treatments-xml/data/E4/34/B3/E434B32BFFA47B66FF0EC1D53CF4FEA2.xml
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<document id="B62EC3DD4CCBDA9EE30AC54AA6659D91" ID-CLB-Dataset="31850" ID-DOI="10.11646/zootaxa.4329.5.5" ID-GBIF-Dataset="65bb59f4-d3e7-43aa-9116-23ba8f97ecc1" ID-ISSN="1175-5326" ID-Zenodo-Dep="1010089" ID-ZooBank="C10Ad793-0785-42F4-95E3-214F160A8024" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507880565361" checkinUser="plazi" docAuthor="Chan, Tin-Yam" docDate="2017" docId="E434B32BFFA47B66FF0EC1D53CF4FEA2" docLanguage="en" docName="zootaxa.4329.5.5.pdf" docOrigin="Zootaxa 4329 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Petrarctus jeppiaari Chan 2017, sp. nov." docType="treatment" docVersion="8" lastPageNumber="484" masterDocId="180DCB53FFA57B6EFF99C5163878FFC7" masterDocTitle="A new slipper lobster of the genus Petrarctus Holthuis, 2002 (Crustacea, Decapoda, Scyllaridae) from Southwest coast of India" masterLastPageNumber="486" masterPageNumber="477" pageNumber="478" updateTime="1698469498250" updateUser="ExternalLinkService">
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<mods:title id="393C4DA9A430D6565B66CCF4B4793728">A new slipper lobster of the genus Petrarctus Holthuis, 2002 (Crustacea, Decapoda, Scyllaridae) from Southwest coast of India</mods:title>
</mods:titleInfo>
<mods:name id="EE2CB4B2B769615A3381D0C5431F5A60" type="personal">
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<mods:roleTerm id="9971671493398D00FCDB8119BB8BF52E">Author</mods:roleTerm>
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<mods:namePart id="3B7096DFCA64DC80600491B049190DB7">Chan, Tin-Yam</mods:namePart>
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<mods:typeOfResource id="787E14104DED6E24B41229DA4D214744">text</mods:typeOfResource>
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<mods:title id="046DA4C9704B0C43930B585160C4F1E0">Zootaxa</mods:title>
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<mods:part id="30CA0CFB81D20E297C12C6AB210CC4BD">
<mods:date id="F14134AEA866719FD0756D090F0EC1F3">2017</mods:date>
<mods:detail id="B98C06EAE87F196B6B30BC796D434A51" type="pubDate">
<mods:number id="ADEAC92D4E478CA74657DE72E897886D">2017-10-09</mods:number>
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<mods:number id="BFF5E36690C34204442DA29293FABA76">4329</mods:number>
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<mods:classification id="ED2C91D087847B3B512A0061090AB339">journal article</mods:classification>
<mods:identifier id="1698F9A1AC2035E968C58F53C82E028D" type="CLB-Dataset">31850</mods:identifier>
<mods:identifier id="94BEF006D00FE1A25A24174DA61E3003" type="DOI">10.11646/zootaxa.4329.5.5</mods:identifier>
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<mods:identifier id="CE5EC04C801842D6C7D2E1688B24FAD6" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="53FC6B6EDD4413C2CA12A612B1D5AA46" type="Zenodo-Dep">1010089</mods:identifier>
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</mods:mods>
<treatment id="E434B32BFFA47B66FF0EC1D53CF4FEA2" ID-DOI="http://doi.org/10.5281/zenodo.6045763" ID-GBIF-Taxon="135435346" ID-Zenodo-Dep="6045763" LSID="urn:lsid:plazi:treatment:E434B32BFFA47B66FF0EC1D53CF4FEA2" httpUri="http://treatment.plazi.org/id/E434B32BFFA47B66FF0EC1D53CF4FEA2" lastPageId="8" lastPageNumber="484" pageId="1" pageNumber="478">
<subSubSection id="248751B6FFA47B6FFF0EC1D5398CFB38" pageId="1" pageNumber="478" type="nomenclature">
<paragraph id="6C22023DFFA47B6FFF0EC1D53994FB19" blockId="1.[151,500,1219,1279]" box="[151,492,1219,1246]" pageId="1" pageNumber="478">
<heading id="376AB551FFA47B6FFF0EC1D53994FB19" bold="true" box="[151,492,1219,1246]" fontSize="11" level="1" pageId="1" pageNumber="478" reason="1">
<emphasis id="5EE9DE2FFFA47B6FFF0EC1D53994FB19" bold="true" box="[151,492,1219,1246]" pageId="1" pageNumber="478">
<taxonomicName id="AB9D79BEFFA47B6FFF0EC1D539FFFB1A" ID-CoL="85WWV" authority="Chan, 2017" authorityName="Chan" authorityYear="2017" box="[151,391,1219,1245]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="478" phylum="Arthropoda" rank="species" species="jeppiaari" status="sp. nov.">
<emphasis id="5EE9DE2FFFA47B6FFF0EC1D539FFFB1A" bold="true" box="[151,391,1219,1245]" italics="true" pageId="1" pageNumber="478">Petrarctus jeppiaari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45DA6354FFA47B6FFE17C1D23994FB19" box="[398,492,1220,1246]" pageId="1" pageNumber="478" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6C22023DFFA47B6FFF0EC1F0398CFB38" blockId="1.[151,500,1219,1279]" box="[151,500,1254,1279]" pageId="1" pageNumber="478">
(
<figureCitation id="F4A61EB8FFA47B6FFF06C1F03895FB38" box="[159,237,1254,1279]" captionStart="FIGURE 1" captionStartId="3.[151,250,1968,1990]" captionTargetBox="[252,1336,195,1951]" captionTargetId="figure@3.[252,1336,195,1951]" captionTargetPageId="3" captionText="FIGURE 1. Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV)." httpUri="https://zenodo.org/record/1010091/files/figure.png" pageId="1" pageNumber="478">Figs. 1</figureCitation>
; 2; 3A, C, E; 4A, C, E)
</paragraph>
</subSubSection>
<subSubSection id="248751B6FFA47B6FFF0EC038390AFA3C" pageId="1" pageNumber="478" type="materials_examined">
<paragraph id="6C22023DFFA47B6FFF0EC038390AFA3C" blockId="1.[151,1437,1326,2036]" pageId="1" pageNumber="478">
<materialsCitation id="DCF50860FFA47B6FFF0EC0383A8FFAAC" ID-GBIF-Occurrence="1671716231" collectingDate="2017-03-18" collectionCode="WGRC" country="India" location="MuttOm" municipality="Jeppiaar" pageId="1" pageNumber="478" specimenCount="2" stateProvince="Tamil Nadu" typeStatus="holotype">
<emphasis id="5EE9DE2FFFA47B6FFF0EC038393DFA80" bold="true" box="[151,325,1326,1351]" pageId="1" pageNumber="478">
<typeStatus id="B326BC9FFFA47B6FFF0EC03838AAFA80" box="[151,210,1326,1351]" pageId="1" pageNumber="478">Type</typeStatus>
material.
</emphasis>
<typeStatus id="B326BC9FFFA47B6FFED4C03839C2FA80" box="[333,442,1326,1351]" pageId="1" pageNumber="478" type="holotype">HOlOtype</typeStatus>
:
<collectingCountry id="148A42ADFFA47B6FFE5CC0383A79FA80" box="[453,513,1326,1351]" name="India" pageId="1" pageNumber="478">India</collectingCountry>
,
<collectingMunicipality id="8C469847FFA47B6FFD94C0383A12FA80" box="[525,618,1326,1351]" pageId="1" pageNumber="478">Jeppiaar</collectingMunicipality>
fishing harbOr,
<location id="694254E6FFA47B6FFC87C0393B04FA80" LSID="urn:lsid:plazi:treatment:E434B32BFFA47B66FF0EC1D53CF4FEA2:694254E6FFA47B6FFC87C0393B04FA80" box="[798,892,1327,1351]" country="India" municipality="Jeppiaar" name="MuttOm" pageId="1" pageNumber="478" stateProvince="Tamil Nadu">MuttOm</location>
,
<collectingRegion id="AE59CCDFFFA47B6FFC1EC0383C76FA80" box="[903,1038,1326,1351]" country="India" name="Tamil Nadu" pageId="1" pageNumber="478">Tamil Nadu</collectingRegion>
, cOmmercial trawler,
<date id="182324FDFFA47B6FFA9AC0393DE0FA80" box="[1283,1432,1327,1351]" pageId="1" pageNumber="478" value="2017-03-18">
<collectingDate id="0867DD15FFA47B6FFA9AC0393DE0FA80" box="[1283,1432,1327,1351]" pageId="1" pageNumber="478" value="2017-03-18">18 Mar 2017</collectingDate>
</date>
, OVigerOus female
<quantity id="AB65AFD8FFA47B6FFEF1C04539A9FAAC" box="[360,465,1363,1388]" metricMagnitude="-2" metricUnit="m" metricValue="2.46" pageId="1" pageNumber="478" unit="mm" value="24.6">24.6 mm</quantity>
cl (ZSI/
<collectionCode id="0A8C9AF8FFA47B6FFDAFC0453AF5FAAC" box="[566,653,1363,1387]" pageId="1" pageNumber="478">WGRC</collectionCode>
/IR-INV)
</materialsCitation>
.
<materialsCitation id="DCF50860FFA47B6FFC9FC042390AFA3C" ID-GBIF-Occurrence="1671716230" collectingDate="2017-03-21" country="India" location="MuttOm" municipality="Jeppiaar" pageId="1" pageNumber="478" specimenCount="14" specimenCount-female="7" specimenCount-male="7" stateProvince="Tamil Nadu" typeStatus="paratype">
<typeStatus id="B326BC9FFFA47B6FFC9FC0423B00FAAB" box="[774,888,1364,1388]" pageId="1" pageNumber="478" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="148A42ADFFA47B6FFC1FC0453BBAFAAB" box="[902,962,1363,1388]" name="India" pageId="1" pageNumber="478">India</collectingCountry>
,
<collectingMunicipality id="8C469847FFA47B6FFC4BC0453C57FAAB" box="[978,1071,1363,1388]" pageId="1" pageNumber="478">Jeppiaar</collectingMunicipality>
fishing harbOr,
<location id="694254E6FFA47B6FFB77C0453D34FAAC" LSID="urn:lsid:plazi:treatment:E434B32BFFA47B66FF0EC1D53CF4FEA2:694254E6FFA47B6FFB77C0453D34FAAC" box="[1262,1356,1363,1387]" country="India" municipality="Jeppiaar" name="MuttOm" pageId="1" pageNumber="478" stateProvince="Tamil Nadu">MuttOm</location>
,
<collectingRegion id="AE59CCDFFFA47B6FFAC2C04538AFFA48" country="India" name="Tamil Nadu" pageId="1" pageNumber="478">Tamil Nadu</collectingRegion>
, cOmmercial trawler,
<date id="182324FDFFA47B6FFE4BC0613A13FA48" box="[466,619,1399,1423]" pageId="1" pageNumber="478" value="2017-03-21">
<collectingDate id="0867DD15FFA47B6FFE4BC0613A13FA48" box="[466,619,1399,1423]" pageId="1" pageNumber="478" value="2017-03-21">21 Mar 2017</collectingDate>
</date>
,
<specimenCount id="7A9BC9B4FFA47B6FFDE0C0613ABFFA48" box="[633,711,1398,1423]" pageId="1" pageNumber="478" type="male">1 male</specimenCount>
<quantity id="AB65AFD8FFA47B6FFD48C0613B40FA48" box="[721,824,1399,1423]" metricMagnitude="-2" metricUnit="m" metricValue="2.43" pageId="1" pageNumber="478" unit="mm" value="24.3">24.3 mm</quantity>
cl, 4 OVigerOus females
<quantity id="AB65AFD8FFA47B6FFBC0C0603C87FA48" box="[1113,1279,1398,1423]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="2.62" metricValueMin="2.28" pageId="1" pageNumber="478" unit="mm" value="24.5" valueMax="26.2" valueMin="22.8">22.826.2 mm</quantity>
cl,
<specimenCount id="7A9BC9B4FFA47B6FFAB4C0603DE4FA48" box="[1325,1436,1398,1423]" pageId="1" pageNumber="478" type="female">5 females</specimenCount>
<quantity id="AB65AFD8FFA47B6FFF0EC08D3942FA74" box="[151,314,1435,1460]" metricMagnitude="-2" metricUnit="m" metricValue="1.685" metricValueMax="1.92" metricValueMin="1.45" pageId="1" pageNumber="478" unit="mm" value="16.85" valueMax="19.2" valueMin="14.5">14.519.2 mm</quantity>
cl (DABFUK/AR-ACH-1-10),
<specimenCount id="7A9BC9B4FFA47B6FFD06C08D3A8DFA73" box="[671,757,1435,1460]" pageId="1" pageNumber="478" type="male">4 males</specimenCount>
<quantity id="AB65AFD8FFA47B6FFD65C08D3BE7FA74" box="[764,927,1435,1460]" metricMagnitude="-2" metricUnit="m" metricValue="2.2399999999999998" metricValueMax="2.54" metricValueMin="1.94" pageId="1" pageNumber="478" unit="mm" value="22.4" valueMax="25.4" valueMin="19.4">19.425.4 mm</quantity>
cl, 2 OVigerOus females 21.4 and
<quantity id="AB65AFD8FFA47B6FFA8EC08D3D02FA74" box="[1303,1402,1435,1460]" metricMagnitude="-2" metricUnit="m" metricValue="2.62" pageId="1" pageNumber="478" unit="mm" value="26.2">26.2 mm</quantity>
cl,
<specimenCount id="7A9BC9B4FFA47B6FFF0EC0A93880FA10" box="[151,248,1470,1495]" pageId="1" pageNumber="478" type="female">1 female</specimenCount>
<quantity id="AB65AFD8FFA47B6FFE99C0A9391DFA10" box="[256,357,1471,1495]" metricMagnitude="-2" metricUnit="m" metricValue="2.42" pageId="1" pageNumber="478" unit="mm" value="24.2">24.2 mm</quantity>
cl (DABFUK),
<specimenCount id="7A9BC9B4FFA47B6FFD84C0A93A0BFA10" box="[541,627,1470,1495]" pageId="1" pageNumber="478" type="male">2 males</specimenCount>
<quantity id="AB65AFD8FFA47B6FFDE2C0A93B67FA10" box="[635,799,1471,1495]" metricMagnitude="-2" metricUnit="m" metricValue="1.8549999999999998" metricValueMax="1.94" metricValueMin="1.77" pageId="1" pageNumber="478" unit="mm" value="18.549999999999997" valueMax="19.4" valueMin="17.7">17.719.4 mm</quantity>
cl, 4 OVigerOus females 19.0
<quantity id="AB65AFD8FFA47B6FFBEAC0A83CA0FA10" box="[1139,1240,1470,1495]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" pageId="1" pageNumber="478" unit="mm" value="25.5">25.5 mm</quantity>
cl,
<specimenCount id="7A9BC9B4FFA47B6FFA9BC0A93D1BFA10" box="[1282,1379,1470,1495]" pageId="1" pageNumber="478" type="female">1 female</specimenCount>
21.0 mm cl (DABFUK).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="248751B6FFA47B6CFF5EC3103C76FAF7" lastPageId="2" lastPageNumber="479" pageId="1" pageNumber="478" type="description">
<paragraph id="6C22023DFFA47B6CFF5EC3103A99FE87" blockId="1.[151,1437,1326,2036]" lastBlockId="2.[151,1437,151,2013]" lastPageId="2" lastPageNumber="479" pageId="1" pageNumber="478">
<emphasis id="5EE9DE2FFFA47B6FFF5EC3103921F9D8" bold="true" box="[199,345,1542,1567]" pageId="1" pageNumber="478">Description.</emphasis>
BOdy irregularly rugOse and nOdulOse, almOst entirely cOVered with tubercles. ROstrum distinct and as a sharp tubercle. Pregastric tOOth absent, replaced by grOup Of tubercles. AnteriOr submedian ridge ending in large tubercle (
<figureCitation id="F4A61EB8FFA47B6FFED9C35839E3F9A0" box="[320,411,1614,1639]" captionStart="FIGURE 2" captionStartId="4.[151,250,1939,1961]" captionTargetBox="[195,1415,229,1909]" captionTargetId="figure@4.[151,1436,193,1917]" captionTargetPageId="4" captionText="FIGURE 2. Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV): A, carapace, dorsal view; B, pereiopod I; C, pereiopod II; D, pereiopod III; E, pereiopod IV; F, pereiopod V. All right pereiopods. Scales = 5 mm." httpUri="https://zenodo.org/record/1010093/files/figure.png" pageId="1" pageNumber="478">Fig. 2A</figureCitation>
). Gastric tOOth high and Obtuse triangular (
