treatments-xml/data/E4/29/4C/E4294C700C395C53AEEBC596FBB8812B.xml

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<mods:title id="9E3444471AA195A2CA41EAE4E9C01333">Original elemets and typification of the names included in the Ononis tridentata complex (Fabaceae)</mods:title>
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<heading id="37774A0A0C395C56AEEBC596FCE287A4" box="[136,784,1460,1487]" fontSize="11" level="2" pageId="1" pageNumber="33" reason="3">
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<treatmentCitation id="ED21DB770C395C56AEEBC596FCE287A4" author="Linnaeus, C." box="[136,784,1460,1487]" page=")" pageId="1" pageNumber="33" year="1753">
<taxonomicName id="AB8086E50C395C56AEEBC596FCE287A4" ID-CoL="49LFF" ID-ENA="798050" authority="Linnaeus (1753: 718)" box="[136,784,1460,1487]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="33" phylum="Tracheophyta" rank="subSpecies" species="tridentata" subSpecies="tridentata">
<emphasis id="5EF421740C395C56AEEBC596FEA187A4" bold="true" box="[136,339,1460,1487]" italics="true" pageId="1" pageNumber="33">Ononis tridentata</emphasis>
<bibRefCitation id="081180970C395C56AF39C596FDB987A5" author="Linnaeus, C." box="[346,587,1460,1487]" pageId="1" pageNumber="33" refId="ref14518" refString="Linnaeus, C. (1753) Species Plantarum 1. Laurentius Salvius, Stockholm, 1200 pp. http: // dx. doi. org / 10.5962 / bhl. title. 669" type="book" year="1753">Linnaeus (1753: 718)</bibRefCitation>
subsp. 
<emphasis id="5EF421740C395C56ACC3C596FCE287A4" bold="true" box="[672,784,1460,1487]" italics="true" pageId="1" pageNumber="33">tridentata</emphasis>
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<paragraph id="6C3FFD660C395C56AEEBC5D4FB22855B" blockId="1.[136,1452,1525,2092]" pageId="1" pageNumber="33">
<treatmentCitation id="ED21DB770C395C56AEEBC5D4FDBF847B" author="Linnaeus, C." box="[136,589,1525,1552]" page=")" pageId="1" pageNumber="33" year="1753">
<taxonomicName id="AB8086E50C395C56AEEBC5D4FDBF847B" ID-CoL="49LFF" authority="Linnaeus (1753: 718)" authorityName="Linnaeus" authorityPageNumber="718" authorityYear="1753" box="[136,589,1525,1552]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="33" phylum="Tracheophyta" rank="species" species="tridentata">
<emphasis id="5EF421740C395C56AEEBC5D4FEA28464" box="[136,336,1525,1552]" italics="true" pageId="1" pageNumber="33">Ononis tridentata</emphasis>
<bibRefCitation id="081180970C395C56AF3AC5D7FDBF847B" author="Linnaeus, C." box="[345,589,1525,1552]" pageId="1" pageNumber="33" refId="ref14518" refString="Linnaeus, C. (1753) Species Plantarum 1. Laurentius Salvius, Stockholm, 1200 pp. http: // dx. doi. org / 10.5962 / bhl. title. 669" type="book" year="1753">Linnaeus (1753: 718)</bibRefCitation>
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was described in the protologue with a phrase-name “Ononis foliis ternatis carnosis sublinearibus tridentatis”. 
<bibRefCitation id="081180970C395C56AC78C63BFD24845F" author="Linnaeus, C." box="[539,726,1561,1588]" pageId="1" pageNumber="33" refId="ref14518" refString="Linnaeus, C. (1753) Species Plantarum 1. Laurentius Salvius, Stockholm, 1200 pp. http: // dx. doi. org / 10.5962 / bhl. title. 669" type="book" year="1753">Linnaeus (1753)</bibRefCitation>
stated about the origin of the plant “Habitat in 
<emphasis id="5EF421740C395C56AB65C638FA9F8458" box="[1286,1389,1562,1587]" italics="true" pageId="1" pageNumber="33">Hispania</emphasis>
” and also cited one synonym “Anonis hispanica frutescens, folio tridentato carnoso” from 
<bibRefCitation id="081180970C395C56AA06C61FFAE1843C" author="Magnol, P." box="[1125,1299,1597,1623]" pageId="1" pageNumber="33" refId="ref14674" refString="Magnol, P. (1697) Hortus regius Monspeliensis. Apud Honoratum Pech, Monspelii, 209 pp. [https: // patrimoniodigital. ucm. es / s / patrimonio / item / 733826]" type="book" year="1697">Magnol (1697)</bibRefCitation>
, 
<bibRefCitation id="081180970C395C56AB41C61CFA558433" author="Ray, J." box="[1314,1447,1598,1624]" pageId="1" pageNumber="33" refId="ref15104" refString="Ray, J. (1704) Historia Plantarum 3. Typis Mariae Clark, prostant apud Henricum Faithorne, London, 666 pp." type="book" year="1704">Ray (1704)</bibRefCitation>
, and 
<bibRefCitation id="081180970C395C56AED9C643FE7B8417" author="Tournefort, J. P. de" box="[186,393,1633,1660]" pageId="1" pageNumber="33" refId="ref15391" refString="Tournefort, J. P. de (1719) Institutiones rei herbariae ed. 3, 1. Typographia Regia, Paris, 252 pp." type="book" year="1719">Tournefort (1719)</bibRefCitation>
. The 
<typeStatus id="B33B43C40C395C56AFAFC643FDC18410" box="[460,563,1633,1659]" pageId="1" pageNumber="33">lectotype</typeStatus>
