245 lines
40 KiB
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245 lines
40 KiB
XML
<document id="C5D611F673FA32ECAA0F4ABD7A016564" ID-DOI="10.11646/zootaxa.5448.3.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="11232043" ID-ZooBank="2435C13A-609B-49F4-AEC3-3B4A0D11DD40" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1716279637972" checkinUser="plazi" docAuthor="Anisyutkin, Leonid N., Vasilenko, Dmitry V. & Perkovsky, Evgeny E." docDate="2024" docId="E2598797FFB3FFD8FF485F69FBA159EA" docLanguage="en" docName="zootaxa.5448.3.7.pdf" docOrigin="Zootaxa 5448 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5360.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Supella eocenica Anisyutkin et Perkovsky 2023" docType="treatment" docVersion="2" lastPageNumber="444" masterDocId="1E60FFEFFFB0FFDDFFDF5C77FFC75B1B" masterDocTitle="New data on the genus Supella Shelford, 1911 (Dictyoptera: Pseudophyllodromiidae) with description of female of S. eocenica Anisyutkin et Perkovsky, 2023 from Rovno amber" masterLastPageNumber="445" masterPageNumber="439" pageNumber="442" updateTime="1716280884736" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="13B2AE6A14C504189A522F70A4ADDB33">New data on the genus Supella Shelford, 1911 (Dictyoptera: Pseudophyllodromiidae) with description of female of S. eocenica Anisyutkin et Perkovsky, 2023 from Rovno amber</mods:title>
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<mods:name id="749EA1E978736BAADECEECD78A80E0BA" type="personal">
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<mods:roleTerm id="4A5D73785B31BBEB03EB79F2F8FD815F">Author</mods:roleTerm>
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<mods:namePart id="8177FB64B9E2E9DE15B4E61CE736BA5A">Anisyutkin, Leonid N.</mods:namePart>
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<mods:affiliation id="EE6AFB5D198036501F97AF6E679B9EE4">Zoological Institute, Russian Academy of Sciences, Universitetskaya Emb. 1, Saint Petersburg, 199034 Russia</mods:affiliation>
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<mods:name id="93F8944A2B90C3E94ED72EED8867C560" type="personal">
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<mods:namePart id="9F0C3449D6C91C196A4851A85CD6D70B">Vasilenko, Dmitry V.</mods:namePart>
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<mods:affiliation id="61741AE75BC2197A0DC93AA1C5E59024">Cherepovets State University, Lunacharsky prospect 5, 162600, Cherepovets, Russia & Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow, 117647 Russia</mods:affiliation>
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<mods:namePart id="A0FD41F859D548831FEA3DB5EBD7FE83">Perkovsky, Evgeny E.</mods:namePart>
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<mods:affiliation id="31FA5903DDE70CBA4F225F1E28E9C68B">Natural History Museum of Denmark, Universitetsparken 15, Copenhagen</mods:affiliation>
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<mods:title id="61360C01F698FD228D7900E701B9877A">Zootaxa</mods:title>
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<mods:part id="D8779CF2E3FA46D84EAB6AD6A1FD9B77">
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<mods:date id="CFC166B93CF18F25FA19F825F38687F0">2024</mods:date>
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<mods:number id="427A8FBF3578030A54C9DC0E1CFB45E0">2024-05-13</mods:number>
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<mods:number id="D9955781DF90F09A8E134D0185C564BA">5448</mods:number>
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<mods:classification id="500DAF6110712FD01D3127E311FDCC39">journal article</mods:classification>
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<treatment id="E2598797FFB3FFD8FF485F69FBA159EA" ID-DOI="http://doi.org/10.5281/zenodo.11232059" ID-Zenodo-Dep="11232059" LSID="urn:lsid:plazi:treatment:E2598797FFB3FFD8FF485F69FBA159EA" httpUri="http://treatment.plazi.org/id/E2598797FFB3FFD8FF485F69FBA159EA" lastPageId="5" lastPageNumber="444" pageId="3" pageNumber="442">
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<subSubSection id="22EA650AFFB3FFDEFF485F69FD225823" box="[151,741,798,825]" pageId="3" pageNumber="442" type="nomenclature">
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<paragraph id="6A4F3681FFB3FFDEFF485F69FD225823" blockId="3.[151,741,798,861]" box="[151,741,798,825]" pageId="3" pageNumber="442">
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<heading id="310781EDFFB3FFDEFF485F69FD225823" bold="true" box="[151,741,798,825]" fontSize="11" level="1" pageId="3" pageNumber="442" reason="1">
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<taxonomicName id="ADF04D02FFB3FFDEFF485F69FD225823" authority="Anisyutkin et Perkovsky, 2023" authorityName="Anisyutkin et Perkovsky" authorityYear="2023" box="[151,741,798,825]" class="Insecta" family="Ectobiidae" genus="Supella" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="442" phylum="Arthropoda" rank="species" species="eocenica">
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<emphasis id="5884EA93FFB3FFDEFF485F69FD225823" bold="true" box="[151,741,798,825]" pageId="3" pageNumber="442">
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<emphasis id="5884EA93FFB3FFDEFF485F69FEA35822" bold="true" box="[151,356,798,825]" italics="true" pageId="3" pageNumber="442">Supella eocenica</emphasis>
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<bibRefCitation id="0E614B70FFB3FFDEFEB65F69FD225823" author="Anisyutkin, L. N. & Perkovsky, E. E." box="[361,741,798,825]" pageId="3" pageNumber="442" pagination="1721 - 1730" refId="ref3586" refString="Anisyutkin, L. N. & Perkovsky, E. E. (2023) Two new cockroaches (Dictyoptera: Ectobiidae) from Rovno amber. Biologia, 78 (6), 1721 - 1730. https: // doi. org / 10.1007 / s 11756 - 023 - 01333 - 6" type="journal article" year="2023">Anisyutkin et Perkovsky, 2023</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="22EA650AFFB3FFDEFF485F34FE915846" box="[151,342,835,861]" pageId="3" pageNumber="442" type="description">
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<paragraph id="6A4F3681FFB3FFDEFF485F34FE915846" blockId="3.[151,741,798,861]" box="[151,342,835,861]" pageId="3" pageNumber="442">
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(
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||
