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<document id="70477FFC52D793B007821E6650DFF290" ID-DOI="10.11646/zootaxa.3670.2.7" ID-GBIF-Dataset="ca02f198-cd57-4096-ad51-bc11ca524d95" ID-ISSN="1175-5326" ID-Zenodo-Dep="215940" ID-ZooBank="A83EF888-DBFF-4320-92D7-252A8DE34E85" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460633811143" checkinUser="plazi" docAuthor="Wiedenbrug, Sofia, Lamas, Carlos E. &amp; Trivinho-Strixino, Susana" docDate="2013" docId="E43287B86D19FFD33085FAF8424246A6" docLanguage="en" docName="zt03670p237.pdf" docOrigin="Zootaxa 3670 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thienemanniella medialis Sublette &amp; Sasa" docType="treatment" docVersion="9" lastPageNumber="221" masterDocId="180BFFC06D1CFFD53012FFE5430D416F" masterDocTitle="A review of Neotropical species in Thienemanniella Kieffer (Diptera, Chironomidae)" masterLastPageNumber="237" masterPageNumber="215" pageNumber="220" updateTime="1698334908411" updateUser="plazi">
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<mods:title id="FBF62739DC208337D3471636DB503F41">A review of Neotropical species in Thienemanniella Kieffer (Diptera, Chironomidae)</mods:title>
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<mods:namePart id="0A2AE03870B0559C631D80428AFA4A26">Wiedenbrug, Sofia</mods:namePart>
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<mods:namePart id="2C31950029C86D92FA9390DF6C73A392">Lamas, Carlos E.</mods:namePart>
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<mods:namePart id="659CFCCE63DFC821CACE298DC392E700">Trivinho-Strixino, Susana</mods:namePart>
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<mods:title id="B91C3E49DB2AB8F01DF79953C76E5AA4">Zootaxa</mods:title>
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<mods:date id="F9A37CCE7D1F1CF4A6B7B30E7A0DA825">2013</mods:date>
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<treatment id="E43287B86D19FFD33085FAF8424246A6" ID-DOI="http://doi.org/10.5281/zenodo.5631806" ID-GBIF-Taxon="119570084" ID-Zenodo-Dep="5631806" LSID="urn:lsid:plazi:treatment:E43287B86D19FFD33085FAF8424246A6" httpUri="http://treatment.plazi.org/id/E43287B86D19FFD33085FAF8424246A6" lastPageId="6" lastPageNumber="221" pageId="5" pageNumber="220">
<subSubSection id="248165256D19FFD03085FAF843E64437" pageId="5" pageNumber="220" type="nomenclature">
<paragraph id="6C2436AE6D19FFD03085FAF841924458" blockId="5.[151,671,1309,1368]" box="[151,671,1309,1335]" pageId="5" pageNumber="220">
<heading id="376C81C26D19FFD03085FAF841924458" bold="true" box="[151,671,1309,1335]" fontSize="11" level="1" pageId="5" pageNumber="220" reason="1">
<taxonomicName id="AB9B4D2D6D19FFD03085FAF841924458" ID-CoL="56GKM" authority="Sublette &amp; Sasa" authorityName="Sublette &amp; Sasa" box="[151,671,1309,1335]" class="Insecta" family="Chironomidae" genus="Thienemanniella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="medialis">
<emphasis id="5EEFEABC6D19FFD03085FAF841924458" bold="true" box="[151,671,1309,1335]" pageId="5" pageNumber="220">
<emphasis id="5EEFEABC6D19FFD03085FAF842DF4458" bold="true" box="[151,466,1309,1335]" italics="true" pageId="5" pageNumber="220">Thienemanniella medialis</emphasis>
Sublette &amp; Sasa
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="6C2436AE6D19FFD03085FAA543E64437" blockId="5.[151,671,1309,1368]" box="[151,235,1344,1368]" pageId="5" pageNumber="220">
(
