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<document id="B5056BC0AA64BEA428FAD87E51814E2D" ID-DOI="10.5281/zenodo.156857" ID-GBIF-Dataset="bc8fbb8b-8101-4fe5-9ffd-0b5aa5db803c" ID-ISSN="1175­5326" ID-Zenodo-Dep="156857" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459605459478" checkinUser="plazi" docAuthor="Marinoni, Luciane, Steyskal, George C. &amp; Knutson, Lloyd" docDate="2003" docId="E43B9566FF9D4746FE9CF94FFD23FD47" docLanguage="en" docName="zt00191.pdf" docOrigin="Zootaxa 191" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Thecomyia Perty" docType="treatment" docVersion="10" lastPageNumber="6" masterDocId="1802ED1EFF9F4743FF94FFB6FFC5FFA9" masterDocTitle="Revision and cladistic analysis of the Neotropical Genus Thecomyia Perty (Diptera: Sciomyzidae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="3" updateTime="1698202859598" updateUser="plazi">
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<mods:title id="67F4640F8841D93F656DD59B4F2160AC">Revision and cladistic analysis of the Neotropical Genus Thecomyia Perty (Diptera: Sciomyzidae)</mods:title>
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<mods:namePart id="6671E7985D5DFEA828E78970C5D972AB">Steyskal, George C.</mods:namePart>
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<heading id="3765931CFF9D4741FE9CF94FFDF5F8BA" bold="true" box="[264,560,1785,1811]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
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Genus
<taxonomicName id="AB925FF3FF9D4741FECBF94FFDF5F8BA" ID-CoL="7VH2" ID-ENA="2174686" authority="Perty" authorityName="Perty" box="[351,560,1785,1811]" class="Insecta" family="Sciomyzidae" genus="Thecomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="5EE6F862FF9D4741FECBF94FFE21F8BA" bold="true" box="[351,484,1785,1811]" italics="true" pageId="2" pageNumber="3">Thecomyia</emphasis>
Perty
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</emphasis>
</heading>
</paragraph>
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<paragraph id="6C2D2470FF9D4740FE9CF88CFE5BFEFA" blockId="2.[264,1322,1850,1905]" lastBlockId="3.[312,1323,284,339]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
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<taxonomicName id="AB925FF3FF9D4741FE9CF88CFDFDF8F8" ID-CoL="7VH2" authority="Perty, 1833: 189" authorityName="Perty" authorityPageNumber="189" authorityYear="1833" box="[264,568,1850,1873]" class="Insecta" family="Sciomyzidae" genus="Thecomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="5EE6F862FF9D4741FE9CF88CFEB8F8F8" box="[264,381,1850,1873]" italics="true" pageId="2" pageNumber="3">Thecomyia</emphasis>
<treatmentCitation id="ED330261FF9D4741FE12F88CFDFDF8F8" author="Perty" box="[390,568,1850,1873]" page="189" pageId="2" pageNumber="3" year="1833">
<bibRefCitation id="08035981FF9D4741FE12F88CFDC1F8F8" author="Perty" box="[390,516,1850,1873]" pageId="2" pageNumber="35" refString="Perty, M. (1833) Delectus animalium articulatorum quae in itinere per Brasiliam annis MDCCCX- VII-MDCCCXX jussu et auspiciis Maximiliani Josephi I. Bavariae regis augustissimi peracto collegerunt Dr. J. B. de Sphix et Dr. C. F. Ph. De Martius. p. 189, pl. 37, Munich." type="journal volume" year="1833">Perty, 1833</bibRefCitation>
: 189
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</taxonomicName>
[
<typeStatus id="B3299AD2FF9D4741FDD3F88CFDBDF8F8" box="[583,632,1850,1873]" pageId="2" pageNumber="3">type</typeStatus>
species:
<taxonomicName id="AB925FF3FF9D4741FD49F88CFBD3F8F8" ID-CoL="7C76X" authority="Perty" authorityName="Perty" box="[733,1046,1850,1873]" class="Insecta" family="Sciomyzidae" genus="Thecomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="longicornis">
<emphasis id="5EE6F862FF9D4741FD49F88CFC16F8F8" box="[733,979,1850,1873]" italics="true" pageId="2" pageNumber="3">Thecomyia longicornis</emphasis>
Perty
</taxonomicName>
, by original designation.­
<treatmentCitation id="ED330261FF9D4741FEACF8ECFD93F8D8" author="Knutson" box="[312,598,1882,1905]" page="13" pageId="2" pageNumber="3" year="1976">
<bibRefCitation id="08035981FF9D4741FEACF8ECFDECF8D8" author="Knutson" box="[312,553,1882,1905]" pageId="2" pageNumber="35" refString="Knutson, L. V., G. C. Steyskal, J. Zuska &amp; J. Abercrombie (1976) 64. Family Sciomyzidae. In N. Papavero ed., A Catalogue of the Diptera of the Americas South of the United States. Museu de Zoologia da Universidade de Sao Paulo. 24 pp." type="book" year="1976">Knutson et al., 1976</bibRefCitation>
: 13
</treatmentCitation>
[Neotropical catalog].­
<treatmentCitation id="ED330261FF9D4741FCCDF8ECFB74F8D8" author="Knutson" box="[857,1201,1882,1905]" page="455" pageId="2" pageNumber="3" year="1999">
<bibRefCitation id="08035981FF9D4741FCCDF8ECFBBDF8D8" author="Knutson" box="[857,1144,1882,1905]" pageId="2" pageNumber="35" refString="Knutson, L. V. &amp; J-C. Vala. (1999) The male of Tetanoptera leucodactyla (Diptera: Sciomyzidae). European Journal of Entomology, 96 (4), 451 - 457." type="journal article" year="1999">Knutson and Vala, 1999</bibRefCitation>
: 455
</treatmentCitation>
[character matrix].­
<treatmentCitation id="ED330261FF9C4740FE06FEAAFD47FE9A" author="Vala" box="[402,642,284,307]" page="253" pageId="3" pageNumber="4" year="2000">
<bibRefCitation id="08035981FF9C4740FE06FEAAFD8BFE9A" author="Vala" box="[402,590,284,307]" pageId="3" pageNumber="35" refString="Vala, J-C., L. Greve &amp; L. V. Knutson (2000) Description of the male of Verbekaria punctipennis (Diptera: Sciomyzidae). Studia Dipterologica, 7 (1), 247 - 255." type="journal article" year="2000">Vala et al., 2000</bibRefCitation>
: 253
</treatmentCitation>
[character matrix].­
<treatmentCitation id="ED330261FF9C4740FCCDFEAAFB05FE9A" author="Marinoni" box="[857,1216,284,307]" page="191" pageId="3" pageNumber="4" year="2000">
<bibRefCitation id="08035981FF9C4740FCCDFEAAFB4EFE9A" author="Marinoni" box="[857,1163,284,307]" pageId="3" pageNumber="35" refString="Marinoni, L. &amp; W. N. Mathis. (2000) A cladistic analysis of Sciomyzidae Fallen (Diptera). Proceedings of Biological Society of Washington 113 (1): 162 - 209." type="journal article" year="2000">Marinoni and Mathis, 2000</bibRefCitation>
: 191
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[cladistic analysis].
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<emphasis id="5EE6F862FF9C4740FE9CFEC8FE5CFE31" bold="true" box="[264,409,382,408]" pageId="3" pageNumber="4">Description</emphasis>
.­General coloration: Body brown with yellow stripe laterad of scutum and vertical stripe from anepisternum to katepisternum. Body length:
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; wing length:
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.
</paragraph>
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<paragraph id="6C2D2470FF9C4740FE9CF895FB02F8F4" blockId="3.[264,1322,1827,1885]" pageId="3" pageNumber="4">
<emphasis id="5EE6F862FF9C4740FE9CF895FE77F892" bold="true" box="[264,434,1827,1851]" pageId="3" pageNumber="4">FIGURES 2­5.</emphasis>
<taxonomicName id="AB925FF3FF9C4740FE29F892FD4FF892" box="[445,650,1828,1851]" class="Insecta" family="Sciomyzidae" genus="Thecomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="mathisi">
<emphasis id="5EE6F862FF9C4740FE29F892FD4FF892" box="[445,650,1828,1851]" italics="true" pageId="3" pageNumber="4">Thecomyia mathisi</emphasis>
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: 2, head, anterolateral view; 3, head anterodorsal view.
<taxonomicName id="AB925FF3FF9C4740FB6DF892FD07F8F4" authority="Marinoni and Steyskal" authorityName="Marinoni and Steyskal" class="Insecta" family="Sciomyzidae" genus="Thecomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="chrysacra">
<emphasis id="5EE6F862FF9C4740FB6DF892FE02F8F4" italics="true" pageId="3" pageNumber="4">Thecomyia chrysacra</emphasis>
Marinoni and Steyskal
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: 4, thorax, dorsal view; 5, abdomen,dorsal view.
</paragraph>
</caption>
<paragraph id="6C2D2470FF9B4747FEACFEADFE08FDE7" blockId="4.[264,1324,283,1875]" pageId="4" pageNumber="5">
<emphasis id="5EE6F862FF9B4747FEACFEADFEBEFE9C" bold="true" box="[312,379,283,309]" pageId="4" pageNumber="5">Head</emphasis>
(
<figureCitation id="F4A938F5FF9B4747FE18FEAAFDEEFE9F" box="[396,555,284,310]" captionStart="FIGURES 2 ­ 5" captionStartId="3.[264,383,1827,1851]" captionTargetBox="[278,1318,532,1805]" captionTargetId="figure@3.[269,1319,532,1808]" captionTargetPageId="3" captionText="FIGURES 2 ­ 5. Thecomyia mathisi: 2, head, anterolateral view; 3, head anterodorsal view. Thecomyia chrysacra Marinoni and Steyskal: 4, thorax, dorsal view; 5, abdomen, dorsal view." httpUri="https://zenodo.org/record/156858/files/figure.png" pageId="4" pageNumber="5">Figs. 2 and 3</figureCitation>
): Ventral portion of head drawn into a conical rostrum; width of gena as much as 0.76 height of eye, reaching
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some females. Palpi reduced, at most represented by a few setae. Eyes very prominent. Head sutures indistinct, ptilinal fissure absent, thus area of lunule greatly exposed, fused with anterior margin of frons. Midfrontal vitta absent. No midfrontal or facial setulae. Fronto­orbital spot present; 0­1 (weak to strong) upper orbital seta; ocellar and postocellar setae lacking; inner and outer vertical setae well developed. Pedicel equal in length to first flagellomere; arista sub­basal, bearing long white hairs.
</paragraph>
<paragraph id="6C2D2470FF9B4747FEACFDEDFD75FAE2" blockId="4.[264,1324,283,1875]" pageId="4" pageNumber="5">
<emphasis id="5EE6F862FF9B4747FEACFDEDFE50FDDC" bold="true" box="[312,405,603,629]" pageId="4" pageNumber="5">Thorax</emphasis>
(
<figureCitation id="F4A938F5FF9B4747FE30FDEAFE28FDDF" box="[420,493,604,630]" captionStart="FIGURES 2 ­ 5" captionStartId="3.[264,383,1827,1851]" captionTargetBox="[278,1318,532,1805]" captionTargetId="figure@3.[269,1319,532,1808]" captionTargetPageId="3" captionText="FIGURES 2 ­ 5. Thecomyia mathisi: 2, head, anterolateral view; 3, head anterodorsal view. Thecomyia chrysacra Marinoni and Steyskal: 4, thorax, dorsal view; 5, abdomen, dorsal view." httpUri="https://zenodo.org/record/156858/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
). Medial mesonotonal stripe simple or tripartite. 0 prosternal, pro­episternal, postpronotal, presutural, subalar, anepisternal, katatergal, katepisternal or anepimeral setae. 1­2 notopleural setae, anterior seta very small and weak; 1 supra­alar; 1 postalar; 1 weak or 0 prescutellar dorsocentral, 1 pair of scutellar. No setae around metathoracic spiracle. Pleura bare. Mesonotal suture complete. Area above hind coxae completely sclerotized. Wing infumate, not patterned; veins R4+5 and M1 not convergent apically, crossvein dm­cu straight; A1+CuA2 complete to margin; anal cell truncate; anal lobe not expanded. Fore coxa bearing a few short antero­apical setae; mid coxa with 1 well developed mid lateral seta and a few short antero­apical setae; hind coxa with posterior inner margin bare, 1 strong posterolateral seta and a few short apical setae; fore and mid trochanters with 1 moderately strong anterodorsal seta; hind trochanter densely covered with short, strong setulae ventrally; mid and hind femora with 4­8 short, strong, spinelike setae ventrally near apex; mid femur with or without small, anteromedial seta; hind femur with or without anteromedial seta; hind femur ventrally with 7­13 subapical, spinelike setae forming lines anteriorly and posteriorly; tibiae with 1 short, dorsal pre­apical seta, that of fore tibia strongest, and 3­4 short, strong apical setae posteriorly; hind tibia with 4­5 pairs of slightly stronger spinelike setae ventrally on each side at apical 1/3; hind tibia straight not arched; fore and hind tibiae on inner apical margin with a pecten (series of many fine, equally sized, short, closely spaced setulae).
</paragraph>
<paragraph id="6C2D2470FF9B4746FEACFAEEFC40FDDF" blockId="4.[264,1324,283,1875]" lastBlockId="5.[264,1324,284,750]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<emphasis id="5EE6F862FF9B4747FEACFAEEFE30FADB" bold="true" box="[312,501,1368,1394]" pageId="4" pageNumber="5">Male abdomen</emphasis>
(
<figureCitation id="F4A938F5FF9B4747FD9DFAEFFD43FADA" box="[521,646,1369,1395]" captionStart-0="FIGURES 6 ­ 8" captionStart-1="FIGURES 9 ­ 11" captionStart-2="FIGURES 12 ­ 14" captionStart-3="FIGURES 15 ­ 17" captionStart-4="FIGURES 18 ­ 23" captionStart-5="FIGURES 24 ­ 29" captionStartId-0="5.[264,383,1718,1742]" captionStartId-1="8.[264,382,1834,1858]" captionStartId-2="9.[264,383,1832,1856]" captionStartId-3="10.[264,383,1836,1860]" captionStartId-4="11.[264,383,1664,1688]" captionStartId-5="12.[264,382,1658,1682]" captionTargetBox-1="[370,1219,320,1823]" captionTargetBox-2="[357,1230,324,1818]" captionTargetBox-3="[386,1202,336,1821]" captionTargetBox-4="[276,1315,322,1585]" captionTargetBox-5="[275,1317,352,1566]" captionTargetId-1="figure@8.[363,1224,314,1823]" captionTargetId-2="figure@9.[351,1235,317,1822]" captionTargetId-3="figure@10.[379,1208,329,1825]" captionTargetId-4="figure@11.[267,1319,319,1611]" captionTargetId-5="figure@12.[267,1319,329,1605]" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionTargetPageId-4="11" captionTargetPageId-5="12" captionText-0="FIGURES 6 ­ 8. Male terminalia, left lateral view: 6, Thecomyia longicornis Perty; 7, T. lateralis (Walker); 8, T. limbata (Wiedemann). App, anterior phallapodeme process; am, anterior margin; as, anterior surstylus; bp, basiphallic process; c, cercus; dis, distiphallus; epa, epandrium; ep, epiphallus; h, hypandrium; p, phallapodeme; pm, posterior margin; ps, posterior surstylus; synst 7 + 8, synsternite 7 + 8; st 6, sternite 6." captionText-1="FIGURES 9 ­ 11. Male terminalia, left lateral view: 9, Thecomyia abercrombiei Marinoni and Steyskal; 10, T. autazensis Marinoni and Styeskal; 11, T. bonattoi Marinoni and Steyskal." captionText-2="FIGURES 12 ­ 14. Male terminalia, left lateral view: 12, Thecomyia chrysacra Marinoni and Steyskal; 13, T. mathisi Marinoni; 14, T. naponica Marinoni and Steyskal." captionText-3="FIGURES 15 ­ 17. Male terminalia, left lateral view: 15, Thecomyia papaveroi Marinoni and Steyskal; 16, T. signorelli Marinoni and Steyskal; 17, T. tricuneata Marinoni and Steyskal." captionText-4="FIGURES 18 ­ 23. Hypandrium, frontal view and right lateral view: 18, Thecomyia longicornis Perty; 19, T. lateralis (Walker); 20, T. limbata (Wiedemann); 21, T. abercrombiei Marinoni and Steyskal; 22, T. autazensis Marinoni and Steyskal; 23, T. bonattoi Marinoni and Steyskal. A, anterior; as, archlike struts; avp, anteroventral plates; d, dorsal; mp, membranous projection; v, ventral." captionText-5="FIGURES 24 ­ 29. Hypandrium, frontal view and right lateral view: 24, Thecomyia chrysacra Marinoni and Steyskal; 25, T. mathisi Marinoni; 26, T. naponica Marinoni and Steyskal; 27, T. papaveroi Marinoni and Steyskal; 28, T. signorelli Marinoni and Steyskal; 29, T. tricuneata Marinoni and Steyskal." httpUri-1="https://zenodo.org/record/156859/files/figure.png" httpUri-2="https://zenodo.org/record/156860/files/figure.png" httpUri-3="https://zenodo.org/record/156861/files/figure.png" httpUri-4="https://zenodo.org/record/156862/files/figure.png" httpUri-5="https://zenodo.org/record/156863/files/figure.png" pageId="4" pageNumber="5">Figs. 6­29</figureCitation>
). Sternite 6 asymmetrical, normally extended from mid dorsum onto left side, where it is expanded and strongly sclerotized; left spiracle 6 mid laterally at beginning of expanded portion; left spiracle
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sclerotized strip lying in membrane below synsternite 7+8. Synsternite 7+8 1/2­2/3 size of epandrium in lateral view. Epandrium in posterior view nearly complete, narrowly membranous below cerci; cercus small, with apico­ventral portion wide and well sclerotized. Ejaculatory apodeme wider basally than phallapodeme, with muscle attachments at base and with short, triangular piece articulated by membrane with stem. Phallapodeme straight to slightly curved distally, with process or spur on anterior surface subapically in some species. Epiphallus covering base of distiphallus. Distiphallus complex, asymmetrical, with acrophallus and associated plates either elongate and curved apically, moderately long and curved, or short with variously shaped plates or processes. Hypandrium with pair of weak, linear struts posteriorly, medial portion very broadly expanded into pair of large, ventrally projecting plates that vary in shape among the species. Anteroventral hypandrial plates projecting below surstylus in some species; a pair of narrow, archlike struts on posterior border of plates, meeting mid dorsally at apex of distiphallus. Hypandrium with margins of anteroventral plates bearing membranous projection varying in shape and size among species; bearing some sort of cilia. Gonopod fused to hypandrium. Aedeagal complex asymmetrical; anterior basiphallic process curved or cuneate and tripartite apically. Posterior and anterior surstyli articulated with well sclerotized bacilliform sclerite, which in turn articulates with epandrium and hypandrium; posterior and anterior surstyli distinct and widely separated or only the posterior well developed.
</paragraph>
<paragraph id="6C2D2470FF9A4746FEACFD35FD23FD47" blockId="5.[264,1324,284,750]" pageId="5" pageNumber="6">
<emphasis id="5EE6F862FF9A4746FEACFD35FE56FD34" bold="true" box="[312,403,643,669]" pageId="5" pageNumber="6">Female</emphasis>
(Figs. 30­40). Sternites 6, 7 and 8 almost fused. With 2 spermatheca; spermathecal duct very wide and well sclerotized at junction with spermatheca; in some species with an apical digitiform projection.
</paragraph>
</subSubSection>
</treatment>
</document>