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<document id="B151459AE531467B66C94104B9D85DD8" ID-DOI="10.5281/zenodo.156857" ID-GBIF-Dataset="bc8fbb8b-8101-4fe5-9ffd-0b5aa5db803c" ID-ISSN="1175­5326" ID-Zenodo-Dep="156857" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459605459478" checkinUser="plazi" docAuthor="Marinoni, Luciane, Steyskal, George C. &amp; Knutson, Lloyd" docDate="2003" docId="E43B9566FF894754FE9CF8B7FEA1FA6F" docLanguage="en" docName="zt00191.pdf" docOrigin="Zootaxa 191" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Thecomyia mathisi Marinoni, new species" docType="treatment" docVersion="10" lastPageNumber="24" masterDocId="1802ED1EFF9F4743FF94FFB6FFC5FFA9" masterDocTitle="Revision and cladistic analysis of the Neotropical Genus Thecomyia Perty (Diptera: Sciomyzidae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="23" updateTime="1698202859598" updateUser="plazi">
<mods:mods id="9B0CD4160A60235EE70470A4435EA0DB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="02EDF99DAE75D764E6C4B833EDD64CCC">Revision and cladistic analysis of the Neotropical Genus Thecomyia Perty (Diptera: Sciomyzidae)</mods:title>
</mods:titleInfo>
<mods:name id="B6591AB1DC78162798F650EB8C29AF20" type="personal">
<mods:role id="03B4BA7EF93EF07D4E2940FCE9BCDDEF">
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<mods:namePart id="D36D6CC4F7F1712D5DEF714DA53D928B">Marinoni, Luciane</mods:namePart>
</mods:name>
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<mods:namePart id="9EB7386AFA8EBD5B3FAC6193659589E0">Steyskal, George C.</mods:namePart>
</mods:name>
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<mods:namePart id="69B7143B43D3B372964CC0326E745431">Knutson, Lloyd</mods:namePart>
</mods:name>
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<mods:title id="7AC99CFB2371EB3B2B129F491E7DD1B5">Zootaxa</mods:title>
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<mods:part id="20E3ADEBA2D7A5C8BA4846F63C289337">
<mods:date id="B0116FEC7EB2224463EF88D0B214C999">2003</mods:date>
<mods:detail id="42D904D57B7AC2DEEF7AC0DA4DC65945" type="volume">
<mods:number id="6BD465A11BED51235FE6BC1B84A2F9F5">191</mods:number>
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<mods:start id="9BE9C3EA8FC13971107B6C80FD6976C0">1</mods:start>
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<mods:classification id="5166F965F047B39D6E1550405E739D53">journal article</mods:classification>
<mods:identifier id="036512FF5216E6699BC5E6D8CDFFDABB" type="DOI">10.5281/zenodo.156857</mods:identifier>
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<mods:identifier id="57782D6C1316D176EFBAF438496B4A0C" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="76C94ED0CEA9A9878A0763D903670F84" type="Zenodo-Dep">156857</mods:identifier>
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<treatment id="E43B9566FF894754FE9CF8B7FEA1FA6F" ID-DOI="http://doi.org/10.5281/zenodo.6273664" ID-GBIF-Taxon="119331432" ID-Zenodo-Dep="6273664" LSID="urn:lsid:plazi:treatment:E43B9566FF894754FE9CF8B7FEA1FA6F" httpUri="http://treatment.plazi.org/id/E43B9566FF894754FE9CF8B7FEA1FA6F" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
<subSubSection id="248877FBFF894755FE9CF8B7FB6EF8B2" box="[264,1195,1793,1819]" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="6C2D2470FF894755FE9CF8B7FB6EF8B2" blockId="22.[264,1195,1793,1819]" box="[264,1195,1793,1819]" pageId="22" pageNumber="23">
<heading id="3765931CFF894755FE9CF8B7FB6EF8B2" bold="true" box="[264,1195,1793,1819]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<emphasis id="5EE6F862FF894755FE9CF8B7FB6EF8B2" bold="true" box="[264,1195,1793,1819]" pageId="22" pageNumber="23">
8.