<figureCitation id="F4A02A2B6D19FFD0308DFAA543EF4437" box="[159,226,1344,1368]" captionStart="FIGURE 3" captionStartId="6.[151,249,1365,1388]" captionTargetBox="[157,1429,203,1324]" captionTargetId="figure@6.[151,1436,193,1344]" captionTargetPageId="6" captionText="FIGURE 3. Thienemanniella medialis Sublette &amp; Sasa. A. Adult male, hypopygium. B – C. Pupa. B. Tergites III – IV. C. Tergites VIII – IX, anal lobes." httpUri="https://zenodo.org/record/215943/files/figure.png" pageId="5" pageNumber="220">Fig. 3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="248165256D19FFD03085FA6D479544F1" box="[151,1176,1416,1438]" pageId="5" pageNumber="220" type="reference_group">
<paragraph id="6C2436AE6D19FFD03085FA6D479544F1" blockId="5.[151,1176,1416,1438]" box="[151,1176,1416,1438]" pageId="5" pageNumber="220">
<treatmentCitationGroup id="4C8B11806D19FFD03085FA6D479544F1" box="[151,1176,1416,1438]" pageId="5" pageNumber="220">
<taxonomicName id="AB9B4D2D6D19FFD03085FA6D419244F1" ID-CoL="56GKM" authority="Sublette &amp; Sasa, 1994: 1" authorityName="Sublette &amp; Sasa" authorityPageNumber="1" authorityYear="1994" box="[151,671,1416,1438]" class="Insecta" family="Chironomidae" genus="Thienemanniella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="medialis">
<emphasis id="5EEFEABC6D19FFD03085FA6D429044F1" box="[151,413,1416,1438]" italics="true" pageId="5" pageNumber="220">Thienemanniella medialis</emphasis>
<treatmentCitation id="ED3A10BF6D19FFD031B1FA6D419244F1" author="Sublette" box="[419,671,1416,1438]" page="1" pageId="5" pageNumber="220" year="1994">
<bibRefCitation id="080A4B5F6D19FFD031B1FA6D418444F1" author="Sublette" box="[419,649,1416,1438]" pageId="5" pageNumber="220" refString="Sublette, J. E. &amp; Sasa, M. (1994) Chironomidae collected in Onchocerciasis endemic areas of Guatemala (Insecta, Diptera). Spixiana, Supplement, 20, 1 - 60." type="journal article" year="1994">Sublette &amp; Sasa, 1994</bibRefCitation>
: 1
</treatmentCitation>
</taxonomicName>
pharate male,
<specimenCount id="7A9DFD276D19FFD03321FA6C408344F1" box="[819,910,1416,1438]" pageId="5" pageNumber="220" type="female">1 female</specimenCount>
.
<typeStatus id="B320880C6D19FFD0338AFA6C40C744F1" box="[920,970,1417,1438]" pageId="5" pageNumber="220">Type</typeStatus>
locality
<collectingCountry id="148C763E6D19FFD03431FA6D479944F1" box="[1059,1172,1416,1438]" name="Guatemala" pageId="5" pageNumber="220">Guatemala</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="248165256D19FFD33085FA2F4125467A" lastPageId="6" lastPageNumber="221" pageId="5" pageNumber="220" type="materials_examined">
<paragraph id="6C2436AE6D19FFD03085FA2F47B2466C" blockId="5.[151,1436,1482,2014]" pageId="5" pageNumber="220">
<emphasis id="5EEFEABC6D19FFD03085FA2F428D448C" bold="true" box="[151,384,1482,1507]" pageId="5" pageNumber="220">Material examined.</emphasis>
<typeStatus id="B320880C6D19FFD03194FA2E42E3448C" box="[390,494,1483,1507]" pageId="5" pageNumber="220" type="holotype">Holotype</typeStatus>
pharate male with pupal exuviae, described by
<bibRefCitation id="080A4B5F6D19FFD033ECFA2E4619448C" author="Sublette" box="[1022,1300,1483,1507]" pageId="5" pageNumber="220" refString="Sublette, J. E. &amp; Sasa, M. (1994) Chironomidae collected in Onchocerciasis endemic areas of Guatemala (Insecta, Diptera). Spixiana, Supplement, 20, 1 - 60." type="journal article" year="1994">Sublette and Sasa (1994)</bibRefCitation>
as “Rincon, leaf litter at small waterfall C, no. I-100” on a single slide with two pupal exuviae of
<taxonomicName id="AB9B4D2D6D19FFD03454FA15466B4768" box="[1094,1382,1519,1543]" class="Insecta" family="Chironomidae" genus="Onconeura" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="semifimbriata">
<emphasis id="5EEFEABC6D19FFD03454FA15466B4768" box="[1094,1382,1519,1543]" italics="true" pageId="5" pageNumber="220">Onconeura semifimbriata</emphasis>
</taxonomicName>
