293 lines
38 KiB
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293 lines
38 KiB
XML
<document id="A7B29FCDABE2ED167589B6BC64F822B5" ID-DOI="10.11646/phytotaxa.598.1.3" ID-ISSN="1179-3163" ID-Zenodo-Dep="7958772" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1684791918802" checkinUser="plazi" docAuthor="Ferrer-Gallego, P. Pablo" docDate="2023" docId="E4294C700C3E5C5FAEEBC5D4FECC8433" docLanguage="en" docName="phytotaxa.598.1.3.pdf" docOrigin="Phytotaxa 598 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.598.1.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.6:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="6" docTitle="Ononis tridentata subsp. arbuscula P. P. Ferrer" docType="treatment" docVersion="1" lastPageNumber="40" masterDocId="181034080C385C57AE63C022FFF2826B" masterDocTitle="Original elemets and typification of the names included in the Ononis tridentata complex (Fabaceae)" masterLastPageNumber="58" masterPageNumber="32" pageNumber="38" updateTime="1684850746924" updateUser="juliana" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="0276C7F255D7517F54202570AD70622B">Original elemets and typification of the names included in the Ononis tridentata complex (Fabaceae)</mods:title>
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<mods:namePart id="1FE12BBADE0746BD9666FCEEF6BDB6B1">Ferrer-Gallego, P. Pablo</mods:namePart>
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<mods:affiliation id="E27B9A5A888EDB1F0EC9E4EAB5324FCC">Servicio de Vida Silvestre y Red Natura 2000, Centro para la Investigación y la Experimentación Forestal (CIEF), Generalitat Valenciana, Avda. Comarques del País Valencià 114, E- 46930 Quart de Poblet, Valencia, Spain. & flora. cief @ gva. es; https: // orcid. org / 0000 - 0001 - 7595 - 9302</mods:affiliation>
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<mods:nameIdentifier id="D9B6AF6938D2FB519A6BBAAACDA06DA0" type="email">flora.cief@gva.es</mods:nameIdentifier>
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<mods:title id="FC28CB711912842351CBD2B72B1AB556">Phytotaxa</mods:title>
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<mods:date id="F2BA21BAF7B1FABBB7E7AE5B89519D1D">2023</mods:date>
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<treatment id="E4294C700C3E5C5FAEEBC5D4FECC8433" LSID="urn:lsid:plazi:treatment:E4294C700C3E5C5FAEEBC5D4FECC8433" httpUri="http://treatment.plazi.org/id/E4294C700C3E5C5FAEEBC5D4FECC8433" lastPageId="8" lastPageNumber="40" pageId="6" pageNumber="38">
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<subSubSection id="249AAEED0C3E5C51AEEBC5D4FCE2847A" box="[136,784,1526,1553]" pageId="6" pageNumber="38" type="nomenclature">
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<paragraph id="6C3FFD660C3E5C51AEEBC5D4FCE2847A" blockId="6.[136,784,1526,1553]" box="[136,784,1526,1553]" pageId="6" pageNumber="38">
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<heading id="37774A0A0C3E5C51AEEBC5D4FCE2847A" box="[136,784,1526,1553]" fontSize="11" level="2" pageId="6" pageNumber="38" reason="3">
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<taxonomicName id="AB8086E50C3E5C51AEEBC5D4FCE2847A" ID-CoL="9DFBB" authority="(Desvaux) P. P. Ferrer" box="[136,784,1526,1553]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="6" pageNumber="38" phylum="Tracheophyta" rank="subSpecies" species="tridentata" subSpecies="arbuscula">
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<emphasis id="5EF421740C3E5C51AEEBC5D4FEA1847A" bold="true" box="[136,339,1526,1553]" italics="true" pageId="6" pageNumber="38">Ononis tridentata</emphasis>
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subsp.
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<emphasis id="5EF421740C3E5C51AFCBC5D4FDE9847A" bold="true" box="[424,539,1526,1553]" italics="true" pageId="6" pageNumber="38">arbuscula</emphasis>
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(Desvaux) P.P. Ferrer
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="249AAEED0C3E5C5FAEEBC61CFECC8433" lastPageId="8" lastPageNumber="40" pageId="6" pageNumber="38" type="description">
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<paragraph id="6C3FFD660C3E5C51AEEBC61CFB8B84CA" blockId="6.[136,1452,1598,2093]" pageId="6" pageNumber="38">
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As mentioned above, due to the epitype designation for the name
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<taxonomicName id="AB8086E50C3E5C51AD38C61DFBD28433" box="[859,1056,1598,1625]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="6" pageNumber="38" phylum="Tracheophyta" rank="species" species="tridentata">
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<emphasis id="5EF421740C3E5C51AD38C61DFBD28433" box="[859,1056,1598,1625]" italics="true" pageId="6" pageNumber="38">Ononis tridentata</emphasis>
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</taxonomicName>
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published by F̂rther (in
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<bibRefCitation id="081180970C3E5C51AB5BC61CFEB48417" author="Turland, N. J. & Jarvis, C. E." pageId="6" pageNumber="38" pagination="457 - 485" refId="ref15471" refString="Turland, N. J. & Jarvis, C. E. (1997) Typification of Linnean specific and varietal names in the Leguminosae (Fabaceae). Taxon 46: 457 - 485. https: // doi. org / 10.2307 / 1224388" type="journal article" year="1997">Turland & Jarvis 1997: 478</bibRefCitation>
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), a new name in subspecies rank should be proposed for the Iberian populations that until now were known with the traditional concept (pre-epitypification) of
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<taxonomicName id="AB8086E50C3E5C51AD7DC6A5FB8384CB" box="[798,1137,1670,1697]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="6" pageNumber="38" phylum="Tracheophyta" rank="subSpecies" species="tridentata" subSpecies="tridentata">
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<emphasis id="5EF421740C3E5C51AD7DC6A5FC5C84CB" box="[798,942,1670,1697]" italics="true" pageId="6" pageNumber="38">O. tridentata</emphasis>
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subsp.