<figureCitation id="F2CB2A04FFB3FFDEFF405F34FF3C5846" box="[159,251,835,861]" captionStart="FIGURE 1" captionStartId="2.[152,255,1552,1576]" captionTargetBox="[158,1427,189,1528]" captionTargetId="figure-35@2.[151,1435,181,1529]" captionTargetPageId="2" captionText="FIGURE 1. A–C—Supella longipalpa (Fabricius, 1798) [male (A, B) and female (C)] and S. eocenica Anisyutkin et Perkovsky, 2023 [female (D)]. Tegmina from above (A, C) and below (D), wing from above (B). Dotted areas show dark colour. Transverse veins are not shown. Abbreviations: 1pl., 2pl., 3pl., CuA, CuP, M, M+CuA, R, R1, RA, Sc—see text. Scale bar 1 mm: a = A, B; b = C; c = D." figureDoi="http://doi.org/10.5281/zenodo.11232047" httpUri="https://zenodo.org/record/11232047/files/figure.png" pageId="3" pageNumber="442">Figs 1D</figureCitation>
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,
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<figureCitation id="F2CB2A04FFB3FFDEFED75F34FE895846" box="[264,334,835,861]" captionStart="FIGURE 2" captionStartId="4.[152,255,1844,1868]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-35@4.[151,1436,181,1821]" captionTargetPageId="4" captionText="FIGURE 2. Supella eocenica Anisyutkin et Perkovsky, 2023, female. A—general view of inclusion (out of scale); B, C— general view from above and right (B) and below (C); D—abdominal apex and ootheca from below." figureDoi="http://doi.org/10.5281/zenodo.11232049" httpUri="https://zenodo.org/record/11232049/files/figure.png" pageId="3" pageNumber="442">2A–D</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="22EA650AFFB3FFDEFF485FFCFD385F0A" pageId="3" pageNumber="442" type="materials_examined">
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<paragraph id="6A4F3681FFB3FFDEFF485FFCFD385F0A" blockId="3.[151,1437,906,1941]" pageId="3" pageNumber="442">
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<emphasis id="5884EA93FFB3FFDEFF485FFCFEF858BF" bold="true" box="[151,319,906,933]" pageId="3" pageNumber="442">New material.</emphasis>
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<collectionCode id="0CE1AE44FFB3FFDEFE985FFCFE4258BE" box="[327,389,907,933]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="3" pageNumber="442">SIZK</collectionCode>
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<collectionCode id="0CE1AE44FFB3FFDEFE535FFCFE7958BE" box="[396,446,907,933]" pageId="3" pageNumber="442">UA-</collectionCode>
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29000,
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<collectingRegion id="A834F863FFB3FFDEFDCD5FFCFD9A58BE" box="[530,605,907,933]" country="Ukraine" name="Rivne" pageId="3" pageNumber="442">Rovno</collectingRegion>
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amber, late Eocene. Syninclusions: Acari:
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<taxonomicName id="ADF04D02FFB3FFDEFBE05FFCFB6958BE" authorityName="A.L.Duges" authorityYear="1834" box="[1087,1198,907,933]" class="Arachnida" family="Bdellidae" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="442" phylum="Arthropoda" rank="family">Bdellidae</taxonomicName>
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; Aranei; Collembola, Entomobryomorpha; Collembola, Poduromorpha;
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<specimenCount id="7CF6FD08FFB3FFDEFD075FD8FC4858D2" box="[728,911,943,969]" count="3" pageId="3" pageNumber="442" type="generic">three specimens</specimenCount>
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of
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<taxonomicName id="ADF04D02FFB3FFDEFC645FD8FADF58D2" authority="(Aphidoidea)" baseAuthorityName="Aphidoidea" box="[955,1304,943,969]" class="Insecta" family="Drepanosiphidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="442" phylum="Arthropoda" rank="family">Drepanosiphidae (Aphidoidea)</taxonomicName>
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from three different species;
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<taxonomicName id="ADF04D02FFB3FFDEFE825FA4FE2958F6" box="[349,494,979,1005]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="442" phylum="Arthropoda" rank="family">Psychodidae</taxonomicName>
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;
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<specimenCount id="7CF6FD08FFB3FFDEFE265FA4FD5858F6" box="[505,671,979,1005]" count="2" pageId="3" pageNumber="442" type="generic">two specimens</specimenCount>
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||
of
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<taxonomicName id="ADF04D02FFB3FFDEFD1B5FA4FCEB58F6" authorityName="Billberg" authorityYear="1820" box="[708,812,979,1005]" class="Insecta" family="Sciaridae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="442" phylum="Arthropoda" rank="family">Sciaridae</taxonomicName>
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from different species; Thysanoptera; worker of the ant
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<taxonomicName id="ADF04D02FFB3FFDEFF485F80FE365F0A" authority="Mayr" authorityName="Mayr" box="[151,497,1015,1041]" class="Insecta" family="Formicidae" genus="Ctenobethylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="442" phylum="Arthropoda" rank="species" species="goepperti">
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<emphasis id="5884EA93FFB3FFDEFF485F80FE6E5F0B" box="[151,425,1015,1041]" italics="true" pageId="3" pageNumber="442">Ctenobethylus goepperti</emphasis>
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Mayr
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</taxonomicName>
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; stellate hairs (
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<figureCitation id="F2CB2A04FFB3FFDEFD475F80FD345F0A" box="[664,755,1015,1041]" captionStart="FIGURE 2" captionStartId="4.[152,255,1844,1868]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-35@4.[151,1436,181,1821]" captionTargetPageId="4" captionText="FIGURE 2. Supella eocenica Anisyutkin et Perkovsky, 2023, female. A—general view of inclusion (out of scale); B, C— general view from above and right (B) and below (C); D—abdominal apex and ootheca from below." figureDoi="http://doi.org/10.5281/zenodo.11232049" httpUri="https://zenodo.org/record/11232049/files/figure.png" pageId="3" pageNumber="442">Fig. 2A</figureCitation>
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).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="22EA650AFFB3FFD8FF18586DFBA159EA" lastPageId="5" lastPageNumber="444" pageId="3" pageNumber="442" type="description">
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<paragraph id="6A4F3681FFB3FFDEFF18586DFCF35FBA" blockId="3.[151,1437,906,1941]" pageId="3" pageNumber="442">
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<emphasis id="5884EA93FFB3FFDEFF18586DFE965F2F" bold="true" box="[199,337,1050,1076]" pageId="3" pageNumber="442">Description</emphasis>
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.