, and one male and one female of
<taxonomicName id="AB9B4D2D6D19FFD031FEF9F1418C4744" box="[492,641,1556,1579]" class="Insecta" family="Chironomidae" genus="Corynoneura" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="genus">
<emphasis id="5EEFEABC6D19FFD031FEF9F1418C4744" box="[492,641,1556,1579]" italics="true" pageId="5" pageNumber="220">Corynoneura</emphasis>
</taxonomicName>
spp. (not
<taxonomicName id="AB9B4D2D6D19FFD032ECF9F140984744" box="[766,917,1555,1579]" class="Insecta" family="Chironomidae" genus="Corynoneura" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="ferelobata">
<emphasis id="5EEFEABC6D19FFD032ECF9F140984744" box="[766,917,1555,1579]" italics="true" pageId="5" pageNumber="220">C. ferelobata</emphasis>
</taxonomicName>
, see
<bibRefCitation id="080A4B5F6D19FFD033C4F9F647E34744" author="Wiedenbrug" box="[982,1262,1555,1579]" pageId="5" pageNumber="220" refString="Wiedenbrug, S., Lamas, C. E. &amp; Trivinho-Strixino, S. (2012) A review of the genus Corynoneura Winnertz (Diptera: Chironomidae) from the Neotropical region. Zootaxa, 3574, 1 - 61." type="journal article" year="2012">
Wiedenbrug
<emphasis id="5EEFEABC6D19FFD03478F9F147AA4744" box="[1130,1191,1555,1579]" italics="true" pageId="5" pageNumber="220">et al.</emphasis>
2012
</bibRefCitation>
). According to
<bibRefCitation id="080A4B5F6D19FFD03085F9D241154720" author="Sublette" box="[151,536,1591,1616]" pageId="5" pageNumber="220" refString="Sublette, J. E. &amp; Sasa, M. (1994) Chironomidae collected in Onchocerciasis endemic areas of Guatemala (Insecta, Diptera). Spixiana, Supplement, 20, 1 - 60." type="journal article" year="1994">Sublette and Sasa (1994, page 28)</bibRefCitation>
the allotype of
<taxonomicName id="AB9B4D2D6D19FFD032D8F9DD40484720" box="[714,837,1591,1615]" class="Insecta" family="Chironomidae" genus="Thienemanniella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="medialis">
<emphasis id="5EEFEABC6D19FFD032D8F9DD40484720" box="[714,837,1591,1615]" italics="true" pageId="5" pageNumber="220">T. medialis</emphasis>
</taxonomicName>
is on slide number
<date id="1825106E6D19FFD03437F9D247454720" box="[1061,1096,1591,1615]" pageId="5" pageNumber="220">I-3</date>
, specimen number 4, together with the
<typeStatus id="B320880C6D19FFD030EEF9BE426C471C" box="[252,353,1627,1651]" pageId="5" pageNumber="220" type="holotype">holotype</typeStatus>
, allotype and
<typeStatus id="B320880C6D19FFD03213F9B94161471B" box="[513,620,1628,1652]" pageId="5" pageNumber="220" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="AB9B4D2D6D19FFD03287F9B94027471C" box="[661,810,1627,1651]" class="Insecta" family="Chironomidae" genus="Corynoneura" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="hirvenojai">
<emphasis id="5EEFEABC6D19FFD03287F9B94027471C" box="[661,810,1627,1651]" italics="true" pageId="5" pageNumber="220">C. hirvenojai</emphasis>
</taxonomicName>
. However, the slide with the
<typeStatus id="B320880C6D19FFD0346EF9BE47D3471C" box="[1148,1246,1627,1651]" pageId="5" pageNumber="220" type="holotype">holotype</typeStatus>
of
<taxonomicName id="AB9B4D2D6D19FFD03514F9B9424047F8" authority="Sublette &amp; Sasa" authorityName="Sublette &amp; Sasa" class="Insecta" family="Chironomidae" genus="Corynoneura" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="hirvenojai">
<emphasis id="5EEFEABC6D19FFD03514F9B94691471C" box="[1286,1436,1627,1651]" italics="true" pageId="5" pageNumber="220">C. hirvenojai</emphasis>
Sublette &amp; Sasa
</taxonomicName>
is number
<date id="1825106E6D19FFD031DFF99A42F647F8" box="[461,507,1663,1687]" pageId="5" pageNumber="220" value="1943-01">I-43</date>
and has only one specimen.
