastPageId="49" lastPageNumber="486" pageId="47" pageNumber="484">
<subSubSection id="24E16572FFDA4337FCE7FEBA87735463" box="[885,1382,334,357]" pageId="47" pageNumber="484" type="nomenclature">
<paragraph id="6C4436F9FFDA4337FCE7FEBA87735463" blockId="47.[885,1382,334,357]" box="[885,1382,334,357]" pageId="47" pageNumber="484">
<heading id="370C8195FFDA4337FCE7FEBA87735463" box="[885,1382,334,357]" centered="true" fontSize="9" level="2" pageId="47" pageNumber="484" reason="2">
<taxonomicName id="ABFB4D7AFFDA4337FCE7FEBA86435465" ID-CoL="6XQBG" authority="Wu &amp; Hwang &amp; Lin, 2008" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[885,1110,334,355]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="47" pageNumber="484" phylum="Mollusca" rank="species" species="pilsbryi" status="sp. nov.">SATSUMA PILSBRYI</taxonomicName>
<emphasis id="5E8FEAEBFFDA4337FBF3FEA686D35462" bold="true" box="[1121,1222,335,356]" pageId="47" pageNumber="484">
<taxonomicNameLabel id="45BC5790FFDA4337FBF3FEA686D35462" box="[1121,1222,335,356]" pageId="47" pageNumber="484" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="F4C02A7CFFDA4337FB4AFEBB87275462" box="[1240,1330,335,356]" captionStart="Figure 37" captionStartId="47.[164,243,1853,1872]" captionTargetBox="[389,1219,1311,1823]" captionTargetId="figure-399@47.[387,1219,1309,1823]" captionTargetPageId="47" captionText="Figure 37. Satsuma pilsbryi sp. nov. AD, holotype (TMMT 0614, shell height = 17.9 mm). E, F, paratype (TMMT 0644, shell height = 17.6 mm). G, living specimen. Arrow indicates the pre-apertural constriction." figureDoi="http://doi.org/10.5281/zenodo.5446784" httpUri="https://zenodo.org/record/5446784/files/figure.png" pageId="47" pageNumber="484">FIGS 37</figureCitation>
,
<figureCitation id="F4C02A7CFFDA4337FAD0FEBB874B5463" box="[1346,1374,335,357]" captionStart="Figure 38" captionStartId="48.[144,223,1795,1814]" captionTargetBox="[148,1421,196,1761]" captionTargetId="figure-0@48.[144,1424,193,1765]" captionTargetPageId="48" captionText="Figure 38. Reproductive system of Satsuma pilsbryi sp. nov. (holotype, TMMT 0614). A, whole genitalia; B, interior of genitalia. Scale bar = 5 mm. See text for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5446786" httpUri="https://zenodo.org/record/5446786/files/figure.png" pageId="47" pageNumber="484">38</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="24E16572FFDA4329FCA8FE8186B65472" lastPageId="49" lastPageNumber="486" pageId="47" pageNumber="484" type="materials_examined">
<paragraph id="6C4436F9FFDA4337FCA8FE818607548C" blockId="47.[826,1441,373,548]" box="[826,1042,373,394]" pageId="47" pageNumber="484">
<emphasis id="5E8FEAEBFFDA4337FCA8FE818607548C" box="[826,1042,373,394]" italics="true" pageId="47" pageNumber="484">Material examined</emphasis>
</paragraph>
<paragraph id="6C4436F9FFDA4337FCA8FE6086F55722" blockId="47.[826,1441,373,548]" pageId="47" pageNumber="484">
<emphasis id="5E8FEAEBFFDA4337FCA8FE6081F954AF" box="[826,1004,404,425]" italics="true" pageId="47" pageNumber="484">Type specimen:</emphasis>
<materialsCitation id="DC933CA4FFDA4337FC6FFE6086F55722" collectionCode="TMMT, BMNH, ANSP" pageId="47" pageNumber="484" specimenCode="TMMT 0614, TMMT 0644, TMMT 0665-0670, BMNH 20060766, ANSP 413684" specimenCount="2" typeStatus="holotype">