<taxonomicName id="AB925FF3FF894755FEB2F8B7FD49F8B2" ID-CoL="5674F" authority="Marinoni" authorityName="Marinoni" box="[294,652,1793,1819]" class="Insecta" family="Sciomyzidae" genus="Thecomyia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="mathisi" status="sp. nov.">
<emphasis id="5EE6F862FF894755FEB2F8B7FDC9F8B2" bold="true" box="[294,524,1793,1819]" italics="true" pageId="22" pageNumber="23">Thecomyia mathisi</emphasis>
Marinoni
</taxonomicName>
,
<taxonomicNameLabel id="45D54519FF894755FD0DF8B7FCECF8B2" box="[665,809,1793,1819]" pageId="22" pageNumber="23" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="F4A938F5FF894755FCACF8B7FC49F8B2" box="[824,908,1793,1819]" captionStart="FIGURE 1" captionStartId="2.[264,368,1607,1631]" captionText="FIGURE 1. Thecomyia mathisi Marinoni ­ male." pageId="22" pageNumber="23">Figs. 1</figureCitation>
,
<figureCitation id="F4A938F5FF894755FC0EF8B7FC02F8B2" box="[922,967,1793,1819]" captionStart="FIGURES 2 ­ 5" captionStartId="3.[264,383,1827,1851]" captionTargetBox="[278,1318,532,1805]" captionTargetId="figure@3.[269,1319,532,1808]" captionTargetPageId="3" captionText="FIGURES 2 ­ 5. Thecomyia mathisi: 2, head, anterolateral view; 3, head anterodorsal view. Thecomyia chrysacra Marinoni and Steyskal: 4, thorax, dorsal view; 5, abdomen, dorsal view." httpUri="https://zenodo.org/record/156858/files/figure.png" pageId="22" pageNumber="23">2, 3</figureCitation>
,
<figureCitation id="F4A938F5FF894755FC41F8B7FC36F8B2" box="[981,1011,1793,1819]" captionStart="FIGURES 12 ­ 14" captionStartId="9.[264,383,1832,1856]" captionTargetBox="[357,1230,324,1818]" captionTargetId="figure@9.[351,1235,317,1822]" captionTargetPageId="9" captionText="FIGURES 12 ­ 14. Male terminalia, left lateral view: 12, Thecomyia chrysacra Marinoni and Steyskal; 13, T. mathisi Marinoni; 14, T. naponica Marinoni and Steyskal." httpUri="https://zenodo.org/record/156860/files/figure.png" pageId="22" pageNumber="23">13</figureCitation>
,
<figureCitation id="F4A938F5FF894755FB95F8B7FBE5F8B2" box="[1025,1056,1793,1819]" captionStart="FIGURES 24 ­ 29" captionStartId="12.[264,382,1658,1682]" captionTargetBox="[275,1317,352,1566]" captionTargetId="figure@12.[267,1319,329,1605]" captionTargetPageId="12" captionText="FIGURES 24 ­ 29. Hypandrium, frontal view and right lateral view: 24, Thecomyia chrysacra Marinoni and Steyskal; 25, T. mathisi Marinoni; 26, T. naponica Marinoni and Steyskal; 27, T. papaveroi Marinoni and Steyskal; 28, T. signorelli Marinoni and Steyskal; 29, T. tricuneata Marinoni and Steyskal." httpUri="https://zenodo.org/record/156863/files/figure.png" pageId="22" pageNumber="23">25</figureCitation>
, 37, 41, 44)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="248877FBFF894754FE9CF8FDFB4CFC1F" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="description">
<paragraph id="6C2D2470FF894754FE9CF8FDFB4CFC1F" blockId="22.[264,1321,1867,1894]" lastBlockId="23.[264,1324,284,1478]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
<emphasis id="5EE6F862FF894755FE9CF8FDFE35F8CC" bold="true" box="[264,496,1867,1893]" pageId="22" pageNumber="23">Description. Adult</emphasis>
. Upper orbital seta absent. Fronto­orbital spot rounded, width approximately 1/3 frontal width. Rostrum
<quantity id="AB6A8995FF884754FD3AFEAAFCF5FE9F" box="[686,816,284,310]" metricMagnitude="-2" metricUnit="m" metricValue="1.5748" metricValueMax="1.6256" metricValueMin="1.524" pageId="23" pageNumber="24" unit="in" value="0.62" valueMax="0.64" valueMin="0.6">0.6­0.64 in</quantity>
males and
<quantity id="AB6A8995FF884754FC2CFEAAFB8CFE9F" box="[952,1097,284,310]" metricMagnitude="-2" metricUnit="m" metricValue="1.6383" metricValueMax="1.6764" metricValueMin="1.6002" pageId="23" pageNumber="24" unit="in" value="0.645" valueMax="0.66" valueMin="0.63">0.63­0.66 in</quantity>