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<emphasis id="5884EA93FFB3FFDEFEBD586CFE715F2E" box="[354,438,1051,1077]" italics="true" pageId="3" pageNumber="442">Female</emphasis>
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(previously unknown). General coloration, as far as can be judged from the cockroach inclusion, light yellowish (
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<figureCitation id="F2CB2A04FFB3FFDEFE135848FD8B5F42" box="[460,588,1087,1113]" captionStart="FIGURE 2" captionStartId="4.[152,255,1844,1868]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-35@4.[151,1436,181,1821]" captionTargetPageId="4" captionText="FIGURE 2. Supella eocenica Anisyutkin et Perkovsky, 2023, female. A—general view of inclusion (out of scale); B, C— general view from above and right (B) and below (C); D—abdominal apex and ootheca from below." figureDoi="http://doi.org/10.5281/zenodo.11232049" httpUri="https://zenodo.org/record/11232049/files/figure.png" pageId="3" pageNumber="442">Fig. 2B–D</figureCitation>
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); head, pronotum from below, thoracal and abdominal sternites, tegmina, wings, coxae and femora from below and ootheca with scattered dark (black?) dots; larger dots present on tegmina, two large dark spots present on trochanter and genital plate.
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</paragraph>
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<paragraph id="6A4F3681FFB3FFDEFF1858DCFA485FF2" blockId="3.[151,1437,906,1941]" pageId="3" pageNumber="442">
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Dense pubescence present on antennae and tarsi (
|
||
<figureCitation id="F2CB2A04FFB3FFDEFD2358DCFCBD5FDE" box="[764,890,1195,1221]" captionStart="FIGURE 2" captionStartId="4.[152,255,1844,1868]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-35@4.[151,1436,181,1821]" captionTargetPageId="4" captionText="FIGURE 2. Supella eocenica Anisyutkin et Perkovsky, 2023, female. A—general view of inclusion (out of scale); B, C— general view from above and right (B) and below (C); D—abdominal apex and ootheca from below." figureDoi="http://doi.org/10.5281/zenodo.11232049" httpUri="https://zenodo.org/record/11232049/files/figure.png" pageId="3" pageNumber="442">Fig. 2B–D</figureCitation>
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); cerci, genital plate and in less degree adjacent abdominal sternites and ootheca densely covered with large chaetae. Surfaces smooth, visible punctuation absent.