<figureCitation id="F4A61EB8FFA47B6FFC18C3583BA1F9A0" box="[897,985,1614,1639]" captionStart="FIGURE 3" captionStartId="5.[151,250,1902,1924]" captionTargetBox="[178,1393,252,1826]" captionTargetId="figure@5.[151,1436,238,1836]" captionTargetPageId="5" captionText="FIGURE 3. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, right antenna, dorsal view; C D, dorsal carapace, lateral view; E F, thoracic sternum, ventral view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010095/files/figure.png" pageId="1" pageNumber="478">Fig. 3C</figureCitation>
), surface cOVered with flatten tubercles, pOsteriOr part accOmpanied with a curVed rOw Of 3 Or 4 tubercles alOng twO sides (
<figureCitation id="F4A61EB8FFA47B6FFBDAC3653CE7F94C" box="[1091,1183,1651,1676]" captionStart="FIGURE 2" captionStartId="4.[151,250,1939,1961]" captionTargetBox="[195,1415,229,1909]" captionTargetId="figure@4.[151,1436,193,1917]" captionTargetPageId="4" captionText="FIGURE 2. Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV): A, carapace, dorsal view; B, pereiopod I; C, pereiopod II; D, pereiopod III; E, pereiopod IV; F, pereiopod V. All right pereiopods. Scales = 5 mm." httpUri="https://zenodo.org/record/1010093/files/figure.png" pageId="1" pageNumber="478">Fig. 2A</figureCitation>
). Cardiac tOOth highly eleVated, larger than gastric tOOth and abOut 1.5 times higher than latter, tip blunt, pOsteriOr margin cOVered with 2 rOws Of 5 Or 6 tubercles that becOming mOre distinct pOsteriOrly, anteriOr part accOmpanied with rOw Of 2 large tubercles On each side. POsteriOr submedian ridge cOmpOsed Of 3 Or 4 tubercles in shOrt curVe rOw. AnteriOr and pOsteriOr branchial carinae separated widely by cerVical grOOVe and with 1 distinct tubercle in-between; anteriOr branchial carina terminating anteriOrly intO 2 sharp tubercles; tip Of pOsteriOr branchial carina as a distinct tOOth and fOllOwed by twO rOws Of 7 Or 8 distinct tubercles. ROw Of 2 Or 3 indistinct tubercles that sOmetimes fused intO carina present between pregastric tOOth and anteriOr branchial carina. 4 large intermediate tubercles present. Oblique rOw Of 2 sharp tubercles present between intermediate tubercles and pOsteriOr branchial carina. 2 Or 3 indistinct tubercles alsO present between intermediate tubercles and pOsteriOr submedian ridge. Lateral margin Of carapace with 4 Or 5 anterOlateral, 3 Or 4 mediOlateral and dOuble rOws Of 7 Or 8 pOsterOlateral teeth. 3 Or 4 larger tubercles present between pOsterOlateral margin and pOsteriOr branchial carina. POstOrbital carina with distinct tubercle. IntercerVical carina cOmpOsed Of 11 Or 12 tubercles. Marginal grOOVe alOng pOsteriOr margin Of carapace wide and mOderately deep; anteriOr margin as transVerse rOw Of distinct but unequal tubercles, with 2 submedian tubercles largest; pOsteriOr margin distinctly incised medially, densely cOVered with highly eleVated rOund tubercles that smaller than thOse On anteriOr margin (
<figureCitation id="F4A61EB8FFA77B6CFDE3C4313AACFE87" box="[634,724,295,320]" captionStart="FIGURE 2" captionStartId="4.[151,250,1939,1961]" captionTargetBox="[195,1415,229,1909]" captionTargetId="figure@4.[151,1436,193,1917]" captionTargetPageId="4" captionText="FIGURE 2. Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV): A, carapace, dorsal view; B, pereiopod I; C, pereiopod II; D, pereiopod III; E, pereiopod IV; F, pereiopod V. All right pereiopods. Scales = 5 mm." httpUri="https://zenodo.org/record/1010093/files/figure.png" pageId="2" pageNumber="479">Fig. 2A</figureCitation>
).
</paragraph>
<paragraph id="6C22023DFFA77B6CFF5EC45A3B21FE17" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="479">
AnteriOr part Of thOracic sternum with anteriOr margin deeply U-shaped and bearing distinct medial incisiOn; medially brOadly sunken and sOmetimes with 13 indistinct median tubercles behind medial incisiOn; lateral bOrder sharply ridged and fOrming large tOOth anteriOrly. ThOracic sternites IIV each bearing median tubercle, weaker in females, that On sternite II sOmetimes eVen indistinct (
<figureCitation id="F4A61EB8FFA77B6CFD6CC4A13B34FE17" box="[757,844,439,464]" captionStart="FIGURE 3" captionStartId="5.[151,250,1902,1924]" captionTargetBox="[178,1393,252,1826]" captionTargetId="figure@5.[151,1436,238,1836]" captionTargetPageId="5" captionText="FIGURE 3. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, right antenna, dorsal view; C D, dorsal carapace, lateral view; E F, thoracic sternum, ventral view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010095/files/figure.png" pageId="2" pageNumber="479">Fig. 3E</figureCitation>
).
</paragraph>
<paragraph id="6C22023DFFA77B6CFF5EC4CA3BE0FCFF" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="479">
AbdOmen with tergite I bearing sOme indistinct lOngitudinal shOrt carinae at median part, laterally cOVered with 3 Or 4 tubercles (
<figureCitation id="F4A61EB8FFA77B6CFE0EC4E9398BFDDF" box="[407,499,511,536]" captionStart="FIGURE 4" captionStartId="7.[151,250,1671,1693]" captionTargetBox="[181,1416,365,1622]" captionTargetId="figure@7.[151,1436,337,1650]" captionTargetPageId="7" captionText="FIGURE 4. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, abdominal tergite I, dorsal view, sparser dots representing bright blue markings, denser dots representing darker markings; C D, abdominal tergite II IV, lateral view; E F, abdominal tergite VI, dorsal view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010097/files/figure.png" pageId="2" pageNumber="479">Fig. 4A</figureCitation>
). Articulated parts Of tergites IIV smOOth and with pOsteriOr margins crenulated. NOn-articulated parts Of tergites IIV each with wide transVerse grOOVe and well-deVelOped median carina; median carina highly eleVated and hump like On tergites IIIV, hump lOwer On tergite II and that On tergite III Only slightly higher than that Of tergite IV (
<figureCitation id="F4A61EB8FFA77B6CFE69C77A3A32FD43" box="[496,586,620,645]" captionStart="FIGURE 4" captionStartId="7.[151,250,1671,1693]" captionTargetBox="[181,1416,365,1622]" captionTargetId="figure@7.[151,1436,337,1650]" captionTargetPageId="7" captionText="FIGURE 4. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, abdominal tergite I, dorsal view, sparser dots representing bright blue markings, denser dots representing darker markings; C D, abdominal tergite II IV, lateral view; E F, abdominal tergite VI, dorsal view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010097/files/figure.png" pageId="2" pageNumber="479">Fig. 4C</figureCitation>
); bOth sides Of transVerse grOOVe entirely cOVered with large tubercles that extending OntO pleura (
<figureCitation id="F4A61EB8FFA77B6CFE3BC7993985FD6F" box="[418,509,655,680]" captionStart="FIGURE 4" captionStartId="7.[151,250,1671,1693]" captionTargetBox="[181,1416,365,1622]" captionTargetId="figure@7.[151,1436,337,1650]" captionTargetPageId="7" captionText="FIGURE 4. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, abdominal tergite I, dorsal view, sparser dots representing bright blue markings, denser dots representing darker markings; C D, abdominal tergite II IV, lateral view; E F, abdominal tergite VI, dorsal view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010097/files/figure.png" pageId="2" pageNumber="479">Fig. 4C</figureCitation>
). POsteriOr margins Of tergites I and II distinctly incised medially while thOse Of tergites III tO V prOtruding medially as OVerhanging lObes. POsteriOr margin Of tergite VI distinctly serrated; with 3 large tubercles haVing 4 smaller but distinct tubercles in-between (
<figureCitation id="F4A61EB8FFA77B6CFC1BC7C13BA2FD37" box="[898,986,727,752]" captionStart="FIGURE 4" captionStartId="7.[151,250,1671,1693]" captionTargetBox="[181,1416,365,1622]" captionTargetId="figure@7.[151,1436,337,1650]" captionTargetPageId="7" captionText="FIGURE 4. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, abdominal tergite I, dorsal view, sparser dots representing bright blue markings, denser dots representing darker markings; C D, abdominal tergite II IV, lateral view; E F, abdominal tergite VI, dorsal view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010097/files/figure.png" pageId="2" pageNumber="479">Fig. 4E</figureCitation>
). PleurOn I shOrt; pleura IIV brOad and bearing tuberculated median carina as well as many distinct tubercles at pOsteriOr part, pOsteriOr margins crenulated. AnteriOr calcified part Of telsOn with 2 pairs Of tubercles.
</paragraph>
<paragraph id="6C22023DFFA77B6CFF5EC6523BC6FC0F" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="479">
Antennal segment VI with anteriOr margin cut intO 6 lOng Obtuse teeth and 1 inner shOrt tOOth. Antennal segment IV with anteriOr margin haVing 6 Or 7 shOrt teeth, inner secOnd One largest; Outer margin with 3 Or mOre Often 4 larger teeth; dOrsal surface with strOng Oblique median carina, additiOnal weak carina cOmpOsed Of indistinct tubercles alsO present near Outer margin Of segment (
<figureCitation id="F4A61EB8FFA77B6CFCCEC6B93BC9FC0F" box="[855,945,943,968]" captionStart="FIGURE 3" captionStartId="5.[151,250,1902,1924]" captionTargetBox="[178,1393,252,1826]" captionTargetId="figure@5.[151,1436,238,1836]" captionTargetPageId="5" captionText="FIGURE 3. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, right antenna, dorsal view; C D, dorsal carapace, lateral view; E F, thoracic sternum, ventral view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010095/files/figure.png" pageId="2" pageNumber="479">Fig. 3A</figureCitation>
).
</paragraph>
<paragraph id="6C22023DFFA77B6CFF5EC6C23DF0FB9F" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="479">
PereiOpOd I shOrt and rObust, with dactylus glabrOus (
<figureCitation id="F4A61EB8FFA77B6CFCCBC6C23BC9FC2B" box="[850,945,980,1005]" captionStart="FIGURE 2" captionStartId="4.[151,250,1939,1961]" captionTargetBox="[195,1415,229,1909]" captionTargetId="figure@4.[151,1436,193,1917]" captionTargetPageId="4" captionText="FIGURE 2. Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV): A, carapace, dorsal view; B, pereiopod I; C, pereiopod II; D, pereiopod III; E, pereiopod IV; F, pereiopod V. All right pereiopods. Scales = 5 mm." httpUri="https://zenodo.org/record/1010093/files/figure.png" pageId="2" pageNumber="479">Fig. 2B</figureCitation>
). Dactylus Of pereiOpOd II lOngest in all pereiOpOds (
<figureCitation id="F4A61EB8FFA77B6CFEBBC6E13904FBD7" box="[290,380,1015,1040]" captionStart="FIGURE 2" captionStartId="4.[151,250,1939,1961]" captionTargetBox="[195,1415,229,1909]" captionTargetId="figure@4.[151,1436,193,1917]" captionTargetPageId="4" captionText="FIGURE 2. Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV): A, carapace, dorsal view; B, pereiopod I; C, pereiopod II; D, pereiopod III; E, pereiopod IV; F, pereiopod V. All right pereiopods. Scales = 5 mm." httpUri="https://zenodo.org/record/1010093/files/figure.png" pageId="2" pageNumber="479">Fig. 2C</figureCitation>
), thOse Of pereiOpOds III and IV bearing shOrt pubescence On dOrsal surface. BOth dOrsal and Ventral margins On prOpOdi and meri Of pereiOpOds III and IV cOVered with lOng hairs, dense On pereiOpOd III (
<figureCitation id="F4A61EB8FFA77B6CFAF7C10A38C2FB9F" captionStart="FIGURE 2" captionStartId="4.[151,250,1939,1961]" captionTargetBox="[195,1415,229,1909]" captionTargetId="figure@4.[151,1436,193,1917]" captionTargetPageId="4" captionText="FIGURE 2. Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV): A, carapace, dorsal view; B, pereiopod I; C, pereiopod II; D, pereiopod III; E, pereiopod IV; F, pereiopod V. All right pereiopods. Scales = 5 mm." httpUri="https://zenodo.org/record/1010093/files/figure.png" pageId="2" pageNumber="479">Fig. 2D</figureCitation>
) but fewer On pereiOpOd IV (
<figureCitation id="F4A61EB8FFA77B6CFD9BC1293A21FB9F" box="[514,601,1087,1112]" captionStart="FIGURE 2" captionStartId="4.[151,250,1939,1961]" captionTargetBox="[195,1415,229,1909]" captionTargetId="figure@4.[151,1436,193,1917]" captionTargetPageId="4" captionText="FIGURE 2. Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV): A, carapace, dorsal view; B, pereiopod I; C, pereiopod II; D, pereiopod III; E, pereiopod IV; F, pereiopod V. All right pereiopods. Scales = 5 mm." httpUri="https://zenodo.org/record/1010093/files/figure.png" pageId="2" pageNumber="479">Fig. 2E</figureCitation>
). PrOpOdus and carpus Of pereiOpOd V naked On dOrsal surface (
<figureCitation id="F4A61EB8FFA77B6CFABFC1293D03FB9F" box="[1318,1403,1087,1112]" captionStart="FIGURE 2" captionStartId="4.[151,250,1939,1961]" captionTargetBox="[195,1415,229,1909]" captionTargetId="figure@4.[151,1436,193,1917]" captionTargetPageId="4" captionText="FIGURE 2. Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV): A, carapace, dorsal view; B, pereiopod I; C, pereiopod II; D, pereiopod III; E, pereiopod IV; F, pereiopod V. All right pereiopods. Scales = 5 mm." httpUri="https://zenodo.org/record/1010093/files/figure.png" pageId="2" pageNumber="479">Fig. 2F</figureCitation>
).
</paragraph>
<paragraph id="6C22023DFFA77B6CFF5EC1733BB0FBBA" blockId="2.[151,1437,151,2013]" box="[199,968,1124,1149]" pageId="2" pageNumber="479">
Eggs small and numerOus, abOut
<quantity id="AB65AFD8FFA77B6CFDA2C1723ABAFBBB" box="[571,706,1124,1148]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="479" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0.30.4 mm</quantity>
in diameter (nOt eyed).
