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<document id="0BB50CE3419DC67FFAF12045CEF2384D" ID-DOI="10.11646/zootaxa.3374.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5253502" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629794947778" checkinUser="felipe" docAuthor="Naderloo, Reza &amp; Türkay, Michael" docDate="2012" docId="E177585FC6361B03FF3CFAA7FBA3F805" docLanguage="en" docName="zt03374p067.pdf" docOrigin="Zootaxa 3374 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.3374.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Opusia indica" docType="treatment" docVersion="3" lastPageNumber="49" masterDocId="1D4E2027C6061B33FFABFF26FF85FFDD" masterDocTitle="Decapod crustaceans of the littoral and shallow sublittoral Iranian coast of the Persian Gulf: Faunistics, Biodiversity and Zoogeography 3374" masterLastPageNumber="67" masterPageNumber="1" pageNumber="49" updateTime="1699112790140" updateUser="plazi" zenodo-license-document="CLOSED">
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<subSubSection id="21C4BAC2C6361B03FF3CFAA7FD70F9D9" pageId="48" pageNumber="49" type="nomenclature">
<paragraph id="6961E949C6361B03FF3CFAA7FE7CFA47" blockId="48.[151,505,1408,1435]" box="[151,505,1408,1435]" pageId="48" pageNumber="49">
<heading id="32295E25C6361B03FF3CFAA7FE7CFA47" bold="true" box="[151,505,1408,1435]" fontSize="11" level="1" pageId="48" pageNumber="49" reason="1">
<taxonomicName id="AEDE92CAC6361B03FF3CFAA7FE7CFA47" ID-CoL="74S2X" authority="(Alcock, 1900)" baseAuthorityName="Alcock" baseAuthorityYear="1900" box="[151,505,1408,1435]" class="Malacostraca" family="Camptandriidae" genus="Opusia" kingdom="Animalia" order="Decapoda" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="5BAA355BC6361B03FF3CFAA7FE7CFA47" bold="true" box="[151,505,1408,1435]" pageId="48" pageNumber="49">
<emphasis id="5BAA355BC6361B03FF3CFAA7FEBAFA46" bold="true" box="[151,319,1409,1435]" italics="true" pageId="48" pageNumber="49">Opusia indica</emphasis>
(
<bibRefCitation id="0D4F94B8C6361B03FEE5FAA6FE74FA47" author="Alcock, A." box="[334,497,1408,1434]" pageId="48" pageNumber="49" pagination="279 - 456" refId="ref44374" refString="Alcock, A. (1900) Material for a carcinological Fauna of India. No. 6: The Brachyura Catometopa, or Grapsoidea. Journal of the Asiatic Society of Bengal, 69 (2), 279 - 456." type="journal article" year="1900">Alcock, 1900</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="6961E949C6361B03FF3CFAE1FD70F9D9" blockId="48.[151,1436,1479,2009]" pageId="48" pageNumber="49">
<emphasis id="5BAA355BC6361B03FF3CFAE1FEB3FA3D" bold="true" box="[151,310,1479,1504]" pageId="48" pageNumber="49">Persian Gulf.</emphasis>
<collectingCountry id="11C9A9D9C6361B03FE96FAEEFEEFFA02" box="[317,362,1480,1503]" name="Iraq" pageId="48" pageNumber="49">Iraq</collectingCountry>
(
<bibRefCitation id="0D4F94B8C6361B03FED2FAE1FDA4FA02" author="Ng, P. K. L. &amp; Rahayu, D. L. &amp; Naser, M. D." box="[377,545,1479,1504]" pageId="48" pageNumber="49" pagination="1 - 26" refId="ref51169" refString="Ng, P. K. L., Rahayu, D. L. &amp; Naser, M. D. (2009) The Campatndriidae of Iraq, with description of a new genus and notes on Leptochryseus Al-khayat &amp; Jones, 1996 (Crustacea: Decapoda: Brachyura). Zootaxa, 2312, 1 - 26." type="journal article" year="2009">