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<emphasis id="5EF421740C3E5C51AA60C6A4FB8384CB" box="[1027,1137,1670,1696]" italics="true" pageId="6" pageNumber="38">tridentata</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="6C3FFD660C3E5C5FAEDEC689FBC3834F" blockId="6.[136,1452,1598,2093]" lastBlockId="8.[136,1452,158,1627]" lastPageId="8" lastPageNumber="40" pageId="6" pageNumber="38">
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In this sense, there is a relevant early name published by
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<bibRefCitation id="081180970C3E5C51AD5DC689FBED84AF" author="Desvaux, A. N." box="[830,1055,1706,1733]" pageId="6" pageNumber="38" pagination="65 - 84" refId="ref13796" refString="Desvaux, A. N. (1814) Memoire et observations sur la Famille des Plantes Legumineuses. Journal de Botanique, Appliquee a l'Agriculture, a la Pharmacie, a la Medecine et aux Arts 3: 65 - 84." type="journal article" year="1814">Desvaux (1814: 76)</bibRefCitation>
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,
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<taxonomicName id="AB8086E50C3E5C51AA48C689FB0184AF" authorityName="Desvaux" authorityYear="1814" box="[1067,1267,1706,1733]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="6" pageNumber="38" phylum="Tracheophyta" rank="species" species="arbuscula">
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<emphasis id="5EF421740C3E5C51AA48C689FB0184AF" box="[1067,1267,1706,1733]" italics="true" pageId="6" pageNumber="38">Ononis arbuscula</emphasis>
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</taxonomicName>
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(see
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<bibRefCitation id="081180970C3E5C51AB4CC689FA5584AE" author="Losa, T. M." box="[1327,1447,1706,1733]" pageId="6" pageNumber="38" pagination="227 - 337" refId="ref14642" refString="Losa, T. M. (1958) El genero Ononis y las Ononis espanolas. Anales del Instituto Botanico A. J. Cavanilles 26: 227 - 337." type="journal article" year="1958">Losa 1958</bibRefCitation>
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,
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<bibRefCitation id="081180970C3E5C51AEEBC6EDFE018483" author="Devesa, J. A. & Lopez Gonzalez, G." box="[136,499,1742,1769]" pageId="6" pageNumber="38" pagination="245 - 260" refId="ref14024" refString="Devesa, J. A. & Lopez Gonzalez, G. (1997) Notas taxonomicas y nomenclaturales sobre el genero Ononis L. (Leguminosae) en la Peninsula Iberica e Islas Baleares. Anales del Jardin Botanico de Madrid 55 (2): 245 - 260." type="journal article" year="1997">Devesa & López González 1997</bibRefCitation>
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). Nicaise Auguste Desvaux (1784-1856) was a French botanist at Angers (
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<collectingCountry id="1497BDF60C3E5C51AB44C6ECFA888483" box="[1319,1402,1742,1768]" name="France" pageId="6" pageNumber="38">France</collectingCountry>
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). He was appointed director of the Angers botanic garden in 1817. Desvaux did not personally collect botanical specimens beyond the region of Angers. He received specimens from many correspondents, but the collector is not noted and the area stated is rarely specific (
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<bibRefCitation id="081180970C3E5C51AC67C718FD1B853F" author="Howard, R. A. & Clausen, K. S. & Gillis, W. T., Jr." box="[516,745,1850,1877]" pageId="6" pageNumber="38" pagination="211 - 242" refId="ref14233" refString="Howard, R. A., Clausen, K. S. & Gillis, W. T., Jr. (1981) William Hamilton (1783 - 1856) and the Prodromus Plantarum Indiae Occidentalis (1825). Journal of the Arnold Arboretum 62: 211 - 242. https: // doi. org / 10.5962 / bhl. part. 11247" type="journal article" year="1981">
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Howard
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<emphasis id="5EF421740C3E5C51AC09C719FD6F853F" box="[618,669,1850,1876]" italics="true" pageId="6" pageNumber="38">et al</emphasis>
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. 1981
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</bibRefCitation>
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||
). In his
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<emphasis id="5EF421740C3E5C51AD2AC718FB49853F" box="[841,1211,1850,1876]" italics="true" pageId="6" pageNumber="38">Journal de Botanique Appliquée</emphasis>
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Desvaux proposed a large number of species. It would not be true to say that his work was neglected; but it was difficult to follow. His descriptions, though accurate as far as they go, are brief and, like so many older ones, omit details later deemed essential. His data of locality are frequently very general and by no means always correct. He never cited collectors; authentic material, where it exists, has been, therefore, hard to identify as such. Until 1896, when it was presented to the Museum d’Histoire Naturelle at Paris, his personal herbarium remained in the possession of his family, presumably more or less inaccesible (