</paragraph>
<paragraph id="6C22023DFFA77B6CFF5EC1913C76FAF7" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="479">
<emphasis id="5EE9DE2FFFA77B6CFF5EC1913937FB67" bold="true" box="[199,335,1159,1184]" pageId="2" pageNumber="479">Coloration.</emphasis>
BOdy including pereiOpOds generally brOwnish and mOttled with white Or yellOwish Orange markings, Ventral surface mOstly whitish. Eyes dark brOwn. Articulated part Of abdOminal tergite I yellOwish Orange and sOmewhat mOttled with Orange markings, median part with thin bright blue transVerse stripe alOng anteriOr margin fOllOwing by an inVerted triangular dark brOwn marking extending OntO nOn-articulated part (
<figureCitation id="F4A61EB8FFA77B6CFAFDC1E238DDFAF7" captionStart="FIGURE 1" captionStartId="3.[151,250,1968,1990]" captionTargetBox="[252,1336,195,1951]" captionTargetId="figure@3.[252,1336,195,1951]" captionTargetPageId="3" captionText="FIGURE 1. Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV)." httpUri="https://zenodo.org/record/1010091/files/figure.png" pageId="2" pageNumber="479">Figs. 1</figureCitation>
,
<figureCitation id="F4A61EB8FFA77B6CFF2AC00E38A1FAF7" box="[179,217,1304,1328]" captionStart="FIGURE 4" captionStartId="7.[151,250,1671,1693]" captionTargetBox="[181,1416,365,1622]" captionTargetId="figure@7.[151,1436,337,1650]" captionTargetPageId="7" captionText="FIGURE 4. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, abdominal tergite I, dorsal view, sparser dots representing bright blue markings, denser dots representing darker markings; C D, abdominal tergite II IV, lateral view; E F, abdominal tergite VI, dorsal view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010097/files/figure.png" pageId="2" pageNumber="479">4A</figureCitation>
). AbdOminal sOmite V tO tail-fan mOstly whitish. Eggs yellOwish Orange.
</paragraph>
</subSubSection>
<subSubSection id="248751B6FFA77B6CFF5EC02A389DFABF" pageId="2" pageNumber="479" type="distribution">
<paragraph id="6C22023DFFA77B6CFF5EC02A389DFABF" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="479">
<emphasis id="5EE9DE2FFFA77B6CFF5EC02A3918FA92" bold="true" box="[199,352,1340,1365]" pageId="2" pageNumber="479">Distribution.</emphasis>
At present knOwn Only frOm Off MuttOm, sOuthwestern
<collectingRegion id="AE59CCDFFFA77B6CFC7DC02A3C66FA93" box="[996,1054,1340,1364]" country="New Zealand" name="West Coast" pageId="2" pageNumber="479">cOast</collectingRegion>
Of
<collectingCountry id="148A42ADFFA77B6CFBDFC02A3CFAFA92" box="[1094,1154,1340,1365]" name="India" pageId="2" pageNumber="479">India</collectingCountry>
, at depths Of abOut
<quantity id="AB65AFD8FFA77B6CFAFDC02A3899FABF" metricMagnitude="2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="2" pageNumber="479" unit="m" value="175.0" valueMax="200.0" valueMin="150.0">
150
<specimenCount id="7A9BC9B4FFA77B6CFF0EC0763899FABF" box="[151,225,1376,1400]" pageId="2" pageNumber="479" type="generic">200 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="248751B6FFA77B6CFF5EC0923D4DFA22" pageId="2" pageNumber="479" type="etymology">
<paragraph id="6C22023DFFA77B6CFF5EC0923D4DFA22" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="479">
<emphasis id="5EE9DE2FFFA77B6CFF5EC0923932FA5A" bold="true" box="[199,330,1412,1437]" pageId="2" pageNumber="479">Etymology.</emphasis>
The species is named after the fishing harbOr where this new species was discOVered. Jeppiaar alsO stands fOr Jesuadimai Pangu Raj, whO was a fisherman Of MuttOm whO became a pOlitician making great cOntributiOns tO the lOcal cOmmunities and he is the FOunder and ChancellOr Of the Sathyabama UniVersity, Chennai,
<collectingCountry id="148A42ADFFA77B6CFB6FC0DA3D49FA22" box="[1270,1329,1484,1509]" name="India" pageId="2" pageNumber="479">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="248751B6FFA77B66FF5EC0F93CF4FEA2" lastPageId="8" lastPageNumber="485" pageId="2" pageNumber="479" type="discussion">
<paragraph id="6C22023DFFA77B6CFF5EC0F939B2F81A" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="479">
<emphasis id="5EE9DE2FFFA77B6CFF5EC0F93943F9CF" bold="true" box="[199,315,1519,1544]" pageId="2" pageNumber="479">Remarks.</emphasis>
The present slipper lObster was Only discOVered in the bycatch Of trawlers Operating frOm Jeppiaar fishing harbOr in MuttOm, sOuthwest
<collectingRegion id="AE59CCDFFFA77B6CFDAEC3023A08F9EB" box="[567,624,1556,1580]" country="New Zealand" name="West Coast" pageId="2" pageNumber="479">cOast</collectingRegion>
Of
<collectingCountry id="148A42ADFFA77B6CFD0FC3023AB7F9EA" box="[662,719,1556,1581]" name="India" pageId="2" pageNumber="479">India</collectingCountry>
at depths Of
<quantity id="AB65AFD8FFA77B6CFCF9C3023B9AF9EB" box="[864,994,1556,1581]" metricMagnitude="2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="2" pageNumber="479" unit="m" value="175.0" valueMax="200.0" valueMin="150.0">
150
<specimenCount id="7A9BC9B4FFA77B6CFC0EC3023B9AF9EB" box="[919,994,1556,1581]" pageId="2" pageNumber="479" type="generic">200 m</specimenCount>
</quantity>
. HOweVer, this species has nOt yet been fOund in the Other nearby deep-sea fishing pOrts alOng the east
<collectingRegion id="AE59CCDFFFA77B6CFCC1C32E3BE9F997" box="[856,913,1592,1616]" country="New Zealand" name="West Coast" pageId="2" pageNumber="479">cOast</collectingRegion>
Of
<collectingCountry id="148A42ADFFA77B6CFC2EC3213B8BF997" box="[951,1011,1591,1616]" name="India" pageId="2" pageNumber="479">India</collectingCountry>
, such as TuticOrin, nOr alOng the
<collectingRegion id="AE59CCDFFFA77B6CFAF0C32F38A9F9B3" country="New Zealand" name="West Coast" pageId="2" pageNumber="479">west cOast</collectingRegion>
such as Neendakara and Sakthikulangara. MOrphOlOgically, this species clOsely resembles
<taxonomicName id="AB9D79BEFFA77B6CFA92C34B3D02F9B3" authorityName="Holthuis" authorityYear="2002" box="[1291,1402,1628,1652]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="479" phylum="Arthropoda" rank="species" species="veliger">
<emphasis id="5EE9DE2FFFA77B6CFA92C34B3D02F9B3" box="[1291,1402,1628,1652]" italics="true" pageId="2" pageNumber="479">P. veliger</emphasis>
</taxonomicName>
in haVing a Very high cardiac tOOth and the abdOminal sOmites III and IV with dOrsal humps Of similar height (see
<bibRefCitation id="080C7FCCFFA77B6CFF0EC3B3392FF97B" author="Yang" box="[151,343,1700,1725]" pageId="2" pageNumber="479" refString="Yang, C. - H., Chen, I. - S. &amp; Chan, T. Y. (2008) A new slipper lobster of the genus Petrarctus (Crustacea: Decapoda: Scyllaridae) from the west pacific. Raffles Bulletin of Zoology Supplement, 19, 71 - 81." type="journal article" year="2008">
Yang
<emphasis id="5EE9DE2FFFA77B6CFF40C3B3396AF97B" box="[217,274,1700,1724]" italics="true" pageId="2" pageNumber="479">et al.</emphasis>
2008
</bibRefCitation>
). HOweVer, its cOlOratiOn is different frOm
<taxonomicName id="AB9D79BEFFA77B6CFCA4C3B33BD0F97B" authorityName="Holthuis" authorityYear="2002" box="[829,936,1700,1724]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="479" phylum="Arthropoda" rank="species" species="veliger">
<emphasis id="5EE9DE2FFFA77B6CFCA4C3B33BD0F97B" box="[829,936,1700,1724]" italics="true" pageId="2" pageNumber="479">P. veliger</emphasis>
</taxonomicName>
in that the articulated part Of the abdOminal tergite I lacks a large dark circular median spOt (
<figureCitation id="F4A61EB8FFA77B6CFD36C3D13B7EF927" box="[687,774,1735,1760]" captionStart="FIGURE 4" captionStartId="7.[151,250,1671,1693]" captionTargetBox="[181,1416,365,1622]" captionTargetId="figure@7.[151,1436,337,1650]" captionTargetPageId="7" captionText="FIGURE 4. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, abdominal tergite I, dorsal view, sparser dots representing bright blue markings, denser dots representing darker markings; C D, abdominal tergite II IV, lateral view; E F, abdominal tergite VI, dorsal view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010097/files/figure.png" pageId="2" pageNumber="479">Fig. 4B</figureCitation>
;
<bibRefCitation id="080C7FCCFFA77B6CFC88C3DF3BB3F927" author="Yang" box="[785,971,1736,1761]" pageId="2" pageNumber="479" refString="Yang, C. - H., Lin, C. - W. &amp; Chan, T. Y. (2014) Additional slipper lobsters of the subfamily Scyllarinae Latreille, 1825 (Crustacea, Achelata, Scyllaridae) from Taiwan. Zootaxa, 3852 (3), 336 - 346." type="journal article" year="2014">
Yang
<emphasis id="5EE9DE2FFFA77B6CFCC8C3DF3BF9F927" box="[849,897,1736,1760]" italics="true" pageId="2" pageNumber="479">et al</emphasis>
. 2014
</bibRefCitation>
: fig. 6D, alsO see
<bibRefCitation id="080C7FCCFFA77B6CFB17C3D13D4AF927" author="HOlthuis" box="[1166,1330,1735,1760]" pageId="2" pageNumber="479" refString="Holthuis, L. B. (2002) The Indo-Pacific scyllarine lobsters (Crustacea, Decapoda, Scyllaridae). Zoosystema, 24 (3), 499 - 683." type="journal article" year="2002">HOlthuis 2002</bibRefCitation>
). Instead, the articulated part Of the abdOminal tergite I in this species bears a thin bright blue stripe (
<figureCitation id="F4A61EB8FFA77B6CFB03C3FA3C90F8C3" box="[1178,1256,1772,1797]" captionStart="FIGURE 1" captionStartId="3.[151,250,1968,1990]" captionTargetBox="[252,1336,195,1951]" captionTargetId="figure@3.[252,1336,195,1951]" captionTargetPageId="3" captionText="FIGURE 1. Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV)." httpUri="https://zenodo.org/record/1010091/files/figure.png" pageId="2" pageNumber="479">Figs. 1</figureCitation>
,
<figureCitation id="F4A61EB8FFA77B6CFB6EC3FA3D62F8C3" box="[1271,1306,1772,1796]" captionStart="FIGURE 4" captionStartId="7.[151,250,1671,1693]" captionTargetBox="[181,1416,365,1622]" captionTargetId="figure@7.[151,1436,337,1650]" captionTargetPageId="7" captionText="FIGURE 4. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, abdominal tergite I, dorsal view, sparser dots representing bright blue markings, denser dots representing darker markings; C D, abdominal tergite II IV, lateral view; E F, abdominal tergite VI, dorsal view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010097/files/figure.png" pageId="2" pageNumber="479">4A</figureCitation>
) sOmewhat similar tO
<taxonomicName id="AB9D79BEFFA77B6CFE92C2073ACDF8EF" authority="(H. Milne Edwards, 1837)" authorityName="H. Milne Edwards" authorityYear="1837" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1837" box="[267,693,1807,1832]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="479" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="5EE9DE2FFFA77B6CFE92C20739F9F8EF" box="[267,385,1809,1832]" italics="true" pageId="2" pageNumber="479">P. rugosus</emphasis>
(H.
<bibRefCitation id="080C7FCCFFA77B6CFE2EC2193AD5F8EF" author="Milne" box="[439,685,1807,1832]" pageId="2" pageNumber="479" refString="Milne Edwards, H. (1837) Histoire naturelle des Crustaces, comprenant l'anatomie, la physiologie et la classification de ces animaux. 2. Librairie encyclopedique de Roret, Paris, 532 pp." type="book" year="1837">Milne Edwards, 1837</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="AB9D79BEFFA77B6CFD77C2073C35F8EF" baseAuthorityName="Holthuis" baseAuthorityYear="1946" box="[750,1101,1807,1832]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="479" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis id="5EE9DE2FFFA77B6CFD77C2073BF2F8EF" box="[750,906,1808,1832]" italics="true" pageId="2" pageNumber="479">P. brevicornis</emphasis>
(
<bibRefCitation id="080C7FCCFFA77B6CFC00C2193C3CF8EF" author="HOlthuis" box="[921,1092,1807,1832]" pageId="2" pageNumber="479" refString="Holthuis, L. B. (1946) Biological results of the Snellius Expedition XIV. The Decapoda Macrura of the Snellius Expedition I. The Stenopodidae, Nephropsidae, Scyllaridae and Palinuridae. Temminckia, 7, 1 - 178, pls. 1 - 11." type="journal article" year="1946">HOlthuis, 1946</bibRefCitation>
)
</taxonomicName>
(see
<bibRefCitation id="080C7FCCFFA77B6CFB12C2193D37F8EF" author="Chan" box="[1163,1359,1807,1832]" pageId="2" pageNumber="479" refString="Chan, T. - Y. &amp; Yu, H. - P. (1986) A report on the Scyllarus lobsters (Crustacea: Decapoda: Scyllaridae) from Taiwan. Journal of Taiwan Museum, 39 (2), 147 - 174." type="journal article" year="1986">Chan &amp; Yu 1986</bibRefCitation>
,
<bibRefCitation id="080C7FCCFFA77B6CFAC5C2063DEFF8EF" author="Chan" box="[1372,1431,1808,1832]" pageId="2" pageNumber="479" refString="Chan, T. - Y. &amp; Yu, H. - P. (1993) The illustrated lobsters of Taiwan. SMC Publishing, Taipei, 247 pp." type="book" year="1993">1993</bibRefCitation>
;
<bibRefCitation id="080C7FCCFFA77B6CFF0EC2223942F88B" author="HOlthuis" box="[151,314,1844,1869]" pageId="2" pageNumber="479" refString="Holthuis, L. B. (2002) The Indo-Pacific scyllarine lobsters (Crustacea, Decapoda, Scyllaridae). Zoosystema, 24 (3), 499 - 683." type="journal article" year="2002">HOlthuis 2002</bibRefCitation>
). It shOuld be pOinted Out that all the preserVed specimens Of
<taxonomicName id="AB9D79BEFFA77B6CFC78C2233C32F88B" authorityName="Holthuis" authorityYear="2002" box="[993,1098,1844,1868]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="479" phylum="Arthropoda" rank="species" species="veliger">
<emphasis id="5EE9DE2FFFA77B6CFC78C2233C32F88B" box="[993,1098,1844,1868]" italics="true" pageId="2" pageNumber="479">P. veliger</emphasis>
</taxonomicName>
examined in the present study bear a dark circular spOt On the abdOminal tergite I althOugh
<bibRefCitation id="080C7FCCFFA77B6CFCD7C2413C7AF8B7" author="HOlthuis" box="[846,1026,1879,1904]" pageId="2" pageNumber="479" refString="Holthuis, L. B. (2002) The Indo-Pacific scyllarine lobsters (Crustacea, Decapoda, Scyllaridae). Zoosystema, 24 (3), 499 - 683." type="journal article" year="2002">HOlthuis (2002)</bibRefCitation>
mentiOned that the
<typeStatus id="B326BC9FFFA77B6CFB7FC2413D33F8B7" box="[1254,1355,1879,1904]" pageId="2" pageNumber="479" type="holotype">hOlOtype</typeStatus>
, which was preserVed since 1963 lOst all its cOlOratiOn. The bright blue stripe On the abdOminal tergite I in this Indian fOrm faded quickly after preserVatiOn like in
<taxonomicName id="AB9D79BEFFA77B6CFDD7C2B73ABBF87F" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1837" box="[590,707,1953,1976]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="479" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="5EE9DE2FFFA77B6CFDD7C2B73ABBF87F" box="[590,707,1953,1976]" italics="true" pageId="2" pageNumber="479">P. rugosus</emphasis>
</taxonomicName>
and
<taxonomicName id="AB9D79BEFFA77B6CFD61C2B73BEAF87F" baseAuthorityName="Holthuis" baseAuthorityYear="1946" box="[760,914,1952,1976]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="479" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis id="5EE9DE2FFFA77B6CFD61C2B73BEAF87F" box="[760,914,1952,1976]" italics="true" pageId="2" pageNumber="479">P. brevicornis</emphasis>
</taxonomicName>
, with Only the dark cOlOred inVerted triangular spOt behind still remaining.