Ng
<emphasis id="5BAA355BC6361B03FE0EFAEFFE53FA3D" box="[421,470,1480,1504]" italics="true" pageId="48" pageNumber="49">et al</emphasis>
. 2009
</bibRefCitation>
),
<collectingCountry id="11C9A9D9C6361B03FD9FFAE1FD03FA02" box="[564,646,1479,1503]" name="Kuwait" pageId="48" pageNumber="49">Kuwait</collectingCountry>
(
<bibRefCitation id="0D4F94B8C6361B03FD3EFAE1FCA0FA02" author="Jones, D. A." box="[661,805,1479,1503]" pageId="48" pageNumber="49" refId="ref48604" refString="Jones, D. A. (1986 a) A field guide to the sea shores of Kuwait and the Persian Gulf. University of Kuwait Blandford Press, Poole, Kuwait, 192 pp." type="book" year="1986">Jones 1986a</bibRefCitation>
;
<bibRefCitation id="0D4F94B8C6361B03FC9BFAE1FB91FA02" author="Snowden, R. J. &amp; Clayton, D. A. &amp; Al-Taher, E. Y." box="[816,1044,1479,1504]" pageId="48" pageNumber="49" pagination="219 - 228" refId="ref52886" refString="Snowden, R. J., Clayton, D. A. &amp; Al-Taher, E. Y. (1994) Aspects of the ecology of Tylodiplax indica Alcock (Brachyura: Ocypodidae) on a Kuwait mudflat. Journal of the University of Kuwait (Science), 21, 219 - 228." type="journal article" year="1994">
Snowden
<emphasis id="5BAA355BC6361B03FC0BFAEFFC57FA3D" box="[928,978,1480,1504]" italics="true" pageId="48" pageNumber="49">et al</emphasis>
1994
</bibRefCitation>
),
<collectingCountry id="11C9A9D9C6361B03FB82FAE1FB39FA02" box="[1065,1212,1479,1503]" name="Saudi Arabia" pageId="48" pageNumber="49">Saudi Arabia</collectingCountry>
(
<bibRefCitation id="0D4F94B8C6361B03FB60FAE1FA0BFA02" author="Stephensen, K." box="[1227,1422,1479,1503]" pageId="48" pageNumber="49" pagination="57 - 237" refId="ref52943" refString="Stephensen, K. (1946) The Brachyura of the Iranian Gulf. Danish Scientific Investigations in Iran, Part IV. E. Munksgaard, Copenhagen, 57 - 237." type="book chapter" year="1946">Stephensen 1946</bibRefCitation>
),
<collectingCountry id="11C9A9D9C6361B03FF3CFACAFF41F9DE" box="[151,196,1516,1539]" name="Iran" pageId="48" pageNumber="49">Iran</collectingCountry>
(
<bibRefCitation id="0D4F94B8C6361B03FF79FACAFE10F9D9" author="Stephensen, K." box="[210,405,1516,1540]" pageId="48" pageNumber="49" pagination="57 - 237" refId="ref52943" refString="Stephensen, K. (1946) The Brachyura of the Iranian Gulf. Danish Scientific Investigations in Iran, Part IV. E. Munksgaard, Copenhagen, 57 - 237." type="book chapter" year="1946">Stephensen 1946</bibRefCitation>
;
<bibRefCitation id="0D4F94B8C6361B03FE0AFACAFDC1F9D9" author="Ng, P. K. L. &amp; Rahayu, D. L. &amp; Naser, M. D." box="[417,580,1516,1540]" pageId="48" pageNumber="49" pagination="1 - 26" refId="ref51169" refString="Ng, P. K. L., Rahayu, D. L. &amp; Naser, M. D. (2009) The Campatndriidae of Iraq, with description of a new genus and notes on Leptochryseus Al-khayat &amp; Jones, 1996 (Crustacea: Decapoda: Brachyura). Zootaxa, 2312, 1 - 26." type="journal article" year="2009">
Ng
<emphasis id="5BAA355BC6361B03FE61FACBFE7EF9D9" box="[458,507,1516,1540]" italics="true" pageId="48" pageNumber="49">et al</emphasis>
. 2009
</bibRefCitation>
; present study).