females as high as eye. White microtomentose infra­ocular marking forming a triangle with margin between gena and postgena emarginate. Mid femur without anteromedial seta. Mid and hind femora wholly yellowish. Mesonotum with medial, gray, tripartite stripe, scutellum with broad medial yellow stripe. Ejaculatory apodeme wide apically, anterior and posterior margins slightly concave (
<figureCitation id="F4A938F5FF884754FE4BFE52FDFFFE57" box="[479,570,484,510]" captionStart="FIGURES 12 ­ 14" captionStartId="9.[264,383,1832,1856]" captionTargetBox="[357,1230,324,1818]" captionTargetId="figure@9.[351,1235,317,1822]" captionTargetPageId="9" captionText="FIGURES 12 ­ 14. Male terminalia, left lateral view: 12, Thecomyia chrysacra Marinoni and Steyskal; 13, T. mathisi Marinoni; 14, T. naponica Marinoni and Steyskal." httpUri="https://zenodo.org/record/156860/files/figure.png" pageId="23" pageNumber="24">Fig. 13</figureCitation>
). Phallapodeme apex broad. Anterior basiphallic process well developed, concave, anterior part bicuspid. Epiphallus well developed, extending to base of distiphallus. Distiphallus with 3 processes, the medial slender, projecting downwards; large membranous projection under distiphallus. Hypandrium in lateral view with dorsal margin slightly concave, anteroventral margin narrow, not rounded (
<figureCitation id="F4A938F5FF884754FBA2FD32FB4AFD37" box="[1078,1167,644,670]" captionStart="FIGURES 24 ­ 29" captionStartId="12.[264,382,1658,1682]" captionTargetBox="[275,1317,352,1566]" captionTargetId="figure@12.[267,1319,329,1605]" captionTargetPageId="12" captionText="FIGURES 24 ­ 29. Hypandrium, frontal view and right lateral view: 24, Thecomyia chrysacra Marinoni and Steyskal; 25, T. mathisi Marinoni; 26, T. naponica Marinoni and Steyskal; 27, T. papaveroi Marinoni and Steyskal; 28, T. signorelli Marinoni and Steyskal; 29, T. tricuneata Marinoni and Steyskal." httpUri="https://zenodo.org/record/156863/files/figure.png" pageId="23" pageNumber="24">Fig. 25</figureCitation>
). Anteroventral hypandrial plates in frontal view narrow, tapered laterally. Membranous projections of margins of anteroventral hypandrial plates broad but not extending to lateral margins of hypandrium, covered with minute hairs (
<figureCitation id="F4A938F5FF884754FD7DFD4AFC84FCBF" box="[745,833,764,790]" captionStart="FIGURES 24 ­ 29" captionStartId="12.[264,382,1658,1682]" captionTargetBox="[275,1317,352,1566]" captionTargetId="figure@12.[267,1319,329,1605]" captionTargetPageId="12" captionText="FIGURES 24 ­ 29. Hypandrium, frontal view and right lateral view: 24, Thecomyia chrysacra Marinoni and Steyskal; 25, T. mathisi Marinoni; 26, T. naponica Marinoni and Steyskal; 27, T. papaveroi Marinoni and Steyskal; 28, T. signorelli Marinoni and Steyskal; 29, T. tricuneata Marinoni and Steyskal." httpUri="https://zenodo.org/record/156863/files/figure.png" pageId="23" pageNumber="24">Fig. 25</figureCitation>
). Anterior surstylus absent, posterior barlike (
<figureCitation id="F4A938F5FF884754FED0FC92FE5EFC97" box="[324,411,804,830]" captionStart="FIGURES 12 ­ 14" captionStartId="9.[264,383,1832,1856]" captionTargetBox="[357,1230,324,1818]" captionTargetId="figure@9.[351,1235,317,1822]" captionTargetPageId="9" captionText="FIGURES 12 ­ 14. Male terminalia, left lateral view: 12, Thecomyia chrysacra Marinoni and Steyskal; 13, T. mathisi Marinoni; 14, T. naponica Marinoni and Steyskal." httpUri="https://zenodo.org/record/156860/files/figure.png" pageId="23" pageNumber="24">Fig. 13</figureCitation>
). Female sternites 6, 7 and 8 almost fused, presence of sternite 6 indicated by line of hairs, longer than wide; anterior margin of sternite 6 very slightly sinuate; mid longitudinal portion less sclerotized. Spermathecal duct with sclerotized region joining spermatheca long; apical, digitiform projection broad at base and short (Fig. 37).