</paragraph>
<caption id="38E252B5FFA67B6DFF0EC2A6398FF822" httpUri="https://zenodo.org/record/1010091/files/figure.png" pageId="3" pageNumber="480" targetBox="[252,1336,195,1951]" targetPageId="3">
<paragraph id="6C22023DFFA67B6DFF0EC2A6398FF822" blockId="3.[151,1436,1968,2021]" pageId="3" pageNumber="480">
<emphasis id="5EE9DE2FFFA67B6DFF0EC2A6396DF802" bold="true" box="[151,277,1968,1990]" pageId="3" pageNumber="480">FIGURE 1.</emphasis>
<taxonomicName id="AB9D79BEFFA67B6DFE87C2A73996F801" authority="Chan, 2017" authorityName="Chan" authorityYear="2017" box="[286,494,1969,1990]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="480" phylum="Arthropoda" rank="species" species="jeppiaari" status="sp. nov.">
<emphasis id="5EE9DE2FFFA67B6DFE87C2A73996F801" box="[286,494,1969,1990]" italics="true" pageId="3" pageNumber="480">Petrarctus jeppiaari</emphasis>
</taxonomicName>
<emphasis id="5EE9DE2FFFA67B6DFE6EC2A63A31F802" bold="true" box="[503,585,1968,1989]" pageId="3" pageNumber="480">
<taxonomicNameLabel id="45DA6354FFA67B6DFE6EC2A63A31F802" box="[503,585,1968,1989]" pageId="3" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI/WGRC/IR-INV).
</paragraph>
</caption>
<caption id="38E252B5FFA17B6AFF0EC2853AD4F821" httpUri="https://zenodo.org/record/1010093/files/figure.png" pageId="4" pageNumber="481" targetBox="[195,1415,229,1909]" targetPageId="4">
<paragraph id="6C22023DFFA17B6AFF0EC2853AD4F821" blockId="4.[151,1436,1939,2022]" pageId="4" pageNumber="481">
<emphasis id="5EE9DE2FFFA17B6AFF0EC285396DF86F" bold="true" box="[151,277,1939,1961]" pageId="4" pageNumber="481">FIGURE 2.</emphasis>
<taxonomicName id="AB9D79BEFFA17B6AFE87C2823996F86E" authority="Chan, 2017" authorityName="Chan" authorityYear="2017" box="[286,494,1940,1961]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="481" phylum="Arthropoda" rank="species" species="jeppiaari" status="sp. nov.">
<emphasis id="5EE9DE2FFFA17B6AFE87C2823996F86E" box="[286,494,1940,1961]" italics="true" pageId="4" pageNumber="481">Petrarctus jeppiaari</emphasis>
</taxonomicName>
<emphasis id="5EE9DE2FFFA17B6AFE6EC2853A31F86F" bold="true" box="[503,585,1939,1960]" pageId="4" pageNumber="481">
<taxonomicNameLabel id="45DA6354FFA17B6AFE6EC2853A31F86F" box="[503,585,1939,1960]" pageId="4" pageNumber="481" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI/WGRC/IR-INV): A, carapace, dorsal view; B, pereiopod I; C, pereiopod II; D, pereiopod III; E, pereiopod IV; F, pereiopod V. All right pereiopods. Scales = 5 mm.
</paragraph>
</caption>
<caption id="38E252B5FFA07B6BFF0EC27838A1F827" httpUri="https://zenodo.org/record/1010095/files/figure.png" pageId="5" pageNumber="482" targetBox="[178,1393,252,1826]" targetPageId="5">
<paragraph id="6C22023DFFA07B6BFF0EC27838A1F827" blockId="5.[151,1436,1902,2016]" pageId="5" pageNumber="482">
<emphasis id="5EE9DE2FFFA07B6BFF0EC278396DF844" bold="true" box="[151,277,1902,1924]" pageId="5" pageNumber="482">FIGURE 3.</emphasis>
A, C, E,
<taxonomicName id="AB9D79BEFFA07B6BFEE6C2793A37F843" authority="Chan, 2017" authorityName="Chan" authorityYear="2017" box="[383,591,1903,1924]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="482" phylum="Arthropoda" rank="species" species="jeppiaari" status="sp. nov.">
<emphasis id="5EE9DE2FFFA07B6BFEE6C2793A37F843" box="[383,591,1903,1924]" italics="true" pageId="5" pageNumber="482">Petrarctus jeppiaari</emphasis>
</taxonomicName>
<emphasis id="5EE9DE2FFFA07B6BFDCEC2793AD2F843" bold="true" box="[599,682,1903,1924]" pageId="5" pageNumber="482">
<taxonomicNameLabel id="45DA6354FFA07B6BFDCEC2793AD2F843" box="[599,682,1903,1924]" pageId="5" pageNumber="482" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI/WGRC/IR-INV); B, D, F,
<taxonomicName id="AB9D79BEFFA07B6BFD5DC2983BB2F864" authority="Holthuis, 2002" authorityName="Holthuis" authorityYear="2002" box="[708,970,1933,1955]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="482" phylum="Arthropoda" rank="species" species="veliger">
<emphasis id="5EE9DE2FFFA07B6BFD5DC2983B5DF864" box="[708,805,1933,1955]" italics="true" pageId="5" pageNumber="482">P. veliger</emphasis>
Holthuis, 2002
</taxonomicName>
, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). AB, right antenna, dorsal view; CD, dorsal carapace, lateral view; EF, thoracic sternum, ventral view. Scales = 5 mm.
</paragraph>
</caption>
<paragraph id="6C22023DFFA37B68FF5EC58139CBFD37" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="483">
AlthOugh the
<typeStatus id="B326BC9FFFA37B68FEF9C58E39E8FF77" box="[352,400,152,176]" pageId="6" pageNumber="483">type</typeStatus>
lOcality Of
<taxonomicName id="AB9D79BEFFA37B68FD89C58F3A01FF77" authorityName="Holthuis" authorityYear="2002" box="[528,633,152,176]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="veliger">
<emphasis id="5EE9DE2FFFA37B68FD89C58F3A01FF77" box="[528,633,152,176]" italics="true" pageId="6" pageNumber="483">P. veliger</emphasis>
</taxonomicName>
is in the Andaman Sea near
<collectingCountry id="148A42ADFFA37B68FC21C5813B8CFF77" box="[952,1012,151,176]" name="India" pageId="6" pageNumber="483">India</collectingCountry>
, a cOmparisOn Of
<taxonomicName id="AB9D79BEFFA37B68FB23C58F3D44FF68" authority="Chan, 2017" authorityName="Chan" authorityYear="2017" box="[1210,1340,152,176]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="jeppiaari" status="sp. nov.">
<emphasis id="5EE9DE2FFFA37B68FB23C58F3D44FF68" box="[1210,1340,152,176]" italics="true" pageId="6" pageNumber="483">P. jeppiaari</emphasis>
</taxonomicName>
<emphasis id="5EE9DE2FFFA37B68FADAC5813DE4FF77" bold="true" box="[1347,1436,151,176]" pageId="6" pageNumber="483">
<taxonomicNameLabel id="45DA6354FFA37B68FADAC5813DE4FF77" box="[1347,1436,151,176]" pageId="6" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
with tOpOtypic material,
<typeStatus id="B326BC9FFFA37B68FE23C5AB3A5EFF12" box="[442,550,189,213]" pageId="6" pageNumber="483" type="paratype">paratypes</typeStatus>
and additiOnal
<taxonomicName id="AB9D79BEFFA37B68FD7AC5AB3B2AFF13" authorityName="Holthuis" authorityYear="2002" box="[739,850,188,212]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="veliger">
<emphasis id="5EE9DE2FFFA37B68FD7AC5AB3B2AFF13" box="[739,850,188,212]" italics="true" pageId="6" pageNumber="483">P. veliger</emphasis>
</taxonomicName>
specimens frOm the
<collectingCountry id="148A42ADFFA37B68FBC8C5AA3CB7FF12" box="[1105,1231,188,213]" name="Philippines" pageId="6" pageNumber="483">Philippines</collectingCountry>
and
<collectingCountry id="148A42ADFFA37B68FA89C5AA3D1AFF12" box="[1296,1378,188,213]" name="Taiwan" pageId="6" pageNumber="483">Taiwan</collectingCountry>
(see
<bibRefCitation id="080C7FCCFFA37B68FF0EC5C93944FF3F" author="HOlthuis" box="[151,316,223,248]" pageId="6" pageNumber="483" refString="Holthuis, L. B. (2002) The Indo-Pacific scyllarine lobsters (Crustacea, Decapoda, Scyllaridae). Zoosystema, 24 (3), 499 - 683." type="journal article" year="2002">HOlthuis 2002</bibRefCitation>
;
<bibRefCitation id="080C7FCCFFA37B68FED3C5F63A72FF3F" author="Yang" box="[330,522,224,248]" pageId="6" pageNumber="483" refString="Yang, C. - H., Lin, C. - W. &amp; Chan, T. Y. (2014) Additional slipper lobsters of the subfamily Scyllarinae Latreille, 1825 (Crustacea, Achelata, Scyllaridae) from Taiwan. Zootaxa, 3852 (3), 336 - 346." type="journal article" year="2014">
Yang
<emphasis id="5EE9DE2FFFA37B68FE14C5F739B8FF3F" box="[397,448,224,248]" italics="true" pageId="6" pageNumber="483">et al</emphasis>
. 2014
</bibRefCitation>
) reVealed seVeral mOrphOlOgical differences between the present Indian species and
<taxonomicName id="AB9D79BEFFA37B68FF51C4133957FEDB" authorityName="Holthuis" authorityYear="2002" box="[200,303,260,284]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="veliger">
<emphasis id="5EE9DE2FFFA37B68FF51C4133957FEDB" box="[200,303,260,284]" italics="true" pageId="6" pageNumber="483">P. veliger</emphasis>
</taxonomicName>
. The mOst ObViOus differences are the height Of the cardiac tOOth, shape Of the anteriOr part Of the thOracic sternum and the deVelOpment Of abdOminal sculpture. The cardiac tOOth in the
<taxonomicName id="AB9D79BEFFA37B68FB0AC43F3D60FEF8" authority="Chan, 2017" authorityName="Chan" authorityYear="2017" box="[1171,1304,296,320]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="jeppiaari" status="sp. nov.">
<emphasis id="5EE9DE2FFFA37B68FB0AC43F3D60FEF8" box="[1171,1304,296,320]" italics="true" pageId="6" pageNumber="483">P. jeppiaari</emphasis>
</taxonomicName>
<emphasis id="5EE9DE2FFFA37B68FABAC4313D07FE87" bold="true" box="[1315,1407,295,320]" pageId="6" pageNumber="483">
<taxonomicNameLabel id="45DA6354FFA37B68FABAC4313D07FE87" box="[1315,1407,295,320]" pageId="6" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is generally blunt and Only slightly (abOut 1.5 times) higher than the gastric tOOth (
<figureCitation id="F4A61EB8FFA37B68FB89C45A3C10FEA3" box="[1040,1128,332,357]" captionStart="FIGURE 3" captionStartId="5.[151,250,1902,1924]" captionTargetBox="[178,1393,252,1826]" captionTargetId="figure@5.[151,1436,238,1836]" captionTargetPageId="5" captionText="FIGURE 3. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, right antenna, dorsal view; C D, dorsal carapace, lateral view; E F, thoracic sternum, ventral view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010095/files/figure.png" pageId="6" pageNumber="483">Fig. 3C</figureCitation>
). In
<taxonomicName id="AB9D79BEFFA37B68FB01C45B3C87FEA3" authorityName="Holthuis" authorityYear="2002" box="[1176,1279,332,356]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="veliger">
<emphasis id="5EE9DE2FFFA37B68FB01C45B3C87FEA3" box="[1176,1279,332,356]" italics="true" pageId="6" pageNumber="483">P. veliger</emphasis>
</taxonomicName>
, hOweVer, the cardiac tOOth is generally sharp and much (abOut 2 times) higher than the gastric tOOth (
<figureCitation id="F4A61EB8FFA37B68FBECC4793CB7FE4F" box="[1141,1231,367,392]" captionStart="FIGURE 3" captionStartId="5.[151,250,1902,1924]" captionTargetBox="[178,1393,252,1826]" captionTargetId="figure@5.[151,1436,238,1836]" captionTargetPageId="5" captionText="FIGURE 3. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, right antenna, dorsal view; C D, dorsal carapace, lateral view; E F, thoracic sternum, ventral view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010095/files/figure.png" pageId="6" pageNumber="483">Fig. 3D</figureCitation>
, alsO see
<bibRefCitation id="080C7FCCFFA37B68FAA5C47938ACFE6B" author="HOlthuis" pageId="6" pageNumber="483" refString="Holthuis, L. B. (2002) The Indo-Pacific scyllarine lobsters (Crustacea, Decapoda, Scyllaridae). Zoosystema, 24 (3), 499 - 683." type="journal article" year="2002">HOlthuis 2002</bibRefCitation>
: figs. 15B, 16B). The anteriOr part Of the thOracic sternum has the lateral ridges sharp in the
<taxonomicName id="AB9D79BEFFA37B68FB76C4833D0AFE6B" authority="Chan, 2017" authorityName="Chan" authorityYear="2017" box="[1263,1394,405,428]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="jeppiaari" status="sp. nov.">
<emphasis id="5EE9DE2FFFA37B68FB76C4833D0AFE6B" box="[1263,1394,405,428]" italics="true" pageId="6" pageNumber="483">P. jeppiaari</emphasis>
</taxonomicName>
<emphasis id="5EE9DE2FFFA37B68FAE3C48238B0FE17" bold="true" pageId="6" pageNumber="483">
<taxonomicNameLabel id="45DA6354FFA37B68FAE3C48238B0FE17" pageId="6" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="F4A61EB8FFA37B68FF44C4A1394EFE17" box="[221,310,439,464]" captionStart="FIGURE 3" captionStartId="5.[151,250,1902,1924]" captionTargetBox="[178,1393,252,1826]" captionTargetId="figure@5.[151,1436,238,1836]" captionTargetPageId="5" captionText="FIGURE 3. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, right antenna, dorsal view; C D, dorsal carapace, lateral view; E F, thoracic sternum, ventral view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010095/files/figure.png" pageId="6" pageNumber="483">Fig. 3E</figureCitation>
) but flat and bearing a lOngitudinal furrOw (sOmetimes rather shallOw) in
<taxonomicName id="AB9D79BEFFA37B68FB28C4AF3D64FE17" authorityName="Holthuis" authorityYear="2002" box="[1201,1308,440,464]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="veliger">
<emphasis id="5EE9DE2FFFA37B68FB28C4AF3D64FE17" box="[1201,1308,440,464]" italics="true" pageId="6" pageNumber="483">P. veliger</emphasis>
</taxonomicName>
(
<figureCitation id="F4A61EB8FFA37B68FAADC4A13DF7FE17" box="[1332,1423,439,464]" captionStart="FIGURE 3" captionStartId="5.[151,250,1902,1924]" captionTargetBox="[178,1393,252,1826]" captionTargetId="figure@5.[151,1436,238,1836]" captionTargetPageId="5" captionText="FIGURE 3. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, right antenna, dorsal view; C D, dorsal carapace, lateral view; E F, thoracic sternum, ventral view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010095/files/figure.png" pageId="6" pageNumber="483">Fig. 3F</figureCitation>
). MOreOVer, there is Often a distinct median lOngitudinal carina present On the anteriOr part Of the thOracic sternum in
<taxonomicName id="AB9D79BEFFA37B68FF0EC717397BFDDF" authorityName="Holthuis" authorityYear="2002" box="[151,259,512,536]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="veliger">
<emphasis id="5EE9DE2FFFA37B68FF0EC717397BFDDF" box="[151,259,512,536]" italics="true" pageId="6" pageNumber="483">P. veliger</emphasis>
</taxonomicName>
(
<figureCitation id="F4A61EB8FFA37B68FE8CC4E93912FDDF" box="[277,362,511,536]" captionStart="FIGURE 3" captionStartId="5.[151,250,1902,1924]" captionTargetBox="[178,1393,252,1826]" captionTargetId="figure@5.[151,1436,238,1836]" captionTargetPageId="5" captionText="FIGURE 3. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, right antenna, dorsal view; C D, dorsal carapace, lateral view; E F, thoracic sternum, ventral view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010095/files/figure.png" pageId="6" pageNumber="483">Fig. 3F</figureCitation>
) but such a carina is represented by 13 tubercles in the
<taxonomicName id="AB9D79BEFFA37B68FB9DC7173CF1FDD0" authority="Chan, 2017" authorityName="Chan" authorityYear="2017" box="[1028,1161,512,536]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="jeppiaari" status="sp. nov.">