<typeStatus id="B340885BFFDA4337FC6FFE60867254AC" box="[1021,1127,404,426]" pageId="47" pageNumber="484" type="holotype">Holotype</typeStatus>
:
<specimenCode id="3C5D9E82FFDA4337FBE9FE60870454AC" box="[1147,1297,404,426]" collectionCode="TMMT" pageId="47" pageNumber="484">TMMT 0614</specimenCode>
(from type locality, dry shell, tissue in alcohol, dissected). Nine
<typeStatus id="B340885BFFDA4337FCA8FE2581BB54E0" box="[826,942,465,486]" pageId="47" pageNumber="484" type="paratype">paratypes</typeStatus>
: all from type locality,
<specimenCode id="3C5D9E82FFDA4337FB27FE25875054E1" box="[1205,1349,465,487]" collectionCode="TMMT" pageId="47" pageNumber="484">TMMT 0644</specimenCode>
,
<specimenCode id="3C5D9E82FFDA4337FAC0FE2581A05703" collectionCode="TMMT" pageId="47" pageNumber="484">TMMT 06650670</specimenCode>
(dry shell, tissue in alcohol);
<specimenCode id="3C5D9E82FFDA4337FADDFE0481BF5722" collectionCode="BMNH" pageId="47" pageNumber="484">BMNH 20060766</specimenCode>
,
<specimenCode id="3C5D9E82FFDA4337FC25FDFA864D5722" box="[951,1112,526,548]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="47" pageNumber="484">ANSP 413684</specimenCode>
(dry shell).
</materialsCitation>
</paragraph>
<paragraph id="6C4436F9FFDA4337FCA8FDB981DE5767" blockId="47.[826,1441,588,672]" box="[826,971,588,610]" pageId="47" pageNumber="484">
<emphasis id="5E8FEAEBFFDA4337FCA8FDB981DE5767" box="[826,971,588,610]" italics="true" pageId="47" pageNumber="484">
<typeStatus id="B340885BFFDA4337FCA8FDB9817B5764" box="[826,878,589,610]" pageId="47" pageNumber="484">Type</typeStatus>
locality
</emphasis>
</paragraph>
<paragraph id="6C4436F9FFDA4337FCA8FD9F818957A6" blockId="47.[826,1441,588,672]" pageId="47" pageNumber="484">
<materialsCitation id="DC933CA4FFDA4337FCA8FD9F818257A6" country="Taiwan" location="Taitung County" pageId="47" pageNumber="484" specimenCount="1" stateProvince="Taitung" typeStatus="holotype">
Taiyuan,
<collectingRegion id="AE3FF81BFFDA4337FC22FD9F867B5786" box="[944,1134,619,641]" country="Taiwan" name="Taitung" pageId="47" pageNumber="484">Taitung County</collectingRegion>
, eastern
<collectingCountry id="14EC7669FFDA4337FB79FD9F872A5787" box="[1259,1343,619,641]" name="Taiwan" pageId="47" pageNumber="484">Taiwan</collectingCountry>
(
<figureCitation id="F4C02A7CFFDA4337FACBFD9F878F5786" box="[1369,1434,619,641]" captionStart="Figure 1" captionStartId="1.[164,243,1882,1901]" captionTargetBox="[387,1219,874,1853]" captionTargetId="graphics-312@1.[386,1220,874,1853]" captionTargetPageId="1" captionText="Figure 1. Sampling localities of all taxa in Taiwan. Bold line, Central Range; thin line, administrative boundary." figureDoi="http://doi.org/10.5281/zenodo.5446702" httpUri="https://zenodo.org/record/5446702/files/figure.png" pageId="47" pageNumber="484">Fig. 1</figureCitation>
,
<tableCitation id="21790342FFDA4337FCA8FD7D819957A6" box="[826,908,649,672]" captionStart="Table 1" captionStartId="2.[144,208,200,219]" captionText="Table 1. List of the sampling localities showing altitude (metres) and WGS84 coordinate" pageId="47" pageNumber="484">Table 1</tableCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="6C4436F9FFDA4337FCA8FD3C81BF57DB" blockId="47.[826,1441,712,887]" box="[826,938,712,733]" pageId="47" pageNumber="484">
<emphasis id="5E8FEAEBFFDA4337FCA8FD3C81BF57DB" box="[826,938,712,733]" italics="true" pageId="47" pageNumber="484">Diagnosis</emphasis>