</paragraph>
</subSubSection>
<subSubSection id="21C4BAC2C6361B03FF6CF936FDFBF965" pageId="48" pageNumber="49" type="materials_examined">
<paragraph id="6961E949C6361B03FF6CF936FBD8F9B2" blockId="48.[151,1436,1479,2009]" pageId="48" pageNumber="49">
<materialsCitation id="D9B6E314C6361B03FF6CF936FE0CF991" collectorName="Qeshm I." country="Iran" location="Hormozgan Province" pageId="48" pageNumber="49" specimenCount="1" stateProvince="Hormozgan">
<emphasis id="5BAA355BC6361B03FF6CF936FE87F9F5" bold="true" box="[199,258,1552,1576]" pageId="48" pageNumber="49">
<collectingCountry id="11C9A9D9C6361B03FF6CF936FF7BF9F5" box="[199,254,1552,1576]" name="Iran" pageId="48" pageNumber="49">Iran</collectingCountry>
.
</emphasis>
<emphasis id="5BAA355BC6361B03FEA7F937FE7AF9FA" box="[268,511,1552,1576]" italics="true" pageId="48" pageNumber="49">
<collectingRegion id="AB1A27ABC6361B03FEA7F937FE7AF9FA" box="[268,511,1552,1576]" country="Iran" name="Hormozgan" pageId="48" pageNumber="49">Hormozgan Province</collectingRegion>
</emphasis>
: Bandar-Tiab,
<collectorName id="C42B8C9FC6361B03FD06F929FC92F9FA" box="[685,791,1551,1575]" pageId="48" pageNumber="49">Qeshm I.</collectorName>
(11 stations around the island), Bandar-Abbas, Bandar- Pohl, Bandar-Khamir
</materialsCitation>
.
<emphasis id="5BAA355BC6361B03FE38F912FDDEF991" box="[403,603,1588,1612]" italics="true" pageId="48" pageNumber="49">
<collectingRegion id="AB1A27ABC6361B03FE38F912FDDEF991" box="[403,603,1588,1612]" country="Iran" name="Bushehr" pageId="48" pageNumber="49">Bushehr Province</collectingRegion>
</emphasis>
: Khalij-Nayband, Bandar-Banak, Delvar, Ramleh, Bandar-Rig.
<emphasis id="5BAA355BC6361B03FA8CF912FF7EF9B2" italics="true" pageId="48" pageNumber="49">
<collectingRegion id="AB1A27ABC6361B03FA8CF912FF7EF9B2" country="Iran" name="Khuzestan" pageId="48" pageNumber="49">Khuzestan Province</collectingRegion>
</emphasis>
: Hendijan, Mahshahr (mangroves of Majidieh Fishery Jetty), Bandar-Emam.
</paragraph>
<paragraph id="6961E949C6361B03FF6CF95DFB3FF949" blockId="48.[151,1436,1479,2009]" box="[199,1210,1659,1684]" pageId="48" pageNumber="49">
<emphasis id="5BAA355BC6361B03FF6CF95DFE46F949" bold="true" box="[199,451,1659,1684]" inLineHeading="true" pageId="48" pageNumber="49" reason="1">General distribution.</emphasis>
Northwestern Indian Ocean: Persian Gulf, Gulf of
<collectingCountry id="11C9A9D9C6361B03FBAAF95AFBCDF949" box="[1025,1096,1660,1684]" name="Oman" pageId="48" pageNumber="49">Oman</collectingCountry>
,
<collectingCountry id="11C9A9D9C6361B03FBFFF95AFB33F949" box="[1108,1206,1660,1684]" name="Pakistan" pageId="48" pageNumber="49">Pakistan</collectingCountry>
.
</paragraph>
<paragraph id="6961E949C6361B03FF6CF9B9FDFBF965" blockId="48.[151,1436,1479,2009]" box="[199,638,1695,1720]" pageId="48" pageNumber="49">
<emphasis id="5BAA355BC6361B03FF6CF9B9FEAFF965" bold="true" box="[199,298,1695,1720]" inLineHeading="true" pageId="48" pageNumber="49" reason="1">Habitat.</emphasis>
Muddy intertidal, mangroves.