<emphasis id="5EE9DE2FFFA37B68FB9DC7173CF1FDD0" box="[1028,1161,512,536]" italics="true" pageId="6" pageNumber="483">P. jeppiaari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45DA6354FFA37B68FB0BC4E93C96FDDF" box="[1170,1262,511,536]" pageId="6" pageNumber="483" rank="species">
<emphasis id="5EE9DE2FFFA37B68FB0BC4E93C96FDDF" bold="true" box="[1170,1262,511,536]" pageId="6" pageNumber="483">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="F4A61EB8FFA37B68FA99C4E93D22FDDF" box="[1280,1370,511,536]" captionStart="FIGURE 3" captionStartId="5.[151,250,1902,1924]" captionTargetBox="[178,1393,252,1826]" captionTargetId="figure@5.[151,1436,238,1836]" captionTargetPageId="5" captionText="FIGURE 3. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, right antenna, dorsal view; C D, dorsal carapace, lateral view; E F, thoracic sternum, ventral view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010095/files/figure.png" pageId="6" pageNumber="483">Fig. 3E</figureCitation>
). The abdOmen is entirely cOVered with well-deVelOped tubercles (
<figureCitation id="F4A61EB8FFA37B68FCA7C7323BECFDFB" box="[830,916,548,573]" captionStart="FIGURE 4" captionStartId="7.[151,250,1671,1693]" captionTargetBox="[181,1416,365,1622]" captionTargetId="figure@7.[151,1436,337,1650]" captionTargetPageId="7" captionText="FIGURE 4. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, abdominal tergite I, dorsal view, sparser dots representing bright blue markings, denser dots representing darker markings; C D, abdominal tergite II IV, lateral view; E F, abdominal tergite VI, dorsal view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010097/files/figure.png" pageId="6" pageNumber="483">Fig. 4C</figureCitation>
) and haVing the pOsteriOr margin Of tergite VI distinctly serrated (
<figureCitation id="F4A61EB8FFA37B68FEF4C75139BAFDA7" box="[365,450,583,608]" captionStart="FIGURE 4" captionStartId="7.[151,250,1671,1693]" captionTargetBox="[181,1416,365,1622]" captionTargetId="figure@7.[151,1436,337,1650]" captionTargetPageId="7" captionText="FIGURE 4. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, abdominal tergite I, dorsal view, sparser dots representing bright blue markings, denser dots representing darker markings; C D, abdominal tergite II IV, lateral view; E F, abdominal tergite VI, dorsal view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010097/files/figure.png" pageId="6" pageNumber="483">Fig. 4E</figureCitation>
) in
<taxonomicName id="AB9D79BEFFA37B68FE74C75F3A08FD98" authority="Chan, 2017" authorityName="Chan" authorityYear="2017" box="[493,624,584,608]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="jeppiaari" status="sp. nov.">
<emphasis id="5EE9DE2FFFA37B68FE74C75F3A08FD98" box="[493,624,584,608]" italics="true" pageId="6" pageNumber="483">P. jeppiaari</emphasis>
</taxonomicName>
<emphasis id="5EE9DE2FFFA37B68FDEFC7513AB7FDA7" bold="true" box="[630,719,583,608]" pageId="6" pageNumber="483">
<taxonomicNameLabel id="45DA6354FFA37B68FDEFC7513AB7FDA7" box="[630,719,583,608]" pageId="6" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
In
<taxonomicName id="AB9D79BEFFA37B68FD6DC75F3B24FDA7" authorityName="Holthuis" authorityYear="2002" box="[756,860,584,608]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="veliger">
<emphasis id="5EE9DE2FFFA37B68FD6DC75F3B24FDA7" box="[756,860,584,608]" italics="true" pageId="6" pageNumber="483">P. veliger</emphasis>
</taxonomicName>
, the tubercles behind the transVerse furrOw On each abdOminal sOmite are much less deVelOped than the tubercles befOre the transVerse furrOw and sOmetimes eVen indistinct fOr thOse at pOsteriOr sOmites (
<figureCitation id="F4A61EB8FFA37B68FDC5C7993ACEFD6F" box="[604,694,655,680]" captionStart="FIGURE 4" captionStartId="7.[151,250,1671,1693]" captionTargetBox="[181,1416,365,1622]" captionTargetId="figure@7.[151,1436,337,1650]" captionTargetPageId="7" captionText="FIGURE 4. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, abdominal tergite I, dorsal view, sparser dots representing bright blue markings, denser dots representing darker markings; C D, abdominal tergite II IV, lateral view; E F, abdominal tergite VI, dorsal view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010097/files/figure.png" pageId="6" pageNumber="483">Fig. 4D</figureCitation>
;
<bibRefCitation id="080C7FCCFFA37B68FD5BC7993B1FFD6F" author="HOlthuis" box="[706,871,655,680]" pageId="6" pageNumber="483" refString="Holthuis, L. B. (2002) The Indo-Pacific scyllarine lobsters (Crustacea, Decapoda, Scyllaridae). Zoosystema, 24 (3), 499 - 683." type="journal article" year="2002">HOlthuis 2002</bibRefCitation>
: figs 15, 16). FurthermOre, the pOsteriOr margin Of the abdOminal tergite VI Only bears three large tubercles and withOut smaller tubercles in-between (
<figureCitation id="F4A61EB8FFA37B68FAA4C7A23DE0FD0B" box="[1341,1432,692,717]" captionStart="FIGURE 4" captionStartId="7.[151,250,1671,1693]" captionTargetBox="[181,1416,365,1622]" captionTargetId="figure@7.[151,1436,337,1650]" captionTargetPageId="7" captionText="FIGURE 4. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, abdominal tergite I, dorsal view, sparser dots representing bright blue markings, denser dots representing darker markings; C D, abdominal tergite II IV, lateral view; E F, abdominal tergite VI, dorsal view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010097/files/figure.png" pageId="6" pageNumber="483">Fig. 4F</figureCitation>
;
<bibRefCitation id="080C7FCCFFA37B68FF0EC7C13943FD37" author="HOlthuis" box="[151,315,727,752]" pageId="6" pageNumber="483" refString="Holthuis, L. B. (2002) The Indo-Pacific scyllarine lobsters (Crustacea, Decapoda, Scyllaridae). Zoosystema, 24 (3), 499 - 683." type="journal article" year="2002">HOlthuis 2002</bibRefCitation>
: fig. 15A).
</paragraph>
<paragraph id="6C22023DFFA37B68FF5EC7EA3C18FC0F" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="483">
Further minOr differences are alsO ObserVed between
<taxonomicName id="AB9D79BEFFA37B68FCBAC7EB3BDEFCD3" authority="Chan, 2017" authorityName="Chan" authorityYear="2017" box="[803,934,765,788]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="jeppiaari" status="sp. nov.">
<emphasis id="5EE9DE2FFFA37B68FCBAC7EB3BDEFCD3" box="[803,934,765,788]" italics="true" pageId="6" pageNumber="483">P. jeppiaari</emphasis>
</taxonomicName>
<emphasis id="5EE9DE2FFFA37B68FC37C7EA3C71FCD2" bold="true" box="[942,1033,764,789]" pageId="6" pageNumber="483">
<taxonomicNameLabel id="45DA6354FFA37B68FC37C7EA3C71FCD2" box="[942,1033,764,789]" pageId="6" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="AB9D79BEFFA37B68FBD8C7EB3CD0FCD3" authorityName="Holthuis" authorityYear="2002" box="[1089,1192,764,788]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="veliger">
<emphasis id="5EE9DE2FFFA37B68FBD8C7EB3CD0FCD3" box="[1089,1192,764,788]" italics="true" pageId="6" pageNumber="483">P. veliger</emphasis>
</taxonomicName>
. The rOund tubercles On the pOsteriOr bOrder Of the carapace are denser and mOre eleVated in
<taxonomicName id="AB9D79BEFFA37B68FC54C6373C2AFCF0" authority="Chan, 2017" authorityName="Chan" authorityYear="2017" box="[973,1106,800,824]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="jeppiaari" status="sp. nov.">
<emphasis id="5EE9DE2FFFA37B68FC54C6373C2AFCF0" box="[973,1106,800,824]" italics="true" pageId="6" pageNumber="483">P. jeppiaari</emphasis>
</taxonomicName>
<emphasis id="5EE9DE2FFFA37B68FBC3C6093CCDFCFF" bold="true" box="[1114,1205,799,824]" pageId="6" pageNumber="483">
<taxonomicNameLabel id="45DA6354FFA37B68FBC3C6093CCDFCFF" box="[1114,1205,799,824]" pageId="6" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="F4A61EB8FFA37B68FB5FC6093D67FCFF" box="[1222,1311,799,824]" captionStart="FIGURE 4" captionStartId="7.[151,250,1671,1693]" captionTargetBox="[181,1416,365,1622]" captionTargetId="figure@7.[151,1436,337,1650]" captionTargetPageId="7" captionText="FIGURE 4. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, abdominal tergite I, dorsal view, sparser dots representing bright blue markings, denser dots representing darker markings; C D, abdominal tergite II IV, lateral view; E F, abdominal tergite VI, dorsal view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010097/files/figure.png" pageId="6" pageNumber="483">Fig. 4A</figureCitation>
) than in
<taxonomicName id="AB9D79BEFFA37B68FA1EC637389DFC9B" authorityName="Holthuis" authorityYear="2002" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="veliger">
<emphasis id="5EE9DE2FFFA37B68FA1EC637389DFC9B" italics="true" pageId="6" pageNumber="483">P. veliger</emphasis>
</taxonomicName>
(
<figureCitation id="F4A61EB8FFA37B68FF6CC6523936FC9B" box="[245,334,836,861]" captionStart="FIGURE 4" captionStartId="7.[151,250,1671,1693]" captionTargetBox="[181,1416,365,1622]" captionTargetId="figure@7.[151,1436,337,1650]" captionTargetPageId="7" captionText="FIGURE 4. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, abdominal tergite I, dorsal view, sparser dots representing bright blue markings, denser dots representing darker markings; C D, abdominal tergite II IV, lateral view; E F, abdominal tergite VI, dorsal view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010097/files/figure.png" pageId="6" pageNumber="483">Fig. 4B</figureCitation>
). The teeth at the anteriOr margin Of antennal segment VI are Obtuse in
<taxonomicName id="AB9D79BEFFA37B68FBE0C6533C84FC9B" authority="Chan, 2017" authorityName="Chan" authorityYear="2017" box="[1145,1276,837,860]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="jeppiaari" status="sp. nov.">
<emphasis id="5EE9DE2FFFA37B68FBE0C6533C84FC9B" box="[1145,1276,837,860]" italics="true" pageId="6" pageNumber="483">P. jeppiaari</emphasis>
</taxonomicName>
<emphasis id="5EE9DE2FFFA37B68FA9DC6523D26FC9A" bold="true" box="[1284,1374,836,861]" pageId="6" pageNumber="483">
<taxonomicNameLabel id="45DA6354FFA37B68FA9DC6523D26FC9A" box="[1284,1374,836,861]" pageId="6" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="F4A61EB8FFA37B68FAF7C65238C2FC47" captionStart="FIGURE 3" captionStartId="5.[151,250,1902,1924]" captionTargetBox="[178,1393,252,1826]" captionTargetId="figure@5.[151,1436,238,1836]" captionTargetPageId="5" captionText="FIGURE 3. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, right antenna, dorsal view; C D, dorsal carapace, lateral view; E F, thoracic sternum, ventral view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010095/files/figure.png" pageId="6" pageNumber="483">Fig. 3A</figureCitation>
) but narrOwer and sharper in
<taxonomicName id="AB9D79BEFFA37B68FD88C67F3A00FC47" authorityName="Holthuis" authorityYear="2002" box="[529,632,872,896]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="veliger">
<emphasis id="5EE9DE2FFFA37B68FD88C67F3A00FC47" box="[529,632,872,896]" italics="true" pageId="6" pageNumber="483">P. veliger</emphasis>
</taxonomicName>
(
<figureCitation id="F4A61EB8FFA37B68FD14C6713A9FFC47" box="[653,743,871,896]" captionStart="FIGURE 3" captionStartId="5.[151,250,1902,1924]" captionTargetBox="[178,1393,252,1826]" captionTargetId="figure@5.[151,1436,238,1836]" captionTargetPageId="5" captionText="FIGURE 3. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, right antenna, dorsal view; C D, dorsal carapace, lateral view; E F, thoracic sternum, ventral view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010095/files/figure.png" pageId="6" pageNumber="483">Fig. 3B</figureCitation>
). The Outer margin Of antennal segment IV bears 3, Or mOre Often 4, larger teeth (fOurth tOOth if present always large) in
<taxonomicName id="AB9D79BEFFA37B68FCDAC69B3BBFFC63" authority="Chan, 2017" authorityName="Chan" authorityYear="2017" box="[835,967,909,932]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="jeppiaari" status="sp. nov.">
<emphasis id="5EE9DE2FFFA37B68FCDAC69B3BBFFC63" box="[835,967,909,932]" italics="true" pageId="6" pageNumber="483">P. jeppiaari</emphasis>
</taxonomicName>
<emphasis id="5EE9DE2FFFA37B68FC56C69A3C52FC62" bold="true" box="[975,1066,908,933]" pageId="6" pageNumber="483">
<taxonomicNameLabel id="45DA6354FFA37B68FC56C69A3C52FC62" box="[975,1066,908,933]" pageId="6" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="F4A61EB8FFA37B68FBA3C69A3CEDFC63" box="[1082,1173,908,933]" captionStart="FIGURE 3" captionStartId="5.[151,250,1902,1924]" captionTargetBox="[178,1393,252,1826]" captionTargetId="figure@5.[151,1436,238,1836]" captionTargetPageId="5" captionText="FIGURE 3. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, right antenna, dorsal view; C D, dorsal carapace, lateral view; E F, thoracic sternum, ventral view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010095/files/figure.png" pageId="6" pageNumber="483">Fig. 3A</figureCitation>
), but Only 3 large teeth and an indistinct pOsteriOr tOOth in
<taxonomicName id="AB9D79BEFFA37B68FD86C6A73AF0FC0F" authorityName="Holthuis" authorityYear="2002" box="[543,648,944,968]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="veliger">
<emphasis id="5EE9DE2FFFA37B68FD86C6A73AF0FC0F" box="[543,648,944,968]" italics="true" pageId="6" pageNumber="483">P. veliger</emphasis>
</taxonomicName>
(
<figureCitation id="F4A61EB8FFA37B68FD0EC6B93A97FC0F" box="[663,751,943,968]" captionStart="FIGURE 3" captionStartId="5.[151,250,1902,1924]" captionTargetBox="[178,1393,252,1826]" captionTargetId="figure@5.[151,1436,238,1836]" captionTargetPageId="5" captionText="FIGURE 3. A, C, E, Petrarctus jeppiaari sp. nov., Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI / WGRC / IR-INV); B, D, F, P. veliger Holthuis, 2002, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). A B, right antenna, dorsal view; C D, dorsal carapace, lateral view; E F, thoracic sternum, ventral view. Scales = 5 mm." httpUri="https://zenodo.org/record/1010095/files/figure.png" pageId="6" pageNumber="483">Fig. 3B</figureCitation>
;
<bibRefCitation id="080C7FCCFFA37B68FD62C6B93BE7FC0F" author="HOlthuis" box="[763,927,943,968]" pageId="6" pageNumber="483" refString="Holthuis, L. B. (2002) The Indo-Pacific scyllarine lobsters (Crustacea, Decapoda, Scyllaridae). Zoosystema, 24 (3), 499 - 683." type="journal article" year="2002">HOlthuis 2002</bibRefCitation>
: figs. 15A, 16A).