<bibRefCitation id="080A4B5F6D19FFD03354F99A47C447F8" author="Sublette" box="[838,1225,1663,1688]" pageId="5" pageNumber="220" refString="Sublette, J. E. &amp; Sasa, M. (1994) Chironomidae collected in Onchocerciasis endemic areas of Guatemala (Insecta, Diptera). Spixiana, Supplement, 20, 1 - 60." type="journal article" year="1994">Sublette and Sasa (1994, page 12)</bibRefCitation>
mention that slide number
<date id="1825106E6D19FFD030E7F946422E47D4" box="[245,291,1699,1723]" pageId="5" pageNumber="220" value="1939-01">I-39</date>
holds several specimens, including the
<typeStatus id="B320880C6D19FFD032F9F946405D47D4" box="[747,848,1699,1723]" pageId="5" pageNumber="220" type="holotype">holotype</typeStatus>
, allotype and
<typeStatus id="B320880C6D19FFD033E3F941475147D3" box="[1009,1116,1700,1724]" pageId="5" pageNumber="220" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="AB9B4D2D6D19FFD03496F94143C047B0" authority="Sublette &amp; Sasa" authorityName="Sublette &amp; Sasa" class="Insecta" family="Chironomidae" genus="Corynoneura" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="ferelobata">
<emphasis id="5EEFEABC6D19FFD03496F941461447D4" box="[1156,1305,1699,1723]" italics="true" pageId="5" pageNumber="220">C. ferelobata</emphasis>
Sublette &amp; Sasa
</taxonomicName>
, and that specimen number 4 is the allotype of
<emphasis id="5EEFEABC6D19FFD032CBF92D405747B0" box="[729,858,1735,1759]" italics="true" pageId="5" pageNumber="220">
<taxonomicName id="AB9B4D2D6D19FFD032CBF92D405A47B0" box="[729,855,1735,1759]" class="Insecta" family="Chironomidae" genus="Thienemanniella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="medialis">T. medialis</taxonomicName>
.
</emphasis>
We have examined this specimen, but since there is no corresponding pupa, that female should be considered as merely tentatively associated only.
</paragraph>
<paragraph id="6C2436AE6D19FFD030D5F8EB4084461E" blockId="5.[151,1436,1482,2014]" pageId="5" pageNumber="220">
<emphasis id="5EEFEABC6D19FFD030D5F8EB42DD4648" bold="true" box="[199,464,1806,1831]" pageId="5" pageNumber="220">Diagnostic characters.</emphasis>
Male antenna with 9 flagellomeres; pupa with tergal conjunctive III/IV carrying strong hooklets; segments III-VIII with 3 taeniate lateral setae; L2 longer and broader than L1 and L3; anal lobe with fringe reduced to about 10 taeniate setae shorter than the anal macrosetae.