</paragraph>
<paragraph id="6C4436F9FFDA4337FCA8FD1281825671" blockId="47.[826,1441,712,887]" pageId="47" pageNumber="484">Shell and soft body yellowish without pedal stripe; shell conical with small diameter; pre-apertural constriction present behind outer lip; base of bursa stalk conical; principal pilaster short; proximal penis short, swollen.</paragraph>
<paragraph id="6C4436F9FFDA4337FCA8FC6B81A456B2" blockId="47.[826,1441,927,1040]" box="[826,945,927,948]" pageId="47" pageNumber="484">
<emphasis id="5E8FEAEBFFDA4337FCA8FC6B81A456B2" box="[826,945,927,948]" italics="true" pageId="47" pageNumber="484">Etymology</emphasis>
</paragraph>
<paragraph id="6C4436F9FFDA4337FCA8FC4986245116" blockId="47.[826,1441,927,1040]" pageId="47" pageNumber="484">This species is named after the American malacologist Henry A. Pilsbry (18621957) who described two species of this group.</paragraph>
<paragraph id="6C4436F9FFDA4337FCA8FBCD81AE5148" blockId="47.[826,1441,1081,1256]" box="[826,955,1081,1102]" pageId="47" pageNumber="484">
<emphasis id="5E8FEAEBFFDA4337FCA8FBCD81AE5148" box="[826,955,1081,1102]" italics="true" pageId="47" pageNumber="484">Description</emphasis>
</paragraph>
<paragraph id="6C4436F9FFDA4329FCA8FBA3830B54B4" blockId="47.[826,1441,1081,1256]" lastBlockId="49.[164,779,198,434]" lastPageId="49" lastPageNumber="486" pageId="47" pageNumber="484">
<emphasis id="5E8FEAEBFFDA4337FCA8FBA38160516A" box="[826,885,1111,1132]" italics="true" pageId="47" pageNumber="484">Shell</emphasis>
(
<figureCitation id="F4C02A7CFFDA4337FC1BFBA381CD516B" box="[905,984,1111,1133]" captionStart="Figure 37" captionStartId="47.[164,243,1853,1872]" captionTargetBox="[389,1219,1311,1823]" captionTargetId="figure-399@47.[387,1219,1309,1823]" captionTargetPageId="47" captionText="Figure 37. Satsuma pilsbryi sp. nov. AD, holotype (TMMT 0614, shell height = 17.9 mm). E, F, paratype (TMMT 0644, shell height = 17.6 mm). G, living specimen. Arrow indicates the pre-apertural constriction." figureDoi="http://doi.org/10.5281/zenodo.5446784" httpUri="https://zenodo.org/record/5446784/files/figure.png" pageId="47" pageNumber="484">Fig. 37</figureCitation>
): Dextral, conical, thin, medium sized to small. Apex obtuse. Whorls inflated. Periphery bluntly angulated, arc-like. Base expanded. Preapertural constriction behind outer lip only. Shell colour light yellow. Periostracum thin. Surface evenly smooth, with fine axial and spiral striae. Aperture diagonal, ovate. Junction between outer lip and inferior lip an angulated curve. Peristome thin, expanded at inferior lip and outer lip. Inferior lip curved downward. Columellar lip vertical, reflected. Umbilicus mostly covered by columellar lip, crevice-like. Junction between columellar lip and inferior lip obtusely angulated.
</paragraph>
<caption id="38846671FFDA4337FF36F8C986A25268" ID-DOI="http://doi.org/10.5281/zenodo.5446784" ID-Zenodo-Dep="5446784" httpUri="https://zenodo.org/record/5446784/files/figure.png" pageId="47" pageNumber="484" startId="47.[164,243,1853,1872]" targetBox="[389,1219,1311,1823]" targetPageId="47">
<paragraph id="6C4436F9FFDA4337FF36F8C986A25268" blockId="47.[164,1441,1852,1902]" pageId="47" pageNumber="484">
<emphasis id="5E8FEAEBFFDA4337FF36F8C980405256" bold="true" box="[164,597,1853,1872]" pageId="47" pageNumber="484">
Figure 37.
<taxonomicName id="ABFB4D7AFFDA4337FEB4F8C983E55256" authority="Wu &amp; Hwang &amp; Lin, 2008" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[294,496,1853,1872]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="47" pageNumber="484" phylum="Mollusca" rank="species" species="pilsbryi" status="sp. nov.">
<emphasis id="5E8FEAEBFFDA4337FEB4F8C983E55256" bold="true" box="[294,496,1853,1872]" italics="true" pageId="47" pageNumber="484">Satsuma pilsbryi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45BC5790FFDA4337FE69F8C980405256" box="[507,597,1853,1872]" pageId="47" pageNumber="484" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
AD, holotype (TMMT 0614, shell height = 17.9 mm). E, F, paratype (TMMT 0644, shell height = 17.6 mm). G, living specimen. Arrow indicates the pre-apertural constriction.
</paragraph>
</caption>
<caption id="38846671FFC54328FF02F8F780EF5235" ID-DOI="http://doi.org/10.5281/zenodo.5446786" ID-Zenodo-Dep="5446786" httpUri="https://zenodo.org/record/5446786/files/figure.png" pageId="48" pageNumber="485" startId="48.[144,223,1795,1814]" targetBox="[148,1421,196,1761]" targetPageId="48">
<paragraph id="6C4436F9FFC54328FF02F8F780EF5235" blockId="48.[144,1421,1794,1843]" pageId="48" pageNumber="485">
<emphasis id="5E8FEAEBFFC54328FF02F8F783105210" bold="true" box="[144,261,1795,1814]" pageId="48" pageNumber="485">Figure 38.</emphasis>
Reproductive system of
<emphasis id="5E8FEAEBFFC54328FD80F8F781295210" bold="true" box="[530,828,1794,1814]" pageId="48" pageNumber="485">
<taxonomicName id="ABFB4D7AFFC54328FD80F8F780CF5213" authority="Wu &amp; Hwang &amp; Lin, 2008" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[530,730,1794,1814]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="48" pageNumber="485" phylum="Mollusca" rank="species" species="pilsbryi" status="sp. nov.">
<emphasis id="5E8FEAEBFFC54328FD80F8F780CF5213" bold="true" box="[530,730,1794,1814]" italics="true" pageId="48" pageNumber="485">Satsuma pilsbryi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45BC5790FFC54328FD71F8F781295210" box="[739,828,1795,1814]" pageId="48" pageNumber="485" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype, TMMT 0614). A, whole genitalia; B, interior of genitalia. Scale bar = 5 mm. See text for abbreviations.
</paragraph>
</caption>
<paragraph id="6C4436F9FFC44329FF36FE2D83485781" blockId="49.[164,779,473,648]" pageId="49" pageNumber="486">
<emphasis id="5E8FEAEBFFC44329FF36FE2D834054E9" box="[164,341,473,495]" italics="true" pageId="49" pageNumber="486">Band or stain:</emphasis>
Polymorphism exists in this taxon. Some individuals lack colour band and stain, whilst some exhibit light red-brown apical spot, subperipheral band or umbilicus stain. Inferior and outer lips are colourless. Band formula = 0 0 0 0 0 0 0 0; 10005070.
</paragraph>
<paragraph id="6C4436F9FFC44329FF36FD5B80F65061" blockId="49.[164,779,687,1383]" pageId="49" pageNumber="486">
<emphasis id="5E8FEAEBFFC44329FF36FD5B839E57C3" box="[164,395,687,709]" italics="true" pageId="49" pageNumber="486">Reproductive system</emphasis>
(
<figureCitation id="F4C02A7CFFC44329FE08FD5B83FC57C0" box="[410,489,687,710]" captionStart="Figure 38" captionStartId="48.[144,223,1795,1814]" captionTargetBox="[148,1421,196,1761]" captionTargetId="figure-0@48.[144,1424,193,1765]" captionTargetPageId="48" captionText="Figure 38. Reproductive system of Satsuma pilsbryi sp. nov. (holotype, TMMT 0614). A, whole genitalia; B, interior of genitalia. Scale bar = 5 mm. See text for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5446786" httpUri="https://zenodo.org/record/5446786/files/figure.png" pageId="49" pageNumber="486">Fig. 38</figureCitation>
): Bursa stalk short, fourfifths spermoviduct, tapering, swollen at base. Bursa copulatrix clavated. Proximal vagina muscular, swollen, smooth, with 14 pilasters internally; middle vagina short, constricted, thick-walled, with 17 weak and wiggly folds inside; distal vagina long, nearly one-third length of vagina, gradually constricted towards atrium, with low folds or fine wrinkles inside. Flagellum short and swollen at base, with apparently short and quickly tapering tip. Epiphallus short, with four pilasters inside. Penial caecum moderate, conical, blunt, smooth; cecal pilaster two in number; remaining inner walls of penial caecum contain several moderate ridges. Proximal penis short, swollen, smooth externally, with one deep groove on epiphallic side corresponding to principal pilaster internally; principal pilaster same length as penial caecum; middle penis constricted suddenly, regularly elongated, smooth externally, with seven to nine strong and smooth pilasters; distal penis short, only weak folds or fine wrinkles inside. Spinules present in vagina, penial caecum and proximal penis. Two individuals from the
<typeStatus id="B340885BFFC44329FE09FAA583D95060" box="[411,460,1361,1382]" pageId="49" pageNumber="486">type</typeStatus>
locality were dissected.
</paragraph>
<paragraph id="6C4436F9FFC44329FF36FA7A833A50A5" blockId="49.[164,778,1422,1536]" box="[164,303,1422,1443]" pageId="49" pageNumber="486">
<emphasis id="5E8FEAEBFFC44329FF36FA7A833A50A5" box="[164,303,1422,1443]" italics="true" pageId="49" pageNumber="486">Distribution</emphasis>
</paragraph>
<paragraph id="6C4436F9FFC44329FF36FA5883A85306" blockId="49.[164,778,1422,1536]" pageId="49" pageNumber="486">
This species was found in lowland forest habitats around the Taiyuan valley,
<collectingRegion id="AE3FF81BFFC44329FE7CFA3F80B350E6" box="[494,678,1483,1505]" country="Taiwan" name="Taitung" pageId="49" pageNumber="486">Taitung County</collectingRegion>
, eastern
<collectingCountry id="14EC7669FFC44329FF36FA1E82ED5306" box="[164,248,1514,1536]" name="Taiwan" pageId="49" pageNumber="486">Taiwan</collectingCountry>
(
<figureCitation id="F4C02A7CFFC44329FE98FA1E835E50F9" box="[266,331,1514,1536]" captionStart="Figure 1" captionStartId="1.[164,243,1882,1901]" captionTargetBox="[387,1219,874,1853]" captionTargetId="graphics-312@1.[386,1220,874,1853]" captionTargetPageId="1" captionText="Figure 1. Sampling localities of all taxa in Taiwan. Bold line, Central Range; thin line, administrative boundary." figureDoi="http://doi.org/10.5281/zenodo.5446702" httpUri="https://zenodo.org/record/5446702/files/figure.png" pageId="49" pageNumber="486">Fig. 1</figureCitation>
,
<tableCitation id="21790342FFC44329FEC9FA1E83BB5306" box="[347,430,1514,1536]" captionStart="Table 1" captionStartId="2.[144,208,200,219]" captionText="Table 1. List of the sampling localities showing altitude (metres) and WGS84 coordinate" pageId="49" pageNumber="486">Table 1</tableCitation>
).
</paragraph>
<paragraph id="6C4436F9FFC44329FF36F9D3831C533A" blockId="49.[164,779,1575,1903]" box="[164,265,1575,1596]" pageId="49" pageNumber="486">
<emphasis id="5E8FEAEBFFC44329FF36F9D3831C533A" box="[164,265,1575,1596]" italics="true" pageId="49" pageNumber="486">Remarks</emphasis>
</paragraph>
<paragraph id="6C4436F9FFC44329FF36F9B18308539F" blockId="49.[164,779,1575,1903]" pageId="49" pageNumber="486">
The perching distance from ground is
<quantity id="AB039B1CFFC44329FDCBF9B28093535D" box="[601,646,1606,1627]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="49" pageNumber="486" unit="m" value="2.0">2 m</quantity>
; also active on low shrubbery. Adults were found in summer (
<figureCitation id="F4C02A7CFFC44329FF3EF9778304539F" box="[172,273,1667,1689]" captionStart="Figure 37" captionStartId="47.[164,243,1853,1872]" captionTargetBox="[389,1219,1311,1823]" captionTargetId="figure-399@47.[387,1219,1309,1823]" captionTargetPageId="47" captionText="Figure 37. Satsuma pilsbryi sp. nov. AD, holotype (TMMT 0614, shell height = 17.9 mm). E, F, paratype (TMMT 0644, shell height = 17.6 mm). G, living specimen. Arrow indicates the pre-apertural constriction." figureDoi="http://doi.org/10.5281/zenodo.5446784" httpUri="https://zenodo.org/record/5446784/files/figure.png" pageId="49" pageNumber="486">Fig. 37G</figureCitation>
).
</paragraph>
<paragraph id="6C4436F9FFC44329FF2EF95586B65472" blockId="49.[164,779,1575,1903]" lastBlockId="49.[826,1441,198,373]" pageId="49" pageNumber="486">
The shell morphology of this species is similar to that of
<taxonomicName id="ABFB4D7AFFC44329FF65F934834153D3" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[247,340,1728,1749]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="49" pageNumber="486" phylum="Mollusca" rank="species" species="vallis">
<emphasis id="5E8FEAEBFFC44329FF65F934834153D3" box="[247,340,1728,1749]" italics="true" pageId="49" pageNumber="486">S. vallis</emphasis>
</taxonomicName>
, but is differs in dimensions and shell colour. The genital morphology of this species is fundamentally similar to that of
<taxonomicName id="ABFB4D7AFFC44329FD83F90980BB5214" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[529,686,1789,1810]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="49" pageNumber="486" phylum="Mollusca" rank="species" species="viridibasis">
<emphasis id="5E8FEAEBFFC44329FD83F90980BB5214" box="[529,686,1789,1810]" italics="true" pageId="49" pageNumber="486">S. viridibasis</emphasis>
</taxonomicName>
and
<taxonomicName id="ABFB4D7AFFC44329FD61F90982F75237" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="49" pageNumber="486" phylum="Mollusca" rank="species" species="vallis">
<emphasis id="5E8FEAEBFFC44329FD61F90982F75237" italics="true" pageId="49" pageNumber="486">S. vallis</emphasis>
</taxonomicName>
with the following exceptions: (1) the proximal vagina is stronger, (2) the base of the bursa stalk is conical and (3) the penial caecum and proximal penis are strong, smooth and muscular.
<taxonomicName id="ABFB4D7AFFC44329FB50FF32879355DD" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[1218,1414,198,219]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="49" pageNumber="486" phylum="Mollusca" rank="species" species="pilsbryi">
<emphasis id="5E8FEAEBFFC44329FB50FF32879355DD" box="[1218,1414,198,219]" italics="true" pageId="49" pageNumber="486">Satsuma pilsbryi</emphasis>
</taxonomicName>
is probably an intraspecific variation from southern populations of
<taxonomicName id="ABFB4D7AFFC44329FC62FEF78646541E" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[1008,1107,259,280]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="49" pageNumber="486" phylum="Mollusca" rank="species" species="vallis">
<emphasis id="5E8FEAEBFFC44329FC62FEF78646541E" box="[1008,1107,259,280]" italics="true" pageId="49" pageNumber="486">S. vallis</emphasis>
</taxonomicName>
and
<taxonomicName id="ABFB4D7AFFC44329FB08FEF7872D541E" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[1178,1336,259,280]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="49" pageNumber="486" phylum="Mollusca" rank="species" species="viridibasis">
<emphasis id="5E8FEAEBFFC44329FB08FEF7872D541E" box="[1178,1336,259,280]" italics="true" pageId="49" pageNumber="486">S. viridibasis</emphasis>
</taxonomicName>
in shell morphology. However, the well-defined geographical isolation and distinguishable reproductive system reveal it merits specific status.
</paragraph>
</subSubSection>
</treatment>
</document>