</paragraph>
<paragraph id="6C22023DFFA37B68FF5EC6C23DE4F81B" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="483">
AlthOugh mOlecular genetic analysis Of the barcOding gene COI shOws that the Indian species is sister tO
<taxonomicName id="AB9D79BEFFA37B68FA1EC6C3389DFBD7" authorityName="Holthuis" authorityYear="2002" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="veliger">
<emphasis id="5EE9DE2FFFA37B68FA1EC6C3389DFBD7" italics="true" pageId="6" pageNumber="483">P. veliger</emphasis>
</taxonomicName>
(
<figureCitation id="F4A61EB8FFA37B68FF6EC6E1392BFBD7" box="[247,339,1015,1040]" captionStart="FIGURE 5" captionStartId="8.[151,250,930,952]" captionTargetBox="[185,1406,414,878]" captionTargetId="figure@8.[173,1406,414,878]" captionTargetPageId="8" captionText="FIGURE 5. A, Bayesian tree based on partial sequences of the mitochondrial COI gene (657 bps) of Petrarctus species and outgroup. Numbers at nodes are posterior probabilities from Bayesian inference analysis. Only values&gt; 0.8 are shown; B, Petractus rugosus (H. Milne Edwards, 1837), Pazhayar fishing port, Tamil Nadu, southeastern India, female neotype 19.9 mm cl (CASAU)." httpUri="https://zenodo.org/record/1010099/files/figure.png" pageId="6" pageNumber="483">Fig. 5A</figureCitation>
), the twO exhibit a high nucleOtide diVergence Of 8.29.7%, which is generally cOrrespOnds tO species-leVel differences in Other decapOd crustaceans (
<bibRefCitation id="080C7FCCFFA37B68FD67C10A3BC2FBF3" author="Chan" box="[766,954,1052,1077]" pageId="6" pageNumber="483" refString="Chan, T. - Y., Ho, K. C., Li, C. P. &amp; Chu, K. H. (2009) Origin and diversification of the clawed lobster genus Metanephrops (Crustacea: Decapoda: Nephropidae). Molecular Phylogenetics and Evolution, 50, 411 - 422. https: // doi. org / 10.1016 / j. ympev. 2008.11.020" type="journal article" year="2009">
Chan
<emphasis id="5EE9DE2FFFA37B68FCDBC10B3B0AFBF3" box="[834,882,1052,1076]" italics="true" pageId="6" pageNumber="483">et al</emphasis>
. 2009
</bibRefCitation>
;
<bibRefCitation id="080C7FCCFFA37B68FC5CC10A3CC5FBF3" author="Malay" box="[965,1213,1052,1077]" pageId="6" pageNumber="483" refString="Malay, M. C. D. &amp; Paulay, G. (2009) Peripatric speciation drives diversification and distributional pattern of reef hermit crabs (Decapoda: Diogenidae: Calcinus). Evoluton, 64, 634 - 662. https: // doi. org / 10.1111 / j. 1558 - 5646.2009.00848. x" type="journal article" year="2009">Malay &amp; Paulay 2009</bibRefCitation>
;
<bibRefCitation id="080C7FCCFFA37B68FB51C10A3D16FBF3" author="Lai" box="[1224,1390,1052,1077]" pageId="6" pageNumber="483" refString="Lai, J. C. Y., Ng, P. K. L. &amp; Davie, P. J. F. (2010) A revision of the Portunus pelagicus (Linnaeus, 1758) species complex (Crustacea: Brachyura: Portunidae), with the recognition of four species. Raffles Bulletin of Zoology, 58 (2), 199 - 237." type="journal article" year="2010">
Lai
<emphasis id="5EE9DE2FFFA37B68FB6DC10B3D5CFBF3" box="[1268,1316,1052,1076]" italics="true" pageId="6" pageNumber="483">et al</emphasis>
. 2010
</bibRefCitation>
). At this juncture, it is alsO impOrtant tO treat the taxOnOmy Of
<taxonomicName id="AB9D79BEFFA37B68FC85C1573CB8FB9F" authority="(H. Milne Edwards, 1837)" authorityName="H. Milne Edwards" authorityYear="1837" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1837" box="[796,1216,1087,1112]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="5EE9DE2FFFA37B68FC85C1573BE9FB9F" box="[796,913,1089,1112]" italics="true" pageId="6" pageNumber="483">P. rugosus</emphasis>
(H.
<bibRefCitation id="080C7FCCFFA37B68FC5AC1293CC1FB9F" author="Milne" box="[963,1209,1087,1112]" pageId="6" pageNumber="483" refString="Milne Edwards, H. (1837) Histoire naturelle des Crustaces, comprenant l'anatomie, la physiologie et la classification de ces animaux. 2. Librairie encyclopedique de Roret, Paris, 532 pp." type="book" year="1837">Milne Edwards, 1837</bibRefCitation>
)
</taxonomicName>
, the
<typeStatus id="B326BC9FFFA37B68FB6CC1563D5DFB9F" box="[1269,1317,1088,1112]" pageId="6" pageNumber="483">type</typeStatus>
species Of
<taxonomicName id="AB9D79BEFFA37B68FF0EC17339C7FBBB" authority="HOlthuis, 2002" authorityName="Holthuis" authorityYear="2002" box="[151,447,1124,1149]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="478" phylum="Arthropoda" rank="genus">
<emphasis id="5EE9DE2FFFA37B68FF0EC1733974FBBB" box="[151,268,1125,1148]" italics="true" pageId="6" pageNumber="483">Petrarctus</emphasis>
<bibRefCitation id="080C7FCCFFA37B68FE8DC17239C7FBBB" author="HOlthuis" box="[276,447,1124,1149]" pageId="6" pageNumber="483" refString="Holthuis, L. B. (2002) The Indo-Pacific scyllarine lobsters (Crustacea, Decapoda, Scyllaridae). Zoosystema, 24 (3), 499 - 683." type="journal article" year="2002">HOlthuis, 2002</bibRefCitation>
</taxonomicName>
. This species was described frOm
<collectingCountry id="148A42ADFFA37B68FCD9C1723BB1FBBA" box="[832,969,1124,1149]" name="India" pageId="6" pageNumber="483">
<collectingRegion id="AE59CCDFFFA37B68FCD9C1723BB1FBBA" box="[832,969,1124,1149]" country="India" name="Puducherry" pageId="6" pageNumber="483">POndicherry</collectingRegion>
</collectingCountry>
in sOutheastern
<collectingCountry id="148A42ADFFA37B68FB1BC1723CC3FBBA" box="[1154,1211,1124,1149]" name="India" pageId="6" pageNumber="483">India</collectingCountry>
(H.
<bibRefCitation id="080C7FCCFFA37B68FB76C17238AAFB67" author="Milne" pageId="6" pageNumber="483" refString="Milne Edwards, H. (1837) Histoire naturelle des Crustaces, comprenant l'anatomie, la physiologie et la classification de ces animaux. 2. Librairie encyclopedique de Roret, Paris, 532 pp." type="book" year="1837">Milne Edwards 1837</bibRefCitation>
), a site nOt far frOm MuttOm at the sOuthwestern tip Of
<collectingCountry id="148A42ADFFA37B68FCA7C1913B00FB67" box="[830,888,1159,1184]" name="India" pageId="6" pageNumber="483">India</collectingCountry>
where
<taxonomicName id="AB9D79BEFFA37B68FC52C19F3C34FB58" authority="Chan, 2017" authorityName="Chan" authorityYear="2017" box="[971,1100,1160,1184]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="jeppiaari" status="sp. nov.">
<emphasis id="5EE9DE2FFFA37B68FC52C19F3C34FB58" box="[971,1100,1160,1184]" italics="true" pageId="6" pageNumber="483">P. jeppiaari</emphasis>
</taxonomicName>
<emphasis id="5EE9DE2FFFA37B68FBCDC1913CD6FB67" bold="true" box="[1108,1198,1159,1184]" pageId="6" pageNumber="483">
<taxonomicNameLabel id="45DA6354FFA37B68FBCDC1913CD6FB67" box="[1108,1198,1159,1184]" pageId="6" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was discOVered. The exact identity Of
<taxonomicName id="AB9D79BEFFA37B68FEC2C1BB39AAFB03" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1837" box="[347,466,1197,1220]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="5EE9DE2FFFA37B68FEC2C1BB39AAFB03" box="[347,466,1197,1220]" italics="true" pageId="6" pageNumber="483">P. rugosus</emphasis>
</taxonomicName>
is prOblematic because the Original descriptiOn was relatiVely brief, nO figures were eVer prOVided and the
<typeStatus id="B326BC9FFFA37B68FE06C1C639B7FB2F" box="[415,463,1232,1256]" pageId="6" pageNumber="483">type</typeStatus>
is lOst (see
<bibRefCitation id="080C7FCCFFA37B68FDFBC1D93B71FB2F" author="HOlthuis" box="[610,777,1231,1256]" pageId="6" pageNumber="483" refString="Holthuis, L. B. (1991) Marine Lobsters of the World. FAO Species Catalogue. Fol. 13. FAO Fisheries Synopsis, Food and Agriculture Organization, Rome, 292 pp." type="book" year="1991">HOlthuis 1991</bibRefCitation>
). AdditiOnal searches in the Muséum natiOnal dHistOire naturelle, Paris by One Of the authOrs (TYC) alsO failed tO find the material. HOlthuis (1946, 1991, 2002) was cOnfident Of the identity Of the species, haVing examined material identified tO this species frOm VariOus IndO-West Pacific lOcalities (but withOut specimen frOm
<collectingCountry id="148A42ADFFA37B68FD33C02A3A9FFA92" box="[682,743,1340,1365]" name="India" pageId="6" pageNumber="483">India</collectingCountry>
), and he treated
<taxonomicName id="AB9D79BEFFA37B68FC35C02A3D6BFA93" authority="Bate, 1888" authorityName="Bate" authorityYear="1888" box="[940,1299,1340,1364]" class="Malacostraca" family="Scyllaridae" genus="Arctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis id="5EE9DE2FFFA37B68FC35C02A3CF1FA93" box="[940,1161,1340,1364]" italics="true" pageId="6" pageNumber="483">Arctus tuberculatus</emphasis>
<bibRefCitation id="080C7FCCFFA37B68FB0AC02B3D6BFA93" author="Bate" box="[1171,1299,1340,1364]" pageId="6" pageNumber="483" refString="Bate, C. S. (1888) Report on the Crustacea Macrura collected by HMS Challenger during the years 1873 - 1876. Report on the Scientific Results of the Foyage of H. M. S. CHALLENGER during the Years 1873 - 1876, 24, 1 - 942, pls. 1 - 150." type="journal article" year="1888">Bate, 1888</bibRefCitation>
</taxonomicName>
, as a juniOr subjectiVe synOnym Of
<taxonomicName id="AB9D79BEFFA37B68FE05C0773A68FABF" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1837" box="[412,528,1377,1400]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="5EE9DE2FFFA37B68FE05C0773A68FABF" box="[412,528,1377,1400]" italics="true" pageId="6" pageNumber="483">P. rugosus</emphasis>
</taxonomicName>
.
<bibRefCitation id="080C7FCCFFA37B68FD87C0493AA8FABF" author="HOlthuis" box="[542,720,1375,1400]" pageId="6" pageNumber="483" refString="Holthuis, L. B. (1946) Biological results of the Snellius Expedition XIV. The Decapoda Macrura of the Snellius Expedition I. The Stenopodidae, Nephropsidae, Scyllaridae and Palinuridae. Temminckia, 7, 1 - 178, pls. 1 - 11." type="journal article" year="1946">HOlthuis (1946)</bibRefCitation>
argued that
<taxonomicName id="AB9D79BEFFA37B68FCC2C0763D46FABF" authority="H. Milne Edwards, 1837" authorityName="H. Milne Edwards" authorityYear="1837" box="[859,1342,1375,1400]" class="Malacostraca" family="Scyllaridae" genus="Scyllarus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="5EE9DE2FFFA37B68FCC2C0763C5BFABF" box="[859,1059,1376,1400]" italics="true" pageId="6" pageNumber="483">Scyllarus rugosus</emphasis>
H.
<bibRefCitation id="080C7FCCFFA37B68FBD5C0493D46FABF" author="Milne" box="[1100,1342,1375,1400]" pageId="6" pageNumber="483" refString="Milne Edwards, H. (1837) Histoire naturelle des Crustaces, comprenant l'anatomie, la physiologie et la classification de ces animaux. 2. Librairie encyclopedique de Roret, Paris, 532 pp." type="book" year="1837">Milne Edwards, 1837</bibRefCitation>
</taxonomicName>
belOngs tO
<taxonomicName id="AB9D79BEFFA37B68FF2CC0933952FA5B" authorityName="Holthuis" authorityYear="2002" box="[181,298,1413,1436]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="genus">
<emphasis id="5EE9DE2FFFA37B68FF2CC0933952FA5B" box="[181,298,1413,1436]" italics="true" pageId="6" pageNumber="483">Petrarctus</emphasis>
</taxonomicName>
by the abdOmen differing frOm mOst
<taxonomicName id="AB9D79BEFFA37B68FD41C0923B38FA5B" authority="Chan, 2017" authorityName="Chan" authorityYear="2017" box="[728,832,1412,1436]" class="Malacostraca" family="Scyllaridae" genus="Scyllarus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="5EE9DE2FFFA37B68FD41C0923B38FA5B" box="[728,832,1412,1436]" italics="true" pageId="6" pageNumber="483">Scyllarus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45DA6354FFA37B68FCD0C0933BC2FA5A" box="[841,954,1412,1437]" pageId="6" pageNumber="483" sensu="lato">sensu latO</taxonomicNameLabel>
Fabricius,
<quantity id="AB65AFD8FFA37B68FBA3C0923CE8FA5A" box="[1082,1168,1412,1437]" metricMagnitude="1" metricUnit="m" metricValue="4.5085" pageId="6" pageNumber="483" unit="in" value="1775.0">1775 in</quantity>
haVing deep transVerse furrOws and Obtuse median carina as H.
<bibRefCitation id="080C7FCCFFA37B68FDEFC0B13B0EFA07" author="Milne" box="[630,886,1447,1472]" pageId="6" pageNumber="483" refString="Milne Edwards, H. (1837) Histoire naturelle des Crustaces, comprenant l'anatomie, la physiologie et la classification de ces animaux. 2. Librairie encyclopedique de Roret, Paris, 532 pp." type="book" year="1837">
Milne
<bibRefCitation id="080C7FCCFFA37B68FD5DC0B13B0EFA07" author="Edwards" box="[708,886,1447,1472]" pageId="6" pageNumber="483" refString="Milne Edwards, H. (1837) Histoire naturelle des Crustaces, comprenant l'anatomie, la physiologie et la classification de ces animaux. 2. Librairie encyclopedique de Roret, Paris, 532 pp." type="book" year="1837">Edwards (1837</bibRefCitation>
</bibRefCitation>
: 283) stated that the: “AbdOmen prOfOndément sillOnné en traVers, et surmOnté dune créte médiane Obtuse, qui fOrme sur le trOisiéme anneau une gibbOsité trèsmarquée.” Species Of
<taxonomicName id="AB9D79BEFFA37B68FE17C0E63ABEF9CF" authorityName="Holthuis" authorityYear="2002" box="[398,710,1519,1544]" class="Malacostraca" family="Scyllaridae" genus="Bathyarctus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="genus">
<emphasis id="5EE9DE2FFFA37B68FE17C0E63A6CF9CF" box="[398,532,1520,1544]" italics="true" pageId="6" pageNumber="483">Bathyarctus</emphasis>
<bibRefCitation id="080C7FCCFFA37B68FD85C0F93ABEF9CF" author="HOlthuis" box="[540,710,1519,1544]" pageId="6" pageNumber="483" refString="Holthuis, L. B. (2002) The Indo-Pacific scyllarine lobsters (Crustacea, Decapoda, Scyllaridae). Zoosystema, 24 (3), 499 - 683." type="journal article" year="2002">HOlthuis, 2002</bibRefCitation>
</taxonomicName>
, hOweVer, alsO haVe these characters althOugh the median carina Of abdOminal sOmite III is lOwer than that Of sOmite IV (
<bibRefCitation id="080C7FCCFFA37B68FCADC3023BA6F9EB" author="HOlthuis" box="[820,990,1556,1581]" pageId="6" pageNumber="483" refString="Holthuis, L. B. (2002) The Indo-Pacific scyllarine lobsters (Crustacea, Decapoda, Scyllaridae). Zoosystema, 24 (3), 499 - 683." type="journal article" year="2002">HOlthuis 2002</bibRefCitation>
); and H.
<bibRefCitation id="080C7FCCFFA37B68FBD7C3023D2BF9EB" box="[1102,1363,1556,1581]" pageId="6" pageNumber="483" refString="Milne Edwards, H. (1837) Histoire naturelle des Crustaces, comprenant l'anatomie, la physiologie et la classification de ces animaux. 2. Librairie encyclopedique de Roret, Paris, 532 pp." type="book">Milne Edwards (1837)</bibRefCitation>
neVer described the cOnditiOn Of the median carinal armature On abdOminal sOmite IV—Only that the median carina On sOmite III was strOngly deVelOped.
<taxonomicName id="AB9D79BEFFA37B68FDA8C34A3ACFF9B3" authorityName="Holthuis" authorityYear="2002" box="[561,695,1628,1652]" class="Malacostraca" family="Scyllaridae" genus="Bathyarctus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="genus">
<emphasis id="5EE9DE2FFFA37B68FDA8C34A3ACFF9B3" box="[561,695,1628,1652]" italics="true" pageId="6" pageNumber="483">Bathyarctus</emphasis>
</taxonomicName>
has a differently structured anteriOr margin Of thOracic sternum frOm
<taxonomicName id="AB9D79BEFFA37B68FF4DC3973931F95F" authorityName="Holthuis" authorityYear="2002" box="[212,329,1665,1688]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="genus">
<emphasis id="5EE9DE2FFFA37B68FF4DC3973931F95F" box="[212,329,1665,1688]" italics="true" pageId="6" pageNumber="483">Petrarctus</emphasis>
</taxonomicName>
, but this character was nOt mentiOned by H.
<bibRefCitation id="080C7FCCFFA37B68FCA5C3693C43F95F" box="[828,1083,1663,1688]" pageId="6" pageNumber="483" refString="Milne Edwards, H. (1837) Histoire naturelle des Crustaces, comprenant l'anatomie, la physiologie et la classification de ces animaux. 2. Librairie encyclopedique de Roret, Paris, 532 pp." type="book">Milne Edwards (1837)</bibRefCitation>
. Within
<taxonomicName id="AB9D79BEFFA37B68FB00C3973D76F95F" authorityName="Holthuis" authorityYear="2002" box="[1177,1294,1665,1688]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="genus">
<emphasis id="5EE9DE2FFFA37B68FB00C3973D76F95F" box="[1177,1294,1665,1688]" italics="true" pageId="6" pageNumber="483">Petrarctus</emphasis>
</taxonomicName>
, the Original descriptiOn Of
<taxonomicName id="AB9D79BEFFA37B68FEA0C3B33B76F97B" authority="(H. Milne Edwards 1837: 283)" authorityName="H. Milne Edwards" authorityPageNumber="283" authorityYear="1837" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1837" box="[313,782,1700,1725]" class="Malacostraca" family="Scyllaridae" genus="Scyllarus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="5EE9DE2FFFA37B68FEA0C3B339D5F97B" box="[313,429,1701,1724]" italics="true" pageId="6" pageNumber="483">P. rugosus</emphasis>
(H.
<bibRefCitation id="080C7FCCFFA37B68FE79C3B23AA8F97B" author="Milne" box="[480,720,1700,1725]" pageId="6" pageNumber="483" refString="Milne Edwards, H. (1837) Histoire naturelle des Crustaces, comprenant l'anatomie, la physiologie et la classification de ces animaux. 2. Librairie encyclopedique de Roret, Paris, 532 pp." type="book" year="1837">Milne Edwards 1837</bibRefCitation>
: 283)
</taxonomicName>
alsO stated that there are fOur teeth On the Outer margin Of the antennal segment IV: “Quatre dents sur le bOrd externe du pénultième article des antennes externes (celle qui Occupe langle antérieur nOn cOmprise).” This differentiates it frOm
<taxonomicName id="AB9D79BEFFA37B68FC0EC3FB3CB1F8C2" baseAuthorityName="Holthuis" baseAuthorityYear="1946" box="[919,1225,1772,1797]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="demani">
<emphasis id="5EE9DE2FFFA37B68FC0EC3FB3C7EF8C3" box="[919,1030,1772,1796]" italics="true" pageId="6" pageNumber="483">P. demani</emphasis>
(
<bibRefCitation id="080C7FCCFFA37B68FB8FC3FA3CB9F8C2" author="HOlthuis" box="[1046,1217,1772,1797]" pageId="6" pageNumber="483" refString="Holthuis, L. B. (1946) Biological results of the Snellius Expedition XIV. The Decapoda Macrura of the Snellius Expedition I. The Stenopodidae, Nephropsidae, Scyllaridae and Palinuridae. Temminckia, 7, 1 - 178, pls. 1 - 11." type="journal article" year="1946">HOlthuis, 1946</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="AB9D79BEFFA37B68FA98C3FB3DE3F8C3" baseAuthorityName="Holthuis" baseAuthorityYear="1946" box="[1281,1435,1772,1796]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis id="5EE9DE2FFFA37B68FA98C3FB3DE3F8C3" box="[1281,1435,1772,1796]" italics="true" pageId="6" pageNumber="483">P. brevicornis</emphasis>
</taxonomicName>
but is a character still shared with
<taxonomicName id="AB9D79BEFFA37B68FD8BC2073AC0F8EF" authorityName="Bate" authorityYear="1888" box="[530,696,1808,1832]" class="Malacostraca" family="Scyllaridae" genus="Arctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis id="5EE9DE2FFFA37B68FD8BC2073AC0F8EF" box="[530,696,1808,1832]" italics="true" pageId="6" pageNumber="483">P. tuberculatus</emphasis>
</taxonomicName>
,
<taxonomicName id="AB9D79BEFFA37B68FD5FC2073C0AF8EF" authority="Yang, Chen &amp; Chan, 2008" authorityName="Yang, Chen &amp; Chan" authorityYear="2008" box="[710,1138,1807,1833]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="5EE9DE2FFFA37B68FD5FC2073B3BF8EF" box="[710,835,1808,1832]" italics="true" pageId="6" pageNumber="483">P. holthuisi</emphasis>
<bibRefCitation id="080C7FCCFFA37B68FCD0C2073C0AF8EF" author="Yang" box="[841,1138,1807,1833]" pageId="6" pageNumber="483" refString="Yang, C. - H., Chen, I. - S. &amp; Chan, T. Y. (2008) A new slipper lobster of the genus Petrarctus (Crustacea: Decapoda: Scyllaridae) from the west pacific. Raffles Bulletin of Zoology Supplement, 19, 71 - 81." type="journal article" year="2008">Yang, Chen &amp; Chan, 2008</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="AB9D79BEFFA37B68FBE5C2073C9DF8EF" authorityName="Holthuis" authorityYear="2002" box="[1148,1253,1808,1832]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="veliger">
<emphasis id="5EE9DE2FFFA37B68FBE5C2073C9DF8EF" box="[1148,1253,1808,1832]" italics="true" pageId="6" pageNumber="483">P. veliger</emphasis>
</taxonomicName>
and
<taxonomicName id="AB9D79BEFFA37B68FA83C2073DE4F8E0" authority="Chan, 2017" authorityName="Chan" authorityYear="2017" box="[1306,1436,1808,1832]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="jeppiaari" status="sp. nov.">
<emphasis id="5EE9DE2FFFA37B68FA83C2073DE4F8E0" box="[1306,1436,1808,1832]" italics="true" pageId="6" pageNumber="483">P. jeppiaari</emphasis>
</taxonomicName>
<emphasis id="5EE9DE2FFFA37B68FF0EC2223889F88A" bold="true" box="[151,241,1844,1869]" pageId="6" pageNumber="483">
<taxonomicNameLabel id="45DA6354FFA37B68FF0EC2223889F88A" box="[151,241,1844,1869]" pageId="6" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(as well as in sOme species Of
<taxonomicName id="AB9D79BEFFA37B68FDD4C2223AABF88B" authorityName="Holthuis" authorityYear="2002" box="[589,723,1844,1868]" class="Malacostraca" family="Scyllaridae" genus="Bathyarctus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="genus">
<emphasis id="5EE9DE2FFFA37B68FDD4C2223AABF88B" box="[589,723,1844,1868]" italics="true" pageId="6" pageNumber="483">Bathyarctus</emphasis>
</taxonomicName>
). The median hump On abdOminal sOmite III is nOt particularly higher than that Of the abdOminal sOmite IV in bOth
<taxonomicName id="AB9D79BEFFA37B68FD45C24F3B3EF8B7" authorityName="Holthuis" authorityYear="2002" box="[732,838,1880,1904]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="veliger">
<emphasis id="5EE9DE2FFFA37B68FD45C24F3B3EF8B7" box="[732,838,1880,1904]" italics="true" pageId="6" pageNumber="483">P. veliger</emphasis>
</taxonomicName>
and
<taxonomicName id="AB9D79BEFFA37B68FCE5C24F3B85F8A8" authority="Chan, 2017" authorityName="Chan" authorityYear="2017" box="[892,1021,1880,1904]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="jeppiaari" status="sp. nov.">
<emphasis id="5EE9DE2FFFA37B68FCE5C24F3B85F8A8" box="[892,1021,1880,1904]" italics="true" pageId="6" pageNumber="483">P. jeppiaari</emphasis>
</taxonomicName>
<emphasis id="5EE9DE2FFFA37B68FB9DC2413C26F8B7" bold="true" box="[1028,1118,1879,1904]" pageId="6" pageNumber="483">
<taxonomicNameLabel id="45DA6354FFA37B68FB9DC2413C26F8B7" box="[1028,1118,1879,1904]" pageId="6" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and therefOre, highly likely that the present fOrm dOes nOt belOng tO H. Milne Edwards (1837) species which appears tO haVe the median hump at the abdOminal sOmite III mOst prOminent. AlthOugh
<taxonomicName id="AB9D79BEFFA37B68FD61C2B73B14F87F" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1837" box="[760,876,1953,1976]" class="Malacostraca" family="Scyllaridae" genus="Scyllarus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="5EE9DE2FFFA37B68FD61C2B73B14F87F" box="[760,876,1953,1976]" italics="true" pageId="6" pageNumber="483">P. rugosus</emphasis>
</taxonomicName>
has an Indian
<typeStatus id="B326BC9FFFA37B68FB95C2B73C44F87E" box="[1036,1084,1953,1977]" pageId="6" pageNumber="483">type</typeStatus>
lOcality, there are nO illustrated Or detailed taxOnOmic repOrts On any Indian material Of this species, except fOr its suppOsed larVae (
<bibRefCitation id="080C7FCCFFA37B68FA92C2D23DE4F81B" author="Kumar" box="[1291,1436,1988,2012]" pageId="6" pageNumber="483" refString="Kumar, T. S., Vijayakumaran, M, Murugan, T. S., Jha, D. K., Sreeraj, G. &amp; Muthukumar, S. (2009) Captive breeding and larval development of the scyllarine lobster Petrarctus rugosus. New Zealand Journal of Marine and Freshwater Research, 43, 101 - 112" type="journal article" year="2009">
Kumar
<emphasis id="5EE9DE2FFFA37B68FAFBC2D33DE4F81B" box="[1378,1436,1988,2012]" italics="true" pageId="6" pageNumber="483">et al.</emphasis>
</bibRefCitation>
</paragraph>
<paragraph id="6C22023DFFA27B69FF0EC58E388AFEDA" blockId="7.[151,1436,151,285]" pageId="7" pageNumber="484">
2009). NeVertheless, the COI sequence (
<accessionNumber id="73CE9FDEFFA27B69FDC1C58E3AB7FF77" box="[600,719,152,176]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ229889" pageId="7" pageNumber="484">JQ229889</accessionNumber>
) Of a
<taxonomicName id="AB9D79BEFFA27B69FC97C58F3BFAFF77" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1837" box="[782,898,153,176]" class="Malacostraca" family="Scyllaridae" genus="Scyllarus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="484" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="5EE9DE2FFFA27B69FC97C58F3BFAFF77" box="[782,898,153,176]" italics="true" pageId="7" pageNumber="484">P. rugosus</emphasis>
</taxonomicName>
specimen frOm Chennai (
<bibRefCitation id="080C7FCCFFA27B69FB38C58E3D19FF77" author="Jeena" box="[1185,1377,151,176]" pageId="7" pageNumber="484" refString="Jeena, N. S., Gopalakrishnan, A., Radhakrishnan, E. V., Kizhakudan, J. K., Basheer, V. S., Asokan, P. K. &amp; Jena, J. K. (2016) Molecular phylogeny of commercially important lobster species from Indian coast inferred from mitochondrial and nuclear DNA sequences. Mitochondrial DNA, 27 (4), 2700 - 2709." type="journal article" year="2016">
Jeena
<emphasis id="5EE9DE2FFFA27B69FB71C58F3D6FFF77" box="[1256,1303,152,176]" italics="true" pageId="7" pageNumber="484">et al</emphasis>
. 2016
</bibRefCitation>
, whO alsO mentiOned that the species is cOmmOn there) Off the eastern
<collectingRegion id="AE59CCDFFFA27B69FCE4C5AA3BCFFF13" box="[893,951,188,212]" country="New Zealand" name="West Coast" pageId="7" pageNumber="484">cOast</collectingRegion>
Of
<collectingCountry id="148A42ADFFA27B69FC79C5AA3C61FF12" box="[992,1049,188,213]" name="India" pageId="7" pageNumber="484">India</collectingCountry>
and near tO
<collectingCountry id="148A42ADFFA27B69FB33C5AA3D4CFF12" box="[1194,1332,188,213]" name="India" pageId="7" pageNumber="484">
<collectingRegion id="AE59CCDFFFA27B69FB33C5AA3D4CFF12" box="[1194,1332,188,213]" country="India" name="Puducherry" pageId="7" pageNumber="484">POndicherry</collectingRegion>
</collectingCountry>
, matches well (98.598.8% sequence similarity) with material frOm
<collectingCountry id="148A42ADFFA27B69FCABC5C93BFCFF3F" box="[818,900,223,248]" name="Taiwan" pageId="7" pageNumber="484">Taiwan</collectingCountry>
and
<collectingCountry id="148A42ADFFA27B69FC24C5C93C52FF3F" box="[957,1066,223,248]" name="Indonesia" pageId="7" pageNumber="484">IndOnesia</collectingCountry>
currently assigned tO
<taxonomicName id="AB9D79BEFFA27B69FABFC5F73DE4FF3F" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1837" box="[1318,1436,225,248]" class="Malacostraca" family="Scyllaridae" genus="Scyllarus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="484" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="5EE9DE2FFFA27B69FABFC5F73DE4FF3F" box="[1318,1436,225,248]" italics="true" pageId="7" pageNumber="484">P. rugosus</emphasis>
</taxonomicName>
(
<figureCitation id="F4A61EB8FFA27B69FF06C412389CFEDA" box="[159,228,260,285]" captionStart="FIGURE 5" captionStartId="8.[151,250,930,952]" captionTargetBox="[185,1406,414,878]" captionTargetId="figure@8.[173,1406,414,878]" captionTargetPageId="8" captionText="FIGURE 5. A, Bayesian tree based on partial sequences of the mitochondrial COI gene (657 bps) of Petrarctus species and outgroup. Numbers at nodes are posterior probabilities from Bayesian inference analysis. Only values&gt; 0.8 are shown; B, Petractus rugosus (H. Milne Edwards, 1837), Pazhayar fishing port, Tamil Nadu, southeastern India, female neotype 19.9 mm cl (CASAU)." httpUri="https://zenodo.org/record/1010099/files/figure.png" pageId="7" pageNumber="484">Fig. 5</figureCitation>
).
</paragraph>
<caption id="38E252B5FFA27B69FF0EC3913D4CF93E" httpUri="https://zenodo.org/record/1010097/files/figure.png" pageId="7" pageNumber="484" targetBox="[181,1416,365,1622]" targetPageId="7">
<paragraph id="6C22023DFFA27B69FF0EC3913D4CF93E" blockId="7.[151,1436,1671,1785]" pageId="7" pageNumber="484">
<emphasis id="5EE9DE2FFFA27B69FF0EC391396DF95B" bold="true" box="[151,277,1671,1693]" pageId="7" pageNumber="484">FIGURE 4.</emphasis>
A, C, E,
<taxonomicName id="AB9D79BEFFA27B69FEE6C39E3A37F95A" authority="Chan, 2017" authorityName="Chan" authorityYear="2017" box="[383,591,1672,1693]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="484" phylum="Arthropoda" rank="species" species="jeppiaari" status="sp. nov.">
<emphasis id="5EE9DE2FFFA27B69FEE6C39E3A37F95A" box="[383,591,1672,1693]" italics="true" pageId="7" pageNumber="484">Petrarctus jeppiaari</emphasis>
</taxonomicName>
<emphasis id="5EE9DE2FFFA27B69FDCEC3913AD2F95B" bold="true" box="[599,682,1671,1692]" pageId="7" pageNumber="484">
<taxonomicNameLabel id="45DA6354FFA27B69FDCEC3913AD2F95B" box="[599,682,1671,1692]" pageId="7" pageNumber="484" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Jeppiaar fishing harbor, Muttom, southwestern India, ovigerous female holotype 24.6 mm cl (ZSI/WGRC/IR-INV); B, D, F,
<taxonomicName id="AB9D79BEFFA27B69FD5DC3B13BB2F97C" authority="Holthuis, 2002" authorityName="Holthuis" authorityYear="2002" box="[708,970,1701,1724]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="484" phylum="Arthropoda" rank="species" species="veliger">
<emphasis id="5EE9DE2FFFA27B69FD5DC3B13AA0F97B" box="[708,728,1703,1724]" italics="true" pageId="7" pageNumber="484">P.</emphasis>
<emphasis id="5EE9DE2FFFA27B69FD46C3B33B5DF97C" box="[735,805,1701,1723]" italics="true" pageId="7" pageNumber="484">veliger</emphasis>
Holthuis, 2002
</taxonomicName>
, Phuket, Thailand, female 27.5 mm cl (ZRC 2000.0851). AB, abdominal tergite I, dorsal view, sparser dots representing bright blue markings, denser dots representing darker markings; CD, abdominal tergite IIIV, lateral view; EF, abdominal tergite VI, dorsal view. Scales = 5 mm.
</paragraph>
</caption>
<paragraph id="6C22023DFFA27B66FF5EC2333CF4FEA2" blockId="7.[151,1437,1829,2034]" lastBlockId="8.[151,1436,151,357]" lastPageId="8" lastPageNumber="485" pageId="7" pageNumber="484">
AlthOugh HOlthuis (1946, 1991, 2002) was cOnfident abOut identifying his material frOm Indian and Pacific Oceans with H. Milne Edwards (1837) specimen Of
<taxonomicName id="AB9D79BEFFA27B69FD75C25C3B18F8A6" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1837" box="[748,864,1866,1889]" class="Malacostraca" family="Scyllaridae" genus="Scyllarus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="484" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="5EE9DE2FFFA27B69FD75C25C3B18F8A6" box="[748,864,1866,1889]" italics="true" pageId="7" pageNumber="484">P. rugosus</emphasis>
</taxonomicName>
, we are less certain. As discussed earlier, there are nOw six species in
<taxonomicName id="AB9D79BEFFA27B69FEE1C2793995F841" authorityName="Holthuis" authorityYear="2002" box="[376,493,1903,1926]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="5EE9DE2FFFA27B69FEE1C2793995F841" box="[376,493,1903,1926]" italics="true" pageId="7" pageNumber="484">Petrarctus</emphasis>
</taxonomicName>
and anOther six species in
<taxonomicName id="AB9D79BEFFA27B69FCA8C2783BCFF841" authorityName="Holthuis" authorityYear="2002" box="[817,951,1902,1926]" class="Malacostraca" family="Scyllaridae" genus="Bathyarctus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="5EE9DE2FFFA27B69FCA8C2783BCFF841" box="[817,951,1902,1926]" italics="true" pageId="7" pageNumber="484">Bathyarctus</emphasis>
</taxonomicName>
, and sOme Of them are separated frOm
<taxonomicName id="AB9D79BEFFA27B69FA1EC2793888F86E" authority="Chan, 2017" authorityName="Chan" authorityYear="2017" class="Malacostraca" family="Scyllaridae" genus="Scyllarus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="484" phylum="Arthropoda" rank="species" sensu="stricto" species="rugosus">
<emphasis id="5EE9DE2FFFA27B69FA1EC2793888F86E" italics="true" pageId="7" pageNumber="484">P. rugosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45DA6354FFA27B69FF6FC28439FAF86D" box="[246,386,1937,1962]" pageId="7" pageNumber="484" sensu="stricto">sensu strictO</taxonomicNameLabel>
by characters that were nOt characterized by H.
<bibRefCitation id="080C7FCCFFA27B69FC07C2873CEEF86D" author="Milne" box="[926,1174,1937,1962]" pageId="7" pageNumber="484" refString="Milne Edwards, H. (1837) Histoire naturelle des Crustaces, comprenant l'anatomie, la physiologie et la classification de ces animaux. 2. Librairie encyclopedique de Roret, Paris, 532 pp." type="book" year="1837">
Milne
<bibRefCitation id="080C7FCCFFA27B69FC71C2873CEEF86D" author="Edwards" box="[1000,1174,1937,1962]" pageId="7" pageNumber="484" refString="Milne Edwards, H. (1837) Histoire naturelle des Crustaces, comprenant l'anatomie, la physiologie et la classification de ces animaux. 2. Librairie encyclopedique de Roret, Paris, 532 pp." type="book" year="1837">Edwards (1837</bibRefCitation>
</bibRefCitation>
; like the relatiVe height Of the median carina On abdOminal sOmite IV); and the shape Of the anteriOr margin Of the thOracic sternum is nOt knOwn (sO it may eVen be a species Of
<taxonomicName id="AB9D79BEFFA27B69FDDCC2CC3AB3F835" authorityName="Holthuis" authorityYear="2002" box="[581,715,2010,2034]" class="Malacostraca" family="Scyllaridae" genus="Bathyarctus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="5EE9DE2FFFA27B69FDDCC2CC3AB3F835" box="[581,715,2010,2034]" italics="true" pageId="7" pageNumber="484">Bathyarctus</emphasis>
</taxonomicName>
). TO stabilize the taxOnOmy Of
<taxonomicName id="AB9D79BEFFA27B69FBBEC2CC3D34F835" authority="HOlthuis, 2002" authorityName="Holthuis" authorityYear="2002" box="[1063,1356,2009,2034]" class="Malacostraca" family="Scyllaridae" genus="Petrarctus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="478" phylum="Arthropoda" rank="genus">
<emphasis id="5EE9DE2FFFA27B69FBBEC2CC3CE4F836" box="[1063,1180,2010,2033]" italics="true" pageId="7" pageNumber="484">Petrarctus</emphasis>
<bibRefCitation id="080C7FCCFFA27B69FB3AC2CF3D34F835" author="HOlthuis" box="[1187,1356,2009,2034]" pageId="7" pageNumber="484" refString="Holthuis, L. B. (2002) The Indo-Pacific scyllarine lobsters (Crustacea, Decapoda, Scyllaridae). Zoosystema, 24 (3), 499 - 683." type="journal article" year="2002">HOlthuis, 2002</bibRefCitation>
</taxonomicName>
, and tO ensure that the identity Of the type species remains as defined by HOlthuis (1946, 1991, 2002), we here designate a female specimen (
<quantity id="AB65AFD8FFAD7B66FEEEC5AA399EFF13" box="[375,486,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.9899999999999998" pageId="8" pageNumber="485" unit="mm" value="19.9">19.9 mm</quantity>
cl) (CASAU) recently cOllected frOm Pazhayar fishing pOrt, 11°2130.28”N, 79°4939.83”E, Tamil Nadu On
<date id="182324FDFFAD7B66FD9FC5C93AE8FF3F" box="[518,656,223,248]" pageId="8" pageNumber="485" value="2017-07-15">15 Jul 2017</date>
, as the neOtype Of
<taxonomicName id="AB9D79BEFFAD7B66FCF5C5F63D25FF3F" authority="H. Milne Edwards, 1837" authorityName="H. Milne Edwards" authorityYear="1837" box="[876,1373,223,248]" class="Malacostraca" family="Scyllaridae" genus="Scyllarus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="485" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="5EE9DE2FFFAD7B66FCF5C5F63C4EFF3F" box="[876,1078,224,248]" italics="true" pageId="8" pageNumber="485">Scyllarus rugosus</emphasis>
H.
<bibRefCitation id="080C7FCCFFAD7B66FBFDC5C93D25FF3F" author="Milne" box="[1124,1373,223,248]" pageId="8" pageNumber="485" refString="Milne Edwards, H. (1837) Histoire naturelle des Crustaces, comprenant l'anatomie, la physiologie et la classification de ces animaux. 2. Librairie encyclopedique de Roret, Paris, 532 pp." type="book" year="1837">Milne Edwards, 1837</bibRefCitation>
</taxonomicName>
. This lOcatiOn is just sOuth Of the Original type lOcality in POndicherry, and the specimen fits well the current mOrphOlOgical cOncept Of
<taxonomicName id="AB9D79BEFFAD7B66FE26C43F3A4CFE87" authorityName="H. Milne Edwards" authorityYear="1837" box="[447,564,297,320]" class="Malacostraca" family="Scyllaridae" genus="Scyllarus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="485" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="5EE9DE2FFFAD7B66FE26C43F3A4CFE87" box="[447,564,297,320]" italics="true" pageId="8" pageNumber="485">P. rugosus</emphasis>
</taxonomicName>
, has a cOlOr pattern (
<figureCitation id="F4A61EB8FFAD7B66FCB9C4313B0EFE87" box="[800,886,295,320]" captionStart="FIGURE 5" captionStartId="8.[151,250,930,952]" captionTargetBox="[185,1406,414,878]" captionTargetId="figure@8.[173,1406,414,878]" captionTargetPageId="8" captionText="FIGURE 5. A, Bayesian tree based on partial sequences of the mitochondrial COI gene (657 bps) of Petrarctus species and outgroup. Numbers at nodes are posterior probabilities from Bayesian inference analysis. Only values&gt; 0.8 are shown; B, Petractus rugosus (H. Milne Edwards, 1837), Pazhayar fishing port, Tamil Nadu, southeastern India, female neotype 19.9 mm cl (CASAU)." httpUri="https://zenodo.org/record/1010099/files/figure.png" pageId="8" pageNumber="485">Fig. 5B</figureCitation>
) that agrees with what is nOw recOgnized fOr the species and has 99.2% sequence similarity (
<accessionNumber id="73CE9FDEFFAD7B66FD18C45A3B79FEA2" box="[641,769,332,357]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF669474" pageId="8" pageNumber="485">MF669474</accessionNumber>
) tO the Chennai material (
<figureCitation id="F4A61EB8FFAD7B66FBBCC45A3CF8FEA2" box="[1061,1152,332,357]" captionStart="FIGURE 5" captionStartId="8.[151,250,930,952]" captionTargetBox="[185,1406,414,878]" captionTargetId="figure@8.[173,1406,414,878]" captionTargetPageId="8" captionText="FIGURE 5. A, Bayesian tree based on partial sequences of the mitochondrial COI gene (657 bps) of Petrarctus species and outgroup. Numbers at nodes are posterior probabilities from Bayesian inference analysis. Only values&gt; 0.8 are shown; B, Petractus rugosus (H. Milne Edwards, 1837), Pazhayar fishing port, Tamil Nadu, southeastern India, female neotype 19.9 mm cl (CASAU)." httpUri="https://zenodo.org/record/1010099/files/figure.png" pageId="8" pageNumber="485">Fig. 5A</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>