</paragraph>
<paragraph id="6C2436AE6D19FFD030D5F898467946D6" blockId="5.[151,1436,1482,2014]" pageId="5" pageNumber="220">
<emphasis id="5EEFEABC6D19FFD030D5F89841A846F9" bold="true" box="[199,677,1917,1942]" pageId="5" pageNumber="220">Additions to previous description. Male</emphasis>
(n = 1). Head. According to
<bibRefCitation id="080A4B5F6D19FFD033E7F898461C46FA" author="Sublette" box="[1013,1297,1917,1941]" pageId="5" pageNumber="220" refString="Sublette, J. E. &amp; Sasa, M. (1994) Chironomidae collected in Onchocerciasis endemic areas of Guatemala (Insecta, Diptera). Spixiana, Supplement, 20, 1 - 60." type="journal article" year="1994">Sublette and Sasa (1994)</bibRefCitation>
the antenna has 10 flagellomeres, but we find the terminal two to be incompletely divided, thus count only 9 flagellomeres.
</paragraph>
<paragraph id="6C2436AE6D19FFD030D5F82347D746B2" blockId="5.[151,1436,1482,2014]" box="[199,1242,1989,2014]" pageId="5" pageNumber="220">
Legs. Dorsal keel of fore leg trochanter resembles that in
<taxonomicName id="AB9B4D2D6D19FFD03359F82240E146B1" box="[843,1004,1990,2014]" class="Insecta" family="Chironomidae" genus="Thienemanniella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="sancticaroli" status="sp. nov.">
<emphasis id="5EEFEABC6D19FFD03359F82240E146B1" box="[843,1004,1990,2014]" italics="true" pageId="5" pageNumber="220">T. sancticaroli</emphasis>
</taxonomicName>
<emphasis id="5EEFEABC6D19FFD033E1F823473F46B1" bold="true" box="[1011,1074,1990,2014]" pageId="5" pageNumber="220">
<taxonomicNameLabel id="45DC57C76D19FFD033E1F823473F46B1" box="[1011,1074,1990,2014]" pageId="5" pageNumber="220" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(see
<figureCitation id="F4A02A2B6D19FFD0347FF82047A346B2" box="[1133,1198,1989,2013]" captionStart="FIGURE 5" captionStartId="9.[151,249,1906,1929]" captionTargetBox="[168,1408,454,1875]" captionTargetId="figure@9.[155,1432,443,1885]" captionTargetPageId="9" captionText="FIGURE 5. Thienemanniella sancticaroli sp. n., adult male. A. Fore trochanter. B. Hind trochanter. C – D. Apex of hind tibia in posterior (C) and anterior view (D). E. Wing. F – G. Hypopygium. F. Tergite IX (part) and gonostylus. G. Tergite IX and gonostyli removed, internal sclerites hatched; left: ventral view, right: dorsal view." httpUri="https://zenodo.org/record/215945/files/figure.png" pageId="5" pageNumber="220">Fig. 5</figureCitation>
A).
</paragraph>
<caption id="38E466266D1AFFD33085FAB042B744E5" httpUri="https://zenodo.org/record/215943/files/figure.png" pageId="6" pageNumber="221" targetBox="[157,1429,203,1324]" targetPageId="6">
<paragraph id="6C2436AE6D1AFFD33085FAB042B744E5" blockId="6.[151,1436,1365,1418]" pageId="6" pageNumber="221">
<emphasis id="5EEFEABC6D1AFFD33085FAB040144405" bold="true" box="[151,793,1365,1388]" pageId="6" pageNumber="221">
FIGURE 3.
<taxonomicName id="AB9B4D2D6D1AFFD33132FAB040144405" authority="Sublette &amp; Sasa. A." authorityName="Sublette &amp; Sasa. A." box="[288,793,1365,1388]" class="Insecta" family="Chironomidae" genus="Thienemanniella" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="221" phylum="Arthropoda" rank="species" species="medialis">
<emphasis id="5EEFEABC6D1AFFD33132FAB0413E4404" bold="true" box="[288,563,1365,1387]" italics="true" pageId="6" pageNumber="221">Thienemanniella medialis</emphasis>
Sublette &amp; Sasa. A.
</taxonomicName>
</emphasis>
Adult male, hypopygium.
<emphasis id="5EEFEABC6D1AFFD3342AFAB347654403" bold="true" box="[1080,1128,1365,1388]" pageId="6" pageNumber="221">B–C</emphasis>
. Pupa. B. Tergites III–IV. C. Tergites VIII–IX, anal lobes.
</paragraph>
</caption>
<paragraph id="6C2436AE6D1AFFD330D5FA53426C449D" blockId="6.[151,1436,1462,1993]" pageId="6" pageNumber="221">Tergites. TI with 4 setae; TII-III impossible to observe; TIV-VI with 3; TVII-VIII with 2 setae. Sternites III- VIII with one seta</paragraph>
<paragraph id="6C2436AE6D1AFFD330D5FA1B40464755" blockId="6.[151,1436,1462,1993]" pageId="6" pageNumber="221">
Hypopygium. (
<figureCitation id="F4A02A2B6D1AFFD33163FA1B42BE4779" box="[369,435,1534,1558]" captionStart="FIGURE 3" captionStartId="6.[151,249,1365,1388]" captionTargetBox="[157,1429,203,1324]" captionTargetId="figure@6.[151,1436,193,1344]" captionTargetPageId="6" captionText="FIGURE 3. Thienemanniella medialis Sublette &amp; Sasa. A. Adult male, hypopygium. B – C. Pupa. B. Tergites III – IV. C. Tergites VIII – IX, anal lobes." httpUri="https://zenodo.org/record/215943/files/figure.png" pageId="6" pageNumber="221">Fig. 3</figureCitation>
A) Laterosternite with 1 seta. Superior volsella difficult to define due to bad condition of pharate material. Sternapodeme and phallapodeme as figured.
</paragraph>
<paragraph id="6C2436AE6D1AFFD330D5F9A242FB479E" blockId="6.[151,1436,1462,1993]" pageId="6" pageNumber="221">
<emphasis id="5EEFEABC6D1AFFD330D5F9A242084730" bold="true" box="[199,261,1607,1631]" pageId="6" pageNumber="221">Pupa</emphasis>
(n = 1). Abdomen (
<figureCitation id="F4A02A2B6D1AFFD331E9F9A3414F4731" box="[507,578,1606,1630]" captionStart="FIGURE 5" captionStartId="9.[151,249,1906,1929]" captionTargetBox="[168,1408,454,1875]" captionTargetId="figure@9.[155,1432,443,1885]" captionTargetPageId="9" captionText="FIGURE 5. Thienemanniella sancticaroli sp. n., adult male. A. Fore trochanter. B. Hind trochanter. C – D. Apex of hind tibia in posterior (C) and anterior view (D). E. Wing. F – G. Hypopygium. F. Tergite IX (part) and gonostylus. G. Tergite IX and gonostyli removed, internal sclerites hatched; left: ventral view, right: dorsal view." httpUri="https://zenodo.org/record/215945/files/figure.png" pageId="6" pageNumber="221">Fig. 5</figureCitation>
B–C). TI-II apparently with few posterior shagreen points; TIII-IX with homogeneous shagreen of elongated spinules. Sternites III-VIII with sparse shagreen of short points. Segments III- VIII with 3 taeniate L-setae; L2 broader and longer than the other two. Anal lobe rounded (
<figureCitation id="F4A02A2B6D1AFFD334F4F96B462347C9" box="[1254,1326,1678,1702]" captionStart="FIGURE 3" captionStartId="6.[151,249,1365,1388]" captionTargetBox="[157,1429,203,1324]" captionTargetId="figure@6.[151,1436,193,1344]" captionTargetPageId="6" captionText="FIGURE 3. Thienemanniella medialis Sublette &amp; Sasa. A. Adult male, hypopygium. B – C. Pupa. B. Tergites III – IV. C. Tergites VIII – IX, anal lobes." httpUri="https://zenodo.org/record/215943/files/figure.png" pageId="6" pageNumber="221">Fig. 3</figureCitation>
C), with incomplete fringe of setae shorter than macrosetae and 3 cylindrical, apically hooked macrosetae on strong bases; medial setae taeniate and short.