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</paragraph>
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<paragraph id="6A4F3681FFB3FFDEFF185884FE185C32" blockId="3.[151,1437,906,1941]" pageId="3" pageNumber="442">
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Structure of head practically indistinguishable. Antennae long, longer than length of body. Pronotum widened caudally, with lateral margins widely rounded and posterior—weakly caudally projected (
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<figureCitation id="F2CB2A04FFB3FFDEFB465960FB345E2A" box="[1177,1267,1303,1329]" captionStart="FIGURE 2" captionStartId="4.[152,255,1844,1868]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-35@4.[151,1436,181,1821]" captionTargetPageId="4" captionText="FIGURE 2. Supella eocenica Anisyutkin et Perkovsky, 2023, female. A—general view of inclusion (out of scale); B, C— general view from above and right (B) and below (C); D—abdominal apex and ootheca from below." figureDoi="http://doi.org/10.5281/zenodo.11232049" httpUri="https://zenodo.org/record/11232049/files/figure.png" pageId="3" pageNumber="442">Fig. 2B</figureCitation>
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). Tegmina and wings approximately equal to length of the abdomen or slightly surpassed abdominal apex (
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<figureCitation id="F2CB2A04FFB3FFDEFB0F594CFAF05E4E" box="[1232,1335,1339,1365]" captionStart="FIGURE 1" captionStartId="2.[152,255,1552,1576]" captionTargetBox="[158,1427,189,1528]" captionTargetId="figure-35@2.[151,1435,181,1529]" captionTargetPageId="2" captionText="FIGURE 1. A–C—Supella longipalpa (Fabricius, 1798) [male (A, B) and female (C)] and S. eocenica Anisyutkin et Perkovsky, 2023 [female (D)]. Tegmina from above (A, C) and below (D), wing from above (B). Dotted areas show dark colour. Transverse veins are not shown. Abbreviations: 1pl., 2pl., 3pl., CuA, CuP, M, M+CuA, R, R1, RA, Sc—see text. Scale bar 1 mm: a = A, B; b = C; c = D." figureDoi="http://doi.org/10.5281/zenodo.11232047" httpUri="https://zenodo.org/record/11232047/files/figure.png" pageId="3" pageNumber="442">Figs. 1D</figureCitation>
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,
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<figureCitation id="F2CB2A04FFB3FFDEFA98594CFA575E4E" box="[1351,1424,1339,1365]" captionStart="FIGURE 2" captionStartId="4.[152,255,1844,1868]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-35@4.[151,1436,181,1821]" captionTargetPageId="4" captionText="FIGURE 2. Supella eocenica Anisyutkin et Perkovsky, 2023, female. A—general view of inclusion (out of scale); B, C— general view from above and right (B) and below (C); D—abdominal apex and ootheca from below." figureDoi="http://doi.org/10.5281/zenodo.11232049" httpUri="https://zenodo.org/record/11232049/files/figure.png" pageId="3" pageNumber="442">2A–D</figureCitation>
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). Tegmina (
|
||
<figureCitation id="F2CB2A04FFB3FFDEFED95928FEA05E62" box="[262,359,1375,1401]" captionStart="FIGURE 1" captionStartId="2.[152,255,1552,1576]" captionTargetBox="[158,1427,189,1528]" captionTargetId="figure-35@2.[151,1435,181,1529]" captionTargetPageId="2" captionText="FIGURE 1. A–C—Supella longipalpa (Fabricius, 1798) [male (A, B) and female (C)] and S. eocenica Anisyutkin et Perkovsky, 2023 [female (D)]. Tegmina from above (A, C) and below (D), wing from above (B). Dotted areas show dark colour. Transverse veins are not shown. Abbreviations: 1pl., 2pl., 3pl., CuA, CuP, M, M+CuA, R, R1, RA, Sc—see text. Scale bar 1 mm: a = A, B; b = C; c = D." figureDoi="http://doi.org/10.5281/zenodo.11232047" httpUri="https://zenodo.org/record/11232047/files/figure.png" pageId="3" pageNumber="442">Figs. 1D</figureCitation>
|
||
,
|
||
<figureCitation id="F2CB2A04FFB3FFDEFEAD5928FE755E62" box="[370,434,1375,1401]" captionStart="FIGURE 2" captionStartId="4.[152,255,1844,1868]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-35@4.[151,1436,181,1821]" captionTargetPageId="4" captionText="FIGURE 2. Supella eocenica Anisyutkin et Perkovsky, 2023, female. A—general view of inclusion (out of scale); B, C— general view from above and right (B) and below (C); D—abdominal apex and ootheca from below." figureDoi="http://doi.org/10.5281/zenodo.11232049" httpUri="https://zenodo.org/record/11232049/files/figure.png" pageId="3" pageNumber="442">2B, C</figureCitation>
|
||
) with costal field narrow; radial field large, about as wide as fused median and cubital ones; anal field about one third of tegmen length. Tegminal venation distinct; Sc simple, nearly straight and thickened; R nearly straight, with more than 19 regular or pectinate anterior apical rami, 1st–3rd anterior rami close to each other; M and CuA not separated, with more than 13 simple or branched oblique veins; bifurcated vein located before CuP; CuP distinct, weakly curved; at least 2 visible anal veins present; intercalated veins numerous, present in radial, median and cubital fields; transverse veins sparse and not numerous. Wings with distinct venation. Legs with well developed armament (
|
||
<figureCitation id="F2CB2A04FFB3FFDEFE4C5A40FDC85D4A" box="[403,527,1591,1617]" captionStart="FIGURE 2" captionStartId="4.[152,255,1844,1868]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-35@4.[151,1436,181,1821]" captionTargetPageId="4" captionText="FIGURE 2. Supella eocenica Anisyutkin et Perkovsky, 2023, female. A—general view of inclusion (out of scale); B, C— general view from above and right (B) and below (C); D—abdominal apex and ootheca from below." figureDoi="http://doi.org/10.5281/zenodo.11232049" httpUri="https://zenodo.org/record/11232049/files/figure.png" pageId="3" pageNumber="442">Fig. 2B, C</figureCitation>