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<bibRefCitation id="081180970C3E5C51AFC9C830FD9B8A47" author="Weatherby, C. A." box="[426,617,2066,2092]" pageId="6" pageNumber="38" pagination="13 - 35" refId="ref15727" refString="Weatherby, C. A. (1936) On the types of Desvaux's American species of Ferns. Contributions from the Gray Herbarium of Harvard University 114: 13 - 35. https: // doi. org / 10.5962 / p. 336179" type="journal article" year="1936">Weatherby 1936</bibRefCitation>
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). Following Desvaux’s death in 1856, his general herbarium was sold by his son to Alphonse Lavallée (
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<bibRefCitation id="081180970C305C5FAFBCC0BCFD8782D3" author="Bureau, Ed." box="[479,629,158,184]" pageId="8" pageNumber="40" pagination="46 - 48" refId="ref13634" refString="Bureau, Ed. (1896) Note sur l'Herbier de A. - N. Desvaux. Bulletin du Muse´um d'histoire naturelle 2: 46 - 48." type="journal article" year="1896">Bureau 1896</bibRefCitation>
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,
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<bibRefCitation id="081180970C305C5FACE2C0BDFCE482D3" author="Bonnet, E." box="[641,790,158,185]" pageId="8" pageNumber="40" pagination="276 - 279" refId="ref13540" refString="Bonnet, E. (1907) Note sur l'herbier de Desvaux. Bulletin du Museum d'Histoire Naturelle 13: 276 - 279." type="journal article" year="1907">Bonnet 1907</bibRefCitation>
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). Lavallée moved the collection to Segrez (Seine-et-Oise). It was given to the Museum of Natural History, Paris, by his widow in 1896 (
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<bibRefCitation id="081180970C305C5FAD96C0E0FB7E82B7" author="Bureau, Ed." box="[1013,1164,194,220]" pageId="8" pageNumber="40" pagination="46 - 48" refId="ref13634" refString="Bureau, Ed. (1896) Note sur l'Herbier de A. - N. Desvaux. Bulletin du Muse´um d'histoire naturelle 2: 46 - 48." type="journal article" year="1896">Bureau 1896</bibRefCitation>
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). The collection has been subsequently integrated into the general herbarium (
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<bibRefCitation id="081180970C305C5FACB0C0C4FC46836B" author="Howard, R. A. & Clausen, K. S. & Gillis, W. T., Jr." box="[723,948,230,257]" pageId="8" pageNumber="40" pagination="211 - 242" refId="ref14233" refString="Howard, R. A., Clausen, K. S. & Gillis, W. T., Jr. (1981) William Hamilton (1783 - 1856) and the Prodromus Plantarum Indiae Occidentalis (1825). Journal of the Arnold Arboretum 62: 211 - 242. https: // doi. org / 10.5962 / bhl. part. 11247" type="journal article" year="1981">
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Howard
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<emphasis id="5EF421740C305C5FAD54C0C5FC9B836B" box="[823,873,230,256]" italics="true" pageId="8" pageNumber="40">et al</emphasis>
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. 1981
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</bibRefCitation>
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). Several
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<typeStatus id="B33B43C40C305C5FAA47C0C5FBAD836A" box="[1060,1119,231,257]" pageId="8" pageNumber="40">types</typeStatus>
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are preserved at P, P-JU, and PC, other material is kept at B (Willdenow), FI, G, and M (Stafleu & Cowan 1976).
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</paragraph>
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<caption id="38FFADEE0C3F5C50AEEBC783FEDB85B0" ID-DOI="http://doi.org/10.5281/zenodo.7958780" ID-Zenodo-Dep="7958780" httpUri="https://zenodo.org/record/7958780/files/figure.png" pageId="7" pageNumber="39" startId="7.[136,229,1953,1975]" targetBox="[252,1335,190,1929]" targetPageId="7" targetType="figure">
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<paragraph id="6C3FFD660C3F5C50AEEBC783FEDB85B0" blockId="7.[136,1451,1953,2011]" pageId="7" pageNumber="39">
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<emphasis id="5EF421740C3F5C50AEEBC783FF0D85DD" bold="true" box="[136,255,1953,1975]" pageId="7" pageNumber="39">FIGURE 4.</emphasis>
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Lectotype of
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<taxonomicName id="AB8086E50C3F5C50AFEAC783FD0985DD" authority="Lange, P" authorityName="Lange, P" box="[393,763,1953,1975]" class="Magnoliopsida" family="Fabaceae" form="latifolia" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="7" pageNumber="39" phylum="Tracheophyta" rank="form" species="tridentata">
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<emphasis id="5EF421740C3F5C50AFEAC783FDC785DD" box="[393,565,1953,1974]" italics="true" pageId="7" pageNumber="39">Ononis tridentata</emphasis>
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f.