</paragraph>
</subSubSection>
<subSubSection id="21C4BAC2C6361B03FF6CF9E2FBA3F805" pageId="48" pageNumber="49" type="discussion">
<paragraph id="6961E949C6361B03FF6CF9E2FBA3F805" blockId="48.[151,1436,1479,2009]" pageId="48" pageNumber="49">
<emphasis id="5BAA355BC6361B03FF6CF9E2FEBEF901" bold="true" box="[199,315,1732,1756]" inLineHeading="true" pageId="48" pageNumber="49" reason="1">Remarks.</emphasis>
<taxonomicName id="AEDE92CAC6361B03FEE1F9E2FD36F901" authority="(Alcock, 1900)" baseAuthorityName="Alcock" baseAuthorityYear="1900" box="[330,691,1732,1756]" class="Malacostraca" family="Camptandriidae" genus="Opusia" kingdom="Animalia" order="Decapoda" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="5BAA355BC6361B03FEE1F9E2FE74F901" box="[330,497,1732,1756]" italics="true" pageId="48" pageNumber="49">Opusia indica</emphasis>
(
<bibRefCitation id="0D4F94B8C6361B03FDA2F9E2FD2EF901" author="Alcock, A." box="[521,683,1732,1756]" pageId="48" pageNumber="49" pagination="279 - 456" refId="ref44374" refString="Alcock, A. (1900) Material for a carcinological Fauna of India. No. 6: The Brachyura Catometopa, or Grapsoidea. Journal of the Asiatic Society of Bengal, 69 (2), 279 - 456." type="journal article" year="1900">Alcock, 1900</bibRefCitation>
)
</taxonomicName>
was described by
<bibRefCitation id="0D4F94B8C6361B03FC0EF9E2FBD5F901" author="Alcock, A." box="[933,1104,1732,1756]" pageId="48" pageNumber="49" pagination="279 - 456" refId="ref44374" refString="Alcock, A. (1900) Material for a carcinological Fauna of India. No. 6: The Brachyura Catometopa, or Grapsoidea. Journal of the Asiatic Society of Bengal, 69 (2), 279 - 456." type="journal article" year="1900">Alcock (1900)</bibRefCitation>
from Karachi,
<collectingCountry id="11C9A9D9C6361B03FABDF9E2FAF0F901" box="[1302,1397,1732,1756]" name="Pakistan" pageId="48" pageNumber="49">Pakistan</collectingCountry>
as
<taxonomicName id="AEDE92CAC6361B03FF3CF9CEFEDFF8DD" authorityName="Alcock" authorityYear="1900" box="[151,346,1768,1792]" class="Malacostraca" family="Camptandriidae" genus="Tylodiplax" kingdom="Animalia" order="Decapoda" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="5BAA355BC6361B03FF3CF9CEFEDFF8DD" box="[151,346,1768,1792]" italics="true" pageId="48" pageNumber="49">Tylodiplax indica</emphasis>
</taxonomicName>
.
<bibRefCitation id="0D4F94B8C6361B03FEC1F9CEFDA5F922" author="Ng, P. K. L. &amp; Rahayu, D. L. &amp; Naser, M. D." box="[362,544,1767,1792]" pageId="48" pageNumber="49" pagination="1 - 26" refId="ref51169" refString="Ng, P. K. L., Rahayu, D. L. &amp; Naser, M. D. (2009) The Campatndriidae of Iraq, with description of a new genus and notes on Leptochryseus Al-khayat &amp; Jones, 1996 (Crustacea: Decapoda: Brachyura). Zootaxa, 2312, 1 - 26." type="journal article" year="2009">
Ng
<emphasis id="5BAA355BC6361B03FE3FF9CFFE43F8DD" box="[404,454,1768,1792]" italics="true" pageId="48" pageNumber="49">et al</emphasis>
. (2009)
</bibRefCitation>
recently have described the monotypic genus
<taxonomicName id="AEDE92CAC6361B03FB86F9CEFBFBF8DD" authorityName="P. K. L. Ng, Rahayu &amp; Naser" authorityYear="2009" box="[1069,1150,1768,1792]" class="Malacostraca" family="Camptandriidae" genus="Opusia" kingdom="Animalia" order="Decapoda" pageId="48" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="5BAA355BC6361B03FB86F9CEFBFBF8DD" box="[1069,1150,1768,1792]" italics="true" pageId="48" pageNumber="49">Opusia</emphasis>
</taxonomicName>
and recorded the species
<taxonomicName id="AEDE92CAC6361B03FF3CF82AFE84F8F9" authorityName="Alcock" authorityYear="1900" box="[151,257,1804,1828]" class="Malacostraca" family="Camptandriidae" genus="Tylodiplax" kingdom="Animalia" order="Decapoda" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="5BAA355BC6361B03FF3CF82AFF2EF8F9" box="[151,171,1804,1828]" italics="true" pageId="48" pageNumber="49">O</emphasis>
.
<emphasis id="5BAA355BC6361B03FF10F82AFE84F8F9" box="[187,257,1804,1828]" italics="true" pageId="48" pageNumber="49">indica</emphasis>
</taxonomicName>
from
<collectingCountry id="11C9A9D9C6361B03FEE1F82AFEFEF8FE" box="[330,379,1804,1827]" name="Iraq" pageId="48" pageNumber="49">Iraq</collectingCountry>
. According to
<bibRefCitation id="0D4F94B8C6361B03FD8DF82AFD64F8F9" author="Ng, P. K. L. &amp; Rahayu, D. L. &amp; Naser, M. D." box="[550,737,1804,1828]" pageId="48" pageNumber="49" pagination="1 - 26" refId="ref51169" refString="Ng, P. K. L., Rahayu, D. L. &amp; Naser, M. D. (2009) The Campatndriidae of Iraq, with description of a new genus and notes on Leptochryseus Al-khayat &amp; Jones, 1996 (Crustacea: Decapoda: Brachyura). Zootaxa, 2312, 1 - 26." type="journal article" year="2009">
Ng
<emphasis id="5BAA355BC6361B03FDFAF82BFD01F8F9" box="[593,644,1804,1828]" italics="true" pageId="48" pageNumber="49">et al</emphasis>
. (2009)
</bibRefCitation>
,
<taxonomicName id="AEDE92CAC6361B03FD44F82AFCC5F8F9" authorityName="P. K. L. Ng, Rahayu &amp; Naser" authorityYear="2009" box="[751,832,1804,1828]" class="Malacostraca" family="Camptandriidae" genus="Opusia" kingdom="Animalia" order="Decapoda" pageId="48" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="5BAA355BC6361B03FD44F82AFCC5F8F9" box="[751,832,1804,1828]" italics="true" pageId="48" pageNumber="49">Opusia</emphasis>
</taxonomicName>
is mainly characterized by a broadly ovate carapace without distinct ridges on dorsal surface (such ridges are seen in
<taxonomicName id="AEDE92CAC6361B03FCC7F816FC67F895" authorityName="de Man, 1895 [in de Man, 1895-" authorityYear="1898" box="[876,994,1840,1864]" class="Malacostraca" family="Camptandriidae" genus="Tylodiplax" kingdom="Animalia" order="Decapoda" pageId="48" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="5BAA355BC6361B03FCC7F816FC67F895" box="[876,994,1840,1864]" italics="true" pageId="48" pageNumber="49">Tylodiplax</emphasis>
</taxonomicName>
), anterolateral margin making no angle with posterolateral margin, the anterior part of the first two thoracic sternites of males are rounded (v.s. triangular in
<taxonomicName id="AEDE92CAC6361B03FF1CF85EFEA8F84D" authorityName="de Man, 1895 [in de Man, 1895-" authorityYear="1898" box="[183,301,1912,1936]" class="Malacostraca" family="Camptandriidae" genus="Tylodiplax" kingdom="Animalia" order="Decapoda" pageId="48" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="5BAA355BC6361B03FF1CF85EFEA8F84D" box="[183,301,1912,1936]" italics="true" pageId="48" pageNumber="49">Tylodiplax</emphasis>
</taxonomicName>
), a distinct complete suture between male abdominal segments, segments 3-5 functionally fused, abdominal segment 5 basically constricted, male G1 ending in a conical apex. Detailed morphological and taxonomic remarks for this genus and allied genera are given by
<bibRefCitation id="0D4F94B8C6361B03FCC7F8E7FBA7F805" author="Ng, P. K. L. &amp; Rahayu, D. L. &amp; Naser, M. D." box="[876,1058,1984,2009]" pageId="48" pageNumber="49" pagination="1 - 26" refId="ref51169" refString="Ng, P. K. L., Rahayu, D. L. &amp; Naser, M. D. (2009) The Campatndriidae of Iraq, with description of a new genus and notes on Leptochryseus Al-khayat &amp; Jones, 1996 (Crustacea: Decapoda: Brachyura). Zootaxa, 2312, 1 - 26." type="journal article" year="2009">
Ng
<emphasis id="5BAA355BC6361B03FC3EF8E7FC42F805" box="[917,967,1984,2008]" italics="true" pageId="48" pageNumber="49">et al</emphasis>
. (2009)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>