</paragraph>
<paragraph id="6C2436AE6D1AFFD330D5F91B4125467A" blockId="6.[151,1436,1462,1993]" box="[199,552,1789,1813]" pageId="6" pageNumber="221">
For measurements, see
<tableCitation id="211903156D1AFFD331DFF918412F467A" box="[461,546,1789,1813]" captionStart="TABLE 3" captionStartId="12.[151,239,915,939]" captionTargetBox="[159,1366,1112,1968]" captionTargetPageId="12" captionText="TABLE 3. Mensural characters of pupae of neotropical Thienemanniella species; measurements in µm; &quot; No. of spines cT &quot; and &quot; No. of spines cS &quot; refer to dorsal and ventral conjunctives, respectively." httpUri="http://table.plazi.org/id/38E466266D10FFD93085FC7640FC42A4" pageId="6" pageNumber="221" tableUuid="38E466266D10FFD93085FC7640FC42A4">Table 3</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="248165256D1AFFD330D5F8C7424246A6" pageId="6" pageNumber="221" type="discussion">
<paragraph id="6C2436AE6D1AFFD330D5F8C742DF46EE" blockId="6.[151,1436,1462,1993]" pageId="6" pageNumber="221">
<emphasis id="5EEFEABC6D1AFFD330D5F8C742364655" bold="true" box="[199,315,1826,1850]" pageId="6" pageNumber="221">Remarks.</emphasis>
<bibRefCitation id="080A4B5F6D1AFFD33151F8C741094656" box="[323,516,1825,1849]" pageId="6" pageNumber="221" refString="Fu, Y., Saether, O. A. &amp; Wang, X. (2010 b) Thienemanniella Kieffer from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa, 2431, 1 - 42." type="journal article">
Fu
<emphasis id="5EEFEABC6D1AFFD3317BF8C742AE4656" box="[361,419,1825,1849]" italics="true" pageId="6" pageNumber="221">et al.</emphasis>
(2010b)
</bibRefCitation>
included
<taxonomicName id="AB9B4D2D6D1AFFD33264F8C741FF4656" box="[630,754,1825,1849]" class="Insecta" family="Chironomidae" genus="Thienemanniella" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="221" phylum="Arthropoda" rank="species" species="medialis">
<emphasis id="5EEFEABC6D1AFFD33264F8C741FF4656" box="[630,754,1825,1849]" italics="true" pageId="6" pageNumber="221">T. medialis</emphasis>
</taxonomicName>
in their key, and accordingly the inferior volsella is present and the superior volsella partly fused with the gonocoxite. More material is necessary to better define the hypopygium of this species.
</paragraph>
<paragraph id="6C2436AE6D1AFFD330D5F86B424246A6" blockId="6.[151,1436,1462,1993]" pageId="6" pageNumber="221">
A very similar pupal exuviae was found in
<collectingCountry id="148C763E6D1AFFD332B4F86B41D146CA" box="[678,732,1934,1957]" name="Peru" pageId="6" pageNumber="221">Peru</collectingCountry>
, deposited in the ZSM as
<taxonomicName id="AB9B4D2D6D1AFFD333E9F86B47B546C9" box="[1019,1208,1934,1958]" class="Insecta" family="Chironomidae" genus="Thienemanniella" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="221" phylum="Arthropoda" rank="genus">
<emphasis id="5EEFEABC6D1AFFD333E9F86B47B546C9" box="[1019,1208,1934,1958]" italics="true" pageId="6" pageNumber="221">Thienemanniella</emphasis>
</taxonomicName>
sp. A2, sample P18, leg. E.J. Fittkau.
</paragraph>
</subSubSection>
</treatment>
</document>
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