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<emphasis id="5EF421740C3F5C50AC31C783FD6C85DD" box="[594,670,1953,1974]" italics="true" pageId="7" pageNumber="39">latifolia</emphasis>
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Lange, P
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</taxonomicName>
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(barcode P02993329). Image courtesy of the herbarium P, reproduced with permission.
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</paragraph>
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</caption>
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<paragraph id="6C3FFD660C305C5FAEDEC10CFEA58396" blockId="8.[136,1452,158,1627]" pageId="8" pageNumber="40">
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An unusually large proportion of Desvaux’s species have been misunderstood or left as frankly doubtful. One of these species could be
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<taxonomicName id="AB8086E50C305C5FAFD1C171FD8E8307" authorityName="Desvaux" authorityYear="1814" box="[434,636,338,365]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="40" phylum="Tracheophyta" rank="species" species="arbuscula">
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<emphasis id="5EF421740C305C5FAFD1C171FD8E8307" box="[434,636,338,365]" italics="true" pageId="8" pageNumber="40">Ononis arbuscula</emphasis>
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</taxonomicName>
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. The identity of this name has long been controversial, it could not be assigned with confidence to any known member of the group due to the ambiguity of the name and the ensuing nomenclatural confusion and uncertainty among users (see
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<bibRefCitation id="081180970C305C5FAD42C1B9FB6083DF" author="Devesa, J. A. & Lopez Gonzalez, G." box="[801,1170,410,437]" pageId="8" pageNumber="40" pagination="245 - 260" refId="ref14024" refString="Devesa, J. A. & Lopez Gonzalez, G. (1997) Notas taxonomicas y nomenclaturales sobre el genero Ononis L. (Leguminosae) en la Peninsula Iberica e Islas Baleares. Anales del Jardin Botanico de Madrid 55 (2): 245 - 260." type="journal article" year="1997">Devesa & López González 1997</bibRefCitation>
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||
). One option would have been to reject this troublesome name altogether. However, a precise typification would allow the use of the name
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<taxonomicName id="AB8086E50C305C5FAEEBC1C1FEA28397" authorityName="Desvaux" authorityYear="1814" box="[136,336,482,509]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="40" phylum="Tracheophyta" rank="species" species="arbuscula">
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<emphasis id="5EF421740C305C5FAEEBC1C1FEA28397" box="[136,336,482,509]" italics="true" pageId="8" pageNumber="40">Ononis arbuscula</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="6C3FFD660C305C5FAEDEC224FA588003" blockId="8.[136,1452,158,1627]" pageId="8" pageNumber="40">
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The protologue of
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<taxonomicName id="AB8086E50C305C5FAFEEC225FCCA804B" authority="Desvaux (1814: 76)" authorityName="Desvaux" authorityPageNumber="76" authorityYear="1814" box="[397,824,518,545]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="40" phylum="Tracheophyta" rank="species" species="arbuscula">
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<emphasis id="5EF421740C305C5FAFEEC225FDA6804B" box="[397,596,518,545]" italics="true" pageId="8" pageNumber="40">Ononis arbuscula</emphasis>
|
||
<bibRefCitation id="081180970C305C5FAC38C225FCCA804B" author="Desvaux, A. N." box="[603,824,518,545]" pageId="8" pageNumber="40" pagination="65 - 84" refId="ref13796" refString="Desvaux, A. N. (1814) Memoire et observations sur la Famille des Plantes Legumineuses. Journal de Botanique, Appliquee a l'Agriculture, a la Pharmacie, a la Medecine et aux Arts 3: 65 - 84." type="journal article" year="1814">Desvaux (1814: 76)</bibRefCitation>
|
||
</taxonomicName>
|
||
includes a description in Latin “Caule fruticoso racemis infra tuberculosis, apicè tomentosis; foliis crassis glabris subspathulatis apicè dentatis; calicibus sub-glabris; leguminibus solitariis pedunculatis hirsutis” followed by the geographical provenance “Habitât in Europeâ australis”.
|
||
</paragraph>
|
||
<paragraph id="6C3FFD660C305C5FAEDEC251FDCF81E3" blockId="8.[136,1452,158,1627]" pageId="8" pageNumber="40">
|
||
In the Willdenow herbarium at B there is a specimen of
|
||
<taxonomicName id="AB8086E50C305C5FAD54C251FC0F80E7" box="[823,1021,626,653]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="40" phylum="Tracheophyta" rank="species" species="tridentata">
|
||
<emphasis id="5EF421740C305C5FAD54C251FC0F80E7" box="[823,1021,626,653]" italics="true" pageId="8" pageNumber="40">Ononis tridentata</emphasis>
|
||
</taxonomicName>
|
||
. The sheet, with barcode B -W 13318 -00 0, bears six poorly preserved plant fragments plant of this species, and is annotated “
|
||
<taxonomicName id="AB8086E50C305C5FAAEAC2B4FAD580DB" box="[1161,1319,662,688]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="40" phylum="Tracheophyta" rank="species" species="tridentata">On. tridentata</taxonomicName>
|
||
/ 1.” at the top of the sheet, and “Roestel. W.” at the base of the sheet. The sheet also bears an original label handwritten by Willdenow “Diadelphia Decandria /
|
||
<taxonomicName id="AB8086E50C305C5FAC4BC2FCFD1E8093" box="[552,748,734,760]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="40" phylum="Tracheophyta" rank="species" species="tridentata">Ononis tridentata</taxonomicName>
|
||
fruticosa, / foliis ternatis cyanosis sub-/ linearibus tridentatis, pedun-/ culis bifloris Lin. Sp. Pl. ed R3. p. 431. / Habitat in
|
||
<taxonomicName id="AB8086E50C305C5FAD43C320FC748177" authorityName="Martínez, Criado-Fornelio, Lanzarot, Fernández-García, Rodríguez-Caabeiro & Merino" authorityYear="2004" box="[800,902,770,796]" class="Maxillopoda" family="Reighardiidae" genus="Hispania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalobaenida" pageId="8" pageNumber="40" phylum="Arthropoda" rank="genus">Hispania</taxonomicName>
|
||
” (image available at https://herbarium.bgbm.org/ object/BW13318000). However, due to the absence of original annotations by Desvaux explicitly linked to his
|
||
<taxonomicName id="AB8086E50C305C5FAB39C305FF0A810F" authorityName="Desvaux" authorityYear="1814" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="40" phylum="Tracheophyta" rank="species" species="arbuscula">
|
||
<emphasis id="5EF421740C305C5FAB39C305FF0A810F" italics="true" pageId="8" pageNumber="40">Ononis arbuscula</emphasis>
|
||
</taxonomicName>
|
||
in the sheet at B and according to Art. 9.4
|
||
<emphasis id="5EF421740C305C5FAC86C369FCF4810E" box="[741,774,843,869]" italics="true" pageId="8" pageNumber="40">(a)</emphasis>
|
||
of the
|
||
<emphasis id="5EF421740C305C5FAD38C368FBFA810F" box="[859,1032,842,868]" italics="true" pageId="8" pageNumber="40">Shenzhen Code</emphasis>
|
||
(
|
||
<bibRefCitation id="081180970C305C5FAA7AC368FB09810F" author="Turland, N. J. & Wiersema, J. H. & Barrie, F. R. & Greuter, W. & Hawksworth, D. L. & Herendeen, P. S. & Knapp, S. & Kusber, W. - H. & Li, D. - Z. & Marhold, K. & May, T. W. & McNeill, J. & Monro, A. M. & Prado, J. & Price, M. J. & Smith, G. F." box="[1049,1275,842,869]" pageId="8" pageNumber="40" refId="ref15518" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. - H., Li, D. - Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. & Smith, G. F. (Eds.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Glash ¸ tten: Koeltz Botanical Books. https: // doi. org / 10.12705 / Code. 2018" type="book" year="2018">
|
||
Turland
|
||
<emphasis id="5EF421740C305C5FAA1FC369FB5D810F" box="[1148,1199,842,868]" italics="true" pageId="8" pageNumber="40">et al</emphasis>
|
||
. 2018
|
||
</bibRefCitation>
|
||
), this specimen cannot be considered original material.
|
||
</paragraph>
|
||
<paragraph id="6C3FFD660C305C5FAEDEC3B0FBAA87EB" blockId="8.[136,1452,158,1627]" pageId="8" pageNumber="40">
|
||
Fortunately, there is a herbarium sheet at P, with barcode P03447770 that bear original material of Desvaux of
|
||
<taxonomicName id="AB8086E50C305C5FAEEBC395FEA281BB" authorityName="Desvaux" authorityYear="1814" box="[136,336,950,977]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="40" phylum="Tracheophyta" rank="species" species="arbuscula">
|
||
<emphasis id="5EF421740C305C5FAEEBC395FEA281BB" box="[136,336,950,977]" italics="true" pageId="8" pageNumber="40">Ononis arbuscula</emphasis>
|
||
</taxonomicName>
|
||
. This sheet bears three stems, with leaves, flowers and fruits, and five labels. The stem mounted on the right half of the herbarium sheet is labeled as “On[onis] tridentara /
|
||
<taxonomicName id="AB8086E50C305C5FADCDC3F8FB83819F" box="[942,1137,986,1012]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="40" phylum="Tracheophyta" rank="species" species="tridentata">Ononis tridentata</taxonomicName>
|
||
/ Cav. / en
|
||
<taxonomicName id="AB8086E50C305C5FAA91C3F8FAA4819F" authorityName="Martínez, Criado-Fornelio, Lanzarot, Fernández-García, Rodríguez-Caabeiro & Merino" authorityYear="2004" box="[1266,1366,986,1012]" class="Maxillopoda" family="Reighardiidae" genus="Hispania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalobaenida" pageId="8" pageNumber="56" phylum="Arthropoda" rank="genus">Hispania</taxonomicName>
|
||
/ prope Granatam. This specimen can be identified as belonging to
|
||
<taxonomicName id="AB8086E50C305C5FAD40C3DCFB728673" authorityName="Nyman" authorityYear="1878" baseAuthorityName="Dufour ex Boiss." box="[803,1152,1022,1048]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="55" phylum="Tracheophyta" rank="subSpecies" species="tridentata" subSpecies="crassifolia">
|
||
O.