of this name was designated by F̂rther (in 
<bibRefCitation id="081180970C395C56AA58C643FAB08417" author="Turland, N. J. &amp; Jarvis, C. E." box="[1083,1346,1633,1660]" pageId="1" pageNumber="33" pagination="457 - 485" refId="ref15471" refString="Turland, N. J. &amp; Jarvis, C. E. (1997) Typification of Linnean specific and varietal names in the Leguminosae (Fabaceae). Taxon 46: 457 - 485. https: // doi. org / 10.2307 / 1224388" type="journal article" year="1997">Turland &amp; Jarvis 1997</bibRefCitation>
[August]: 478) from an illustration “Anonis Hispanica frutescens folio tridentato carnoso” published by 
<bibRefCitation id="081180970C395C56AAC8C6A7FABD84F4" author="Magnol, P." box="[1195,1359,1669,1695]" pageId="1" pageNumber="33" refId="ref14674" refString="Magnol, P. (1697) Hortus regius Monspeliensis. Apud Honoratum Pech, Monspelii, 209 pp. [https: // patrimoniodigital. ucm. es / s / patrimonio / item / 733826]" type="book" year="1697">Magnol (1697</bibRefCitation>
: tab. [4] unnumbered). The lectotypification excluded top right-hand plant, which belongs to another species (
<figureCitation id="F4BBE1E30C395C56AB79C68BFAAD84AF" box="[1306,1375,1705,1732]" captionStart="FIGURE 1" captionStartId="2.[136,229,1240,1262]" captionTargetBox="[466,1122,190,1215]" captionTargetId="figure-15@2.[466,1122,190,1215]" captionTargetPageId="2" captionText="FIGURE 1. Lectotype of Ononis tridentata L. subsp. tridentata, illustration “Anonis Hispanica frutescens folio tridentato carnoso” published by in Magnol (1697: tab. [4] unnumbered) [excluding top right-hand plant]." figureDoi="http://doi.org/10.5281/zenodo.7958774" httpUri="https://zenodo.org/record/7958774/files/figure.png" pageId="1" pageNumber="33">Fig. 1</figureCitation>
). This illustration match with the traditional concept and current use of the name as a species with leaflets blade obovate-cuneate, less than six times longer than wider, and with (0–)3–5(–9) teeth in the upper half (see e.g., 
<bibRefCitation id="081180970C395C56AB4FC6D0FA558567" author="Losa, T. M." box="[1324,1447,1777,1804]" pageId="1" pageNumber="33" pagination="227 - 337" refId="ref14642" refString="Losa, T. M. (1958) El genero Ononis y las Ononis espanolas. Anales del Instituto Botanico A. J. Cavanilles 26: 227 - 337." type="journal article" year="1958">Losa 1958</bibRefCitation>
, 
<bibRefCitation id="081180970C395C56AEEBC737FEAD8544" author="Ivimey-Cook, R. B." box="[136,351,1813,1839]" pageId="1" pageNumber="33" refId="ref14338" refString="Ivimey-Cook, R. B. (1968) Ononis L. In: Tutin, T. G., Heywoood, V. H., Burges, N. A., Valentine, D. H., Walters, S. M. &amp; Webb, D. A. (eds.) Flora Europaea, Vol. 2. Cambridge University Press, Cambridge, pp. 143 - 148." type="book" year="1968">Ivimey-Cook 1968</bibRefCitation>
, 
<bibRefCitation id="081180970C395C56AF09C734FD2B855B" author="Devesa, J. A. &amp; Lopez Gonzalez, G." box="[362,729,1813,1840]" pageId="1" pageNumber="33" pagination="245 - 260" refId="ref14024" refString="Devesa, J. A. &amp; Lopez Gonzalez, G. (1997) Notas taxonomicas y nomenclaturales sobre el genero Ononis L. (Leguminosae) en la Peninsula Iberica e Islas Baleares. Anales del Jardin Botanico de Madrid 55 (2): 245 - 260." type="journal article" year="1997">Devesa &amp; López González 1997</bibRefCitation>
, 
<bibRefCitation id="081180970C395C56AC87C734FC8B855B" author="Devesa, J. A." box="[740,889,1814,1840]" pageId="1" pageNumber="33" pagination="590 - 646" refId="ref13871" refString="Devesa, J. A. (2000) Ononis L. In: Talavera, S., Aedo, C., Castroviejo, S., Herrero, A., Romero Zarco, C., Salgueiro, F. J. &amp; Velayos, M. (eds.) Flora iberica, Vol. 7 (2), Leguminosae (partim). Servicio de Publicaciones del CSIC, Madrid, pp. 590 - 646." type="book chapter" year="2000">Devesa 2000</bibRefCitation>
, 
<bibRefCitation id="081180970C395C56ADE7C734FC4D855B" author="Devesa, J. A." box="[900,959,1814,1840]" pageId="1" pageNumber="33" refId="ref13955" refString="Devesa, J. A. (2009) Ononis L. In: Blanca, G., Cabezudo, B., Cueto, M., Fernandez-Lopez, C. &amp; Morales-Torres, C. (eds.) Flora Vascular de Andalucia Oriental, Vol. 2. Consejeria de Medio Ambiente, Junta de Andalucia, Sevilla, pp. 387 - 398." type="book" year="2009">2009</bibRefCitation>
, 
<bibRefCitation id="081180970C395C56ADA9C734FB31855B" author="Lopez Gonzalez, G." box="[970,1219,1813,1840]" pageId="1" pageNumber="33" refId="ref14608" refString="Lopez Gonzalez, G. (2001) Los arboles y arbustos de la Peninsula Iberica e Islas Baleares. Tomo I &amp; II. Ed. Mundi-Prensa, Madrid, 1728 pp." type="book" year="2001">López González 2001</bibRefCitation>
).
</paragraph>
<paragraph id="6C3FFD660C395C56AEDEC718FAD48A47" blockId="1.[136,1452,1525,2092]" pageId="1" pageNumber="33">
On the other hand, 
<bibRefCitation id="081180970C395C56AFF6C718FCF88538" author="Devesa, J. A. &amp; Lopez Gonzalez, G." box="[405,778,1849,1876]" pageId="1" pageNumber="33" pagination="245 - 260" refId="ref14024" refString="Devesa, J. A. &amp; Lopez Gonzalez, G. (1997) Notas taxonomicas y nomenclaturales sobre el genero Ononis L. (Leguminosae) en la Peninsula Iberica e Islas Baleares. Anales del Jardin Botanico de Madrid 55 (2): 245 - 260." type="journal article" year="1997">Devesa &amp; López González (1997</bibRefCitation>
[December]: 254) designated as the “ 
<typeStatus id="B33B43C40C395C56AAC9C71BFAE48538" box="[1194,1302,1849,1875]" pageId="1" pageNumber="33">lectotype</typeStatus>
” of the name the same illustration, as “ 
<typeStatus id="B33B43C40C395C56AFC9C77CFDDB8513" box="[426,553,1886,1912]" pageId="1" pageNumber="33">Lectotypus</typeStatus>
(Iconotypus): Pierre Magnol, Hortus Regius Monspeliensis, t. [4], 1697, excl. ram. ster. [Designado en este trabajo]”. Although this designation is later than the one published by F̂rther, and therefore ineffective, these authors make an exhaustive exposition about the possible origin of the material used as voucher for Magnol’s illustration (1697: tab. [4] unnumbered). These authors concluded that the origin of the plant cultivated in the Botanical Garden of Montpelier used as voucher could come from seeds collected by Tournefort in Cardona (
<collectingRegion id="AE4433840C395C56AE95C833FE948A40" box="[246,358,2065,2091]" country="Spain" name="Catalunya" pageId="1" pageNumber="33">Catalonia</collectingRegion>
, 
<collectingCountry id="1497BDF60C395C56AF12C833FE408A40" box="[369,434,2065,2091]" name="Spain" pageId="1" pageNumber="33">Spain</collectingCountry>
) during his trip to 
<collectingCountry id="1497BDF60C395C56ACE0C833FD318A40" box="[643,707,2065,2091]" name="Spain" pageId="1" pageNumber="33">Spain</collectingCountry>
in 1687 (see 
<bibRefCitation id="081180970C395C56AD39C833FBFD8A47" author="Henriques, J." box="[858,1039,2065,2092]" pageId="1" pageNumber="33" pagination="108 - 174" refId="ref14209" refString="Henriques, J. (1898) Exploracies botanicas em Hespanha por Tournefort. Boletim da Sociedade Broteriana 15: 108 - 174." type="journal article" year="1898">Henriques 1898</bibRefCitation>
, 
<bibRefCitation id="081180970C395C56AA79C830FAEB8A47" author="Devesa, J. A. &amp; Lopez Gonzalez, G." box="[1050,1305,2066,2092]" pageId="1" pageNumber="33" pagination="245 - 260" refId="ref14024" refString="Devesa, J. A. &amp; Lopez Gonzalez, G. (1997) Notas taxonomicas y nomenclaturales sobre el genero Ononis L. (Leguminosae) en la Peninsula Iberica e Islas Baleares. Anales del Jardin Botanico de Madrid 55 (2): 245 - 260." type="journal article" year="1997">Devesa &amp; López 1997</bibRefCitation>
).
</paragraph>
<caption id="38FFADEE0C3A5C55AEEBC4FAFC318779" ID-DOI="http://doi.org/10.5281/zenodo.7958774" ID-Zenodo-Dep="7958774" httpUri="https://zenodo.org/record/7958774/files/figure.png" pageId="2" pageNumber="34" startId="2.[136,229,1240,1262]" targetBox="[466,1122,190,1215]" targetPageId="2" targetType="figure">
<paragraph id="6C3FFD660C3A5C55AEEBC4FAFC318779" blockId="2.[136,1451,1240,1298]" pageId="2" pageNumber="34">
<emphasis id="5EF421740C3A5C55AEEBC4FAFEF38686" bold="true" box="[136,257,1240,1262]" pageId="2" pageNumber="34">FIGURE 1.</emphasis>
Lectotype of 
<taxonomicName id="AB8086E50C3A5C55AFF2C4FBFDCD8686" box="[401,575,1240,1262]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="34" phylum="Tracheophyta" rank="species" species="tridentata">
<emphasis id="5EF421740C3A5C55AFF2C4FBFDCD8686" box="[401,575,1240,1262]" italics="true" pageId="2" pageNumber="34">Ononis tridentata</emphasis>
</taxonomicName>
L. 
<taxonomicName id="AB8086E50C3A5C55AC05C4FAFCFE8686" box="[614,780,1240,1262]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="34" phylum="Tracheophyta" rank="subSpecies" species="tridentata" subSpecies="tridentata">
subsp. 
<emphasis id="5EF421740C3A5C55ACCEC4FAFCFE8686" box="[685,780,1240,1261]" italics="true" pageId="2" pageNumber="34">tridentata</emphasis>
</taxonomicName>
, illustration “Anonis Hispanica frutescens folio tridentato carnoso” published by in 
<bibRefCitation id="081180970C3A5C55AF41C4DEFE5C8779" author="Magnol, P." box="[290,430,1276,1298]" pageId="2" pageNumber="34" refId="ref14674" refString="Magnol, P. (1697) Hortus regius Monspeliensis. Apud Honoratum Pech, Monspelii, 209 pp. [https: // patrimoniodigital. ucm. es / s / patrimonio / item / 733826]" type="book" year="1697">Magnol (1697</bibRefCitation>
: tab. [4] unnumbered) [excluding top right-hand plant].