</paragraph>
</subSubSection>
<subSubSection id="248877FBFF884754FEACFC75FC45FB07" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="6C2D2470FF884754FEACFC75FD26FBF7" blockId="23.[264,1324,284,1478]" pageId="23" pageNumber="24">
<emphasis id="5EE6F862FF884754FEACFC75FDC7FC74" bold="true" box="[312,514,963,989]" pageId="23" pageNumber="24">
<typeStatus id="B3299AD2FF884754FEACFC75FEB0FC74" box="[312,373,963,989]" pageId="23" pageNumber="24">Type</typeStatus>
specimens
</emphasis>
.
<typeStatus id="B3299AD2FF884754FDB7FC72FD7BFC77" box="[547,702,964,990]" pageId="23" pageNumber="24" type="holotype">HOLOTYPE</typeStatus>
MALE. 1. White rectangular label: [
<collectingCountry id="148564E0FF884754FB04FC72FAE1FC77" box="[1168,1316,964,990]" name="Ecuador" pageId="23" pageNumber="24">ECUADOR</collectingCountry>
] &quot;
<collectingCountry id="148564E0FF884754FE85FC42FEBDFBA7" box="[273,376,1012,1038]" name="Ecuador" pageId="23" pageNumber="24">Ecuador</collectingCountry>
. Prt. Orllna: Rio Tiputini (0O 38. 2'S, 76O 8. 9'W)
<date id="182C02B0FF884754FC55FC42FB46FBA7" box="[961,1155,1012,1038]" pageId="23" pageNumber="24" value="1999-08-12" valueMax="1999-08-26" valueMin="1999-08-12">12­26 Aug 1999</date>
, W.N. Mathis, A. Baptista, M. Kotrba&quot; (
<collectionCode id="0A83BCB5FF884754FDADFBAAFD51FB9F" LSID="urn:lsid:biocol.org:col:34871" box="[569,660,1052,1078]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="23" pageNumber="24">USNM</collectionCode>
).
<typeStatus id="B3299AD2FF884754FD33FBAAFC88FB9F" box="[679,845,1052,1078]" pageId="23" pageNumber="24" type="paratype">PARATYPES</typeStatus>
. The same label as on the
<typeStatus id="B3299AD2FF884754FB15FBAAFB2FFB9F" box="[1153,1258,1052,1078]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
,
<specimenCount id="7A94EFF9FF884754FB63FBAAFAEEFB9F" box="[1271,1323,1052,1078]" pageId="23" pageNumber="24" type="generic">
<quantity id="AB6A8995FF884754FB63FBAAFAE2FB9F" box="[1271,1319,1052,1078]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="23" pageNumber="24" unit="m" value="8.0">8 m</quantity>
.
</specimenCount>
and
<specimenCount id="7A94EFF9FF884754FEAEFBF2FEA4FBF7" box="[314,353,1092,1118]" pageId="23" pageNumber="24" type="generic">2 f.</specimenCount>
(
<collectionCode id="0A83BCB5FF884754FEE4FBF2FE0EFBF7" LSID="urn:lsid:biocol.org:col:34871" box="[368,459,1092,1118]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="23" pageNumber="24">USNM</collectionCode>
);
<specimenCount id="7A94EFF9FF884754FE4BFBF2FDD1FBF7" box="[479,532,1092,1118]" pageId="23" pageNumber="24" type="generic">
<quantity id="AB6A8995FF884754FE4BFBF2FDD5FBF7" box="[479,528,1092,1118]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="23" pageNumber="24" unit="m" value="5.0">5 m</quantity>
.
</specimenCount>
and
<specimenCount id="7A94EFF9FF884754FDD9FBF2FDB1FBF7" box="[589,628,1092,1118]" pageId="23" pageNumber="24" type="generic">2 f.</specimenCount>
(
<collectionCode id="0A83BCB5FF884754FD17FBF2FD13FBF7" box="[643,726,1092,1118]" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="23" pageNumber="24">DZUP</collectionCode>
).
