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<mods:title id="A3E74D5258AAB11991A07FFE7BC022E3">New Mixaderus species from the Mascarene Islands (Coleoptera: Aderidae)</mods:title>
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<treatment id="E4241704FFD9B140A8B7DF50FB00A8A3" ID-DOI="http://doi.org/10.5281/zenodo.4811358" ID-GBIF-Taxon="182398104" ID-Zenodo-Dep="4811358" LSID="urn:lsid:zoobank.org:act:49978B14-8773-4E28-87C1-32CE269E7BFD" httpUri="http://treatment.plazi.org/id/E4241704FFD9B140A8B7DF50FB00A8A3" lastPageId="3" lastPageNumber="169" pageId="2" pageNumber="168">
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<heading id="377A117EFFD9B141A8B7DF50FE71ABA7" bold="true" box="[151,490,1662,1689]" fontSize="11" level="1" pageId="2" pageNumber="168" reason="1">
<emphasis id="5EF97A00FFD9B141A8B7DF50FE71ABA7" bold="true" box="[151,490,1662,1689]" pageId="2" pageNumber="168">
<taxonomicName id="AB8DDD91FFD9B141A8B7DF50FE3BABA7" authority="Gompel, 2021" authorityName="Gompel" authorityYear="2021" box="[151,416,1662,1689]" class="Insecta" family="Aderidae" genus="Mixaderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="168" phylum="Arthropoda" rank="species" species="lemagneni" status="sp. nov.">
<emphasis id="5EF97A00FFD9B141A8B7DF50FE3BABA7" bold="true" box="[151,416,1662,1689]" italics="true" pageId="2" pageNumber="168">Mixaderus lemagneni</emphasis>
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<taxonomicNameLabel id="45CAC77BFFD9B141A987DF51FE71ABA7" box="[423,490,1663,1689]" pageId="2" pageNumber="168" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="2497F599FFD9B141A8B7DF8DFE19AB80" box="[151,386,1699,1726]" pageId="2" pageNumber="168" type="description">
<paragraph id="6C32A612FFD9B141A8B7DF8DFE19AB80" blockId="2.[151,490,1662,1726]" box="[151,386,1699,1726]" pageId="2" pageNumber="168">
(
<figureCitation id="F4B6BA97FFD9B141A8BFDF8DFE8CAB83" box="[159,279,1699,1725]" captionStart="FIGURE 1" captionStartId="5.[151,250,1828,1852]" captionTargetBox="[164,1423,561,1798]" captionTargetId="figure-197@5.[163,1423,561,1805]" captionTargetPageId="5" captionText="FIGURE 1. Habitus of new species of Aderidae from the Mascarene Islands. a, Mixaderus reunionensis n. sp. (♂ holotype from La Réunion); b, Mixaderus tamisieri n. sp. (♂ paratype from La Réunion); c, Mixaderus mauritiensis n. sp. (♀ holotype from Mauritius); d, Mixaderus chassaini n. sp. (♂ holotype from La Réunion); e, Mixaderus lemagneni n. sp. (♂ paratype from La Réunion); f, Mixaderus voisini n. sp. (♀ paratype from La Réunion). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4746057" httpUri="https://zenodo.org/record/4746057/files/figure.png" pageId="2" pageNumber="168">Figures 1e</figureCitation>
,
<figureCitation id="F4B6BA97FFD9B141A904DF8DFEE1AB80" box="[292,378,1699,1726]" captionStart="FIGURE 2" captionStartId="6.[151,250,1597,1621]" captionTargetBox="[252,1354,215,1574]" captionTargetId="figure-16@6.[233,1355,181,1574]" captionTargetPageId="6" captionText="FIGURE 2. Anatomy of new species of Aderidae from the Mascarene Islands. ag: aedeagus of a, Mixaderus reunionensis n. sp. (♂ holotype from La Réunion); b, Mixaderus chassaini n. sp. (♂ holotype from La Réunion); c, Mixaderus tamisieri n. sp. (♂ paratype from La Réunion); d, Mixaderus lemagneni n. sp. (♂ paratype from La Réunion); e, Mixaderus gomyi Gompel, 2012 (♂ holotype from La Réunion); fg, Mixaderus voisini n. sp.: f, anterior struts retracted (♂ holotype from La Réunion); g, anterior struts extended from phallobase (♂ paratype from La Réunion). ho: antennae of h, Mixaderus reunionensis n. sp. (♂ holotype from La Réunion); i, Mixaderus chassaini n. sp. (♂ holotype from La Réunion); j, Mixaderus tamisieri n. sp. (♂ paratype from La Réunion); k, Mixaderus lemagneni n. sp. (♂ paratype from La Réunion); l, Mixaderus gomyi Gompel, 2012 (♂ holotype from La Réunion); m, Mixaderus mauritiensis n. sp. (♀ holotype from Mauritius); n, Mixaderus voisini n. sp. (♀ paratype from La Réunion). o,p: male right prothoracic tarsae of o, Mixaderus gomyi Gompel, 2012 (♂ holotype from La Réun- ion); p, Mixaderus lemagneni n. sp. (♂ paratype from La Réunion). qv: Mixaderus head schematics in side view. q, Mixaderus reunionensis n. sp.; r, Mixaderus chassaini n. sp.; s, Mixaderus tamisieri n. sp.; t, Mixaderus lemagneni n. sp.; u, Mixaderus voisini n. sp.; v, Mixaderus mauritiensis n. sp. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4746059" httpUri="https://zenodo.org/record/4746059/files/figure.png" pageId="2" pageNumber="168">2d,k,p,t</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="2497F599FFD9B141A8B7DFC5FD55AA17" pageId="2" pageNumber="168" type="etymology">
<paragraph id="6C32A612FFD9B141A8B7DFC5FD55AA17" blockId="2.[151,1436,1771,2013]" pageId="2" pageNumber="168">
<emphasis id="5EF97A00FFD9B141A8B7DFC5FE85AA3B" bold="true" box="[151,286,1771,1797]" pageId="2" pageNumber="168">Etymology.</emphasis>
This taxon is named after my colleague Emmanuel Lemagnen in appreciation of his active support of my work on
<taxonomicName id="AB8DDD91FFD9B141A904DE21FE10AA17" box="[292,395,1807,1833]" class="Insecta" family="Aderidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="168" phylum="Arthropoda" rank="family">Aderidae</taxonomicName>
from the Mascarene islands.
</paragraph>
</subSubSection>
<subSubSection id="2497F599FFD9B141A8E7DE1DFB70AA73" box="[199,1259,1842,1869]" pageId="2" pageNumber="168" type="materials_examined">
<paragraph id="6C32A612FFD9B141A8E7DE1DFB70AA73" blockId="2.[151,1436,1771,2013]" box="[199,1259,1842,1869]" pageId="2" pageNumber="168">
<materialsCitation id="DCE5AC4FFFD9B141A8E7DE1DFB7CAA73" ID-GBIF-Occurrence="3118911306" box="[199,1255,1842,1869]" collectorName="La Ressource" country="France" elevation="310" latitude="-20.933332" location="Reunion" longLatPrecision="1266" longitude="55.516666" pageId="2" pageNumber="168" specimenCount="1" stateProvince="Reunion" typeStatus="holotype">
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<typeStatus id="B33618B0FFD9B141A8E7DE1DFE99AA73" box="[199,258,1843,1869]" pageId="2" pageNumber="168">Type</typeStatus>
locality.
<collectingCountry id="149AE682FFD9B141A94CDE1DFE5BAA73" box="[364,448,1843,1869]" name="France" pageId="2" pageNumber="168">France</collectingCountry>
</emphasis>
: La
<collectingRegion id="AE4968F0FFD9B141A9D4DE1DFDCDAA73" box="[500,598,1843,1869]" country="France" name="Reunion" pageId="2" pageNumber="168">Réunion</collectingRegion>
, Sainte-Marie,
<collectorName id="C178C3C4FFD9B141AB21DE1DFC00AA73" box="[769,923,1843,1869]" pageId="2" pageNumber="168">La Ressource</collectorName>
, (
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<elevation id="E7A04121FFD9B141AB8EDE1DFC63AA73" box="[942,1016,1843,1869]" metricMagnitude="2" metricUnit="m" metricValue="3.1" pageId="2" pageNumber="168" unit="m" value="310.0">310 m</elevation>
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) [
<geoCoordinate id="09B9C0D5FFD9B141AC2EDE1CFBEAAA73" box="[1038,1137,1842,1869]" degrees="20" direction="south" minutes="56" orientation="latitude" pageId="2" pageNumber="168" precision="925" value="-20.933332">20°56 S</geoCoordinate>
<geoCoordinate id="09B9C0D5FFD9B141AC58DE1CFB44AA73" box="[1144,1247,1842,1869]" degrees="55" direction="east" minutes="31" orientation="longitude" pageId="2" pageNumber="168" precision="925" value="55.516666">55°31 E</geoCoordinate>
]
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.
</paragraph>
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<paragraph id="6C32A612FFD9B141A8E7DE78FDD5AA4F" blockId="2.[151,1436,1771,2013]" box="[199,590,1878,1905]" pageId="2" pageNumber="168">
<emphasis id="5EF97A00FFD9B141A8E7DE78FEFAAA4E" bold="true" box="[199,353,1878,1904]" pageId="2" pageNumber="168">Distribution.</emphasis>
<collectingCountry id="149AE682FFD9B141A947DE79FE23AA4F" box="[359,440,1879,1905]" name="France" pageId="2" pageNumber="168">France</collectingCountry>
:
<collectingCountry id="149AE682FFD9B141A9E2DE79FDD1AA4F" box="[450,586,1879,1905]" name="Réunion" pageId="2" pageNumber="168">La Réunion</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="2497F599FFD9B140A8E7DE54FC8EA99F" lastPageId="3" lastPageNumber="169" pageId="2" pageNumber="168" type="description">
<paragraph id="6C32A612FFD9B141A8E7DE54FB3DAA87" blockId="2.[151,1436,1771,2013]" pageId="2" pageNumber="168">
<emphasis id="5EF97A00FFD9B141A8E7DE54FEC2AAAA" bold="true" box="[199,345,1914,1940]" pageId="2" pageNumber="168">Description.</emphasis>
Body length.
<specimenCount id="7A8B6D9BFFD9B141A9DADE55FD89AAAB" box="[506,530,1915,1941]" pageId="2" pageNumber="168" type="male"></specimenCount>
: 1.63±
<quantity id="AB750BF7FFD9B141AA7DDE55FD59AAAB" box="[605,706,1915,1941]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="168" unit="mm" value="0.11">0.11 mm</quantity>
(n=5);
<specimenCount id="7A8B6D9BFFD9B141AB38DE55FCB4AAAB" box="[792,815,1915,1941]" pageId="2" pageNumber="168" type="female"></specimenCount>
: 1.77±
<quantity id="AB750BF7FFD9B141AB5BDE55FC7AAAAB" box="[891,993,1915,1941]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="168" unit="mm" value="0.17">0.17 mm</quantity>
(n=3). Body bicolour, head black, pronotum dark brown, elytra brown with broad white markings. Appendages yellow and brown.
</paragraph>
<paragraph id="6C32A612FFD9B140A8E7DEEDFD83ADC7" blockId="2.[151,1436,1771,2013]" lastBlockId="3.[151,1437,151,1437]" lastPageId="3" lastPageNumber="169" pageId="2" pageNumber="168">
<emphasis id="5EF97A00FFD9B141A8E7DEEDFE9FAAE3" box="[199,260,1987,2013]" italics="true" pageId="2" pageNumber="168">Head</emphasis>
. Transverse, black, slightly wider than pronotum. Frons weakly convex. Palpi yellow. Punctuation dense and fine. Pubescence double, dense, wholly, white, homogeneous, largely covering the tegument. Eyes nearly reaching the posterior edge of the head in side view, or separated from it by a strip of tegument narrower than the width of a row of ommatidia (
<figureCitation id="F4B6BA97FFD8B140A980D9F1FD90ADC7" box="[416,523,223,249]" captionStart="FIGURE 2" captionStartId="6.[151,250,1597,1621]" captionTargetBox="[252,1354,215,1574]" captionTargetId="figure-16@6.[233,1355,181,1574]" captionTargetPageId="6" captionText="FIGURE 2. Anatomy of new species of Aderidae from the Mascarene Islands. ag: aedeagus of a, Mixaderus reunionensis n. sp. (♂ holotype from La Réunion); b, Mixaderus chassaini n. sp. (♂ holotype from La Réunion); c, Mixaderus tamisieri n. sp. (♂ paratype from La Réunion); d, Mixaderus lemagneni n. sp. (♂ paratype from La Réunion); e, Mixaderus gomyi Gompel, 2012 (♂ holotype from La Réunion); fg, Mixaderus voisini n. sp.: f, anterior struts retracted (♂ holotype from La Réunion); g, anterior struts extended from phallobase (♂ paratype from La Réunion). ho: antennae of h, Mixaderus reunionensis n. sp. (♂ holotype from La Réunion); i, Mixaderus chassaini n. sp. (♂ holotype from La Réunion); j, Mixaderus tamisieri n. sp. (♂ paratype from La Réunion); k, Mixaderus lemagneni n. sp. (♂ paratype from La Réunion); l, Mixaderus gomyi Gompel, 2012 (♂ holotype from La Réunion); m, Mixaderus mauritiensis n. sp. (♀ holotype from Mauritius); n, Mixaderus voisini n. sp. (♀ paratype from La Réunion). o,p: male right prothoracic tarsae of o, Mixaderus gomyi Gompel, 2012 (♂ holotype from La Réun- ion); p, Mixaderus lemagneni n. sp. (♂ paratype from La Réunion). qv: Mixaderus head schematics in side view. q, Mixaderus reunionensis n. sp.; r, Mixaderus chassaini n. sp.; s, Mixaderus tamisieri n. sp.; t, Mixaderus lemagneni n. sp.; u, Mixaderus voisini n. sp.; v, Mixaderus mauritiensis n. sp. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4746059" httpUri="https://zenodo.org/record/4746059/files/figure.png" pageId="3" pageNumber="169">
Figure
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).
</paragraph>
<paragraph id="6C32A612FFD8B140A8E7D82DFF60AC93" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="169">
<emphasis id="5EF97A00FFD8B140A8E7D82DFEAAAC23" box="[199,305,259,285]" italics="true" pageId="3" pageNumber="169">Antennae</emphasis>
(
<figureCitation id="F4B6BA97FFD8B140A960D82DFE28AC20" box="[320,435,259,286]" captionStart="FIGURE 2" captionStartId="6.[151,250,1597,1621]" captionTargetBox="[252,1354,215,1574]" captionTargetId="figure-16@6.[233,1355,181,1574]" captionTargetPageId="6" captionText="FIGURE 2. Anatomy of new species of Aderidae from the Mascarene Islands. ag: aedeagus of a, Mixaderus reunionensis n. sp. (♂ holotype from La Réunion); b, Mixaderus chassaini n. sp. (♂ holotype from La Réunion); c, Mixaderus tamisieri n. sp. (♂ paratype from La Réunion); d, Mixaderus lemagneni n. sp. (♂ paratype from La Réunion); e, Mixaderus gomyi Gompel, 2012 (♂ holotype from La Réunion); fg, Mixaderus voisini n. sp.: f, anterior struts retracted (♂ holotype from La Réunion); g, anterior struts extended from phallobase (♂ paratype from La Réunion). ho: antennae of h, Mixaderus reunionensis n. sp. (♂ holotype from La Réunion); i, Mixaderus chassaini n. sp. (♂ holotype from La Réunion); j, Mixaderus tamisieri n. sp. (♂ paratype from La Réunion); k, Mixaderus lemagneni n. sp. (♂ paratype from La Réunion); l, Mixaderus gomyi Gompel, 2012 (♂ holotype from La Réunion); m, Mixaderus mauritiensis n. sp. (♀ holotype from Mauritius); n, Mixaderus voisini n. sp. (♀ paratype from La Réunion). o,p: male right prothoracic tarsae of o, Mixaderus gomyi Gompel, 2012 (♂ holotype from La Réun- ion); p, Mixaderus lemagneni n. sp. (♂ paratype from La Réunion). qv: Mixaderus head schematics in side view. q, Mixaderus reunionensis n. sp.; r, Mixaderus chassaini n. sp.; s, Mixaderus tamisieri n. sp.; t, Mixaderus lemagneni n. sp.; u, Mixaderus voisini n. sp.; v, Mixaderus mauritiensis n. sp. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4746059" httpUri="https://zenodo.org/record/4746059/files/figure.png" pageId="3" pageNumber="169">Figure 2k</figureCitation>
). Uniformly yellow, or at most orange in distal half, contrasting with
<taxonomicName id="AB8DDD91FFD8B140ACE8D82DFF49AC7F" authority="Gompel, 2012" authorityName="Gompel" authorityYear="2012" class="Insecta" family="Aderidae" genus="Mixaderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="169" phylum="Arthropoda" rank="species" species="gomyi">
<emphasis id="5EF97A00FFD8B140ACE8D82DFAA8AC22" box="[1224,1331,259,284]" italics="true" pageId="3" pageNumber="169">M. gomyi</emphasis>
<bibRefCitation id="081CDBE3FFD8B140AD1BD82DFF49AC7F" author="Gompel, N." pageId="3" pageNumber="169" pagination="77 - 88" refId="ref8049" refString="Gompel, N. (2012) The genus Mixaderus Collado &amp; Alonso-Zaragoza, 1996 in Madagascar and the Mascareignes Archipelago, with description of three new species (Coleoptera: Aderidae). Boletiin de la Sociedad Entomoloigica Aragonesa (S. E. A.), 50, 77 - 88." type="journal article" year="2012">Gompel, 2012</bibRefCitation>
</taxonomicName>
, in which only the first 34 antennomeres are pale and the rest of the antenna is dark brown. Thin, almost as long as elytra in males, about a third shorter than elytra in females, pubescent. Antennomeres 35 regularly cylindrical, 810 conical with apex broader, and antennomere 11 than twice longer than penultimate, asymmetrical and rounded.
</paragraph>
<paragraph id="6C32A612FFD8B140A8E7D899FC2CAF03" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="169">
<emphasis id="5EF97A00FFD8B140A8E7D899FEADACEE" box="[199,310,439,464]" italics="true" pageId="3" pageNumber="169">Pronotum</emphasis>
. Convex, rounded, marked by a pair of dimples just anterior to the posterior border. Sides straight. Surface densely punctuated, the space between punctures equivalent to the puncture diameters, punctures of intermediate size, deep. Pubescence dense, double, golden anteriorly and white in the posterior third, including in the dimples, with a sharp, sinuate boundary between anterior and posterior.
</paragraph>
<paragraph id="6C32A612FFD8B140A8E7DB69FD45AE07" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="169">
<emphasis id="5EF97A00FFD8B140A8E7DB69FE96AF5F" box="[199,269,583,609]" italics="true" pageId="3" pageNumber="169">Elytra</emphasis>
. Length:
<specimenCount id="7A8B6D9BFFD8B140A95ADB69FE09AF5F" box="[378,402,583,609]" pageId="3" pageNumber="169" type="male"></specimenCount>
: 1.15±
<quantity id="AB750BF7FFD8B140A9FDDB68FDD8AF5F" box="[477,579,582,609]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="169" unit="mm" value="0.08">0.08 mm</quantity>
(n=5);
<specimenCount id="7A8B6D9BFFD8B140AAB8DB69FD34AF5F" box="[664,687,583,609]" pageId="3" pageNumber="169" type="female"></specimenCount>
: 1.24±
<quantity id="AB750BF7FFD8B140AADCDB69FCFBAF5F" box="[764,864,583,609]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="169" unit="mm" value="0.11">0.11 mm</quantity>
(n=3). Sides subparallel, rounded posteriorly, with humeral callus well marked. Punctuation stronger and deeper and as dense as on the pronotum, space between punctures equal to the puncture diameters on disc. Disc depressed. Pubescence short, golden, comprised of longer setae borne from the punctuation and interspersed pruinosity, with a distinct pattern of white pubescence on each elytron, formed of three elements: a weak and small humeral spot, a broad anterior transversal band at the level of the disc, hardly reaching the elytral suture and broadening on the elytral disc, and a second posterior transversal band on the declivity, curved and not reaching the elytral suture.
</paragraph>
<paragraph id="6C32A612FFD8B140A8E7DA6DFEF0AE9B" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="169">
<emphasis id="5EF97A00FFD8B140A8E7DA6DFF67AE62" box="[199,252,835,860]" italics="true" pageId="3" pageNumber="169">Legs</emphasis>
. Simple, tibiae and tarsae of fore- and midlegs yellow, femora darker in the proximal half; hindlegs with femora brown, except their apex, which is yellow, hindtibiae brown in their proximal half and yellow in their distal half, tarsae yellow.
</paragraph>
<paragraph id="6C32A612FFD8B140A8E7DA81FD24A92F" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="169">
<emphasis id="5EF97A00FFD8B140A8E7DA81FE05AEF7" box="[199,414,943,969]" italics="true" pageId="3" pageNumber="169">Sexual dimorphism</emphasis>
. First article of the male foretarsae very broad and longer than the next two articles together. This character is also more pronounced than in the closely related species
<taxonomicName id="AB8DDD91FFD8B140ABC4DAFDFB63AED3" authority="Gompel, 2012" authorityName="Gompel" authorityYear="2012" box="[996,1272,979,1005]" class="Insecta" family="Aderidae" genus="Mixaderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="169" phylum="Arthropoda" rank="species" species="gomyi">
<emphasis id="5EF97A00FFD8B140ABC4DAFDFBD5AED2" box="[996,1102,979,1004]" italics="true" pageId="3" pageNumber="169">M. gomyi</emphasis>
<bibRefCitation id="081CDBE3FFD8B140AC77DAFDFB63AED3" author="Gompel, N." box="[1111,1272,979,1005]" pageId="3" pageNumber="169" pagination="77 - 88" refId="ref8049" refString="Gompel, N. (2012) The genus Mixaderus Collado &amp; Alonso-Zaragoza, 1996 in Madagascar and the Mascareignes Archipelago, with description of three new species (Coleoptera: Aderidae). Boletiin de la Sociedad Entomoloigica Aragonesa (S. E. A.), 50, 77 - 88." type="journal article" year="2012">Gompel, 2012</bibRefCitation>
</taxonomicName>
(
<figureCitation id="F4B6BA97FFD8B140AD28DAFDFA14AED3" box="[1288,1423,979,1005]" captionStart="FIGURE 2" captionStartId="6.[151,250,1597,1621]" captionTargetBox="[252,1354,215,1574]" captionTargetId="figure-16@6.[233,1355,181,1574]" captionTargetPageId="6" captionText="FIGURE 2. Anatomy of new species of Aderidae from the Mascarene Islands. ag: aedeagus of a, Mixaderus reunionensis n. sp. (♂ holotype from La Réunion); b, Mixaderus chassaini n. sp. (♂ holotype from La Réunion); c, Mixaderus tamisieri n. sp. (♂ paratype from La Réunion); d, Mixaderus lemagneni n. sp. (♂ paratype from La Réunion); e, Mixaderus gomyi Gompel, 2012 (♂ holotype from La Réunion); fg, Mixaderus voisini n. sp.: f, anterior struts retracted (♂ holotype from La Réunion); g, anterior struts extended from phallobase (♂ paratype from La Réunion). ho: antennae of h, Mixaderus reunionensis n. sp. (♂ holotype from La Réunion); i, Mixaderus chassaini n. sp. (♂ holotype from La Réunion); j, Mixaderus tamisieri n. sp. (♂ paratype from La Réunion); k, Mixaderus lemagneni n. sp. (♂ paratype from La Réunion); l, Mixaderus gomyi Gompel, 2012 (♂ holotype from La Réunion); m, Mixaderus mauritiensis n. sp. (♀ holotype from Mauritius); n, Mixaderus voisini n. sp. (♀ paratype from La Réunion). o,p: male right prothoracic tarsae of o, Mixaderus gomyi Gompel, 2012 (♂ holotype from La Réun- ion); p, Mixaderus lemagneni n. sp. (♂ paratype from La Réunion). qv: Mixaderus head schematics in side view. q, Mixaderus reunionensis n. sp.; r, Mixaderus chassaini n. sp.; s, Mixaderus tamisieri n. sp.; t, Mixaderus lemagneni n. sp.; u, Mixaderus voisini n. sp.; v, Mixaderus mauritiensis n. sp. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4746059" httpUri="https://zenodo.org/record/4746059/files/figure.png" pageId="3" pageNumber="169">Figure 2o,p</figureCitation>
). Antennae a third longer in males than in females.
</paragraph>
<paragraph id="6C32A612FFD8B140A8E7DD35FC8EA99F" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="169">
<emphasis id="5EF97A00FFD8B140A8E7DD35FEAFA90B" box="[199,308,1051,1077]" italics="true" pageId="3" pageNumber="169">Aedeagus</emphasis>
(
<figureCitation id="F4B6BA97FFD8B140A965DD35FE23A908" box="[325,440,1051,1078]" captionStart="FIGURE 2" captionStartId="6.[151,250,1597,1621]" captionTargetBox="[252,1354,215,1574]" captionTargetId="figure-16@6.[233,1355,181,1574]" captionTargetPageId="6" captionText="FIGURE 2. Anatomy of new species of Aderidae from the Mascarene Islands. ag: aedeagus of a, Mixaderus reunionensis n. sp. (♂ holotype from La Réunion); b, Mixaderus chassaini n. sp. (♂ holotype from La Réunion); c, Mixaderus tamisieri n. sp. (♂ paratype from La Réunion); d, Mixaderus lemagneni n. sp. (♂ paratype from La Réunion); e, Mixaderus gomyi Gompel, 2012 (♂ holotype from La Réunion); fg, Mixaderus voisini n. sp.: f, anterior struts retracted (♂ holotype from La Réunion); g, anterior struts extended from phallobase (♂ paratype from La Réunion). ho: antennae of h, Mixaderus reunionensis n. sp. (♂ holotype from La Réunion); i, Mixaderus chassaini n. sp. (♂ holotype from La Réunion); j, Mixaderus tamisieri n. sp. (♂ paratype from La Réunion); k, Mixaderus lemagneni n. sp. (♂ paratype from La Réunion); l, Mixaderus gomyi Gompel, 2012 (♂ holotype from La Réunion); m, Mixaderus mauritiensis n. sp. (♀ holotype from Mauritius); n, Mixaderus voisini n. sp. (♀ paratype from La Réunion). o,p: male right prothoracic tarsae of o, Mixaderus gomyi Gompel, 2012 (♂ holotype from La Réun- ion); p, Mixaderus lemagneni n. sp. (♂ paratype from La Réunion). qv: Mixaderus head schematics in side view. q, Mixaderus reunionensis n. sp.; r, Mixaderus chassaini n. sp.; s, Mixaderus tamisieri n. sp.; t, Mixaderus lemagneni n. sp.; u, Mixaderus voisini n. sp.; v, Mixaderus mauritiensis n. sp. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4746059" httpUri="https://zenodo.org/record/4746059/files/figure.png" pageId="3" pageNumber="169">Figure 2d</figureCitation>
). Oblong, apicale tapered in apical fifth in dorsal view, ending in a small semi-circular knob (inset on
<figureCitation id="F4B6BA97FFD8B140A962DD11FE29A967" box="[322,434,1087,1113]" captionStart="FIGURE 2" captionStartId="6.[151,250,1597,1621]" captionTargetBox="[252,1354,215,1574]" captionTargetId="figure-16@6.[233,1355,181,1574]" captionTargetPageId="6" captionText="FIGURE 2. Anatomy of new species of Aderidae from the Mascarene Islands. ag: aedeagus of a, Mixaderus reunionensis n. sp. (♂ holotype from La Réunion); b, Mixaderus chassaini n. sp. (♂ holotype from La Réunion); c, Mixaderus tamisieri n. sp. (♂ paratype from La Réunion); d, Mixaderus lemagneni n. sp. (♂ paratype from La Réunion); e, Mixaderus gomyi Gompel, 2012 (♂ holotype from La Réunion); fg, Mixaderus voisini n. sp.: f, anterior struts retracted (♂ holotype from La Réunion); g, anterior struts extended from phallobase (♂ paratype from La Réunion). ho: antennae of h, Mixaderus reunionensis n. sp. (♂ holotype from La Réunion); i, Mixaderus chassaini n. sp. (♂ holotype from La Réunion); j, Mixaderus tamisieri n. sp. (♂ paratype from La Réunion); k, Mixaderus lemagneni n. sp. (♂ paratype from La Réunion); l, Mixaderus gomyi Gompel, 2012 (♂ holotype from La Réunion); m, Mixaderus mauritiensis n. sp. (♀ holotype from Mauritius); n, Mixaderus voisini n. sp. (♀ paratype from La Réunion). o,p: male right prothoracic tarsae of o, Mixaderus gomyi Gompel, 2012 (♂ holotype from La Réun- ion); p, Mixaderus lemagneni n. sp. (♂ paratype from La Réunion). qv: Mixaderus head schematics in side view. q, Mixaderus reunionensis n. sp.; r, Mixaderus chassaini n. sp.; s, Mixaderus tamisieri n. sp.; t, Mixaderus lemagneni n. sp.; u, Mixaderus voisini n. sp.; v, Mixaderus mauritiensis n. sp. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4746059" httpUri="https://zenodo.org/record/4746059/files/figure.png" pageId="3" pageNumber="169">Figure 2d</figureCitation>
; compare to the simple tip of
<taxonomicName id="AB8DDD91FFD8B140AB27DD11FCE9A966" authorityName="Gompel" authorityYear="2012" box="[775,882,1087,1112]" class="Insecta" family="Aderidae" genus="Mixaderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="169" phylum="Arthropoda" rank="species" species="gomyi">
<emphasis id="5EF97A00FFD8B140AB27DD11FCE9A966" box="[775,882,1087,1112]" italics="true" pageId="3" pageNumber="169">M. gomyi</emphasis>
</taxonomicName>
phallus: inset on
<figureCitation id="F4B6BA97FFD8B140AC63DD11FB29A967" box="[1091,1202,1087,1113]" captionStart="FIGURE 2" captionStartId="6.[151,250,1597,1621]" captionTargetBox="[252,1354,215,1574]" captionTargetId="figure-16@6.[233,1355,181,1574]" captionTargetPageId="6" captionText="FIGURE 2. Anatomy of new species of Aderidae from the Mascarene Islands. ag: aedeagus of a, Mixaderus reunionensis n. sp. (♂ holotype from La Réunion); b, Mixaderus chassaini n. sp. (♂ holotype from La Réunion); c, Mixaderus tamisieri n. sp. (♂ paratype from La Réunion); d, Mixaderus lemagneni n. sp. (♂ paratype from La Réunion); e, Mixaderus gomyi Gompel, 2012 (♂ holotype from La Réunion); fg, Mixaderus voisini n. sp.: f, anterior struts retracted (♂ holotype from La Réunion); g, anterior struts extended from phallobase (♂ paratype from La Réunion). ho: antennae of h, Mixaderus reunionensis n. sp. (♂ holotype from La Réunion); i, Mixaderus chassaini n. sp. (♂ holotype from La Réunion); j, Mixaderus tamisieri n. sp. (♂ paratype from La Réunion); k, Mixaderus lemagneni n. sp. (♂ paratype from La Réunion); l, Mixaderus gomyi Gompel, 2012 (♂ holotype from La Réunion); m, Mixaderus mauritiensis n. sp. (♀ holotype from Mauritius); n, Mixaderus voisini n. sp. (♀ paratype from La Réunion). o,p: male right prothoracic tarsae of o, Mixaderus gomyi Gompel, 2012 (♂ holotype from La Réun- ion); p, Mixaderus lemagneni n. sp. (♂ paratype from La Réunion). qv: Mixaderus head schematics in side view. q, Mixaderus reunionensis n. sp.; r, Mixaderus chassaini n. sp.; s, Mixaderus tamisieri n. sp.; t, Mixaderus lemagneni n. sp.; u, Mixaderus voisini n. sp.; v, Mixaderus mauritiensis n. sp. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4746059" httpUri="https://zenodo.org/record/4746059/files/figure.png" pageId="3" pageNumber="169">Figure 2e</figureCitation>
). Phallobase short, a fifth of total phallus length (a third of phallus length in
<taxonomicName id="AB8DDD91FFD8B140AB28DD4DFCE9A942" authorityName="Gompel" authorityYear="2012" box="[776,882,1123,1148]" class="Insecta" family="Aderidae" genus="Mixaderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="169" phylum="Arthropoda" rank="species" species="gomyi">
<emphasis id="5EF97A00FFD8B140AB28DD4DFCE9A942" box="[776,882,1123,1148]" italics="true" pageId="3" pageNumber="169">M. gomyi</emphasis>
</taxonomicName>
,
<figureCitation id="F4B6BA97FFD8B140ABA1DD4DFB9EA943" box="[897,1029,1123,1149]" captionStart="FIGURE 2" captionStartId="6.[151,250,1597,1621]" captionTargetBox="[252,1354,215,1574]" captionTargetId="figure-16@6.[233,1355,181,1574]" captionTargetPageId="6" captionText="FIGURE 2. Anatomy of new species of Aderidae from the Mascarene Islands. ag: aedeagus of a, Mixaderus reunionensis n. sp. (♂ holotype from La Réunion); b, Mixaderus chassaini n. sp. (♂ holotype from La Réunion); c, Mixaderus tamisieri n. sp. (♂ paratype from La Réunion); d, Mixaderus lemagneni n. sp. (♂ paratype from La Réunion); e, Mixaderus gomyi Gompel, 2012 (♂ holotype from La Réunion); fg, Mixaderus voisini n. sp.: f, anterior struts retracted (♂ holotype from La Réunion); g, anterior struts extended from phallobase (♂ paratype from La Réunion). ho: antennae of h, Mixaderus reunionensis n. sp. (♂ holotype from La Réunion); i, Mixaderus chassaini n. sp. (♂ holotype from La Réunion); j, Mixaderus tamisieri n. sp. (♂ paratype from La Réunion); k, Mixaderus lemagneni n. sp. (♂ paratype from La Réunion); l, Mixaderus gomyi Gompel, 2012 (♂ holotype from La Réunion); m, Mixaderus mauritiensis n. sp. (♀ holotype from Mauritius); n, Mixaderus voisini n. sp. (♀ paratype from La Réunion). o,p: male right prothoracic tarsae of o, Mixaderus gomyi Gompel, 2012 (♂ holotype from La Réun- ion); p, Mixaderus lemagneni n. sp. (♂ paratype from La Réunion). qv: Mixaderus head schematics in side view. q, Mixaderus reunionensis n. sp.; r, Mixaderus chassaini n. sp.; s, Mixaderus tamisieri n. sp.; t, Mixaderus lemagneni n. sp.; u, Mixaderus voisini n. sp.; v, Mixaderus mauritiensis n. sp. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4746059" httpUri="https://zenodo.org/record/4746059/files/figure.png" pageId="3" pageNumber="169">Figure 2d,e</figureCitation>
: double arrows). Accessory lobes of the apicale present, not extending beyond the phallus tip.
</paragraph>
</subSubSection>
<subSubSection id="2497F599FFD8B140A8E7DD85FB00A8A3" pageId="3" pageNumber="169" type="materials_examined">
<paragraph id="6C32A612FFD8B140A8E7DD85FB00A8A3" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="169">
<emphasis id="5EF97A00FFD8B140A8E7DD85FEE3A9FA" bold="true" box="[199,376,1194,1221]" pageId="3" pageNumber="169">Type material.</emphasis>
<materialsCitation id="DCE5AC4FFFD8B140A9A3DD85FAEBA9FB" ID-GBIF-Occurrence="3118911313" box="[387,1392,1195,1221]" collectingDate="1937-10-29" collectionCode="BMNH" collectorName="J. Vinson" country="France" location="Cote Sud-Est" municipality="La" pageId="3" pageNumber="169" specimenCount="1" specimenCount-male="1" stateProvince="Reunion" typeStatus="paratype">
<emphasis id="5EF97A00FFD8B140A9A3DD85FE4CA9FB" bold="true" box="[387,471,1195,1221]" pageId="3" pageNumber="169">
<collectingCountry id="149AE682FFD8B140A9A3DD85FE4CA9FB" box="[387,471,1195,1221]" name="France" pageId="3" pageNumber="169">France</collectingCountry>
</emphasis>
:
<collectingMunicipality id="8C563C68FFD8B140A9C9DD85FD9CA9FB" box="[489,519,1195,1221]" pageId="3" pageNumber="169">La</collectingMunicipality>
<collectingRegion id="AE4968F0FFD8B140AA31DD85FDE8A9FB" box="[529,627,1195,1221]" country="France" name="Reunion" pageId="3" pageNumber="169">Réunion</collectingRegion>
,
<location id="6952F0C9FFD8B140AAA1DD85FC86A9FB" LSID="urn:lsid:plazi:treatment:E4241704FFD9B140A8B7DF50FB00A8A3:6952F0C9FFD8B140AAA1DD85FC86A9FB" box="[641,797,1195,1221]" country="France" municipality="La" name="Cote Sud-Est" pageId="3" pageNumber="169" stateProvince="Reunion">Côte Sud-Est</location>
,
<date id="183380D2FFD8B140AB0BDD85FC38A9FB" box="[811,931,1195,1221]" pageId="3" pageNumber="169" value="1937-10-29">
<collectingDate id="0877793AFFD8B140AB0BDD85FC38A9FB" box="[811,931,1195,1221]" pageId="3" pageNumber="169" value="1937-10-29">29.X.1937</collectingDate>
</date>
,
<collectorName id="C178C3C4FFD8B140AB93DD85FB85A9FB" box="[947,1054,1195,1221]" pageId="3" pageNumber="169">J. Vinson</collectorName>
leg. (
<specimenCount id="7A8B6D9BFFD8B140AC47DD85FB08A9FB" box="[1127,1171,1195,1221]" pageId="3" pageNumber="169" type="male">1 ♂</specimenCount>
<typeStatus id="B33618B0FFD8B140ACBDDD85FA9BA9FB" box="[1181,1280,1195,1221]" pageId="3" pageNumber="169" type="paratype">paratype</typeStatus>
,
<collectionCode id="0A9C3ED7FFD8B140AD2FDD85FAF3A9FB" box="[1295,1384,1195,1221]" pageId="3" pageNumber="169">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="DCE5AC4FFFD8B140AD5FDD85FF74A833" ID-GBIF-Occurrence="3118911323" collectingDate="2006-03-02" collectionCode="NGPC" collectorName="J. Poussereau" country="France" county="La" latitude="-20.883333" location="La Grande Chaloupe" longLatPrecision="1266" longitude="55.366665" municipality="Saint-Denis" pageId="3" pageNumber="169" specimenCount="1" specimenCount-female="1" stateProvince="Reunion" typeStatus="paratype">
<collectingCounty id="8553DE9EFFD8B140AD5FDD85FA06A9FB" box="[1407,1437,1195,1221]" pageId="3" pageNumber="169">La</collectingCounty>
<collectingRegion id="AE4968F0FFD8B140A8B7DDE1FF62A9D7" box="[151,249,1231,1257]" country="France" name="Reunion" pageId="3" pageNumber="169">Réunion</collectingRegion>
,
<collectingMunicipality id="8C563C68FFD8B140A924DDE1FE10A9D7" box="[260,395,1231,1257]" pageId="3" pageNumber="169">Saint-Denis</collectingMunicipality>
,
<location id="6952F0C9FFD8B140A9B6DDE1FD19A9D7" LSID="urn:lsid:plazi:treatment:E4241704FFD9B140A8B7DF50FB00A8A3:6952F0C9FFD8B140A9B6DDE1FD19A9D7" box="[406,642,1231,1257]" country="France" county="La" latitude="-20.883333" longLatPrecision="1266" longitude="55.366665" municipality="Saint-Denis" name="La Grande Chaloupe" pageId="3" pageNumber="169" stateProvince="Reunion">La Grande Chaloupe</location>
, N18 [
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<geoCoordinate id="09B9C0D5FFD8B140AB15DDE0FC07A9D7" box="[821,924,1230,1257]" degrees="55" direction="east" minutes="22" orientation="longitude" pageId="3" pageNumber="169" precision="925" value="55.366665">55°22 E</geoCoordinate>
],
<date id="183380D2FFD8B140AB8FDDE1FBBAA9D7" box="[943,1057,1231,1257]" pageId="3" pageNumber="169" value="2006-03-02">
<collectingDate id="0877793AFFD8B140AB8FDDE1FBBAA9D7" box="[943,1057,1231,1257]" pageId="3" pageNumber="169" value="2006-03-02">2.III.2006</collectingDate>
</date>
,
<collectorName id="C178C3C4FFD8B140AC0DDDE1FB5FA9D7" box="[1069,1220,1231,1257]" pageId="3" pageNumber="169">J. Poussereau</collectorName>
leg. (
<specimenCount id="7A8B6D9BFFD8B140AD26DDE1FAB5A9D7" box="[1286,1326,1231,1257]" pageId="3" pageNumber="169" type="female">1 ♀</specimenCount>
<typeStatus id="B33618B0FFD8B140AD15DDE1FA03A9D7" box="[1333,1432,1231,1257]" pageId="3" pageNumber="169" type="paratype">paratype</typeStatus>
,
<collectionCode id="0A9C3ED7FFD8B140A8B7DDDDFF7CA833" box="[151,231,1267,1293]" pageId="3" pageNumber="169">NGPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="DCE5AC4FFFD8B140A8DADDDDFEFBA80F" ID-GBIF-Occurrence="3118911307" collectingDate="2009-02-19" collectionCode="NGPC" collectorName="S. Gasnier" country="France" county="La" latitude="-20.883333" location="La Grande Chaloupe" longLatPrecision="1266" longitude="55.366665" municipality="Saint-Denis" pageId="3" pageNumber="169" specimenCount="1" specimenCount-female="1" stateProvince="Reunion" typeStatus="paratype">
<collectingCounty id="8553DE9EFFD8B140A8DADDDDFE83A833" box="[250,280,1267,1293]" pageId="3" pageNumber="169">La</collectingCounty>
<collectingRegion id="AE4968F0FFD8B140A93FDDDDFE1AA833" box="[287,385,1267,1293]" country="France" name="Reunion" pageId="3" pageNumber="169">Réunion</collectingRegion>
,
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,
<location id="6952F0C9FFD8B140AB15DC15FC4BA86B" LSID="urn:lsid:plazi:treatment:E4241704FFD9B140A8B7DF50FB00A8A3:6952F0C9FFD8B140AB15DC15FC4BA86B" box="[821,976,1339,1365]" country="France" county="La" latitude="-20.933332" longLatPrecision="1266" longitude="55.516666" municipality="Sainte-Marie" name="La Ressource" pageId="3" pageNumber="169" stateProvince="Reunion">La Ressource</location>
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<collectingDate id="0877793AFFD8B140AA77DCADFD5FA8A3" box="[599,708,1411,1437]" pageId="3" pageNumber="169" value="2018-01-26">26.I.2018</collectingDate>
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,
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leg. (
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)
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.
</paragraph>
</subSubSection>
</treatment>
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