</paragraph>
</caption>
<paragraph id="6C3FFD660C3A5C55AEDEC563FCFD8458" blockId="2.[136,1458,1345,2093]" pageId="2" pageNumber="34">
On the other hand, F̂rther (in 
<bibRefCitation id="081180970C3A5C55AC4BC563FC9F8730" author="Turland, N. J. &amp; Jarvis, C. E." box="[552,877,1345,1371]" pageId="2" pageNumber="34" pagination="457 - 485" refId="ref15471" refString="Turland, N. J. &amp; Jarvis, C. E. (1997) Typification of Linnean specific and varietal names in the Leguminosae (Fabaceae). Taxon 46: 457 - 485. https: // doi. org / 10.2307 / 1224388" type="journal article" year="1997">Turland &amp; Jarvis 1997: 478</bibRefCitation>
) designates an epitype of 
<taxonomicName id="AB8086E50C3A5C55AAC2C563FA988730" box="[1185,1386,1345,1371]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="34" phylum="Tracheophyta" rank="species" species="tridentata">
<emphasis id="5EF421740C3A5C55AAC2C563FA988730" box="[1185,1386,1345,1371]" italics="true" pageId="2" pageNumber="34">Ononis tridentata</emphasis>
</taxonomicName>
from a specimen preserved at M, as “ 
<collectingCountry id="1497BDF60C3A5C55AF99C547FDCC8714" box="[506,574,1381,1407]" name="Spain" pageId="2" pageNumber="34">Spain</collectingCountry>
, Granada, in salsis et collibus argillosis La Mala prope Granatam, 
<date id="183EDBA60C3A5C55AB21C547FA558714" box="[1346,1447,1381,1407]" pageId="2" pageNumber="34" value="1837-07">Jul 1837</date>
, 
<emphasis id="5EF421740C3A5C55AEEBC5ABFF1687C9" box="[136,228,1417,1442]" italics="true" pageId="2" pageNumber="34">Boissier</emphasis>
(M)”. Unfortunately, this epitypification of 
<taxonomicName id="AB8086E50C3A5C55ACA8C5ABFCA887C8" box="[715,858,1417,1443]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="34" phylum="Tracheophyta" rank="species" species="tridentata">
<emphasis id="5EF421740C3A5C55ACA8C5ABFD2D87C8" box="[715,735,1417,1443]" italics="true" pageId="2" pageNumber="34">O</emphasis>
. 
<emphasis id="5EF421740C3A5C55AC8FC5ABFCA887C8" box="[748,858,1417,1443]" italics="true" pageId="2" pageNumber="34">tridentata</emphasis>
</taxonomicName>
published by F̂rther (in 
<bibRefCitation id="081180970C3A5C55AA11C5ABFA5687C8" author="Turland, N. J. &amp; Jarvis, C. E." box="[1138,1444,1417,1443]" pageId="2" pageNumber="34" pagination="457 - 485" refId="ref15471" refString="Turland, N. J. &amp; Jarvis, C. E. (1997) Typification of Linnean specific and varietal names in the Leguminosae (Fabaceae). Taxon 46: 457 - 485. https: // doi. org / 10.2307 / 1224388" type="journal article" year="1997">Turland &amp; Jarvis 1997: 478</bibRefCitation>
) has gone unnoticed (see e.g., 
<bibRefCitation id="081180970C3A5C55AFB3C58FFD9487AC" author="Devesa, J. A." box="[464,614,1453,1479]" pageId="2" pageNumber="34" pagination="590 - 646" refId="ref13871" refString="Devesa, J. A. (2000) Ononis L. In: Talavera, S., Aedo, C., Castroviejo, S., Herrero, A., Romero Zarco, C., Salgueiro, F. J. &amp; Velayos, M. (eds.) Flora iberica, Vol. 7 (2), Leguminosae (partim). Servicio de Publicaciones del CSIC, Madrid, pp. 590 - 646." type="book chapter" year="2000">Devesa 2000</bibRefCitation>
). In the herbarium M there are 
<specimenCount id="7A8636EF0C3A5C55ADDCC58FFB9787AC" box="[959,1125,1453,1479]" pageId="2" pageNumber="34" type="generic">two specimens</specimenCount>
of the gathering indicated by F̂rther, barcoded as M0232961 and M0232962 (
<figureCitation id="F4BBE1E30C3A5C55ACC4C5F3FD198780" box="[679,747,1489,1515]" captionStart="FIGURE 2" captionStartId="3.[136,229,1952,1974]" captionTargetBox="[231,1356,190,1928]" captionTargetId="figure-25@3.[231,1356,190,1928]" captionTargetPageId="3" captionText="FIGURE 2. Epitype of Ononis tridentata L. subsp. tridentata, M (barcode M0232962). Image courtesy of the herbarium M, reproduced with permission." figureDoi="http://doi.org/10.5281/zenodo.7958776" httpUri="https://zenodo.org/record/7958776/files/figure.png" pageId="2" pageNumber="34">Fig. 2</figureCitation>
). Therefore, the epitypification proposed by F̂rther (in 
<bibRefCitation id="081180970C3A5C55AB37C5F3FED78464" author="Turland, N. J. &amp; Jarvis, C. E." pageId="2" pageNumber="34" pagination="457 - 485" refId="ref15471" refString="Turland, N. J. &amp; Jarvis, C. E. (1997) Typification of Linnean specific and varietal names in the Leguminosae (Fabaceae). Taxon 46: 457 - 485. https: // doi. org / 10.2307 / 1224388" type="journal article" year="1997">Turland &amp; Jarvis 1997</bibRefCitation>
) may be further narrowed to a single specimen by a “second-step” epitypification according to Art. 9.17 of the 
<emphasis id="5EF421740C3A5C55AEB3C63BFF0D8458" box="[208,255,1561,1587]" italics="true" pageId="2" pageNumber="34">ICN</emphasis>
(
<bibRefCitation id="081180970C3A5C55AF6EC63BFE148458" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. - H. &amp; Li, D. - Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." box="[269,486,1561,1587]" pageId="2" pageNumber="34" refId="ref15518" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. - H., Li, D. - Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (Eds.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Glash ¸ tten: Koeltz Botanical Books. https: // doi. org / 10.12705 / Code. 2018" type="book" year="2018">
Turland 
<emphasis id="5EF421740C3A5C55AF0EC63BFE6C8458" box="[365,414,1561,1587]" italics="true" pageId="2" pageNumber="34">et al</emphasis>
. 2018
</bibRefCitation>
) (see more details below).
</paragraph>
<paragraph id="6C3FFD660C3A5C55AEDEC61FFAFE8544" blockId="2.[136,1458,1345,2093]" pageId="2" pageNumber="34">
Consequently, under Art. 9.20, first sentence, and Art. 9.9, the epitype selected by F̂rther (in 
<bibRefCitation id="081180970C3A5C55AA8FC61FFF338410" author="Turland, N. J. &amp; Jarvis, C. E." pageId="2" pageNumber="34" pagination="457 - 485" refId="ref15471" refString="Turland, N. J. &amp; Jarvis, C. E. (1997) Typification of Linnean specific and varietal names in the Leguminosae (Fabaceae). Taxon 46: 457 - 485. https: // doi. org / 10.2307 / 1224388" type="journal article" year="1997">Turland &amp; Jarvis 1997</bibRefCitation>
) must determine the precise application of 
<taxonomicName id="AB8086E50C3A5C55ACC5C643FC9E8410" box="[678,876,1633,1659]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="34" phylum="Tracheophyta" rank="species" sensu="stricto" species="tridentata">
<emphasis id="5EF421740C3A5C55ACC5C643FC9E8410" box="[678,876,1633,1659]" italics="true" pageId="2" pageNumber="34">Ononis tridentata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45C79C0F0C3A5C55AD10C643FC558410" box="[883,935,1633,1659]" pageId="2" pageNumber="34" sensu="stricto">s.str.</taxonomicNameLabel>
Although in the epityping it is not mentioned, we assume that F̂rther select an epitype because the 
<typeStatus id="B33B43C40C3A5C55AC8BC6A7FCBD84F4" box="[744,847,1669,1695]" pageId="2" pageNumber="34">lectotype</typeStatus>
“is demonstrably ambiguous and cannot be critically identified for purposes of the precise application of the name to a taxon” (see 
<emphasis id="5EF421740C3A5C55AD93C68BFBED84A8" box="[1008,1055,1705,1731]" italics="true" pageId="2" pageNumber="34">ICN</emphasis>
Art. 9.9). However, in our opinion, the epitification was not a good choice, since the 
<typeStatus id="B33B43C40C3A5C55ACD0C6EFFCE8848C" box="[691,794,1741,1767]" pageId="2" pageNumber="34">lectotype</typeStatus>
of the name, the illustration “Anonis Hispanica frutescens folio tridentato carnoso” in 
<bibRefCitation id="081180970C3A5C55AFABC6D3FD9C8560" author="Magnol, P." box="[456,622,1777,1803]" pageId="2" pageNumber="34" refId="ref14674" refString="Magnol, P. (1697) Hortus regius Monspeliensis. Apud Honoratum Pech, Monspelii, 209 pp. [https: // patrimoniodigital. ucm. es / s / patrimonio / item / 733826]" type="book" year="1697">Magnol (1697</bibRefCitation>
: tab. [4] unnumbered) can be critically identified for the purposes of its taxonomic identification as was clearly and explicitly mentioned by 
<bibRefCitation id="081180970C3A5C55ADE5C737FAF58544" author="Devesa, J. A. &amp; Lopez Gonzalez, G." box="[902,1287,1813,1839]" pageId="2" pageNumber="34" pagination="245 - 260" refId="ref14024" refString="Devesa, J. A. &amp; Lopez Gonzalez, G. (1997) Notas taxonomicas y nomenclaturales sobre el genero Ononis L. (Leguminosae) en la Peninsula Iberica e Islas Baleares. Anales del Jardin Botanico de Madrid 55 (2): 245 - 260." type="journal article" year="1997">Devesa &amp; López González (1997)</bibRefCitation>
.
</paragraph>
<paragraph id="6C3FFD660C3A5C53AEDEC71BFBFD836A" blockId="2.[136,1458,1345,2093]" lastBlockId="4.[136,1452,158,833]" lastPageId="4" lastPageNumber="36" pageId="2" pageNumber="34">
Once an epitype is designated in accordance with the provisions of the 
<emphasis id="5EF421740C3A5C55AD88C71BFB6A8538" box="[1003,1176,1849,1875]" italics="true" pageId="2" pageNumber="34">Shenzhen Code</emphasis>
, the identity of the type it supports (in this case the 
<typeStatus id="B33B43C40C3A5C55AFAEC77FFDC4851C" box="[461,566,1885,1911]" pageId="2" pageNumber="34">lectotype</typeStatus>
) is totally irrelevant for the purpose of application of the name. In addition, that although the type being supported by an epitype should be “demonstrably ambiguous and cannot be critically identified for purposes of the precise application of the name to a taxon” (Art. 9.9), even if there is evidence that this is demonstrably not the case (hardly the situation here), there is no provision in the 
<emphasis id="5EF421740C3A5C55AA65C7EBFBB38588" box="[1030,1089,1993,2019]" italics="true" pageId="2" pageNumber="34">Code</emphasis>
for the designated epitype not to continue to determine the application of the name. Therefore, under the current rules 
<taxonomicName id="AB8086E50C3A5C55AA27C7CFFB278A6C" box="[1092,1237,2029,2055]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="34" phylum="Tracheophyta" rank="species" species="tridentata">
<emphasis id="5EF421740C3A5C55AA27C7CFFBAA8A6C" box="[1092,1112,2029,2055]" italics="true" pageId="2" pageNumber="34">O</emphasis>
. 
<emphasis id="5EF421740C3A5C55AA04C7CFFB278A6C" box="[1127,1237,2029,2055]" italics="true" pageId="2" pageNumber="34">tridentata</emphasis>
</taxonomicName>
must be applied to the species to which F̂rther’s epitype belongs (Art. 9.9 (“An epitype is a specimen or illustration selected to serve as an interpretative type ...”) and Art. 9.20 (“The author who first designates (Art. 7.10, 7.11, and F.5.4) an epitype must be followed.”). Only if the type that an epitype supports is displaced (lost, destroyed, or supersedable under Art. 9.19) does an epitype cease to be the element determining the application of the name.
</paragraph>
<caption id="38FFADEE0C3B5C54AEEBC782FEDB85B0" ID-DOI="http://doi.org/10.5281/zenodo.7958776" ID-Zenodo-Dep="7958776" httpUri="https://zenodo.org/record/7958776/files/figure.png" pageId="3" pageNumber="35" startId="3.[136,229,1952,1974]" targetBox="[231,1356,190,1928]" targetPageId="3" targetType="figure">
<paragraph id="6C3FFD660C3B5C54AEEBC782FEDB85B0" blockId="3.[136,1451,1952,2011]" pageId="3" pageNumber="35">
<emphasis id="5EF421740C3B5C54AEEBC782FF0C85DE" bold="true" box="[136,254,1952,1974]" pageId="3" pageNumber="35">FIGURE 2.</emphasis>
Epitype of 
<taxonomicName id="AB8086E50C3B5C54AF0CC783FDE885DD" box="[367,538,1953,1974]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="35" phylum="Tracheophyta" rank="species" species="tridentata">
<emphasis id="5EF421740C3B5C54AF0CC783FDE885DD" box="[367,538,1953,1974]" italics="true" pageId="3" pageNumber="35">Ononis tridentata</emphasis>
</taxonomicName>
L. 
<taxonomicName id="AB8086E50C3B5C54AC58C783FD2F85DD" box="[571,733,1953,1975]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="35" phylum="Tracheophyta" rank="subSpecies" species="tridentata" subSpecies="tridentata">
subsp. 
<emphasis id="5EF421740C3B5C54AC1DC783FD2F85DD" box="[638,733,1953,1974]" italics="true" pageId="3" pageNumber="35">tridentata</emphasis>
</taxonomicName>
, M (barcode M0232962). Image courtesy of the herbarium M, reproduced with permission.
</paragraph>
</caption>
<paragraph id="6C3FFD660C3C5C53AEDEC128FB4F83B3" blockId="4.[136,1452,158,833]" pageId="4" pageNumber="36">
The epitype designated by F̂rther (in 
<bibRefCitation id="081180970C3C5C53AC0EC128FC99834F" author="Turland, N. J. &amp; Jarvis, C. E." box="[621,875,266,292]" pageId="4" pageNumber="36" pagination="457 - 485" refId="ref15471" refString="Turland, N. J. &amp; Jarvis, C. E. (1997) Typification of Linnean specific and varietal names in the Leguminosae (Fabaceae). Taxon 46: 457 - 485. https: // doi. org / 10.2307 / 1224388" type="journal article" year="1997">Turland &amp; Jarvis 1997</bibRefCitation>
) “ 
<collectingCountry id="1497BDF60C3C5C53ADE6C128FC3B834F" box="[901,969,266,292]" name="Spain" pageId="4" pageNumber="36">Spain</collectingCountry>
, Granada, in salsis et collibus argillosis La Mala prope Granatam, 
<date id="183EDBA60C3C5C53AFE9C10CFE1E8323" box="[394,492,302,328]" pageId="4" pageNumber="36" value="1837-07">Jul 1837</date>
, 
<emphasis id="5EF421740C3C5C53AF94C10DFDA18323" box="[503,595,303,328]" italics="true" pageId="4" pageNumber="36">Boissier</emphasis>
(M)” is undoubtedly identifiable with the traditional concept and current use to which the name 
<taxonomicName id="AB8086E50C3C5C53AF38C171FD478307" authorityName="Nyman" authorityYear="1878" baseAuthorityName="Dufour ex Boiss." box="[347,693,338,365]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="36" phylum="Tracheophyta" rank="subSpecies" species="tridentata" subSpecies="crassifolia">
<emphasis id="5EF421740C3C5C53AF38C171FE198307" box="[347,491,338,365]" italics="true" pageId="4" pageNumber="36">O. tridentata</emphasis>
subsp. 
<emphasis id="5EF421740C3C5C53AC5CC170FD478307" box="[575,693,338,364]" italics="true" pageId="4" pageNumber="36">crassifolia</emphasis>
</taxonomicName>
is generally applied (leaflets blade lanceolate–cuneate, less than six times longer than wider, with 3–9 teeth in the upper half, sepals entire, inflorescence peduncle shorter than 
<quantity id="AB7850830C3C5C53AB37C154FA5583FA" box="[1364,1447,374,401]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="36" unit="mm" value="10.0">10 mm</quantity>
, usually with an inconspicuous arista, inflorescence 1–2-flowered and raceme-like, indumenta of short glandular hairs and sessile glands) (see 
<bibRefCitation id="081180970C3C5C53AFF6C19DFCF683B3" author="Devesa, J. A. &amp; Lopez Gonzalez, G." box="[405,772,446,473]" pageId="4" pageNumber="36" pagination="245 - 260" refId="ref14024" refString="Devesa, J. A. &amp; Lopez Gonzalez, G. (1997) Notas taxonomicas y nomenclaturales sobre el genero Ononis L. (Leguminosae) en la Peninsula Iberica e Islas Baleares. Anales del Jardin Botanico de Madrid 55 (2): 245 - 260." type="journal article" year="1997">Devesa &amp; López González 1997</bibRefCitation>
, 
<bibRefCitation id="081180970C3C5C53AD6CC19DFC5183B3" author="Devesa, J. A." box="[783,931,446,473]" pageId="4" pageNumber="36" pagination="590 - 646" refId="ref13871" refString="Devesa, J. A. (2000) Ononis L. In: Talavera, S., Aedo, C., Castroviejo, S., Herrero, A., Romero Zarco, C., Salgueiro, F. J. &amp; Velayos, M. (eds.) Flora iberica, Vol. 7 (2), Leguminosae (partim). Servicio de Publicaciones del CSIC, Madrid, pp. 590 - 646." type="book chapter" year="2000">Devesa 2000</bibRefCitation>
, 
<bibRefCitation id="081180970C3C5C53ADCDC19CFC1B83B3" author="Devesa, J. A." box="[942,1001,446,472]" pageId="4" pageNumber="36" refId="ref13955" refString="Devesa, J. A. (2009) Ononis L. In: Blanca, G., Cabezudo, B., Cueto, M., Fernandez-Lopez, C. &amp; Morales-Torres, C. (eds.) Flora Vascular de Andalucia Oriental, Vol. 2. Consejeria de Medio Ambiente, Junta de Andalucia, Sevilla, pp. 387 - 398." type="book" year="2009">2009</bibRefCitation>
, 
<bibRefCitation id="081180970C3C5C53AD97C19DFB4383B3" author="Mota, J. F. &amp; Martinez-Hernandez, F. &amp; Guirado, J. S." box="[1012,1201,446,473]" pageId="4" pageNumber="36" refId="ref14885" refString="Mota, J. F., Martinez-Hernandez, F. &amp; Guirado, J. S. (2011) Diversidad Vegetal de las Yeseras Ibericas. El reto de los archipielagos edaficos para la biologia de la conservacion. ADIF-Mediterraneo Asesores Consultores, Almeria, 629 pp." type="book" year="2011">
Mota 
<emphasis id="5EF421740C3C5C53AA55C19DFB9A83B3" box="[1078,1128,446,472]" italics="true" pageId="4" pageNumber="36">et al</emphasis>
. 2011
</bibRefCitation>
).
</paragraph>
<paragraph id="6C3FFD660C3C5C53AEDEC1C1FBB8812B" blockId="4.[136,1452,158,833]" pageId="4" pageNumber="36">
However, the epitypification from the specimen at M generates an abrupt and relevant change in the nomenclature of 
<taxonomicName id="AB8086E50C3C5C53AECBC225FEC8804B" box="[168,314,518,545]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="36" phylum="Tracheophyta" rank="species" species="tridentata">
<emphasis id="5EF421740C3C5C53AECBC225FF4E804A" box="[168,188,519,545]" italics="true" pageId="4" pageNumber="36">O</emphasis>
. 
<emphasis id="5EF421740C3C5C53AEAFC224FEC8804B" box="[204,314,518,544]" italics="true" pageId="4" pageNumber="36">tridentata</emphasis>
</taxonomicName>
complex, because the name 
<taxonomicName id="AB8086E50C3C5C53ACE7C225FC2F804B" box="[644,989,518,545]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="36" phylum="Tracheophyta" rank="subSpecies" species="tridentata" subSpecies="tridentata">
<emphasis id="5EF421740C3C5C53ACE7C225FD6A804A" box="[644,664,519,545]" italics="true" pageId="4" pageNumber="36">O</emphasis>
. 
<emphasis id="5EF421740C3C5C53ACCBC224FCE4804B" box="[680,790,518,544]" italics="true" pageId="4" pageNumber="36">tridentata</emphasis>
subsp. 
<emphasis id="5EF421740C3C5C53AD0CC224FC2F804B" box="[879,989,518,544]" italics="true" pageId="4" pageNumber="36">tridentata</emphasis>
</taxonomicName>
must be applied to populations endemic to the southeastern Iberian Peninsula (with diagnostic characters mentioned above), and another name (in subspecies rank, see below) should be proposed to name the Iberian Peninsula populations traditionally called as 
<taxonomicName id="AB8086E50C3C5C53AB7AC26DFEB680E7" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="36" phylum="Tracheophyta" rank="subSpecies" species="tridentata" subSpecies="tridentata">
<emphasis id="5EF421740C3C5C53AB7AC26DFADF8002" box="[1305,1325,591,617]" italics="true" pageId="4" pageNumber="36">O</emphasis>
. 
<emphasis id="5EF421740C3C5C53AB5EC26CFA598003" box="[1341,1451,590,616]" italics="true" pageId="4" pageNumber="36">tridentata</emphasis>
subsp. 
<emphasis id="5EF421740C3C5C53AEB5C250FEB680E7" box="[214,324,626,652]" italics="true" pageId="4" pageNumber="36">tridentata</emphasis>
</taxonomicName>
, with leaflets blade lanceolate–cuneate, less than six times longer than wider, with (0–)3(–5) teeth in the upper half, inflorescence most times 1–2-flowered, axillary; indumenta only of short glandular hairs (see 
<bibRefCitation id="081180970C3C5C53AB5EC2B5FE8D80BF" author="Devesa, J. A. &amp; Lopez Gonzalez, G." pageId="4" pageNumber="36" pagination="245 - 260" refId="ref14024" refString="Devesa, J. A. &amp; Lopez Gonzalez, G. (1997) Notas taxonomicas y nomenclaturales sobre el genero Ononis L. (Leguminosae) en la Peninsula Iberica e Islas Baleares. Anales del Jardin Botanico de Madrid 55 (2): 245 - 260." type="journal article" year="1997">Devesa &amp; López González 1997</bibRefCitation>
, 
<bibRefCitation id="081180970C3C5C53AFEAC299FDEE80BF" author="Devesa, J. A." box="[393,540,698,725]" pageId="4" pageNumber="36" pagination="590 - 646" refId="ref13871" refString="Devesa, J. A. (2000) Ononis L. In: Talavera, S., Aedo, C., Castroviejo, S., Herrero, A., Romero Zarco, C., Salgueiro, F. J. &amp; Velayos, M. (eds.) Flora iberica, Vol. 7 (2), Leguminosae (partim). Servicio de Publicaciones del CSIC, Madrid, pp. 590 - 646." type="book chapter" year="2000">Devesa 2000</bibRefCitation>
, 
<bibRefCitation id="081180970C3C5C53AC45C298FD9380BF" author="Devesa, J. A." box="[550,609,698,724]" pageId="4" pageNumber="36" refId="ref13955" refString="Devesa, J. A. (2009) Ononis L. In: Blanca, G., Cabezudo, B., Cueto, M., Fernandez-Lopez, C. &amp; Morales-Torres, C. (eds.) Flora Vascular de Andalucia Oriental, Vol. 2. Consejeria de Medio Ambiente, Junta de Andalucia, Sevilla, pp. 387 - 398." type="book" year="2009">2009</bibRefCitation>
, 
<bibRefCitation id="081180970C3C5C53AC08C298FC9080BF" author="Lopez Gonzalez, G." box="[619,866,698,724]" pageId="4" pageNumber="36" refId="ref14608" refString="Lopez Gonzalez, G. (2001) Los arboles y arbustos de la Peninsula Iberica e Islas Baleares. Tomo I &amp; II. Ed. Mundi-Prensa, Madrid, 1728 pp." type="book" year="2001">López González 2001</bibRefCitation>
, 
<bibRefCitation id="081180970C3C5C53AD0FC299FBD680BF" author="Mota, J. F. &amp; Martinez-Hernandez, F. &amp; Guirado, J. S." box="[876,1060,698,725]" pageId="4" pageNumber="36" refId="ref14885" refString="Mota, J. F., Martinez-Hernandez, F. &amp; Guirado, J. S. (2011) Diversidad Vegetal de las Yeseras Ibericas. El reto de los archipielagos edaficos para la biologia de la conservacion. ADIF-Mediterraneo Asesores Consultores, Almeria, 629 pp." type="book" year="2011">
Mota 
<emphasis id="5EF421740C3C5C53ADCEC299FC2C80BF" box="[941,990,698,724]" italics="true" pageId="4" pageNumber="36">et al</emphasis>
. 2011
</bibRefCitation>
, 
<bibRefCitation id="081180970C3C5C53AA4EC298FB0980BF" author="Agullo, J. C. &amp; Juan, A. &amp; Alonso, M. A. &amp; Terrones, A. &amp; Crespo, M. B." box="[1069,1275,698,725]" pageId="4" pageNumber="36" pagination="645 - 653" refId="ref13142" refString="Agullo, J. C., Juan, A., Alonso, M. A., Terrones, A. &amp; Crespo, M. B. (2013) Taxonomic status of Ononis tridentata (Fabaceae) from Morocco, resolved by multivariate morphometric analyses. Plant Biosystems 147 (3): 645 - 653. http: // dx. doi. org / 10.1080 / 11263504.2013.776650" type="journal article" year="2013">
Agulló 
<emphasis id="5EF421740C3C5C53AAE2C299FB4380BF" box="[1153,1201,698,724]" italics="true" pageId="4" pageNumber="36">et al</emphasis>
. 2013
</bibRefCitation>
). Consequently, the names 
<taxonomicName id="AB8086E50C3C5C53AF62C2FDFE628093" authorityName="Dufour, BORD" authorityYear="1860" box="[257,400,734,761]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="36" phylum="Tracheophyta" rank="species" species="barrelieri">
<emphasis id="5EF421740C3C5C53AF62C2FDFE628093" box="[257,400,734,761]" italics="true" pageId="4" pageNumber="36">O. barrelieri</emphasis>
</taxonomicName>
, 
<taxonomicName id="AB8086E50C3C5C53AFFDC2FDFD3D8093" box="[414,719,734,761]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="36" phylum="Tracheophyta" rank="variety" species="crassifolia" variety="intricata">
<emphasis id="5EF421740C3C5C53AFFDC2FDFDC58093" box="[414,567,734,761]" italics="true" pageId="4" pageNumber="36">O. crassifolia</emphasis>
var. 
<emphasis id="5EF421740C3C5C53AC0CC2FDFD3D8093" box="[623,719,735,760]" italics="true" pageId="4" pageNumber="36">intricata</emphasis>
</taxonomicName>
, 
<taxonomicName id="AB8086E50C3C5C53ACBDC2FDFBE88093" box="[734,1050,734,761]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="36" phylum="Tracheophyta" rank="variety" species="tridentata" variety="canescens">
<emphasis id="5EF421740C3C5C53ACBDC2FDFC9D8093" box="[734,879,734,761]" italics="true" pageId="4" pageNumber="36">O. tridentata</emphasis>
var. 
<emphasis id="5EF421740C3C5C53ADCBC2FDFBE88093" box="[936,1050,735,760]" italics="true" pageId="4" pageNumber="36">canescens</emphasis>
</taxonomicName>
, 
<taxonomicName id="AB8086E50C3C5C53AA4BC2FDFAA38093" box="[1064,1361,734,761]" class="Magnoliopsida" family="Fabaceae" form="intermedia" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="36" phylum="Tracheophyta" rank="form" species="tridentata">
<emphasis id="5EF421740C3C5C53AA4BC2FDFB4A8093" box="[1064,1208,734,761]" italics="true" pageId="4" pageNumber="36">O. tridentata</emphasis>
f. 
<emphasis id="5EF421740C3C5C53AABBC2FCFAA38093" box="[1240,1361,734,760]" italics="true" pageId="4" pageNumber="36">intermedia</emphasis>
</taxonomicName>
, and 
<taxonomicName id="AB8086E50C3C5C53ABF3C2FDFDF88177" class="Magnoliopsida" family="Fabaceae" form="erecta" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="36" phylum="Tracheophyta" rank="form" species="tridentata" variety="intermedia">
<emphasis id="5EF421740C3C5C53ABF3C2FDFF048177" italics="true" pageId="4" pageNumber="36">O. tridentata</emphasis>
var. 
<emphasis id="5EF421740C3C5C53AF4DC320FE558177" box="[302,423,770,796]" italics="true" pageId="4" pageNumber="36">intermedia</emphasis>
f. 
<emphasis id="5EF421740C3C5C53AFA6C321FDF88177" box="[453,522,771,796]" italics="true" pageId="4" pageNumber="36">erecta</emphasis>
</taxonomicName>
, traditionally considered as synonyms of 
<taxonomicName id="AB8086E50C3C5C53ADBEC321FAC38177" box="[989,1329,770,797]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="36" phylum="Tracheophyta" rank="subSpecies" species="tridentata" subSpecies="tridentata">
<emphasis id="5EF421740C3C5C53ADBEC321FC038176" box="[989,1009,771,797]" italics="true" pageId="4" pageNumber="36">O</emphasis>
. 
<emphasis id="5EF421740C3C5C53AA63C320FB9C8177" box="[1024,1134,770,796]" italics="true" pageId="4" pageNumber="36">tridentata</emphasis>
subsp. 
<emphasis id="5EF421740C3C5C53AAA0C320FAC38177" box="[1219,1329,770,796]" italics="true" pageId="4" pageNumber="36">tridentata</emphasis>
</taxonomicName>
, should be treated as synonyms of 
<taxonomicName id="AB8086E50C3C5C53AFF3C305FD14812B" box="[400,742,806,833]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="36" phylum="Tracheophyta" rank="subSpecies" species="tridentata" subSpecies="arbuscula">
<emphasis id="5EF421740C3C5C53AFF3C305FE56812A" box="[400,420,807,833]" italics="true" pageId="4" pageNumber="36">O</emphasis>
. 
<emphasis id="5EF421740C3C5C53AFD0C304FDD3812B" box="[435,545,806,832]" italics="true" pageId="4" pageNumber="36">tridentata</emphasis>
subsp. 
<emphasis id="5EF421740C3C5C53AC15C304FD14812B" box="[630,742,806,832]" italics="true" pageId="4" pageNumber="36">arbuscula</emphasis>
</taxonomicName>
, comb. &amp; stat nov. (see below).
</paragraph>
</subSubSection>
</treatment>
</document>