|
||
). Anterior margin of fore femora armed according to
|
||
<typeStatus id="B54B8823FFB3FFDEFBAC5A40FB645D4A" box="[1139,1187,1591,1617]" pageId="3" pageNumber="442">type</typeStatus>
|
||
A (
|
||
<emphasis id="5884EA93FFB3FFDEFB115A40FACB5D4B" box="[1230,1292,1591,1616]" italics="true" pageId="3" pageNumber="442">sensu</emphasis>
|
||
<bibRefCitation id="0E614B70FFB3FFDEFACA5A40FF155D6E" author="Bey-Bienko, G. Ya." pageId="3" pageNumber="442" refId="ref3707" refString="Bey-Bienko, G. Ya. (1950) Nasekomye Tarakanovye [Cockroach insects]. Fauna of the USSR. New Series 40. Nauka Publishers, Leningrad, Moscow, 343 pp [in Russian]" type="book" year="1950">Bey-Bienko 1950</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="0E614B70FFB3FFDEFF015A2CFE9F5D6E" author="Roth, L. M." box="[222,344,1627,1653]" pageId="3" pageNumber="442" pagination="1 - 186" refId="ref4471" refString="Roth, L. M. (2003) Systematics and phylogeny of cockroaches (Dictyoptera: Blattaria). Oriental Insects, 37, 1 - 186. https: // doi. org / 10.1080 / 00305316.2003.10417344" type="journal article" year="2003">Roth 2003</bibRefCitation>
|
||
), with numerous spines of equal size and 2 (?—difficult to distinguish on inclusion) larger apical spines. Hind tarsus (
|
||
<figureCitation id="F2CB2A04FFB3FFDEFEA55A08FE085D82" box="[378,463,1663,1689]" captionStart="FIGURE 2" captionStartId="4.[152,255,1844,1868]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-35@4.[151,1436,181,1821]" captionTargetPageId="4" captionText="FIGURE 2. Supella eocenica Anisyutkin et Perkovsky, 2023, female. A—general view of inclusion (out of scale); B, C— general view from above and right (B) and below (C); D—abdominal apex and ootheca from below." figureDoi="http://doi.org/10.5281/zenodo.11232049" httpUri="https://zenodo.org/record/11232049/files/figure.png" pageId="3" pageNumber="442">Fig. 2C</figureCitation>
|
||
) with metatarsus longer than other segments combined, 2nd–4th segments short; claws symmetrical; arolium large, about as long as claw length. Abdominal sternites without visible specializations (
|
||
<figureCitation id="F2CB2A04FFB3FFDEFAB05AD4FF7C5DFA" captionStart="FIGURE 2" captionStartId="4.[152,255,1844,1868]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-35@4.[151,1436,181,1821]" captionTargetPageId="4" captionText="FIGURE 2. Supella eocenica Anisyutkin et Perkovsky, 2023, female. A—general view of inclusion (out of scale); B, C— general view from above and right (B) and below (C); D—abdominal apex and ootheca from below." figureDoi="http://doi.org/10.5281/zenodo.11232049" httpUri="https://zenodo.org/record/11232049/files/figure.png" pageId="3" pageNumber="442">Fig. 2B</figureCitation>
|
||
). Cerci long, apically acute, slightly longer than genital plate (
|
||
<figureCitation id="F2CB2A04FFB3FFDEFC4E5AB0FC2A5DFA" box="[913,1005,1735,1761]" captionStart="FIGURE 2" captionStartId="4.[152,255,1844,1868]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-35@4.[151,1436,181,1821]" captionTargetPageId="4" captionText="FIGURE 2. Supella eocenica Anisyutkin et Perkovsky, 2023, female. A—general view of inclusion (out of scale); B, C— general view from above and right (B) and below (C); D—abdominal apex and ootheca from below." figureDoi="http://doi.org/10.5281/zenodo.11232049" httpUri="https://zenodo.org/record/11232049/files/figure.png" pageId="3" pageNumber="442">Fig. 2D</figureCitation>
|
||
), with at least 10 distinctly separated segments. Genital plate subtriangular in shape, weakly incised caudally (
|
||
<figureCitation id="F2CB2A04FFB3FFDEFC135A9CFBE05C1E" box="[972,1063,1771,1797]" captionStart="FIGURE 2" captionStartId="4.[152,255,1844,1868]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-35@4.[151,1436,181,1821]" captionTargetPageId="4" captionText="FIGURE 2. Supella eocenica Anisyutkin et Perkovsky, 2023, female. A—general view of inclusion (out of scale); B, C— general view from above and right (B) and below (C); D—abdominal apex and ootheca from below." figureDoi="http://doi.org/10.5281/zenodo.11232049" httpUri="https://zenodo.org/record/11232049/files/figure.png" pageId="3" pageNumber="442">Fig. 2D</figureCitation>
|
||
). Ootheca short and bulked, with distinct dorsal keel (
|
||
<figureCitation id="F2CB2A04FFB3FFDEFEA55B78FE155C32" box="[378,466,1807,1833]" captionStart="FIGURE 2" captionStartId="4.[152,255,1844,1868]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-35@4.[151,1436,181,1821]" captionTargetPageId="4" captionText="FIGURE 2. Supella eocenica Anisyutkin et Perkovsky, 2023, female. A—general view of inclusion (out of scale); B, C— general view from above and right (B) and below (C); D—abdominal apex and ootheca from below." figureDoi="http://doi.org/10.5281/zenodo.11232049" httpUri="https://zenodo.org/record/11232049/files/figure.png" pageId="3" pageNumber="442">Fig. 2B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6A4F3681FFB3FFDEFF185B44FC565C8E" blockId="3.[151,1437,906,1941]" pageId="3" pageNumber="442">Measurements (mm). Total length 11.4 (as preserved, from head to ootheca apex); pronotum width 3.1; tegmen length 8.0; tegmen width 2.3; hind femur length 3.6; hind tibia length 3.6; ootheca width 1.6. All measurements are approximate due to inclusion distortion and cockroach deformation.</paragraph>
|
||