|
||
<emphasis id="5EF421740C305C5FAD25C3DCFC468673" box="[838,948,1022,1048]" italics="true" pageId="8" pageNumber="40">tridentata</emphasis>
|
||
subsp.
|
||
<emphasis id="5EF421740C305C5FAA69C3DCFB728673" box="[1034,1152,1022,1048]" italics="true" pageId="8" pageNumber="40">crassifolia</emphasis>
|
||
</taxonomicName>
|
||
. The stem mounted on the center of the sheet is labeled as “
|
||
<taxonomicName id="AB8086E50C305C5FAF8BC400FD078657" authority="Desv." authorityName="Desvaux" authorityYear="1814" box="[488,757,1058,1084]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="40" phylum="Tracheophyta" rank="species" species="arbuscula">Ononis arbuscula Desv.</taxonomicName>
|
||
”. The stem mounted on the left half of the sheet is accompanied by two handwritten labels by Desvaux, annotated as: 1)
|
||
<taxonomicName id="AB8086E50C305C5FAD60C464FBD2860B" authority="L. Cav." authorityName="L. Cav." box="[771,1056,1094,1121]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="40" phylum="Tracheophyta" rank="species" species="tridentata">Ononis tridentata L. Cav.</taxonomicName>
|
||
ic. / 2. t. 152 / -
|
||
<taxonomicName id="AB8086E50C305C5FAA83C464FF0786EF" authority="Desv. J. Bot. 1814" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="40" phylum="Tracheophyta" rank="species" species="arbuscula">arbuscula Desv. J. Bot. 1814</taxonomicName>
|
||
/
|
||
<specimenCount id="7A8636EF0C305C5FAF6AC448FEC086EE" box="[265,306,1130,1157]" pageId="8" pageNumber="40" type="generic">1 p.</specimenCount>
|
||
76 / Mag. hort. t. 16 / hab. in
|
||
<taxonomicName id="AB8086E50C305C5FAC1BC448FD1386EF" authorityName="Martínez, Criado-Fornelio, Lanzarot, Fernández-García, Rodríguez-Caabeiro & Merino" authorityYear="2004" box="[632,737,1130,1156]" class="Maxillopoda" family="Reighardiidae" genus="Hispania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalobaenida" pageId="8" pageNumber="40" phylum="Arthropoda" rank="genus">Hispania</taxonomicName>
|
||
”, and 2) “Spec. nova / hab. Hisp.”. The sheet also bears a printed label “Herb. Mus. Paris / Herbier de A. N. Desvaux / Donné par
|
||
<superScript id="9BF5502E0C305C5FAD17C4ADFC3F86F7" attach="left" box="[884,973,1166,1193]" fontSize="6" pageId="8" pageNumber="40">Mme Vve</superScript>
|
||
Lavallée en 1896” (
|
||
<figureCitation id="F4BBE1E30C305C5FAADAC4ACFB0D86C2" box="[1209,1279,1166,1193]" captionStart="FIGURE 5" captionStartId="9.[136,229,1953,1975]" captionTargetBox="[237,1350,190,1928]" captionTargetId="figure-25@9.[237,1350,190,1928]" captionTargetPageId="9" captionText="FIGURE 5. Lectotype of Ononis tridentata subsp. arbuscula (Desvaux) P.P. Ferrer, P (barcode P03447770 [the lectotype is the stem mounted on the left half of the sheet, accompanied by two labels handwritten by Desvaux]). Image courtesy of the herbarium P, reproduced with permission." figureDoi="http://doi.org/10.5281/zenodo.7958782" httpUri="https://zenodo.org/record/7958782/files/figure.png" pageId="8" pageNumber="40">Fig. 5</figureCitation>
|
||
). Among these three stems, I designate as the
|
||
<typeStatus id="B33B43C40C305C5FAFBCC490FDB486A7" box="[479,582,1202,1228]" pageId="8" pageNumber="40">lectotype</typeStatus>
|
||
of the name
|
||
<taxonomicName id="AB8086E50C305C5FACBAC491FC5386A7" box="[729,929,1202,1229]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="40" phylum="Tracheophyta" rank="species" species="arbuscula">
|
||
<emphasis id="5EF421740C305C5FACBAC491FC5386A7" box="[729,929,1202,1229]" italics="true" pageId="8" pageNumber="40">Ononis arbuscula</emphasis>
|
||
</taxonomicName>
|
||
the stem mounted on the left half of the sheet, which bears two original labels handwritten by Desvaux. This specimen shows important diagnostic features (leaflets blade lanceolate-cuneate, less than six times longer than wider, with (0–)3(–5) teeth in the upper half, inflorescence most times 1–2-flowered, axillary; indumenta only of short glandular hairs) and matches both the historical and current concepts of this taxon (until now named under
|
||
<taxonomicName id="AB8086E50C305C5FACECC561FC128737" box="[655,992,1346,1373]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="40" phylum="Tracheophyta" rank="subSpecies" species="tridentata" subSpecies="tridentata">
|
||
<emphasis id="5EF421740C305C5FACECC561FD518736" box="[655,675,1347,1373]" italics="true" pageId="8" pageNumber="40">O</emphasis>
|
||
.
|
||
<emphasis id="5EF421740C305C5FACD3C560FCEC8737" box="[688,798,1346,1372]" italics="true" pageId="8" pageNumber="40">tridentata</emphasis>
|
||
subsp.
|
||
<emphasis id="5EF421740C305C5FAD11C560FC128737" box="[882,992,1346,1372]" italics="true" pageId="8" pageNumber="40">tridentata</emphasis>
|
||
</taxonomicName>
|
||
) (see e.g., see
|
||
<bibRefCitation id="081180970C305C5FAAE1C561FF3187EB" author="Devesa, J. A. & Lopez Gonzalez, G." pageId="8" pageNumber="40" pagination="245 - 260" refId="ref14024" refString="Devesa, J. A. & Lopez Gonzalez, G. (1997) Notas taxonomicas y nomenclaturales sobre el genero Ononis L. (Leguminosae) en la Peninsula Iberica e Islas Baleares. Anales del Jardin Botanico de Madrid 55 (2): 245 - 260." type="journal article" year="1997">Devesa & López González 1997</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="081180970C305C5FAEADC545FE9087EB" author="Devesa, J. A." box="[206,354,1382,1409]" pageId="8" pageNumber="40" pagination="590 - 646" refId="ref13871" refString="Devesa, J. A. (2000) Ononis L. In: Talavera, S., Aedo, C., Castroviejo, S., Herrero, A., Romero Zarco, C., Salgueiro, F. J. & Velayos, M. (eds.) Flora iberica, Vol. 7 (2), Leguminosae (partim). Servicio de Publicaciones del CSIC, Madrid, pp. 590 - 646." type="book chapter" year="2000">Devesa 2000</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="081180970C305C5FAF0EC544FE5A87EB" author="Devesa, J. A." box="[365,424,1382,1408]" pageId="8" pageNumber="40" refId="ref13955" refString="Devesa, J. A. (2009) Ononis L. In: Blanca, G., Cabezudo, B., Cueto, M., Fernandez-Lopez, C. & Morales-Torres, C. (eds.) Flora Vascular de Andalucia Oriental, Vol. 2. Consejeria de Medio Ambiente, Junta de Andalucia, Sevilla, pp. 387 - 398." type="book" year="2009">2009</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="081180970C305C5FAFD0C544FD5E87EB" author="Lopez Gonzalez, G." box="[435,684,1382,1408]" pageId="8" pageNumber="40" refId="ref14608" refString="Lopez Gonzalez, G. (2001) Los arboles y arbustos de la Peninsula Iberica e Islas Baleares. Tomo I & II. Ed. Mundi-Prensa, Madrid, 1728 pp." type="book" year="2001">López González 2001</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="081180970C305C5FACD4C545FC8087EB" author="Mota, J. F. & Martinez-Hernandez, F. & Guirado, J. S." box="[695,882,1382,1409]" pageId="8" pageNumber="40" refId="ref14885" refString="Mota, J. F., Martinez-Hernandez, F. & Guirado, J. S. (2011) Diversidad Vegetal de las Yeseras Ibericas. El reto de los archipielagos edaficos para la biologia de la conservacion. ADIF-Mediterraneo Asesores Consultores, Almeria, 629 pp." type="book" year="2011">
|
||
Mota
|
||
<emphasis id="5EF421740C305C5FAC9AC545FCD887EB" box="[761,810,1382,1408]" italics="true" pageId="8" pageNumber="40">et al</emphasis>
|
||
. 2011
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="081180970C305C5FAD1FC544FBBE87EB" author="Agullo, J. C. & Juan, A. & Alonso, M. A. & Terrones, A. & Crespo, M. B." box="[892,1100,1382,1409]" pageId="8" pageNumber="40" pagination="645 - 653" refId="ref13142" refString="Agullo, J. C., Juan, A., Alonso, M. A., Terrones, A. & Crespo, M. B. (2013) Taxonomic status of Ononis tridentata (Fabaceae) from Morocco, resolved by multivariate morphometric analyses. Plant Biosystems 147 (3): 645 - 653. http: // dx. doi. org / 10.1080 / 11263504.2013.776650" type="journal article" year="2013">
|
||
Agulló
|
||
<emphasis id="5EF421740C305C5FADB3C545FBF087EB" box="[976,1026,1382,1408]" italics="true" pageId="8" pageNumber="40">et al</emphasis>
|
||
. 2013
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6C3FFD660C305C5FAEDEC5A9FECC8433" blockId="8.[136,1452,158,1627]" pageId="8" pageNumber="40">
|
||
<taxonomicName id="AB8086E50C305C5FAEDEC5A9FE7687CF" box="[189,388,1418,1445]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="40" phylum="Tracheophyta" rank="species" sensu="lato" species="tridentata">
|
||
<emphasis id="5EF421740C305C5FAEDEC5A9FE7687CF" box="[189,388,1418,1445]" italics="true" pageId="8" pageNumber="40">Ononis tridentata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="45C79C0F0C305C5FAFEEC5A8FE5C87CF" box="[397,430,1418,1444]" pageId="8" pageNumber="40" sensu="lato">s.l.</taxonomicNameLabel>
|
||
includes several names validly published as nomenclatural and taxonomic synonyms, and the typification of the most relevant names is necessary for their correct application in this taxonomically difficult group. In this sense, the names
|
||
<taxonomicName id="AB8086E50C305C5FAF85C5F1FD868787" authorityName="Dufour, BORD" authorityYear="1860" box="[486,628,1490,1517]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="40" phylum="Tracheophyta" rank="species" species="barrelieri">
|
||
<emphasis id="5EF421740C305C5FAF85C5F1FD868787" box="[486,628,1490,1517]" italics="true" pageId="8" pageNumber="40">O. barrelieri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="AB8086E50C305C5FACE2C5F1FC428787" box="[641,944,1490,1517]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="40" phylum="Tracheophyta" rank="variety" species="crassifolia" variety="intricata">
|
||
<emphasis id="5EF421740C305C5FACE2C5F1FCEB8787" box="[641,793,1490,1517]" italics="true" pageId="8" pageNumber="40">O. crassifolia</emphasis>
|
||
var.
|
||
<emphasis id="5EF421740C305C5FAD33C5F1FC428787" box="[848,944,1491,1516]" italics="true" pageId="8" pageNumber="40">intricata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="AB8086E50C305C5FADDDC5F1FB058787" box="[958,1271,1490,1517]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="40" phylum="Tracheophyta" rank="variety" species="tridentata" variety="canescens">
|
||
<emphasis id="5EF421740C305C5FADDDC5F1FBBC8787" box="[958,1102,1490,1517]" italics="true" pageId="8" pageNumber="40">O. tridentata</emphasis>
|
||
var.
|
||
<emphasis id="5EF421740C305C5FAAE6C5F1FB058787" box="[1157,1271,1491,1516]" italics="true" pageId="8" pageNumber="40">canescens</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="AB8086E50C305C5FAB67C5F1FEF3847B" class="Magnoliopsida" family="Fabaceae" form="intermedia" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="40" phylum="Tracheophyta" rank="form" species="tridentata">
|
||
<emphasis id="5EF421740C305C5FAB67C5F1FA668787" box="[1284,1428,1490,1517]" italics="true" pageId="8" pageNumber="40">O. tridentata</emphasis>
|
||
f.
|
||
<emphasis id="5EF421740C305C5FAEEBC5D4FEF3847B" box="[136,257,1526,1552]" italics="true" pageId="8" pageNumber="40">intermedia</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="AB8086E50C305C5FAF5FC5D5FD28847B" box="[316,730,1526,1553]" class="Magnoliopsida" family="Fabaceae" form="erecta" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="40" phylum="Tracheophyta" rank="form" species="tridentata" variety="intermedia">
|
||
<emphasis id="5EF421740C305C5FAF5FC5D5FE39847B" box="[316,459,1526,1553]" italics="true" pageId="8" pageNumber="40">O. tridentata</emphasis>
|
||
var.
|
||
<emphasis id="5EF421740C305C5FAC63C5D4FD8B847B" box="[512,633,1526,1552]" italics="true" pageId="8" pageNumber="40">intermedia</emphasis>
|
||
f.
|
||
<emphasis id="5EF421740C305C5FACF6C5D5FD28847B" box="[661,730,1527,1552]" italics="true" pageId="8" pageNumber="40">erecta</emphasis>
|
||
</taxonomicName>
|
||
should be treated as synonyms of
|
||
<taxonomicName id="AB8086E50C305C5FAA30C5D5FA56847B" box="[1107,1444,1526,1553]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="40" phylum="Tracheophyta" rank="subSpecies" species="tridentata" subSpecies="arbuscula">
|
||
<emphasis id="5EF421740C305C5FAA30C5D5FB95847A" box="[1107,1127,1527,1553]" italics="true" pageId="8" pageNumber="40">O</emphasis>
|
||
.
|
||
<emphasis id="5EF421740C305C5FAA17C5D4FB10847B" box="[1140,1250,1526,1552]" italics="true" pageId="8" pageNumber="40">tridentata</emphasis>
|
||
subsp.
|
||
<emphasis id="5EF421740C305C5FAB57C5D4FA56847B" box="[1332,1444,1526,1552]" italics="true" pageId="8" pageNumber="40">arbuscula</emphasis>
|
||
</taxonomicName>
|
||
, comb. & stat nov., and not as synonyms of
|
||
<taxonomicName id="AB8086E50C305C5FAC0DC639FC9F845F" authority="Linnaeus" authorityName="Linnaeus" box="[622,877,1562,1589]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="40" phylum="Tracheophyta" rank="species" species="tridentata">
|
||
<emphasis id="5EF421740C305C5FAC0DC639FD0C845F" box="[622,766,1562,1589]" italics="true" pageId="8" pageNumber="40">O. tridentata</emphasis>
|
||
Linnaeus
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="45C79C0F0C305C5FAD17C639FC55845E" box="[884,935,1563,1589]" pageId="8" pageNumber="40" sensu="stricto">s.str.</taxonomicNameLabel>
|
||
due to the epitype of this name designated by F̂rther in 1997.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |