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<document id="5F34610F4484C0D936D771C6A79C6FFD" ID-CLB-Dataset="8862" ID-DOI="10.11646/zootaxa.4908.1.4" ID-GBIF-Dataset="25f021d5-c95b-441a-b014-cae55dace166" ID-ISSN="1175-5326" ID-Zenodo-Dep="4435769" ID-ZooBank="B9622350-0A3E-4C86-A1E3-6316690004A1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610524185345" checkinUser="plazi" docAuthor="Stauffer, Jay R., Bills, Roger &amp; Skelton, Paul H." docDate="2021" docId="E449C714AA40FFAB3EC6FF1CFD15F172" docLanguage="en" docName="zootaxa.4908.1.4.pdf" docOrigin="Zootaxa 4908 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Serranochromis alvum Stauffer &amp; Bills &amp; Skelton 2021, new species" docType="treatment" docUuid="15F2840B-9A1A-4264-89F2-C8253C3D06CB" docUuidSource="ZooBank" docVersion="8" lastPageNumber="72" masterDocId="1870BF6CAA43FFAD3E51FF8BFFA3F06F" masterDocTitle="Four new species of Serranochromis (Teleostei: Cichlidae) from the Cuanza and Okavango river systems in Angola, including a preliminary key for the genus" masterLastPageNumber="84" masterPageNumber="66" pageNumber="69" updateTime="1698924308383" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="634D81412A6506B65D762F3960AA7DA3">Four new species of Serranochromis (Teleostei: Cichlidae) from the Cuanza and Okavango river systems in Angola, including a preliminary key for the genus</mods:title>
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</mods:role>
<mods:namePart id="6C8CC0D36EE24DEDC2A268EF05E4BB98">Stauffer, Jay R.</mods:namePart>
<mods:affiliation id="5BE008D19DD78723598C120CECB3A9EE">Penn State University, 432 Forest Resources Building, University Park, Pennsylvania 16802, USA., Honorary Research Associate, South African Institute for Aquatic Biodiversity, Makhanda, RSA. &amp; South African Institute for Aquatic Biodiversity, Private Bag 1015, Makhanda, 6140, RSA</mods:affiliation>
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<mods:namePart id="88F40727A423C8B202619F85F4D71206">Bills, Roger</mods:namePart>
<mods:nameIdentifier id="E21CE41D78976129C465F7BDD5E631D7" type="ORCID">0000-0001-6034-4196</mods:nameIdentifier>
<mods:affiliation id="482F85DA09D05A9CFA3B51764066EF43">South African Institute for Aquatic Biodiversity, Private Bag 1015, Makhanda, 6140, RSA &amp; https: // orcid. org / 0000 - 0001 - 6034 - 4196</mods:affiliation>
</mods:name>
<mods:name id="22C83DDF486DF80A20CFD870D4F0A0A7" type="personal">
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</mods:role>
<mods:namePart id="FBF1B176C85BD34D56E72D978E22F085">Skelton, Paul H.</mods:namePart>
<mods:nameIdentifier id="3E501215AAE0BC84769A79E2D64A83F3" type="ORCID">0000-0001-9587-2802</mods:nameIdentifier>
<mods:affiliation id="D0D7A6177838780D251B3172F48B26CB">South African Institute for Aquatic Biodiversity, Private Bag 1015, Makhanda, 6140, RSA &amp; National Geographic Okavango Wilderness Project, Wild Bird Trust. &amp; https: // orcid. org / 0000 - 0001 - 9587 - 2802</mods:affiliation>
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<mods:title id="802C0B7625CF3C5E709B41808921F281">Zootaxa</mods:title>
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<mods:part id="C2B5BC35122660AB5C67B10E68D9D9D3">
<mods:date id="A6347725FEBBDCBF55671012C9872346">2021</mods:date>
<mods:detail id="5D33B702A7C330D25CA77F5FB01B60ED" type="pubDate">
<mods:number id="6EED78EA7FE1D89B037EB09D72BE5E69">2021-01-13</mods:number>
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<mods:number id="6E5118D0D3C2D252820E91B70CED9CF9">4908</mods:number>
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<mods:identifier id="15DBD93FB502A28DE49399A6428C1C7D" type="CLB-Dataset">8862</mods:identifier>
<mods:identifier id="7637B5E4FB4FA30FC7BD658F52DB7FD3" type="DOI">10.11646/zootaxa.4908.1.4</mods:identifier>
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<treatment id="E449C714AA40FFAB3EC6FF1CFD15F172" ID-DOI="http://doi.org/10.5281/zenodo.4442684" ID-GBIF-Taxon="176596412" ID-Zenodo-Dep="4442684" LSID="urn:lsid:zoobank.org:act:15F2840B-9A1A-4264-89F2-C8253C3D06CB" httpUri="http://treatment.plazi.org/id/E449C714AA40FFAB3EC6FF1CFD15F172" lastPageId="6" lastPageNumber="72" pageId="3" pageNumber="69">
<subSubSection id="24FA2589AA40FFAE3EC6FF1CFC98F0B9" pageId="3" pageNumber="69" type="nomenclature">
<paragraph id="6C5F7602AA40FFAE3EC6FF1CFDE7F0DE" blockId="3.[151,827,151,249]" box="[151,580,151,178]" pageId="3" pageNumber="69">
<heading id="3717C16EAA40FFAE3EC6FF1CFDE7F0DE" bold="true" box="[151,580,151,178]" fontSize="11" level="1" pageId="3" pageNumber="69" reason="1">
<emphasis id="5E94AA10AA40FFAE3EC6FF1CFDE7F0DE" bold="true" box="[151,580,151,178]" pageId="3" pageNumber="69">
<taxonomicName id="ABE00D81AA40FFAE3EC6FF1CFE0EF0DD" authority="Stauffer &amp; Bills &amp; Skelton, 2021" authorityName="Stauffer &amp; Bills &amp; Skelton" authorityYear="2021" box="[151,429,151,178]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="69" phylum="Chordata" rank="species" species="alvum" status="sp. nov.">
<emphasis id="5E94AA10AA40FFAE3EC6FF1CFE0EF0DD" bold="true" box="[151,429,151,178]" italics="true" pageId="3" pageNumber="69">Serranochromis alvum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45A7176BAA40FFAE3FE5FF1CFDE7F0DE" box="[436,580,151,177]" pageId="3" pageNumber="69" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6C5F7602AA40FFAE3EC6FF30FC98F0B9" blockId="3.[151,827,151,249]" box="[151,827,187,214]" pageId="3" pageNumber="69">Suggested common name: Blue-Headed Largemouth Bream</paragraph>
</subSubSection>
<subSubSection id="24FA2589AA40FFAE3EC6FF54FF51F096" box="[151,242,223,249]" pageId="3" pageNumber="69" type="description">
<paragraph id="6C5F7602AA40FFAE3EC6FF54FF51F096" blockId="3.[151,827,151,249]" box="[151,242,223,249]" pageId="3" pageNumber="69">
(
<figureCitation id="F4DB6A87AA40FFAE3EF1FF54FF46F096" box="[160,229,223,249]" captionStart="FIGURE 2" captionStartId="3.[151,250,1976,2000]" captionTargetBox="[204,1383,1317,1952]" captionTargetId="figure-509@3.[203,1383,1317,1952]" captionTargetPageId="3" captionText="FIGURE 2. Serranochromis alvum, Holotype SAIAB 186842, 151.8 mm SL, collected from Cuito Canavale bridge, Angola; 15º10’16.4”S 19º11’39.3”E, 18 May 2012." figureDoi="http://doi.org/10.5281/zenodo.4435773" httpUri="https://zenodo.org/record/4435773/files/figure.png" pageId="3" pageNumber="69">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="24FA2589AA40FFAE3EC6FEACFC5FF1C2" pageId="3" pageNumber="69" type="materials_examined">
<paragraph id="6C5F7602AA40FFAE3EC6FEACFDFAF1E5" blockId="3.[151,1437,295,1294]" pageId="3" pageNumber="69">
<materialsCitation id="DC887C5FAA40FFAE3EC6FEACFDFAF1E5" ID-GBIF-Occurrence="3017348302" collectingDate="2012-05-18" collectionCode="SAIAB, R" collectorName="Bills, P. H &amp; Skelton, F." country="Angola" county="Okavango River" latitude="-15.171223" location="Cuito-Cuanavale" longLatPrecision="1" longitude="19.194248" municipality="Cuito River" pageId="3" pageNumber="69" specimenCount="1" typeStatus="holotype">
<emphasis id="5E94AA10AA40FFAE3EC6FEACFEA8F12E" bold="true" box="[151,267,295,321]" pageId="3" pageNumber="69">
<typeStatus id="B35BC8A0AA40FFAE3EC6FEACFEA5F12E" box="[151,262,295,321]" pageId="3" pageNumber="69" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="0AF1EEC7AA40FFAE3F42FEACFEC4F12E" box="[275,359,295,321]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="3" pageNumber="69" type="Other type of research institution/biorepository">SAIAB</collectionCode>
<quantity id="AB18DBE7AA40FFAE3F21FEACFDE5F12D" box="[368,582,295,322]" metricMagnitude="5" metricUnit="m" metricValue="1.868421518" pageId="3" pageNumber="69" unit="mm" value="1.868421518E8">186842, 151.8 mm</quantity>
SL (male), collected from
<location id="693F20D9AA40FFAE3D2BFEACFB9BF12E" LSID="urn:lsid:plazi:treatment:E449C714AA40FFAB3EC6FF1CFD15F172:693F20D9AA40FFAE3D2BFEACFB9BF12E" box="[890,1080,295,321]" country="Angola" county="Okavango River" latitude="-15.171223" longLatPrecision="1" longitude="19.194248" municipality="Cuito River" name="Cuito-Cuanavale" pageId="3" pageNumber="69">Cuito-Cuanavale</location>
bridge,
<collectingMunicipality id="8C3BEC78AA40FFAE3AC7FEACFABEF12E" box="[1174,1309,295,321]" pageId="3" pageNumber="69">Cuito River</collectingMunicipality>
,
<collectingCounty id="853E0E8EAA40FFAE3B78FEACFF76F10A" pageId="3" pageNumber="69">Okavango River</collectingCounty>
system,
<collectingCountry id="14F73692AA40FFAE3F68FEC0FE2CF10A" box="[313,399,331,357]" name="Angola" pageId="3" pageNumber="69">Angola</collectingCountry>
;
<geoCoordinate id="09D410C5AA40FFAE3FCAFEC0FD98F10A" box="[411,571,331,358]" degrees="15" direction="south" minutes="10" orientation="latitude" pageId="3" pageNumber="69" precision="1" seconds="16.4" value="-15.171223">15º10’16.4” S</geoCoordinate>
,
<geoCoordinate id="09D410C5AA40FFAE3C18FEC0FD48F109" box="[585,747,331,358]" degrees="19" direction="east" minutes="11" orientation="longitude" pageId="3" pageNumber="69" precision="1" seconds="39.3" value="19.194248">19º11’39.3” E</geoCoordinate>
. Collected by
<collectionCode id="0AF1EEC7AA40FFAE3DDEFEC7FC06F109" box="[911,933,332,358]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="69">R</collectionCode>
.
<collectorName id="C11513D4AA40FFAE3DE0FEC0FB84F109" box="[945,1063,331,358]" pageId="3" pageNumber="69">Bills, P. H</collectorName>
.
<collectorName id="C11513D4AA40FFAE3A62FEC0FB0EF109" box="[1075,1197,331,358]" pageId="3" pageNumber="69">Skelton, F.</collectorName>
de Almedia,
<date id="185E50C2AA40FFAE3B14FEC0FF70F1E6" pageId="3" pageNumber="69" value="2012-05-18">
<collectingDate id="081AA92AAA40FFAE3B14FEC0FF70F1E6" pageId="3" pageNumber="69" value="2012-05-18">18 May 2012</collectingDate>
</date>
. Collected with gill and seine nets.
</materialsCitation>
</paragraph>
<paragraph id="6C5F7602AA40FFAE3E96FE18FC5FF1C2" blockId="3.[151,1437,295,1294]" box="[199,1020,403,430]" pageId="3" pageNumber="69">
<materialsCitation id="DC887C5FAA40FFAE3E96FE18FC5BF1C2" ID-GBIF-Occurrence="3017348308" box="[199,1016,403,430]" collectingDate="2012-05-18" collectionCode="SAIAB" collectorName="Bills, P. H &amp; Skelton, F." country="Angola" county="Okavango River" latitude="-15.171223" location="Cuito-Cuanavale" longLatPrecision="1" longitude="19.194248" municipality="Cuito River" pageId="3" pageNumber="69" specimenCode="SAIAB 205219" specimenCount="1" typeStatus="paratype">
<emphasis id="5E94AA10AA40FFAE3E96FE18FEE5F1C2" bold="true" box="[199,326,403,429]" pageId="3" pageNumber="69">
<typeStatus id="B35BC8A0AA40FFAE3E96FE18FEE2F1C2" box="[199,321,403,429]" pageId="3" pageNumber="69" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCode id="3C46DE79AA40FFAE3F1CFE18FE5FF1C1" box="[333,508,403,430]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="3" pageNumber="69" type="Other type of research institution/biorepository">SAIAB 205219</specimenCode>
(5),
<quantity id="AB18DBE7AA40FFAE3C63FE18FD41F1C1" box="[562,738,403,430]" metricMagnitude="-2" metricUnit="m" metricValue="9.3" metricValueMax="11.45" metricValueMin="7.15" pageId="3" pageNumber="69" unit="mm" value="93.0" valueMax="114.5" valueMin="71.5">71.5–114.5 mm</quantity>
SL, data as for holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="24FA2589AA40FFAE3E96FE3CFC7FF356" pageId="3" pageNumber="69" type="diagnosis">
<paragraph id="6C5F7602AA40FFAE3E96FE3CFC7FF356" blockId="3.[151,1437,295,1294]" pageId="3" pageNumber="69">
<emphasis id="5E94AA10AA40FFAE3E96FE3CFEE1F1BE" bold="true" box="[199,322,439,465]" pageId="3" pageNumber="69">Diagnosis.</emphasis>
The presence of ocelli throughout the anal fin of breeding males distinguishes
<taxonomicName id="ABE00D81AA40FFAE3A8BFE3CFA9EF1BE" authorityName="Stauffer &amp; Bills &amp; Skelton" authorityYear="2021" box="[1242,1341,439,465]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="69" phylum="Chordata" rank="species" species="alvum">
<emphasis id="5E94AA10AA40FFAE3A8BFE3CFA9EF1BE" box="[1242,1341,439,465]" italics="true" pageId="3" pageNumber="69">S. alvum</emphasis>
</taxonomicName>
from
<taxonomicName id="ABE00D81AA40FFAE3BD6FE3CFF55F19A" baseAuthorityName="Gunther" baseAuthorityYear="1864" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="69" phylum="Chordata" rank="species" species="robustus">
<emphasis id="5E94AA10AA40FFAE3BD6FE3CFF55F19A" italics="true" pageId="3" pageNumber="69">S. robustus</emphasis>
</taxonomicName>
and
<taxonomicName id="ABE00D81AA40FFAE3F7DFE50FD17F199" authority="(Boulenger)" baseAuthorityName="Boulenger" baseAuthorityYear="1896" box="[300,692,475,502]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="69" phylum="Chordata" rank="species" species="jallae">
<emphasis id="5E94AA10AA40FFAE3F7DFE50FD85F19A" box="[300,550,475,501]" italics="true" pageId="3" pageNumber="69">Serranochromis jallae</emphasis>
(Boulenger)
</taxonomicName>
in which the ocelli in breeding males are restricted to the posterior 4–5 membranes of the anal fin. The exposed teeth of
<taxonomicName id="ABE00D81AA40FFAE3CBBFE74FCE9F276" authorityName="Stauffer &amp; Bills &amp; Skelton" authorityYear="2021" box="[746,842,511,537]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="69" phylum="Chordata" rank="species" species="alvum">
<emphasis id="5E94AA10AA40FFAE3CBBFE74FCE9F276" box="[746,842,511,537]" italics="true" pageId="3" pageNumber="69">S. alvum</emphasis>
</taxonomicName>
differs from those of
<taxonomicName id="ABE00D81AA40FFAE3A6EFE74FB61F276" box="[1087,1218,511,537]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="69" phylum="Chordata" rank="species" species="stappersi">
<emphasis id="5E94AA10AA40FFAE3A6EFE74FB61F276" box="[1087,1218,511,537]" italics="true" pageId="3" pageNumber="69">S. stappersi</emphasis>
</taxonomicName>
and
<taxonomicName id="ABE00D81AA40FFAE3AA6FE74FAEAF276" box="[1271,1353,511,537]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="69" phylum="Chordata" rank="species" species="altus">
<emphasis id="5E94AA10AA40FFAE3AA6FE74FAEAF276" box="[1271,1353,511,537]" italics="true" pageId="3" pageNumber="69">S. altus</emphasis>
</taxonomicName>
, which possess small teeth that are buried in the lips.
<taxonomicName id="ABE00D81AA40FFAE3CC9FDA8FC35F252" authorityName="Stauffer &amp; Bills &amp; Skelton" authorityYear="2021" box="[664,918,547,573]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="69" phylum="Chordata" rank="species" species="alvum">
<emphasis id="5E94AA10AA40FFAE3CC9FDA8FC35F252" box="[664,918,547,573]" italics="true" pageId="3" pageNumber="69">Serranochromis alvum</emphasis>
</taxonomicName>
has an emarginate caudal fin, while
<taxonomicName id="ABE00D81AA40FFAE3B61FDA8FF58F20E" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="69" phylum="Chordata" rank="species" species="macrocephalus">
<emphasis id="5E94AA10AA40FFAE3B61FDA8FF58F20E" italics="true" pageId="3" pageNumber="69">S. macrocephalus</emphasis>
</taxonomicName>
,
<taxonomicName id="ABE00D81AA40FFAE3F59FDCCFE4BF20E" authority="(Trewavas)" baseAuthorityName="Trewavas" box="[264,488,583,609]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="69" phylum="Chordata" rank="species" species="janus">
<emphasis id="5E94AA10AA40FFAE3F59FDCCFEC3F20E" box="[264,352,583,609]" italics="true" pageId="3" pageNumber="69">S. janus</emphasis>
(Trewavas)
</taxonomicName>
, and
<taxonomicName id="ABE00D81AA40FFAE3C70FDCCFD63F20E" baseAuthorityName="Boulenger" baseAuthorityYear="1907" box="[545,704,583,609]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="69" phylum="Chordata" rank="species" species="angusticeps">
<emphasis id="5E94AA10AA40FFAE3C70FDCCFD63F20E" box="[545,704,583,609]" italics="true" pageId="3" pageNumber="69">S. angusticeps</emphasis>
</taxonomicName>
have rounded caudal fins. The shorter jaw of
<taxonomicName id="ABE00D81AA40FFAE3AECFDCCFABFF20E" authorityName="Stauffer &amp; Bills &amp; Skelton" authorityYear="2021" box="[1213,1308,583,609]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="69" phylum="Chordata" rank="species" species="alvum">
<emphasis id="5E94AA10AA40FFAE3AECFDCCFABFF20E" box="[1213,1308,583,609]" italics="true" pageId="3" pageNumber="69">S. alvum</emphasis>
</taxonomicName>
(43.4–53.5 % HL) separates it from
<taxonomicName id="ABE00D81AA40FFAE3FEAFDE0FC8DF2EA" authority="(Trewavas)" baseAuthorityName="Trewavas" box="[443,814,619,645]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="69" phylum="Chordata" rank="species" species="spei">
<emphasis id="5E94AA10AA40FFAE3FEAFDE0FD06F2EA" box="[443,677,619,645]" italics="true" pageId="3" pageNumber="69">Serranochromis spei</emphasis>
(Trewavas)
</taxonomicName>
(53.5–57.2 % HL).
<taxonomicName id="ABE00D81AA40FFAE3A4AFDE0FABEF2EA" authorityName="Stauffer &amp; Bills &amp; Skelton" authorityYear="2021" box="[1051,1309,619,645]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="69" phylum="Chordata" rank="species" species="alvum">
<emphasis id="5E94AA10AA40FFAE3A4AFDE0FABEF2EA" box="[1051,1309,619,645]" italics="true" pageId="3" pageNumber="69">Serranochromis alvum</emphasis>
</taxonomicName>
has 34–36 lateral-line scales, while
<taxonomicName id="ABE00D81AA40FFAE3FE3FD04FDE2F2C6" baseAuthorityName="Castelnau" baseAuthorityYear="1861" box="[434,577,655,681]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="69" phylum="Chordata" rank="species" species="thumbergi">
<emphasis id="5E94AA10AA40FFAE3FE3FD04FDE2F2C6" box="[434,577,655,681]" italics="true" pageId="3" pageNumber="69">S. thumbergi</emphasis>
</taxonomicName>
possesses greater than 39 lateral-line scales. The acute angle of the cleft of the mouth (50–60º of horizontal) of
<taxonomicName id="ABE00D81AA40FFAE3C7BFD38FD60F2A2" baseAuthorityName="Jubb" baseAuthorityYear="1967" box="[554,707,691,717]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="69" phylum="Chordata" rank="species" species="meridianus">
<emphasis id="5E94AA10AA40FFAE3C7BFD38FD60F2A2" box="[554,707,691,717]" italics="true" pageId="3" pageNumber="69">S. meridianus</emphasis>
</taxonomicName>
delimits it from the 30º angle of
<taxonomicName id="ABE00D81AA40FFAE3A63FD38FB31F2A2" authorityName="Stauffer &amp; Bills &amp; Skelton" authorityYear="2021" box="[1074,1170,691,717]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="69" phylum="Chordata" rank="species" species="alvum">
<emphasis id="5E94AA10AA40FFAE3A63FD38FB31F2A2" box="[1074,1170,691,717]" italics="true" pageId="3" pageNumber="69">S. alvum</emphasis>
</taxonomicName>
. The long dark pectoral fins which reach past the middle of the dorsal fin separates
<taxonomicName id="ABE00D81AA40FFAE3D16FD5CFC4BF29E" authorityName="Trewavas" authorityYear="1964" baseAuthorityName="Boulenger" baseAuthorityYear="1911" box="[839,1000,727,753]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="69" phylum="Chordata" rank="species" species="longimanus">
<emphasis id="5E94AA10AA40FFAE3D16FD5CFC4BF29E" box="[839,1000,727,753]" italics="true" pageId="3" pageNumber="69">S. longimanus</emphasis>
</taxonomicName>
from
<emphasis id="5E94AA10AA40FFAE3A61FD5CFB39F29E" box="[1072,1178,727,753]" italics="true" pageId="3" pageNumber="69">
<taxonomicName id="ABE00D81AA40FFAE3A61FD5CFB35F29E" authorityName="Stauffer &amp; Bills &amp; Skelton" authorityYear="2021" box="[1072,1174,727,753]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="69" phylum="Chordata" rank="species" species="alvum">S. alvum</taxonomicName>
.