<caption id="3E8F6609FFB4FFD9FF475B43FB575C6A" ID-DOI="http://doi.org/10.5281/zenodo.11232049" ID-Zenodo-Dep="11232049" httpUri="https://zenodo.org/record/11232049/files/figure.png" pageId="4" pageNumber="443" startId="4.[152,255,1844,1868]" targetBox="[151,1436,181,1821]" targetPageId="4" targetType="figure">
|
||
<paragraph id="6A4F3681FFB4FFD9FF475B43FB575C6A" blockId="4.[151,1436,1844,1905]" pageId="4" pageNumber="443">
|
||
<emphasis id="5884EA93FFB4FFD9FF475B43FEDC5C56" bold="true" box="[152,283,1844,1869]" pageId="4" pageNumber="443">FIGURE 2.</emphasis>
|
||
<taxonomicName id="ADF04D02FFB4FFD9FEFB5B42FCDB5C56" authority="Anisyutkin et Perkovsky, 2023" authorityName="Anisyutkin et Perkovsky" authorityYear="2023" box="[292,796,1845,1869]" class="Insecta" family="Ectobiidae" genus="Supella" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="443" phylum="Arthropoda" rank="species" species="eocenica">
|
||
<emphasis id="5884EA93FFB4FFD9FEFB5B42FE155C57" box="[292,466,1845,1869]" italics="true" pageId="4" pageNumber="443">Supella eocenica</emphasis>
|
||
<bibRefCitation id="0E614B70FFB4FFD9FE055B42FCDB5C56" author="Anisyutkin, L. N. & Perkovsky, E. E." box="[474,796,1845,1869]" pageId="4" pageNumber="443" pagination="1721 - 1730" refId="ref3586" refString="Anisyutkin, L. N. & Perkovsky, E. E. (2023) Two new cockroaches (Dictyoptera: Ectobiidae) from Rovno amber. Biologia, 78 (6), 1721 - 1730. https: // doi. org / 10.1007 / s 11756 - 023 - 01333 - 6" type="journal article" year="2023">Anisyutkin et Perkovsky, 2023</bibRefCitation>
|
||
</taxonomicName>
|
||
, female. A—general view of inclusion (out of scale); B, C— general view from above and right (B) and below (C); D—abdominal apex and ootheca from below.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="6A4F3681FFB5FFD8FF185CE0FCC25A92" blockId="5.[151,1437,151,753]" pageId="5" pageNumber="444">
|
||
<emphasis id="5884EA93FFB5FFD8FF185CE0FECD5BAA" bold="true" box="[199,266,151,177]" pageId="5" pageNumber="444">Notes</emphasis>
|
||
. The remarkable similarity in the structure of the tegmina of the above-described specimen (
|
||
<figureCitation id="F2CB2A04FFB5FFD8FAE55CE0FA545BAA" box="[1338,1427,151,177]" captionStart="FIGURE 1" captionStartId="2.[152,255,1552,1576]" captionTargetBox="[158,1427,189,1528]" captionTargetId="figure-35@2.[151,1435,181,1529]" captionTargetPageId="2" captionText="FIGURE 1. A–C—Supella longipalpa (Fabricius, 1798) [male (A, B) and female (C)] and S. eocenica Anisyutkin et Perkovsky, 2023 [female (D)]. Tegmina from above (A, C) and below (D), wing from above (B). Dotted areas show dark colour. Transverse veins are not shown. Abbreviations: 1pl., 2pl., 3pl., CuA, CuP, M, M+CuA, R, R1, RA, Sc—see text. Scale bar 1 mm: a = A, B; b = C; c = D." figureDoi="http://doi.org/10.5281/zenodo.11232047" httpUri="https://zenodo.org/record/11232047/files/figure.png" pageId="5" pageNumber="444">Fig. 1D</figureCitation>
|
||
) and the brachypterous female of
|
||
<taxonomicName id="ADF04D02FFB5FFD8FDCF5CCCFD605BCE" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[528,679,187,213]" class="Insecta" family="Ectobiidae" genus="Supella" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="longipalpa">
|
||
<emphasis id="5884EA93FFB5FFD8FDCF5CCCFDD95BCE" box="[528,542,187,213]" italics="true" pageId="5" pageNumber="444">S</emphasis>
|
||
.
|
||
<emphasis id="5884EA93FFB5FFD8FDF15CCCFD605BCE" box="[558,679,187,213]" italics="true" pageId="5" pageNumber="444">longipalpa</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="F2CB2A04FFB5FFD8FD665CCCFCF35BCE" box="[697,820,187,213]" captionStart="FIGURE 1" captionStartId="2.[152,255,1552,1576]" captionTargetBox="[158,1427,189,1528]" captionTargetId="figure-35@2.[151,1435,181,1529]" captionTargetPageId="2" captionText="FIGURE 1. A–C—Supella longipalpa (Fabricius, 1798) [male (A, B) and female (C)] and S. eocenica Anisyutkin et Perkovsky, 2023 [female (D)]. Tegmina from above (A, C) and below (D), wing from above (B). Dotted areas show dark colour. Transverse veins are not shown. Abbreviations: 1pl., 2pl., 3pl., CuA, CuP, M, M+CuA, R, R1, RA, Sc—see text. Scale bar 1 mm: a = A, B; b = C; c = D." figureDoi="http://doi.org/10.5281/zenodo.11232047" httpUri="https://zenodo.org/record/11232047/files/figure.png" pageId="5" pageNumber="444">Fig. 1C, D</figureCitation>
|
||
) leaves no doubt that it belongs to the genus
|
||
<taxonomicName id="ADF04D02FFB5FFD8FA9E5CCCFA525BCE" authorityName="Shelford" authorityYear="1911" box="[1345,1429,187,213]" class="Insecta" family="Ectobiidae" genus="Supella" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="444" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5884EA93FFB5FFD8FA9E5CCCFA525BCE" box="[1345,1429,187,213]" italics="true" pageId="5" pageNumber="444">Supella</emphasis>
|
||
</taxonomicName>
|
||
. An extinct species of the genus
|
||
<taxonomicName id="ADF04D02FFB5FFD8FDDF5CA8FD935BE2" authorityName="Shelford" authorityYear="1911" box="[512,596,223,249]" class="Insecta" family="Ectobiidae" genus="Supella" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="444" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5884EA93FFB5FFD8FDDF5CA8FD935BE2" box="[512,596,223,249]" italics="true" pageId="5" pageNumber="444">Supella</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="ADF04D02FFB5FFD8FDBC5CA8FD255BE3" authorityName="Anisyutkin et Perkovsky" authorityYear="2023" box="[611,738,223,249]" class="Insecta" family="Ectobiidae" genus="Supella" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="eocenica">
|
||
<emphasis id="5884EA93FFB5FFD8FDBC5CA8FDB65BE2" box="[611,625,223,249]" italics="true" pageId="5" pageNumber="444">S</emphasis>
|
||
.