</paragraph>
<paragraph id="6C2D2470FF884754FEACFBDDFC45FB07" blockId="23.[264,1324,284,1478]" pageId="23" pageNumber="24">
<emphasis id="5EE6F862FF884754FEACFBDDFD61FB2C" bold="true" box="[312,676,1131,1157]" pageId="23" pageNumber="24">Other specimens examined:</emphasis>
<collectingCountry id="148564E0FF884754FD20FBDAFCF5FB2F" box="[692,816,1132,1158]" name="Bolivia" pageId="23" pageNumber="24">BOLIVIA</collectingCountry>
. Provincia Andres Ibanez Potrerillo del Guenda,
<date id="182C02B0FF884754FEE6FB22FDC8FB07" box="[370,525,1172,1198]" pageId="23" pageNumber="24" value="1993-11-12">12 Nov 1993</date>
, P. Bettella col.,
<specimenCount id="7A94EFF9FF884754FD45FB22FCC3FB07" box="[721,774,1172,1198]" pageId="23" pageNumber="24" type="generic">
<quantity id="AB6A8995FF884754FD45FB22FCC7FB07" box="[721,770,1172,1198]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="24" unit="m" value="1.0">1 m</quantity>
.
</specimenCount>
(
<collectionCode id="0A83BCB5FF884754FC81FB22FCB1FB07" box="[789,884,1172,1198]" pageId="23" pageNumber="24">BMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="248877FBFF884754FEACFB0DFC62FA8F" pageId="23" pageNumber="24" type="etymology">
<paragraph id="6C2D2470FF884754FEACFB0DFC62FA8F" blockId="23.[264,1324,284,1478]" pageId="23" pageNumber="24">
<emphasis id="5EE6F862FF884754FEACFB0DFE78FB7C" bold="true" box="[312,445,1211,1237]" pageId="23" pageNumber="24">Etymology</emphasis>
.­ The species epithet,
<emphasis id="5EE6F862FF884754FD50FB0AFCE4FB7C" box="[708,801,1212,1237]" italics="true" pageId="23" pageNumber="24">
<taxonomicName id="AB925FF3FF884754FD50FB0AFCD8FB7C" box="[708,797,1212,1237]" class="Insecta" family="Sciomyzidae" genus="Thecomyia" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="mathisi">mathisi</taxonomicName>
,
</emphasis>
is a Latin genitive patronym to the collector of the
<typeStatus id="B3299AD2FF884754FEC2FB52FE4DFB57" box="[342,392,1252,1278]" pageId="23" pageNumber="24">type</typeStatus>
species and to recognize the numerous and varied contributions of Dr. Wayne N. Mathis to the study of
<taxonomicName id="AB925FF3FF884754FDA1FABAFD55FA8F" box="[565,656,1292,1318]" kingdom="Plantae" order="Diptera" pageId="23" pageNumber="24" rank="order">Diptera</taxonomicName>
, especially
<taxonomicName id="AB925FF3FF884754FC83FABAFC67FA8F" box="[791,930,1292,1318]" class="Insecta" family="Ephydridae" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="family">Ephydridae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="248877FBFF884754FEACFA85FEA1FA6F" pageId="23" pageNumber="24" type="discussion">
<paragraph id="6C2D2470FF884754FEACFA85FEA1FA6F" blockId="23.[264,1324,284,1478]" pageId="23" pageNumber="24">
<emphasis id="5EE6F862FF884754FEACFA85FE74FAE4" bold="true" box="[312,433,1331,1357]" pageId="23" pageNumber="24">Remarks.</emphasis>
One male specimen from the BMNH, collected in
<collectingCountry id="148564E0FF884754FB9EFA82FBA4FAE7" box="[1034,1121,1332,1358]" name="Bolivia" pageId="23" pageNumber="24">Bolivia</collectingCountry>
has all the external characters and the genitalia very similar to
<taxonomicName id="AB925FF3FF884754FCBDFAEAFC5EFADC" box="[809,923,1372,1397]" class="Insecta" family="Sciomyzidae" genus="Thecomyia" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="mathisi">
<emphasis id="5EE6F862FF884754FCBDFAEAFC5EFADC" box="[809,923,1372,1397]" italics="true" pageId="23" pageNumber="24">T. mathisi</emphasis>
</taxonomicName>
with some variation in the genitalia. It would be necessary to study more specimens to confirm if it is or not another species.
</paragraph>
</subSubSection>
</treatment>
</document>
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