</emphasis>
The interorbital width (14.3–15.9 % HL) of
<taxonomicName id="ABE00D81AA40FFAE3FDBFD70FE48F37A" authorityName="Stauffer &amp; Bills &amp; Skelton" authorityYear="2021" box="[394,491,763,789]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="69" phylum="Chordata" rank="species" species="alvum">
<emphasis id="5E94AA10AA40FFAE3FDBFD70FE48F37A" box="[394,491,763,789]" italics="true" pageId="3" pageNumber="69">S. alvum</emphasis>
</taxonomicName>
is narrower than that of
<taxonomicName id="ABE00D81AA40FFAE3CAEFD70FBA8F37A" authority="Stauffer &amp; Bills &amp; Skelton, 2021" authorityName="Stauffer &amp; Bills &amp; Skelton" authorityYear="2021" box="[767,1035,763,789]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="69" phylum="Chordata" rank="species" species="swartzi" status="sp. nov.">
<emphasis id="5E94AA10AA40FFAE3CAEFD70FBA8F37A" box="[767,1035,763,789]" italics="true" pageId="3" pageNumber="69">Serranochromis swartzi</emphasis>
</taxonomicName>
<emphasis id="5E94AA10AA40FFAE3A43FD70FBF2F37A" bold="true" box="[1042,1105,763,789]" pageId="3" pageNumber="69">
<taxonomicNameLabel id="45A7176BAA40FFAE3A43FD70FBF2F37A" box="[1042,1105,763,789]" pageId="3" pageNumber="69" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(17.6–19.8),
<taxonomicName id="ABE00D81AA40FFAE3AB8FD70FF45F356" authority="Stauffer &amp; Bills &amp; Skelton, 2021" authorityName="Stauffer &amp; Bills &amp; Skelton" authorityYear="2021" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="69" phylum="Chordata" rank="species" species="cuanza" status="sp. nov.">
<emphasis id="5E94AA10AA40FFAE3AB8FD70FF45F356" italics="true" pageId="3" pageNumber="69">Serranochromis cuanza</emphasis>
</taxonomicName>
<emphasis id="5E94AA10AA40FFAE3EBFFC94FE8FF356" bold="true" box="[238,300,799,825]" pageId="3" pageNumber="69">
<taxonomicNameLabel id="45A7176BAA40FFAE3EBFFC94FE8FF356" box="[238,300,799,825]" pageId="3" pageNumber="69" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(16.3–18.0), and
<taxonomicName id="ABE00D81AA40FFAE3FA5FC94FCA6F356" authority="Stauffer &amp; Bills &amp; Skelton, 2021" authorityName="Stauffer &amp; Bills &amp; Skelton" authorityYear="2021" box="[500,773,799,825]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="69" phylum="Chordata" rank="species" species="cacuchi" status="sp. nov.">
<emphasis id="5E94AA10AA40FFAE3FA5FC94FCA6F356" box="[500,773,799,825]" italics="true" pageId="3" pageNumber="69">Serranochromis cacuchi</emphasis>
</taxonomicName>
<emphasis id="5E94AA10AA40FFAE3D5DFC94FCE8F356" bold="true" box="[780,843,799,825]" pageId="3" pageNumber="69">
<taxonomicNameLabel id="45A7176BAA40FFAE3D5DFC94FCE8F356" box="[780,843,799,825]" pageId="3" pageNumber="69" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(20.0–21.7).
</paragraph>
</subSubSection>
<subSubSection id="24FA2589AA40FFA83E96FCC8FAB3F45A" lastPageId="5" lastPageNumber="71" pageId="3" pageNumber="69" type="description">
<paragraph id="6C5F7602AA40FFAE3E96FCC8FAC8F332" blockId="3.[151,1437,295,1294]" box="[199,1387,835,862]" pageId="3" pageNumber="69">
<emphasis id="5E94AA10AA40FFAE3E96FCC8FEFAF332" bold="true" box="[199,345,835,861]" pageId="3" pageNumber="69">Description.</emphasis>
Morphometric ratios and meristic data in
<tableCitation id="216243B9AA40FFAE3D61FCC8FC27F332" box="[816,900,835,861]" captionStart="TABLE 1" captionStartId="4.[151,239,151,175]" captionText="TABLE 1. Morphometric and meristic values of Serranochromis alvum (n=6). The mean, standard deviation and range include holotype and paratypes." pageId="3" pageNumber="69">Table 1</tableCitation>
. Body shape and pigment patterns in
<figureCitation id="F4DB6A87AA40FFAE3B7BFCC8FAC4F332" box="[1322,1383,835,861]" captionStart="FIGURE 2" captionStartId="3.[151,250,1976,2000]" captionTargetBox="[204,1383,1317,1952]" captionTargetId="figure-509@3.[203,1383,1317,1952]" captionTargetPageId="3" captionText="FIGURE 2. Serranochromis alvum, Holotype SAIAB 186842, 151.8 mm SL, collected from Cuito Canavale bridge, Angola; 15º10’16.4”S 19º11’39.3”E, 18 May 2012." figureDoi="http://doi.org/10.5281/zenodo.4435773" httpUri="https://zenodo.org/record/4435773/files/figure.png" pageId="3" pageNumber="69">Fig 2</figureCitation>
.
</paragraph>
<paragraph id="6C5F7602AA40FFAE3E96FCECFB5DF562" blockId="3.[151,1437,295,1294]" pageId="3" pageNumber="69">
Body slender, fusiform, and compressed, deepest at origin of dorsal fin. Dorsal-fin origin at vertical through posterior edge of operculum, dorsal-fin spines to maximum length over 5–6 spines, spine lappets prominent. Soft dorsal rounded behind, extending to base of caudal fin. Caudal peduncle short (13–18 % SL) 7.5 times in SL, slightly longer than deep (CPL/CPD 1.1–1.2). Caudal fin broad, relatively short (little more than half the head length), rectangular and truncate. Anal-fin origin closer to caudal-fin base than tip of snout, below vertical through first dorsal soft-ray; soft-rayed section with bluntly rounded hind edge, extending to caudal-fin base. Pectoral fin close behind gill slit, ventral on flanks, base sub-vertical, short, not extending beyond pelvic fins, postero-ventral margin bluntly rounded. Pelvic fins ventral, origin narrowly behind base of pectoral fin, short, not reaching origin to the anal fin, strong leading spine 2/3 length of fin. Scales small, in 15 even rows across flanks from dorsal-fin origin to anal-fin origin, 16 rows of scales around caudal peduncle,
<quantity id="AB18DBE7AA40FFAE3D17FB20FC0AF4AA" box="[838,937,1195,1221]" metricMagnitude="-1" metricUnit="m" metricValue="8.89" metricValueMax="9.144" metricValueMin="8.636" pageId="3" pageNumber="69" unit="in" value="35.0" valueMax="36.0" valueMin="34.0">34–36 in</quantity>
lateral line; upper lateral line gently curved, lower lateral line straight through mid-caudal peduncle; five scale rows between anterior dorsal and lateral line, two scale rows between soft dorsal and posterior end of upper lateral line, nape and chest scales reduced.
</paragraph>
<caption id="389F268AAA40FFAE3EC6F833FDE9F79B" ID-DOI="http://doi.org/10.5281/zenodo.4435773" ID-Zenodo-Dep="4435773" httpUri="https://zenodo.org/record/4435773/files/figure.png" pageId="3" pageNumber="69" startId="3.[151,250,1976,2000]" targetBox="[204,1383,1317,1952]" targetPageId="3">
<paragraph id="6C5F7602AA40FFAE3EC6F833FDE9F79B" blockId="3.[151,1436,1976,2036]" pageId="3" pageNumber="69">
<emphasis id="5E94AA10AA40FFAE3EC6F833FEB7F7BF" bold="true" box="[151,276,1976,2000]" pageId="3" pageNumber="69">FIGURE 2.</emphasis>
<taxonomicName id="ABE00D81AA40FFAE3F4AF833FDA0F7BF" authorityName="Stauffer &amp; Bills &amp; Skelton" authorityYear="2021" box="[283,515,1976,2000]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="69" phylum="Chordata" rank="species" species="alvum">
<emphasis id="5E94AA10AA40FFAE3F4AF833FDA0F7BF" box="[283,515,1976,2000]" italics="true" pageId="3" pageNumber="69">Serranochromis alvum</emphasis>
</taxonomicName>
, Holotype SAIAB 186842, 151.8 mm SL, collected from Cuito Canavale bridge, Angola; 15º10’16.4”S 19º11’39.3”E, 18 May 2012.
</paragraph>
</caption>
<caption id="389F268AAA47FFA93EC6FF1CFE48F0BB" pageId="4" pageNumber="70">
<paragraph id="6C5F7602AA47FFA93EC6FF1CFE48F0BB" blockId="4.[151,1436,151,212]" pageId="4" pageNumber="70">
<emphasis id="5E94AA10AA47FFA93EC6FF1CFEA8F0C0" bold="true" box="[151,267,151,175]" pageId="4" pageNumber="70">TABLE 1.</emphasis>
Morphometric and meristic values of
<taxonomicName id="ABE00D81AA47FFA93CF8FF1CFC3EF0C0" authorityName="Stauffer &amp; Bills &amp; Skelton" authorityYear="2021" box="[681,925,151,175]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="alvum">
<emphasis id="5E94AA10AA47FFA93CF8FF1CFC3EF0C0" box="[681,925,151,175]" italics="true" pageId="4" pageNumber="70">Serranochromis alvum</emphasis>
</taxonomicName>
(n=6). The mean, standard deviation and range include holotype and paratypes.
</paragraph>
</caption>
<paragraph id="6C5F7602AA47FFA93EF7FF68FB07F7D1" pageId="4" pageNumber="70">
<table id="1EE084A2AA4700523EF7FF68FACBF7D1" box="[166,1384,227,1982]" colsContinueIn="12.[166,1381,227,1982]" gridcols="5" gridrows="44" pageId="4" pageNumber="70">
<tr id="D2D07440AA4700523EF7FF68FACBF094" box="[166,1384,227,251]" gridrow="0" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FF68FDAFF094" box="[166,524,227,251]" gridcol="0" gridrow="0" pageId="4" pageNumber="70">
<emphasis id="5E94AA10AA47FFA93EF7FF68FEA1F094" bold="true" box="[166,258,227,251]" pageId="4" pageNumber="70">Variable</emphasis>
</th>
<th id="91011D3CAA4700523D60FF68FC37F094" box="[817,916,227,251]" gridcol="1" gridrow="0" pageId="4" pageNumber="70">
<emphasis id="5E94AA10AA47FFA93D60FF68FC37F094" bold="true" box="[817,916,227,251]" pageId="4" pageNumber="70">Holotype</emphasis>
</th>
<th id="91011D3CAA4700523DBCFF68FB90F094" box="[1005,1075,227,251]" gridcol="2" gridrow="0" pageId="4" pageNumber="70">
<emphasis id="5E94AA10AA47FFA93DA0FF68FB8CF094" bold="true" box="[1009,1071,227,251]" pageId="4" pageNumber="70">Mean</emphasis>
</th>
<th id="91011D3CAA4700523A2AFF68FB04F094" box="[1147,1191,227,251]" gridcol="3" gridrow="0" pageId="4" pageNumber="70">
<emphasis id="5E94AA10AA47FFA93AD0FF68FB02F094" bold="true" box="[1153,1185,227,251]" pageId="4" pageNumber="70">SD</emphasis>
</th>
<th id="91011D3CAA4700523AAAFF68FACBF094" box="[1275,1384,227,251]" gridcol="4" gridrow="0" pageId="4" pageNumber="70">
<emphasis id="5E94AA10AA47FFA93B5EFF68FAF7F094" bold="true" box="[1295,1364,227,251]" pageId="4" pageNumber="70">Range</emphasis>
</th>
</tr>
<tr id="D2D07440AA4700523EF7FE80FACBF14B" box="[166,1384,267,292]" gridrow="1" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FE80FDAFF14B" box="[166,524,267,292]" gridcol="0" gridrow="1" pageId="4" pageNumber="70">Standard length, mm</th>
<td id="91011D3CAA4700523D60FE80FC37F14B" box="[817,916,267,292]" gridcol="1" gridrow="1" pageId="4" pageNumber="70">151.8</td>
<td id="91011D3CAA4700523DBCFE80FB90F14B" box="[1005,1075,267,292]" gridcol="2" gridrow="1" pageId="4" pageNumber="70">103.27</td>
<td id="91011D3CAA4700523A2AFE80FB04F14B" box="[1147,1191,267,292]" gridcol="3" gridrow="1" pageId="4" pageNumber="70">29.0</td>
<td id="91011D3CAA4700523AAAFE80FACBF14B" box="[1275,1384,267,292]" gridcol="4" gridrow="1" pageId="4" pageNumber="70">71.5-151.8</td>
</tr>
<tr id="D2D07440AA4700523EF7FEBFFACBF123" box="[166,1384,308,332]" gridrow="2" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FEBFFDAFF123" box="[166,524,308,332]" gridcol="0" gridrow="2" pageId="4" pageNumber="70">Head length, mm</th>
<td id="91011D3CAA4700523D60FEBFFC37F123" box="[817,916,308,332]" gridcol="1" gridrow="2" pageId="4" pageNumber="70">57.9</td>
<td id="91011D3CAA4700523DBCFEBFFB90F123" box="[1005,1075,308,332]" gridcol="2" gridrow="2" pageId="4" pageNumber="70">39.07</td>
<td id="91011D3CAA4700523A2AFEBFFB04F123" box="[1147,1191,308,332]" gridcol="3" gridrow="2" pageId="4" pageNumber="70">11.0</td>
<td id="91011D3CAA4700523AAAFEBFFACBF123" box="[1275,1384,308,332]" gridcol="4" gridrow="2" pageId="4" pageNumber="70">28-57.9</td>
</tr>
<tr id="D2D07440AA4700523EF7FED0FACBF11B" box="[166,1384,347,372]" gridrow="3" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FED0FACBF11B" box="[166,1384,347,372]" colspan="5" colspanRight="4" gridcol="0" gridrow="3" pageId="4" pageNumber="70">
<emphasis id="5E94AA10AA47FFA93EF7FED7FE66F11B" bold="true" box="[166,453,347,372]" pageId="4" pageNumber="70">Percent of standard length</emphasis>
</th>
</tr>
<tr id="D2D07440AA4700523EF7FE0FFACBF1F3" box="[166,1384,388,412]" gridrow="4" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FE0FFDAFF1F3" box="[166,524,388,412]" gridcol="0" gridrow="4" pageId="4" pageNumber="70">Head length</th>
<td id="91011D3CAA4700523D60FE0FFC37F1F3" box="[817,916,388,412]" gridcol="1" gridrow="4" pageId="4" pageNumber="70">38.1</td>
<td id="91011D3CAA4700523DBCFE0FFB90F1F3" box="[1005,1075,388,412]" gridcol="2" gridrow="4" pageId="4" pageNumber="70">37.8</td>
<td id="91011D3CAA4700523A2AFE0FFB04F1F3" box="[1147,1191,388,412]" gridcol="3" gridrow="4" pageId="4" pageNumber="70">0.68</td>
<td id="91011D3CAA4700523AAAFE0FFACBF1F3" box="[1275,1384,388,412]" gridcol="4" gridrow="4" pageId="4" pageNumber="70">37–39.0</td>
</tr>
<tr id="D2D07440AA4700523EF7FE27FACBF1AA" box="[166,1384,428,453]" gridrow="5" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FE27FDAFF1AA" box="[166,524,428,453]" gridcol="0" gridrow="5" pageId="4" pageNumber="70">Body depth</th>
<td id="91011D3CAA4700523D60FE27FC37F1AA" box="[817,916,428,453]" gridcol="1" gridrow="5" pageId="4" pageNumber="70">33.6</td>
<td id="91011D3CAA4700523DBCFE27FB90F1AA" box="[1005,1075,428,453]" gridcol="2" gridrow="5" pageId="4" pageNumber="70">31.7</td>
<td id="91011D3CAA4700523A2AFE27FB04F1AA" box="[1147,1191,428,453]" gridcol="3" gridrow="5" pageId="4" pageNumber="70">1.39</td>
<td id="91011D3CAA4700523AAAFE27FACBF1AA" box="[1275,1384,428,453]" gridcol="4" gridrow="5" pageId="4" pageNumber="70">30.1–33.6</td>
</tr>
<tr id="D2D07440AA4700523EF7FE5EFACBF182" box="[166,1384,469,493]" gridrow="6" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FE5EFDAFF182" box="[166,524,469,493]" gridcol="0" gridrow="6" pageId="4" pageNumber="70">Snout to dorsal-fin origin</th>
<td id="91011D3CAA4700523D60FE5EFC37F182" box="[817,916,469,493]" gridcol="1" gridrow="6" pageId="4" pageNumber="70">40.2</td>
<td id="91011D3CAA4700523DBCFE5EFB90F182" box="[1005,1075,469,493]" gridcol="2" gridrow="6" pageId="4" pageNumber="70">39.5</td>
<td id="91011D3CAA4700523A2AFE5EFB04F182" box="[1147,1191,469,493]" gridcol="3" gridrow="6" pageId="4" pageNumber="70">0.93</td>
<td id="91011D3CAA4700523AAAFE5EFACBF182" box="[1275,1384,469,493]" gridcol="4" gridrow="6" pageId="4" pageNumber="70">38–40.4</td>
</tr>
<tr id="D2D07440AA4700523EF7FE76FACBF27A" box="[166,1384,509,533]" gridrow="7" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FE76FDAFF27A" box="[166,524,509,533]" gridcol="0" gridrow="7" pageId="4" pageNumber="70">Snout to pelvic-fin origin</th>
<td id="91011D3CAA4700523D60FE76FC37F27A" box="[817,916,509,533]" gridcol="1" gridrow="7" pageId="4" pageNumber="70">43.5</td>
<td id="91011D3CAA4700523DBCFE76FB90F27A" box="[1005,1075,509,533]" gridcol="2" gridrow="7" pageId="4" pageNumber="70">42.4</td>
<td id="91011D3CAA4700523A2AFE76FB04F27A" box="[1147,1191,509,533]" gridcol="3" gridrow="7" pageId="4" pageNumber="70">1.30</td>
<td id="91011D3CAA4700523AAAFE76FACBF27A" box="[1275,1384,509,533]" gridcol="4" gridrow="7" pageId="4" pageNumber="70">40.2–43.6</td>
</tr>
<tr id="D2D07440AA4700523EF7FDAEFACBF252" box="[166,1384,549,573]" gridrow="8" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FDAEFDAFF252" box="[166,524,549,573]" gridcol="0" gridrow="8" pageId="4" pageNumber="70">Dorsal fin base length</th>
<td id="91011D3CAA4700523D60FDAEFC37F252" box="[817,916,549,573]" gridcol="1" gridrow="8" pageId="4" pageNumber="70">54.1</td>
<td id="91011D3CAA4700523DBCFDAEFB90F252" box="[1005,1075,549,573]" gridcol="2" gridrow="8" pageId="4" pageNumber="70">53.4</td>
<td id="91011D3CAA4700523A2AFDAEFB04F252" box="[1147,1191,549,573]" gridcol="3" gridrow="8" pageId="4" pageNumber="70">0.83</td>
<td id="91011D3CAA4700523AAAFDAEFACBF252" box="[1275,1384,549,573]" gridcol="4" gridrow="8" pageId="4" pageNumber="70">52.4–54.2</td>
</tr>
<tr id="D2D07440AA4700523EF7FDC6FACBF209" box="[166,1384,589,614]" gridrow="9" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FDC6FDAFF209" box="[166,524,589,614]" gridcol="0" gridrow="9" pageId="4" pageNumber="70">Anterior dorsal to anterior anal</th>
<td id="91011D3CAA4700523D60FDC6FC37F209" box="[817,916,589,614]" gridcol="1" gridrow="9" pageId="4" pageNumber="70">48.0</td>
<td id="91011D3CAA4700523DBCFDC6FB90F209" box="[1005,1075,589,614]" gridcol="2" gridrow="9" pageId="4" pageNumber="70">46.1</td>
<td id="91011D3CAA4700523A2AFDC6FB04F209" box="[1147,1191,589,614]" gridcol="3" gridrow="9" pageId="4" pageNumber="70">1.3</td>
<td id="91011D3CAA4700523AAAFDC6FACBF209" box="[1275,1384,589,614]" gridcol="4" gridrow="9" pageId="4" pageNumber="70">44.4–48</td>
</tr>
<tr id="D2D07440AA4700523EF7FDFDFACBF2E1" box="[166,1384,630,654]" gridrow="10" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FDFDFDAFF2E1" box="[166,524,630,654]" gridcol="0" gridrow="10" pageId="4" pageNumber="70">Anterior dorsal to posterior anal</th>
<td id="91011D3CAA4700523D60FDFDFC37F2E1" box="[817,916,630,654]" gridcol="1" gridrow="10" pageId="4" pageNumber="70">57.9</td>
<td id="91011D3CAA4700523DBCFDFDFB90F2E1" box="[1005,1075,630,654]" gridcol="2" gridrow="10" pageId="4" pageNumber="70">57.3</td>
<td id="91011D3CAA4700523A2AFDFDFB04F2E1" box="[1147,1191,630,654]" gridcol="3" gridrow="10" pageId="4" pageNumber="70">0.67</td>
<td id="91011D3CAA4700523AAAFDFDFACBF2E1" box="[1275,1384,630,654]" gridcol="4" gridrow="10" pageId="4" pageNumber="70">56.4–58.1</td>
</tr>
<tr id="D2D07440AA4700523EF7FD15FACBF2D9" box="[166,1384,670,694]" gridrow="11" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FD15FDAFF2D9" box="[166,524,670,694]" gridcol="0" gridrow="11" pageId="4" pageNumber="70">Posterior dorsal to posterior anal</th>
<td id="91011D3CAA4700523D60FD15FC37F2D9" box="[817,916,670,694]" gridcol="1" gridrow="11" pageId="4" pageNumber="70">14.4</td>
<td id="91011D3CAA4700523DBCFD15FB90F2D9" box="[1005,1075,670,694]" gridcol="2" gridrow="11" pageId="4" pageNumber="70">14.5</td>
<td id="91011D3CAA4700523A2AFD15FB04F2D9" box="[1147,1191,670,694]" gridcol="3" gridrow="11" pageId="4" pageNumber="70">0.89</td>
<td id="91011D3CAA4700523AAAFD15FACBF2D9" box="[1275,1384,670,694]" gridcol="4" gridrow="11" pageId="4" pageNumber="70">13.4–15.9</td>
</tr>
<tr id="D2D07440AA4700523EF7FD4DFACBF2B1" box="[166,1384,710,734]" gridrow="12" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FD4DFDAFF2B1" box="[166,524,710,734]" gridcol="0" gridrow="12" pageId="4" pageNumber="70">Posterior dorsal to anterior anal</th>
<td id="91011D3CAA4700523D60FD4DFC37F2B1" box="[817,916,710,734]" gridcol="1" gridrow="12" pageId="4" pageNumber="70">27.9</td>
<td id="91011D3CAA4700523DBCFD4DFB90F2B1" box="[1005,1075,710,734]" gridcol="2" gridrow="12" pageId="4" pageNumber="70">28.4</td>
<td id="91011D3CAA4700523A2AFD4DFB04F2B1" box="[1147,1191,710,734]" gridcol="3" gridrow="12" pageId="4" pageNumber="70">0.41</td>
<td id="91011D3CAA4700523AAAFD4DFACBF2B1" box="[1275,1384,710,734]" gridcol="4" gridrow="12" pageId="4" pageNumber="70">27.9–28.9</td>
</tr>
<tr id="D2D07440AA4700523EF7FD65FACBF368" box="[166,1384,750,775]" gridrow="13" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FD65FDAFF368" box="[166,524,750,775]" gridcol="0" gridrow="13" pageId="4" pageNumber="70">Posterior dorsal ventral caudal</th>
<td id="91011D3CAA4700523D60FD65FC37F368" box="[817,916,750,775]" gridcol="1" gridrow="13" pageId="4" pageNumber="70">18.4</td>
<td id="91011D3CAA4700523DBCFD65FB90F368" box="[1005,1075,750,775]" gridcol="2" gridrow="13" pageId="4" pageNumber="70">18.8</td>
<td id="91011D3CAA4700523A2AFD65FB04F368" box="[1147,1191,750,775]" gridcol="3" gridrow="13" pageId="4" pageNumber="70">1.2</td>
<td id="91011D3CAA4700523AAAFD65FACBF368" box="[1275,1384,750,775]" gridcol="4" gridrow="13" pageId="4" pageNumber="70">17.1–20.2</td>
</tr>
<tr id="D2D07440AA4700523EF7FC9CFACBF340" box="[166,1384,791,815]" gridrow="14" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FC9CFDAFF340" box="[166,524,791,815]" gridcol="0" gridrow="14" pageId="4" pageNumber="70">Posterior anal to dorsal caudal</th>
<td id="91011D3CAA4700523D60FC9CFC37F340" box="[817,916,791,815]" gridcol="1" gridrow="14" pageId="4" pageNumber="70">20.1</td>
<td id="91011D3CAA4700523DBCFC9CFB90F340" box="[1005,1075,791,815]" gridcol="2" gridrow="14" pageId="4" pageNumber="70">20.6</td>
<td id="91011D3CAA4700523A2AFC9CFB04F340" box="[1147,1191,791,815]" gridcol="3" gridrow="14" pageId="4" pageNumber="70">1.0</td>
<td id="91011D3CAA4700523AAAFC9CFACBF340" box="[1275,1384,791,815]" gridcol="4" gridrow="14" pageId="4" pageNumber="70">19.4–22.4</td>
</tr>
<tr id="D2D07440AA4700523EF7FCB4FACBF338" box="[166,1384,831,855]" gridrow="15" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FCB4FDAFF338" box="[166,524,831,855]" gridcol="0" gridrow="15" pageId="4" pageNumber="70">Posterior dorsal to pelvic-fin origin</th>
<td id="91011D3CAA4700523D60FCB4FC37F338" box="[817,916,831,855]" gridcol="1" gridrow="15" pageId="4" pageNumber="70">53.1</td>
<td id="91011D3CAA4700523DBCFCB4FB90F338" box="[1005,1075,831,855]" gridcol="2" gridrow="15" pageId="4" pageNumber="70">51.9</td>
<td id="91011D3CAA4700523A2AFCB4FB04F338" box="[1147,1191,831,855]" gridcol="3" gridrow="15" pageId="4" pageNumber="70">1.2</td>
<td id="91011D3CAA4700523AAAFCB4FACBF338" box="[1275,1384,831,855]" gridcol="4" gridrow="15" pageId="4" pageNumber="70">50.6–53.4</td>
</tr>
<tr id="D2D07440AA4700523EF7FCECFACBF310" box="[166,1384,871,895]" gridrow="16" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FCECFDAFF310" box="[166,524,871,895]" gridcol="0" gridrow="16" pageId="4" pageNumber="70">Anterior dorsal to pelvic-fin origin</th>
<td id="91011D3CAA4700523D60FCECFC37F310" box="[817,916,871,895]" gridcol="1" gridrow="16" pageId="4" pageNumber="70">35.2</td>
<td id="91011D3CAA4700523DBCFCECFB90F310" box="[1005,1075,871,895]" gridcol="2" gridrow="16" pageId="4" pageNumber="70">32.8</td>
<td id="91011D3CAA4700523A2AFCECFB04F310" box="[1147,1191,871,895]" gridcol="3" gridrow="16" pageId="4" pageNumber="70">1.7</td>
<td id="91011D3CAA4700523AAAFCECFACBF310" box="[1275,1384,871,895]" gridcol="4" gridrow="16" pageId="4" pageNumber="70">30.7–35.2</td>
</tr>
<tr id="D2D07440AA4700523EF7FC04FACBF3C7" box="[166,1384,911,936]" gridrow="17" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FC04FDAFF3C7" box="[166,524,911,936]" gridcol="0" gridrow="17" pageId="4" pageNumber="70">Caudal peduncle length</th>
<td id="91011D3CAA4700523D60FC04FC37F3C7" box="[817,916,911,936]" gridcol="1" gridrow="17" pageId="4" pageNumber="70">15.0</td>
<td id="91011D3CAA4700523DBCFC04FB90F3C7" box="[1005,1075,911,936]" gridcol="2" gridrow="17" pageId="4" pageNumber="70">15.8</td>
<td id="91011D3CAA4700523A2AFC04FB04F3C7" box="[1147,1191,911,936]" gridcol="3" gridrow="17" pageId="4" pageNumber="70">1.7</td>
<td id="91011D3CAA4700523AAAFC04FACBF3C7" box="[1275,1384,911,936]" gridcol="4" gridrow="17" pageId="4" pageNumber="70">13–18</td>
</tr>
<tr id="D2D07440AA4700523EF7FC33FACBF3BF" box="[166,1384,952,976]" gridrow="18" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FC33FDAFF3BF" box="[166,524,952,976]" gridcol="0" gridrow="18" pageId="4" pageNumber="70">Least caudal peduncle depth</th>
<td id="91011D3CAA4700523D60FC33FC37F3BF" box="[817,916,952,976]" gridcol="1" gridrow="18" pageId="4" pageNumber="70">12.3</td>
<td id="91011D3CAA4700523DBCFC33FB90F3BF" box="[1005,1075,952,976]" gridcol="2" gridrow="18" pageId="4" pageNumber="70">11.7</td>
<td id="91011D3CAA4700523A2AFC33FB04F3BF" box="[1147,1191,952,976]" gridcol="3" gridrow="18" pageId="4" pageNumber="70">0.67</td>
<td id="91011D3CAA4700523AAAFC33FACBF3BF" box="[1275,1384,952,976]" gridcol="4" gridrow="18" pageId="4" pageNumber="70">10.5–12.3</td>
</tr>
<tr id="D2D07440AA4700523EF7FC54FACBF397" box="[166,1384,991,1016]" gridrow="19" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FC54FACBF397" box="[166,1384,991,1016]" colspan="5" colspanRight="4" gridcol="0" gridrow="19" pageId="4" pageNumber="70">
<emphasis id="5E94AA10AA47FFA93EF7FC6BFE3BF397" bold="true" box="[166,408,991,1016]" pageId="4" pageNumber="70">Percent of head length</emphasis>
</th>
</tr>
<tr id="D2D07440AA4700523EF7FB83FACBF44F" box="[166,1384,1032,1056]" gridrow="20" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FB83FDAFF44F" box="[166,524,1032,1056]" gridcol="0" gridrow="20" pageId="4" pageNumber="70">Snout length</th>
<td id="91011D3CAA4700523D60FB83FC37F44F" box="[817,916,1032,1056]" gridcol="1" gridrow="20" pageId="4" pageNumber="70">34.9</td>
<td id="91011D3CAA4700523DBCFB83FB90F44F" box="[1005,1075,1032,1056]" gridcol="2" gridrow="20" pageId="4" pageNumber="70">32.4</td>
<td id="91011D3CAA4700523A2AFB83FB04F44F" box="[1147,1191,1032,1056]" gridcol="3" gridrow="20" pageId="4" pageNumber="70">1.69</td>
<td id="91011D3CAA4700523AAAFB83FACBF44F" box="[1275,1384,1032,1056]" gridcol="4" gridrow="20" pageId="4" pageNumber="70">30.3–34.9</td>
</tr>
<tr id="D2D07440AA4700523EF7FBBBFACBF426" box="[166,1384,1072,1097]" gridrow="21" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FBBBFDAFF426" box="[166,524,1072,1097]" gridcol="0" gridrow="21" pageId="4" pageNumber="70">Postorbital head length</th>
<td id="91011D3CAA4700523D60FBBBFC37F426" box="[817,916,1072,1097]" gridcol="1" gridrow="21" pageId="4" pageNumber="70">45.0</td>
<td id="91011D3CAA4700523DBCFBBBFB90F426" box="[1005,1075,1072,1097]" gridcol="2" gridrow="21" pageId="4" pageNumber="70">43.2</td>
<td id="91011D3CAA4700523A2AFBBBFB04F426" box="[1147,1191,1072,1097]" gridcol="3" gridrow="21" pageId="4" pageNumber="70">1.49</td>
<td id="91011D3CAA4700523AAAFBBBFACBF426" box="[1275,1384,1072,1097]" gridcol="4" gridrow="21" pageId="4" pageNumber="70">41.1–45.0</td>
</tr>
<tr id="D2D07440AA4700523EF7FBD2FACBF41E" box="[166,1384,1113,1137]" gridrow="22" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FBD2FDAFF41E" box="[166,524,1113,1137]" gridcol="0" gridrow="22" pageId="4" pageNumber="70">Horizontal eye diameter</th>
<td id="91011D3CAA4700523D60FBD2FC37F41E" box="[817,916,1113,1137]" gridcol="1" gridrow="22" pageId="4" pageNumber="70">23.4</td>
<td id="91011D3CAA4700523DBCFBD2FB90F41E" box="[1005,1075,1113,1137]" gridcol="2" gridrow="22" pageId="4" pageNumber="70">25.0</td>
<td id="91011D3CAA4700523A2AFBD2FB04F41E" box="[1147,1191,1113,1137]" gridcol="3" gridrow="22" pageId="4" pageNumber="70">2.30</td>
<td id="91011D3CAA4700523AAAFBD2FACBF41E" box="[1275,1384,1113,1137]" gridcol="4" gridrow="22" pageId="4" pageNumber="70">21–26.7</td>
</tr>
<tr id="D2D07440AA4700523EF7FB0AFACBF4F6" box="[166,1384,1153,1177]" gridrow="23" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FB0AFDAFF4F6" box="[166,524,1153,1177]" gridcol="0" gridrow="23" pageId="4" pageNumber="70">Vertical eye diameter</th>
<td id="91011D3CAA4700523D60FB0AFC37F4F6" box="[817,916,1153,1177]" gridcol="1" gridrow="23" pageId="4" pageNumber="70">21.3</td>
<td id="91011D3CAA4700523DBCFB0AFB90F4F6" box="[1005,1075,1153,1177]" gridcol="2" gridrow="23" pageId="4" pageNumber="70">23.4</td>
<td id="91011D3CAA4700523A2AFB0AFB04F4F6" box="[1147,1191,1153,1177]" gridcol="3" gridrow="23" pageId="4" pageNumber="70">2.06</td>
<td id="91011D3CAA4700523AAAFB0AFACBF4F6" box="[1275,1384,1153,1177]" gridcol="4" gridrow="23" pageId="4" pageNumber="70">21.2–25.7</td>
</tr>
<tr id="D2D07440AA4700523EF7FB22FACBF4AE" box="[166,1384,1193,1217]" gridrow="24" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FB22FDAFF4AE" box="[166,524,1193,1217]" gridcol="0" gridrow="24" pageId="4" pageNumber="70">Interorbital width</th>
<td id="91011D3CAA4700523D60FB22FC37F4AE" box="[817,916,1193,1217]" gridcol="1" gridrow="24" pageId="4" pageNumber="70">15.3</td>
<td id="91011D3CAA4700523DBCFB22FB90F4AE" box="[1005,1075,1193,1217]" gridcol="2" gridrow="24" pageId="4" pageNumber="70">15.1</td>
<td id="91011D3CAA4700523A2AFB22FB04F4AE" box="[1147,1191,1193,1217]" gridcol="3" gridrow="24" pageId="4" pageNumber="70">0.54</td>
<td id="91011D3CAA4700523AAAFB22FACBF4AE" box="[1275,1384,1193,1217]" gridcol="4" gridrow="24" pageId="4" pageNumber="70">14.3–15.9</td>
</tr>
<tr id="D2D07440AA4700523EF7FB5AFACBF485" box="[166,1384,1233,1258]" gridrow="25" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FB5AFDAFF485" box="[166,524,1233,1258]" gridcol="0" gridrow="25" pageId="4" pageNumber="70">Head depth</th>
<td id="91011D3CAA4700523D60FB5AFC37F485" box="[817,916,1233,1258]" gridcol="1" gridrow="25" pageId="4" pageNumber="70">70.0</td>
<td id="91011D3CAA4700523DBCFB5AFB90F485" box="[1005,1075,1233,1258]" gridcol="2" gridrow="25" pageId="4" pageNumber="70">70.8</td>
<td id="91011D3CAA4700523A2AFB5AFB04F485" box="[1147,1191,1233,1258]" gridcol="3" gridrow="25" pageId="4" pageNumber="70">5.1</td>
<td id="91011D3CAA4700523AAAFB5AFACBF485" box="[1275,1384,1233,1258]" gridcol="4" gridrow="25" pageId="4" pageNumber="70">63.3–78</td>
</tr>
<tr id="D2D07440AA4700523EF7FB72FACBF57D" box="[166,1384,1273,1298]" gridrow="26" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FB72FDAFF57D" box="[166,524,1273,1298]" gridcol="0" gridrow="26" pageId="4" pageNumber="70">Pre-orbital depth</th>
<td id="91011D3CAA4700523D60FB72FC37F57D" box="[817,916,1273,1298]" gridcol="1" gridrow="26" pageId="4" pageNumber="70">19.7</td>
<td id="91011D3CAA4700523DBCFB72FB90F57D" box="[1005,1075,1273,1298]" gridcol="2" gridrow="26" pageId="4" pageNumber="70">17.1</td>
<td id="91011D3CAA4700523A2AFB72FB04F57D" box="[1147,1191,1273,1298]" gridcol="3" gridrow="26" pageId="4" pageNumber="70">1.52</td>
<td id="91011D3CAA4700523AAAFB72FACBF57D" box="[1275,1384,1273,1298]" gridcol="4" gridrow="26" pageId="4" pageNumber="70">15.2–19.7</td>
</tr>
<tr id="D2D07440AA4700523EF7FAA9FACBF555" box="[166,1384,1314,1338]" gridrow="27" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FAA9FDAFF555" box="[166,524,1314,1338]" gridcol="0" gridrow="27" pageId="4" pageNumber="70">Cheek depth</th>
<td id="91011D3CAA4700523D60FAA9FC37F555" box="[817,916,1314,1338]" gridcol="1" gridrow="27" pageId="4" pageNumber="70">27.6</td>
<td id="91011D3CAA4700523DBCFAA9FB90F555" box="[1005,1075,1314,1338]" gridcol="2" gridrow="27" pageId="4" pageNumber="70">25.7</td>
<td id="91011D3CAA4700523A2AFAA9FB04F555" box="[1147,1191,1314,1338]" gridcol="3" gridrow="27" pageId="4" pageNumber="70">1.97</td>
<td id="91011D3CAA4700523AAAFAA9FACBF555" box="[1275,1384,1314,1338]" gridcol="4" gridrow="27" pageId="4" pageNumber="70">23.5–28.1</td>
</tr>
<tr id="D2D07440AA4700523EF7FAC1FACBF50D" box="[166,1384,1354,1378]" gridrow="28" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FAC1FDAFF50D" box="[166,524,1354,1378]" gridcol="0" gridrow="28" pageId="4" pageNumber="70">Lower jaw length</th>
<td id="91011D3CAA4700523D60FAC1FC37F50D" box="[817,916,1354,1378]" gridcol="1" gridrow="28" pageId="4" pageNumber="70">51.4</td>
<td id="91011D3CAA4700523DBCFAC1FB90F50D" box="[1005,1075,1354,1378]" gridcol="2" gridrow="28" pageId="4" pageNumber="70">48.7</td>
<td id="91011D3CAA4700523A2AFAC1FB04F50D" box="[1147,1191,1354,1378]" gridcol="3" gridrow="28" pageId="4" pageNumber="70">3.73</td>
<td id="91011D3CAA4700523AAAFAC1FACBF50D" box="[1275,1384,1354,1378]" gridcol="4" gridrow="28" pageId="4" pageNumber="70">43.4–53.5</td>
</tr>
<tr id="D2D07440AA4700523EF7FAF9FACBF5E5" box="[166,1384,1394,1418]" gridrow="29" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FAF9FACBF5E5" box="[166,1384,1394,1418]" colspan="5" colspanRight="4" gridcol="0" gridrow="29" pageId="4" pageNumber="70">
<emphasis id="5E94AA10AA47FFA93EF7FAF9FF50F5E5" bold="true" box="[166,243,1394,1418]" pageId="4" pageNumber="70">Counts</emphasis>
</th>
</tr>
<tr id="D2D07440AA4700523EF7FA11FACBF5DC" box="[166,1384,1434,1459]" gridrow="30" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FA11FDAFF5DC" box="[166,524,1434,1459]" gridcol="0" gridrow="30" pageId="4" pageNumber="70">Dorsal-fin spines</th>
<td id="91011D3CAA4700523D60FA11FC37F5DC" box="[817,916,1434,1459]" gridcol="1" gridrow="30" pageId="4" pageNumber="70">15</td>
<td id="91011D3CAA4700523DBCFA11FB90F5DC" box="[1005,1075,1434,1459]" gridcol="2" gridrow="30" pageId="4" pageNumber="70">15</td>
<td id="91011D3CAA4700523A2AFA11FB04F5DC" box="[1147,1191,1434,1459]" gridcol="3" gridrow="30" pageId="4" pageNumber="70">83.3</td>
<td id="91011D3CAA4700523AAAFA11FACBF5DC" box="[1275,1384,1434,1459]" gridcol="4" gridrow="30" pageId="4" pageNumber="70">14-15</td>
</tr>
<tr id="D2D07440AA4700523EF7FA48FACBF5B4" box="[166,1384,1475,1499]" gridrow="31" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7FA48FDAFF5B4" box="[166,524,1475,1499]" gridcol="0" gridrow="31" pageId="4" pageNumber="70">Dorsal-fin rays</th>
<td id="91011D3CAA4700523D60FA48FC37F5B4" box="[817,916,1475,1499]" gridcol="1" gridrow="31" pageId="4" pageNumber="70">14</td>
<td id="91011D3CAA4700523DBCFA48FB90F5B4" box="[1005,1075,1475,1499]" gridcol="2" gridrow="31" pageId="4" pageNumber="70">14</td>
<td id="91011D3CAA4700523A2AFA48FB04F5B4" box="[1147,1191,1475,1499]" gridcol="3" gridrow="31" pageId="4" pageNumber="70">50</td>
<td id="91011D3CAA4700523AAAFA48FACBF5B4" box="[1275,1384,1475,1499]" gridcol="4" gridrow="31" pageId="4" pageNumber="70">13–15</td>
</tr>
<tr id="D2D07440AA4700523EF7FA60FACBF66C" box="[166,1384,1515,1539]" gridrow="32" pageId="4" pageNumber="70" rowspan-4="1">
<th id="91011D3CAA4700523EF7FA60FDAFF66C" box="[166,524,1515,1539]" gridcol="0" gridrow="32" pageId="4" pageNumber="70">Anal-fin spines</th>
<td id="91011D3CAA4700523D60FA60FC37F66C" box="[817,916,1515,1539]" gridcol="1" gridrow="32" pageId="4" pageNumber="70">3</td>
<td id="91011D3CAA4700523DBCFA60FB90F66C" box="[1005,1075,1515,1539]" gridcol="2" gridrow="32" pageId="4" pageNumber="70">3</td>
<td id="91011D3CAA4700523A2AFA60FB04F66C" box="[1147,1191,1515,1539]" gridcol="3" gridrow="32" pageId="4" pageNumber="70">100</td>
</tr>
<tr id="D2D07440AA4700523EF7F998FACBF643" box="[166,1384,1555,1580]" gridrow="33" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7F998FDAFF643" box="[166,524,1555,1580]" gridcol="0" gridrow="33" pageId="4" pageNumber="70">Anal-fin rays</th>
<td id="91011D3CAA4700523D60F998FC37F643" box="[817,916,1555,1580]" gridcol="1" gridrow="33" pageId="4" pageNumber="70">10</td>
<td id="91011D3CAA4700523DBCF998FB90F643" box="[1005,1075,1555,1580]" gridcol="2" gridrow="33" pageId="4" pageNumber="70">10</td>
<td id="91011D3CAA4700523A2AF998FB04F643" box="[1147,1191,1555,1580]" gridcol="3" gridrow="33" pageId="4" pageNumber="70">83.3</td>
<td id="91011D3CAA4700523AAAF998FACBF643" box="[1275,1384,1555,1580]" gridcol="4" gridrow="33" pageId="4" pageNumber="70">9–10</td>
</tr>
<tr id="D2D07440AA4700523EF7F9B0FACBF63B" box="[166,1384,1595,1620]" gridrow="34" pageId="4" pageNumber="70" rowspan-4="1">
<th id="91011D3CAA4700523EF7F9B0FDAFF63B" box="[166,524,1595,1620]" gridcol="0" gridrow="34" pageId="4" pageNumber="70">Pelvic-fin rays</th>
<td id="91011D3CAA4700523D60F9B0FC37F63B" box="[817,916,1595,1620]" gridcol="1" gridrow="34" pageId="4" pageNumber="70">5</td>
<td id="91011D3CAA4700523DBCF9B0FB90F63B" box="[1005,1075,1595,1620]" gridcol="2" gridrow="34" pageId="4" pageNumber="70">5</td>
<td id="91011D3CAA4700523A2AF9B0FB04F63B" box="[1147,1191,1595,1620]" gridcol="3" gridrow="34" pageId="4" pageNumber="70">100</td>
</tr>
<tr id="D2D07440AA4700523EF7F9EFFACBF613" box="[166,1384,1636,1660]" gridrow="35" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7F9EFFDAFF613" box="[166,524,1636,1660]" gridcol="0" gridrow="35" pageId="4" pageNumber="70">Pectoral-fin rays</th>
<td id="91011D3CAA4700523D60F9EFFC37F613" box="[817,916,1636,1660]" gridcol="1" gridrow="35" pageId="4" pageNumber="70">15</td>
<td id="91011D3CAA4700523DBCF9EFFB90F613" box="[1005,1075,1636,1660]" gridcol="2" gridrow="35" pageId="4" pageNumber="70">15</td>
<td id="91011D3CAA4700523A2AF9EFFB04F613" box="[1147,1191,1636,1660]" gridcol="3" gridrow="35" pageId="4" pageNumber="70">50</td>
<td id="91011D3CAA4700523AAAF9EFFACBF613" box="[1275,1384,1636,1660]" gridcol="4" gridrow="35" pageId="4" pageNumber="70">13–15</td>
</tr>
<tr id="D2D07440AA4700523EF7F907FACBF6CB" box="[166,1384,1676,1700]" gridrow="36" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7F907FDAFF6CB" box="[166,524,1676,1700]" gridcol="0" gridrow="36" pageId="4" pageNumber="70">Lateral line scales</th>
<td id="91011D3CAA4700523D60F907FC37F6CB" box="[817,916,1676,1700]" gridcol="1" gridrow="36" pageId="4" pageNumber="70">35</td>
<td id="91011D3CAA4700523DBCF907FB90F6CB" box="[1005,1075,1676,1700]" gridcol="2" gridrow="36" pageId="4" pageNumber="70">35</td>
<td id="91011D3CAA4700523A2AF907FB04F6CB" box="[1147,1191,1676,1700]" gridcol="3" gridrow="36" pageId="4" pageNumber="70">66.7</td>
<td id="91011D3CAA4700523AAAF907FACBF6CB" box="[1275,1384,1676,1700]" gridcol="4" gridrow="36" pageId="4" pageNumber="70">34–36</td>
</tr>
<tr id="D2D07440AA4700523EF7F93FFACBF6A3" box="[166,1384,1716,1740]" gridrow="37" pageId="4" pageNumber="70" rowspan-4="1">
<th id="91011D3CAA4700523EF7F93FFDAFF6A3" box="[166,524,1716,1740]" gridcol="0" gridrow="37" pageId="4" pageNumber="70">Pored scales posterior to LL</th>
<td id="91011D3CAA4700523D60F93FFC37F6A3" box="[817,916,1716,1740]" gridcol="1" gridrow="37" pageId="4" pageNumber="70">2</td>
<td id="91011D3CAA4700523DBCF93FFB90F6A3" box="[1005,1075,1716,1740]" gridcol="2" gridrow="37" pageId="4" pageNumber="70">2</td>
<td id="91011D3CAA4700523A2AF93FFB04F6A3" box="[1147,1191,1716,1740]" gridcol="3" gridrow="37" pageId="4" pageNumber="70">100</td>
</tr>
<tr id="D2D07440AA4700523EF7F957FACBF69A" box="[166,1384,1756,1781]" gridrow="38" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7F957FDAFF69A" box="[166,524,1756,1781]" gridcol="0" gridrow="38" pageId="4" pageNumber="70">Cheek scales</th>
<td id="91011D3CAA4700523D60F957FC37F69A" box="[817,916,1756,1781]" gridcol="1" gridrow="38" pageId="4" pageNumber="70">9</td>
<td id="91011D3CAA4700523DBCF957FB90F69A" box="[1005,1075,1756,1781]" gridcol="2" gridrow="38" pageId="4" pageNumber="70">8</td>
<td id="91011D3CAA4700523A2AF957FB04F69A" box="[1147,1191,1756,1781]" gridcol="3" gridrow="38" pageId="4" pageNumber="70">50</td>
<td id="91011D3CAA4700523AAAF957FACBF69A" box="[1275,1384,1756,1781]" gridcol="4" gridrow="38" pageId="4" pageNumber="70">7–10</td>
</tr>
<tr id="D2D07440AA4700523EF7F88EFACBF772" box="[166,1384,1797,1821]" gridrow="39" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7F88EFDAFF772" box="[166,524,1797,1821]" gridcol="0" gridrow="39" pageId="4" pageNumber="70">Gill rakers on first epibranchial</th>
<td id="91011D3CAA4700523D60F88EFC37F772" box="[817,916,1797,1821]" gridcol="1" gridrow="39" pageId="4" pageNumber="70">3</td>
<td id="91011D3CAA4700523DBCF88EFB90F772" box="[1005,1075,1797,1821]" gridcol="2" gridrow="39" pageId="4" pageNumber="70">3/4</td>
<td id="91011D3CAA4700523A2AF88EFB04F772" box="[1147,1191,1797,1821]" gridcol="3" gridrow="39" pageId="4" pageNumber="70">50</td>
<td id="91011D3CAA4700523AAAF88EFACBF772" box="[1275,1384,1797,1821]" gridcol="4" gridrow="39" pageId="4" pageNumber="70">3–4</td>
</tr>
<tr id="D2D07440AA4700523EF7F8A6FACBF72A" box="[166,1384,1837,1861]" gridrow="40" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7F8A6FDAFF72A" box="[166,524,1837,1861]" gridcol="0" gridrow="40" pageId="4" pageNumber="70">Gill rakers on first ceratobranchial</th>
<td id="91011D3CAA4700523D60F8A6FC37F72A" box="[817,916,1837,1861]" gridcol="1" gridrow="40" pageId="4" pageNumber="70">11</td>
<td id="91011D3CAA4700523DBCF8A6FB90F72A" box="[1005,1075,1837,1861]" gridcol="2" gridrow="40" pageId="4" pageNumber="70">9</td>
<td id="91011D3CAA4700523A2AF8A6FB04F72A" box="[1147,1191,1837,1861]" gridcol="3" gridrow="40" pageId="4" pageNumber="70">50</td>
<td id="91011D3CAA4700523AAAF8A6FACBF72A" box="[1275,1384,1837,1861]" gridcol="4" gridrow="40" pageId="4" pageNumber="70">9–11</td>
</tr>
<tr id="D2D07440AA4700523EF7F8DEFACBF702" box="[166,1384,1877,1901]" gridrow="41" pageId="4" pageNumber="70">
<th id="91011D3CAA4700523EF7F8DEFDAFF702" box="[166,524,1877,1901]" gridcol="0" gridrow="41" pageId="4" pageNumber="70">Teeth outer row of left lower jaw</th>
<td id="91011D3CAA4700523D60F8DEFC37F702" box="[817,916,1877,1901]" gridcol="1" gridrow="41" pageId="4" pageNumber="70">2</td>
<td id="91011D3CAA4700523DBCF8DEFB90F702" box="[1005,1075,1877,1901]" gridcol="2" gridrow="41" pageId="4" pageNumber="70">8</td>
<td id="91011D3CAA4700523A2AF8DEFB04F702" box="[1147,1191,1877,1901]" gridcol="3" gridrow="41" pageId="4" pageNumber="70">50</td>
<td id="91011D3CAA4700523AAAF8DEFACBF702" box="[1275,1384,1877,1901]" gridcol="4" gridrow="41" pageId="4" pageNumber="70">7–12</td>
</tr>
<tr id="D2D07440AA4700523EF7F8F6FACBF7F9" box="[166,1384,1917,1942]" gridrow="42" pageId="4" pageNumber="70" rowspan-4="1">
<th id="91011D3CAA4700523EF7F8F6FDAFF7F9" box="[166,524,1917,1942]" gridcol="0" gridrow="42" pageId="4" pageNumber="70">Teeth rows on upper jaw</th>
<td id="91011D3CAA4700523D60F8F6FC37F7F9" box="[817,916,1917,1942]" gridcol="1" gridrow="42" pageId="4" pageNumber="70">11</td>
<td id="91011D3CAA4700523DBCF8F6FB90F7F9" box="[1005,1075,1917,1942]" gridcol="2" gridrow="42" pageId="4" pageNumber="70">2</td>
<td id="91011D3CAA4700523A2AF8F6FB04F7F9" box="[1147,1191,1917,1942]" gridcol="3" gridrow="42" pageId="4" pageNumber="70">100</td>
</tr>
<tr id="D2D07440AA4700523EF7F82DFACBF7D1" box="[166,1384,1958,1982]" gridrow="43" pageId="4" pageNumber="70" rowspan-4="1">
<th id="91011D3CAA4700523EF7F82DFDAFF7D1" box="[166,524,1958,1982]" gridcol="0" gridrow="43" pageId="4" pageNumber="70">Teeth rows on lower jaw</th>
<td id="91011D3CAA4700523D60F82DFC37F7D1" box="[817,916,1958,1982]" gridcol="1" gridrow="43" pageId="4" pageNumber="70">2</td>
<td id="91011D3CAA4700523DBCF82DFB90F7D1" box="[1005,1075,1958,1982]" gridcol="2" gridrow="43" pageId="4" pageNumber="70">2</td>
<td id="91011D3CAA4700523A2AF82DFB04F7D1" box="[1147,1191,1958,1982]" gridcol="3" gridrow="43" pageId="4" pageNumber="70">100</td>
</tr>
</table>
</paragraph>
<paragraph id="6C5F7602AA46FFA83E96FF1CFAD4F1BE" blockId="5.[151,1437,150,1077]" pageId="5" pageNumber="71">
Head elongate 2.5–2.7 times in SL (37–39%SL), length greater than body depth, pointed with straight predorsal profile extending 20˚ above horizontal. Eyes large (HED 21–26.7%HL; VED 21.2–25.7% HL), dorso-lateral in anterior half of head, entirely above level of the mouth and before the dorsal edge of the operculum. Interorbit narrow, less than half orbit diameter. Snout relatively long, greater than orbit diameter, nares in mid-snout before orbits. Lachrymal (pre-orbit) depth equal to orbit diameter. Cheek below and behind orbits deep with 7–10 rows of scales. Post orbit half-length of head, interorbit 14.3–5.9% HL. Mouth terminal, large, protrusible; angle of closed jaw of
<typeStatus id="B35BC8A0AA46FFA83EC6FEE4FF5AF1E6" box="[151,249,367,393]" pageId="5" pageNumber="71" type="holotype">holotype</typeStatus>
30˚ below horizontal, posterior premaxilla to below anterior orbit, lips well developed, even. Teeth canini-form, exposed, widely spaced in 2 series on upper and lower jaws. First branchial arch with 3–4-1-9–11 gill-rakers. Gill-slit large, extending from upper flank opposite dorsal orbit rim in smooth arch to mid-ventral below orbits.
</paragraph>
<paragraph id="6C5F7602AA46FFA83E96FE51FD4FF3A6" blockId="5.[151,1437,150,1077]" pageId="5" pageNumber="71">
<emphasis id="5E94AA10AA46FFA83E96FE51FEEBF19B" bold="true" box="[199,328,474,500]" pageId="5" pageNumber="71">Coloration</emphasis>
. Preserved specimens yellowish-brown with darker greyish brown interorbital and body bars, ventral surfaces yellow-brown. Fins light yellowish with grey markings; head with broad greyish band across lachrymal through eye to upper operculum and opercular spot. Body with two lateral black pigment bands and six broad dark brown bars to caudal-fin base reaching from dorsal fin to lower flanks, dark brown spots at base of scales. Live males with broad black bar from lachrymal through lower eye across post-orbit to large opercular spot; iridescent pale blue and white patches on cheeks and above post-orbital bar, white lower cheeks and gular, opercle dusted with grey and black below dorsal black bar, pale blue opercular membrane. Body bluish-black above, prominent black bars on body with white infused with blue interspaces between bars, below pectoral fins, and on lower flanks and ventral side. Dorsal fin with grey-black blotched membranes between anterior spines; dark grey to black blotches with light blocks on membranes between posterior 5–6 spines; soft dorsal with dark grey-black blotches and clear spots on membranes, edges grey. Caudal fin with elongate dark grey-black rectangles and white spots on membranes, a lighter creamish brown submarginal band and greyish black marginal band. Anal fin with 3– 4 rows of deep greyish-red spots between pale yellow bands. Pectoral fins with grey rays and translucent hyaline membranes. Pelvic fins with clear rays and a sooty dust over rays.
</paragraph>
<paragraph id="6C5F7602AA46FFA83E96FC59FAB3F45A" blockId="5.[151,1437,150,1077]" pageId="5" pageNumber="71">
<emphasis id="5E94AA10AA46FFA83E96FC59FE89F383" bold="true" box="[199,298,978,1004]" pageId="5" pageNumber="71">Habitat.</emphasis>
Collected in gill nets set along the fringing
<taxonomicName id="ABE00D81AA46FFA83D7EFC58FC13F382" box="[815,944,979,1005]" class="Liliopsida" family="Poaceae" genus="Phragmites" kingdom="Plantae" order="Poales" pageId="5" pageNumber="71" phylum="Tracheophyta" rank="genus">
<emphasis id="5E94AA10AA46FFA83D7EFC58FC13F382" box="[815,944,979,1005]" italics="true" pageId="5" pageNumber="71">Phragmites</emphasis>
</taxonomicName>
reed bed in the main Cuito channel at the Cuito-Cuanavale main road bridge (
<figureCitation id="F4DB6A87AA46FFA83C65FC7CFDD8F47E" box="[564,635,1015,1041]" captionStart="FIGURE 3" captionStartId="5.[151,250,2013,2037]" captionTargetBox="[162,1424,1101,1988]" captionTargetId="figure-467@5.[162,1424,1101,1989]" captionTargetPageId="5" captionText="FIGURE 3. The main Cuito channel at the Cuito-Cuanavale main road bridge, where Serranochromis alvum was collected." figureDoi="http://doi.org/10.5281/zenodo.4435775" httpUri="https://zenodo.org/record/4435775/files/figure.png" pageId="5" pageNumber="71">Fig. 3</figureCitation>
). Water depths were approximately
<quantity id="AB18DBE7AA46FFA83A4AFC7DFBD0F47E" box="[1051,1139,1014,1041]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="5" pageNumber="71" unit="m" value="1.75" valueMax="2.0" valueMin="1.5">
1.5–
<specimenCount id="7AE6BD8BAA46FFA83A1CFC7DFBD0F47E" box="[1101,1139,1014,1041]" pageId="5" pageNumber="71" type="generic">2m</specimenCount>
</quantity>
. The riverbed was mostly sand substrate with occasional marginal aquatic weed beds and fringing emergent beds of
<taxonomicName id="ABE00D81AA46FFA83AD9FB90FAAAF45A" box="[1160,1289,1051,1077]" class="Liliopsida" family="Poaceae" genus="Phragmites" kingdom="Plantae" order="Poales" pageId="5" pageNumber="71" phylum="Tracheophyta" rank="genus">
<emphasis id="5E94AA10AA46FFA83AD9FB90FAAAF45A" box="[1160,1289,1051,1077]" italics="true" pageId="5" pageNumber="71">Phragmites</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="389F268AAA46FFA83EC6F856FADCF79A" ID-DOI="http://doi.org/10.5281/zenodo.4435775" ID-Zenodo-Dep="4435775" box="[151,1407,2013,2037]" httpUri="https://zenodo.org/record/4435775/files/figure.png" pageId="5" pageNumber="71" startId="5.[151,250,2013,2037]" targetBox="[162,1424,1101,1988]" targetPageId="5">
<paragraph id="6C5F7602AA46FFA83EC6F856FADCF79A" blockId="5.[151,1407,2013,2037]" box="[151,1407,2013,2037]" pageId="5" pageNumber="71">
<emphasis id="5E94AA10AA46FFA83EC6F856FEB0F79A" bold="true" box="[151,275,2013,2037]" pageId="5" pageNumber="71">FIGURE 3.</emphasis>
The main Cuito channel at the Cuito-Cuanavale main road bridge, where
<taxonomicName id="ABE00D81AA46FFA83A50F856FB4BF79A" authorityName="Stauffer &amp; Bills &amp; Skelton" authorityYear="2021" box="[1025,1256,2013,2037]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="71" phylum="Chordata" rank="species" species="alvum">
<emphasis id="5E94AA10AA46FFA83A50F856FB4BF79A" box="[1025,1256,2013,2037]" italics="true" pageId="5" pageNumber="71">Serranochromis alvum</emphasis>
</taxonomicName>
was collected.
</paragraph>
</caption>
<subSubSection id="24FA2589AA45FFAB3E96FF1DFCE0F0BA" pageId="6" pageNumber="72" type="distribution">
<paragraph id="6C5F7602AA45FFAB3E96FF1DFCE0F0BA" blockId="6.[151,1436,150,285]" pageId="6" pageNumber="72">
<emphasis id="5E94AA10AA45FFAB3E96FF1DFEC2F0DF" bold="true" box="[199,353,150,176]" pageId="6" pageNumber="72">Distribution.</emphasis>
Currently known only from the
<typeStatus id="B35BC8A0AA45FFAB3C9CFF1CFD5EF0DE" box="[717,765,151,177]" pageId="6" pageNumber="72">type</typeStatus>
locality at Cuito-Cuanavale at the junction of the Cuito and Cuanavale rivers, tributary to the Okavango River in
<collectingCountry id="14F73692AA45FFAB3CBBFF30FC9FF0BA" box="[746,828,187,213]" name="Angola" pageId="6" pageNumber="72">Angola</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="24FA2589AA45FFAB3E96FF54FD15F172" pageId="6" pageNumber="72" type="etymology">
<paragraph id="6C5F7602AA45FFAB3E96FF54FD15F172" blockId="6.[151,1436,150,285]" pageId="6" pageNumber="72">
<emphasis id="5E94AA10AA45FFAB3E96FF54FEEDF096" bold="true" box="[199,334,223,249]" pageId="6" pageNumber="72">Etymology.</emphasis>
The specific name,
<emphasis id="5E94AA10AA45FFAB3C62FF54FDDDF096" box="[563,638,223,249]" italics="true" pageId="6" pageNumber="72">
<taxonomicName id="ABE00D81AA45FFAB3C62FF54FDD9F096" authorityName="Stauffer &amp; Bills &amp; Skelton" authorityYear="2021" box="[563,634,223,249]" class="Actinopterygii" family="Cichlidae" genus="Serranochromis" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="72" phylum="Chordata" rank="species" species="alvum">alvum</taxonomicName>
,
</emphasis>
a noun in apposition, is derived from the Latin, which means blotches and refers to the prominent lateral pigmentation.
</paragraph>
</subSubSection>
</treatment>
</document>
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