|
||
<emphasis id="5884EA93FFB5FFD8FDA05CA8FD255BE3" box="[639,738,223,248]" italics="true" pageId="5" pageNumber="444">eocenica</emphasis>
|
||
</taxonomicName>
|
||
, was previously described by us from
|
||
<collectingRegion id="A834F863FFB5FFD8FB465CA8FB235BE2" box="[1177,1252,223,249]" country="Ukraine" name="Rivne" pageId="5" pageNumber="444">Rovno</collectingRegion>
|
||
amber based on a single male (
|
||
<bibRefCitation id="0E614B70FFB5FFD8FEE65D74FD6F5A06" author="Anisyutkin, L. N. & Perkovsky, E. E." box="[313,680,259,285]" pageId="5" pageNumber="444" pagination="1721 - 1730" refId="ref3586" refString="Anisyutkin, L. N. & Perkovsky, E. E. (2023) Two new cockroaches (Dictyoptera: Ectobiidae) from Rovno amber. Biologia, 78 (6), 1721 - 1730. https: // doi. org / 10.1007 / s 11756 - 023 - 01333 - 6" type="journal article" year="2023">Anisyutkin and Perkovsky 2023</bibRefCitation>
|
||
). Attribution of males and females of cockroaches, especially in the case of developed sexual dimorphism, is complex even for extant cockroaches. Nevertheless, we preferred to refrain from the new species erection, and provisionally classify the female described above as
|
||
<taxonomicName id="ADF04D02FFB5FFD8FBA55D3CFB3E5A7F" authorityName="Anisyutkin et Perkovsky" authorityYear="2023" box="[1146,1273,331,357]" class="Insecta" family="Ectobiidae" genus="Supella" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="eocenica">
|
||
<emphasis id="5884EA93FFB5FFD8FBA55D3CFB4F5A7E" box="[1146,1160,331,357]" italics="true" pageId="5" pageNumber="444">S</emphasis>
|
||
.
|
||
<emphasis id="5884EA93FFB5FFD8FB495D3CFB3E5A7F" box="[1174,1273,331,356]" italics="true" pageId="5" pageNumber="444">eocenica</emphasis>
|
||
</taxonomicName>
|
||
. We hope that further research of
|
||
<collectingRegion id="A834F863FFB5FFD8FEB25D18FE7F5A92" box="[365,440,367,393]" country="Ukraine" name="Rivne" pageId="5" pageNumber="444">Rovno</collectingRegion>
|
||
amber will resolve this issue.
|
||
</paragraph>
|
||
<paragraph id="6A4F3681FFB5FFD8FF185DE4FE7A5902" blockId="5.[151,1437,151,753]" pageId="5" pageNumber="444">
|
||
It can be supposed that the process of shortening of the tegmina and wings has begun in the female of
|
||
<taxonomicName id="ADF04D02FFB5FFD8FAC75DE4FA525AB7" authorityName="Anisyutkin et Perkovsky" authorityYear="2023" box="[1304,1429,403,429]" class="Insecta" family="Ectobiidae" genus="Supella" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="eocenica">
|
||
<emphasis id="5884EA93FFB5FFD8FAC75DE4FAE15AB6" box="[1304,1318,403,429]" italics="true" pageId="5" pageNumber="444">S</emphasis>
|
||
.
|
||
<emphasis id="5884EA93FFB5FFD8FAED5DE4FA525AB7" box="[1330,1429,403,428]" italics="true" pageId="5" pageNumber="444">eocenica</emphasis>
|
||
</taxonomicName>
|
||
. This is evidenced by the relatively short and wide tegmina, although exceeding the length of the abdomen, and the convex anterior margin of the tegmen (compare
|
||
<figureCitation id="F2CB2A04FFB5FFD8FD655DACFCDA5AEE" box="[698,797,475,501]" captionStart="FIGURE 1" captionStartId="2.[152,255,1552,1576]" captionTargetBox="[158,1427,189,1528]" captionTargetId="figure-35@2.[151,1435,181,1529]" captionTargetPageId="2" captionText="FIGURE 1. A–C—Supella longipalpa (Fabricius, 1798) [male (A, B) and female (C)] and S. eocenica Anisyutkin et Perkovsky, 2023 [female (D)]. Tegmina from above (A, C) and below (D), wing from above (B). Dotted areas show dark colour. Transverse veins are not shown. Abbreviations: 1pl., 2pl., 3pl., CuA, CuP, M, M+CuA, R, R1, RA, Sc—see text. Scale bar 1 mm: a = A, B; b = C; c = D." figureDoi="http://doi.org/10.5281/zenodo.11232047" httpUri="https://zenodo.org/record/11232047/files/figure.png" pageId="5" pageNumber="444">Figs. 1D</figureCitation>
|
||
,
|
||
<figureCitation id="F2CB2A04FFB5FFD8FCF55DACFC8C5AEE" box="[810,843,475,501]" captionStart="FIGURE 2" captionStartId="4.[152,255,1844,1868]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-35@4.[151,1436,181,1821]" captionTargetPageId="4" captionText="FIGURE 2. Supella eocenica Anisyutkin et Perkovsky, 2023, female. A—general view of inclusion (out of scale); B, C— general view from above and right (B) and below (C); D—abdominal apex and ootheca from below." figureDoi="http://doi.org/10.5281/zenodo.11232049" httpUri="https://zenodo.org/record/11232049/files/figure.png" pageId="5" pageNumber="444">2C</figureCitation>
|
||
with
|
||
<figureCitation id="F2CB2A04FFB5FFD8FC525DACFBCF5AEE" box="[909,1032,475,501]" captionStart="FIGURE 1" captionStartId="2.[152,255,1552,1576]" captionTargetBox="[158,1427,189,1528]" captionTargetId="figure-35@2.[151,1435,181,1529]" captionTargetPageId="2" captionText="FIGURE 1. A–C—Supella longipalpa (Fabricius, 1798) [male (A, B) and female (C)] and S. eocenica Anisyutkin et Perkovsky, 2023 [female (D)]. Tegmina from above (A, C) and below (D), wing from above (B). Dotted areas show dark colour. Transverse veins are not shown. Abbreviations: 1pl., 2pl., 3pl., CuA, CuP, M, M+CuA, R, R1, RA, Sc—see text. Scale bar 1 mm: a = A, B; b = C; c = D." figureDoi="http://doi.org/10.5281/zenodo.11232047" httpUri="https://zenodo.org/record/11232047/files/figure.png" pageId="5" pageNumber="444">Fig. 1A, C</figureCitation>
|
||
). The last character is characteristic of brachypterous tegmina.
|
||
</paragraph>
|
||
<paragraph id="6A4F3681FFB5FFD8FF185E54FBD1599E" blockId="5.[151,1437,151,753]" pageId="5" pageNumber="444">The homologation of the bifurcated vein located immediately anterior to CuP is unclear. This vein may be the last branch of M+CuA, but it may also be an independent CuA. In the latter case, the vein system lying anterior of this vein is M, and not M+CuA. Further research is needed to resolve this issue.</paragraph>
|
||
<paragraph id="6A4F3681FFB5FFD8FF185EF8FBA159EA" blockId="5.[151,1437,151,753]" pageId="5" pageNumber="444">
|
||
It is also necessary to note an interesting character present in females of
|
||
<taxonomicName id="ADF04D02FFB5FFD8FBD45EF8FB6559B2" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[1035,1186,655,681]" class="Insecta" family="Ectobiidae" genus="Supella" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="longipalpa">
|
||
<emphasis id="5884EA93FFB5FFD8FBD45EF8FBDE59B2" box="[1035,1049,655,681]" italics="true" pageId="5" pageNumber="444">S</emphasis>
|
||
.
|
||
<emphasis id="5884EA93FFB5FFD8FBF65EF8FB6559B2" box="[1065,1186,655,681]" italics="true" pageId="5" pageNumber="444">longipalpa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="ADF04D02FFB5FFD8FB025EF8FA9959B3" authorityName="Anisyutkin et Perkovsky" authorityYear="2023" box="[1245,1374,655,681]" class="Insecta" family="Ectobiidae" genus="Supella" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="eocenica">
|
||
<emphasis id="5884EA93FFB5FFD8FB025EF8FB2C59B2" box="[1245,1259,655,681]" italics="true" pageId="5" pageNumber="444">S</emphasis>
|
||
.
|
||
<emphasis id="5884EA93FFB5FFD8FB245EF8FA9959B3" box="[1275,1374,655,680]" italics="true" pageId="5" pageNumber="444">eocenica</emphasis>
|
||
</taxonomicName>
|
||
—the closeness of the first anterior rami of R in the tegmina (
|
||
<figureCitation id="F2CB2A04FFB5FFD8FCF65EC4FC6C59D6" box="[809,939,691,717]" captionStart="FIGURE 1" captionStartId="2.[152,255,1552,1576]" captionTargetBox="[158,1427,189,1528]" captionTargetId="figure-35@2.[151,1435,181,1529]" captionTargetPageId="2" captionText="FIGURE 1. A–C—Supella longipalpa (Fabricius, 1798) [male (A, B) and female (C)] and S. eocenica Anisyutkin et Perkovsky, 2023 [female (D)]. Tegmina from above (A, C) and below (D), wing from above (B). Dotted areas show dark colour. Transverse veins are not shown. Abbreviations: 1pl., 2pl., 3pl., CuA, CuP, M, M+CuA, R, R1, RA, Sc—see text. Scale bar 1 mm: a = A, B; b = C; c = D." figureDoi="http://doi.org/10.5281/zenodo.11232047" httpUri="https://zenodo.org/record/11232047/files/figure.png" pageId="5" pageNumber="444">Fig. 1C, D</figureCitation>
|
||
). In the future, it is necessary to study the distribution of this character in representatives of the genus
|
||
<taxonomicName id="ADF04D02FFB5FFD8FCE95EA0FC4D59EA" authorityName="Shelford" authorityYear="1911" box="[822,906,727,753]" class="Insecta" family="Ectobiidae" genus="Supella" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="444" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5884EA93FFB5FFD8FCE95EA0FC4D59EA" box="[822,906,727,753]" italics="true" pageId="5" pageNumber="444">Supella</emphasis>
|
||
</taxonomicName>
|
||
and related genera.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |