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<document id="BE8892312F06BAF8567F88960F3ADAAB" ID-DOI="10.11646/zootaxa.3721.1.2" ID-GBIF-Dataset="d9159fd2-66d7-46dc-8ddd-0f9777c9fb1a" ID-ISSN="1175-5326" ID-Zenodo-Dep="285105" ID-ZooBank="935F1F1B-F09B-4BD0-A102-8FE8515CDC04" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460688313893" checkinUser="plazi" docAuthor="Jantarit, Sopark, Satasook, Chutamas &amp; Deharveng, Louis" docDate="2013" docId="E170A43E025DFFD3588D4876FC8AFD3C" docLanguage="en" docName="zt03721p070.pdf" docOrigin="Zootaxa 3721 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyphoderopsis thachana Jantarit, Satasook &amp; Deharveng, 2013, sp. nov." docType="treatment" docVersion="6" lastPageNumber="61" masterDocId="1D49DC460254FFDF581A4D4AFFAFFFF1" masterDocTitle="The genus Cyphoderopsis Carpenter (Collembola: Paronellidae) in Thailand and a faunal transition at the Isthmus of Kra in Troglopedetinae" masterLastPageNumber="70" masterPageNumber="49" pageNumber="58" updateTime="1698351439593" updateUser="plazi">
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<mods:title id="1090307490590F57863BA58A122221CE">The genus Cyphoderopsis Carpenter (Collembola: Paronellidae) in Thailand and a faunal transition at the Isthmus of Kra in Troglopedetinae</mods:title>
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<mods:namePart id="244AA75D0B6A0B8580AD6FF6F69F27AA">Jantarit, Sopark</mods:namePart>
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<mods:namePart id="D33441AB8309FF1B5B63CE1A54A6E28B">Satasook, Chutamas</mods:namePart>
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<mods:namePart id="C175F06B6C931B9511A55392AA70D56B">Deharveng, Louis</mods:namePart>
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<mods:title id="85143979B6FDF6F1E9B3617639311F63">Zootaxa</mods:title>
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<mods:date id="EDF5873415BA053D2ABA9CBAF2D1804F">2013</mods:date>
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<mods:number id="8342EBF7722CA9029CA660FA8815DC1B">3721</mods:number>
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<treatment id="E170A43E025DFFD3588D4876FC8AFD3C" ID-DOI="http://doi.org/10.5281/zenodo.6155353" ID-GBIF-Taxon="127666106" ID-Zenodo-Dep="6155353" LSID="urn:lsid:plazi:treatment:E170A43E025DFFD3588D4876FC8AFD3C" httpUri="http://treatment.plazi.org/id/E170A43E025DFFD3588D4876FC8AFD3C" lastPageId="12" lastPageNumber="61" pageId="9" pageNumber="58">
<subSubSection id="21C346A3025DFFD6588D4876FF44FA86" pageId="9" pageNumber="58" type="nomenclature">
<paragraph id="69661528025DFFD6588D4876FD89FAA7" blockId="9.[151,550,1340,1399]" box="[151,550,1340,1366]" pageId="9" pageNumber="58">
<heading id="322EA244025DFFD6588D4876FD89FAA7" bold="true" box="[151,550,1340,1366]" fontSize="11" level="1" pageId="9" pageNumber="58" reason="1">
<emphasis id="5BADC93A025DFFD6588D4876FD89FAA7" bold="true" box="[151,550,1340,1366]" pageId="9" pageNumber="58">
<taxonomicName id="AED96EAB025DFFD6588D4876FE6DFAA7" box="[151,450,1340,1366]" class="Entognatha" family="Paronellidae" genus="Cyphoderopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="9" pageNumber="58" phylum="Arthropoda" rank="species" species="thachana" status="sp. nov.">
<emphasis id="5BADC93A025DFFD6588D4876FE6DFAA7" bold="true" box="[151,450,1340,1366]" italics="true" pageId="9" pageNumber="58">Cyphoderopsis thachana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="409E7441025DFFD659D34876FD89FAA7" box="[457,550,1340,1366]" pageId="9" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="69661528025DFFD6588D4814FF44FA86" blockId="9.[151,550,1340,1399]" box="[151,235,1374,1399]" pageId="9" pageNumber="58">
(
<figureCitation id="F1E209AD025DFFD658854814FF4DFA86" box="[159,226,1374,1399]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[229,1357,256,1867]" captionTargetId="figure@10.[211,1376,173,1874]" captionTargetPageId="10" captionText="FIGURE 5. Cyphoderopsis thachana sp. nov. (A) Chaetotaxy of basal area of labium; (B) Outer maxillary lobe, right side; (C) Labrum, distal part in ventral view with two lateral combs and two central curved rods; (D) Different types of antennal chaetae (see text for chaeta type numbering); (E) Tergite chaetotaxy, left side (mes number not complete); (F) Two types of S-chaetae on Th. II; (G) Foot complex and distal part of tibiotarsus III; (H) Claw and distal part of tibiotarsus III; (I) Anterior side of ventral tube; (J) Dens and mucro; (K) Mucro." httpUri="https://zenodo.org/record/285110/files/figure.png" pageId="9" pageNumber="58">Fig. 5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="21C346A3025DFFD6588D48ECFB36F9BF" pageId="9" pageNumber="58" type="materials_examined">
<paragraph id="69661528025DFFD6588D48ECFCEBF9DB" blockId="9.[151,1437,1446,1975]" pageId="9" pageNumber="58">
<emphasis id="5BADC93A025DFFD6588D48ECFEECFA4E" bold="true" box="[151,323,1446,1471]" pageId="9" pageNumber="58">
<typeStatus id="B662AB8A025DFFD6588D48ECFF7DFA4E" box="[151,210,1446,1471]" pageId="9" pageNumber="58">Type</typeStatus>
material
</emphasis>
(point C
<quantity id="AE21B8CD025DFFD659AE48EDFE4EFA4E" box="[436,481,1446,1471]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="9" pageNumber="58" unit="in" value="1.0">1 in</quantity>
<figureCitation id="F1E209AD025DFFD659F748ECFD9DFA4E" box="[493,562,1446,1471]" captionStart="FIGURE 1" captionStartId="4.[151,250,1704,1726]" captionTargetBox="[348,1244,195,1687]" captionTargetId="figure@4.[348,1245,195,1687]" captionTargetPageId="4" captionText="FIGURE 1. Distribution of Paronellidae in sampled caves of Thailand. Black circles, caves with Cyphoderopsis. C 1, C. thachana sp. nov.; C 2, C. khaophang sp. nov. and C. cavicola sp. nov.; C 3, C. phangnga sp. nov.; C 4, Cyphoderopsis unidentified species (Tham Khao Noi, Phatthalung prov.); C 5, Cyphoderopsis unidentified species (Tham Peung, Surat Thani prov.). Half plain red circles, Troglopedetes records. T 1, T. maungonensis Deharveng &amp; Gers, 1993; T 2, T. maffrei Deharveng &amp; Gers, 1993; T 3, T. microps Deharveng &amp; Gers, 1993; T 4, T. longicornis Deharveng &amp; Gers, 1993; T 5, T. centralis Deharveng &amp; Gers, 1993; T 6, T. dispersus Deharveng &amp; Gers, 1993; T 7, T. multispinosus Deharveng &amp; Gers, 1993; T 8, T. convergens Deharveng &amp; Gers, 1993; T 9, T. paucisetosus Deharveng &amp; Gers, 1993; T 10, T. calvus Deharveng &amp; Gers, 1993; T 11, T. fredstonei Deharveng, 1988 and T. leclerci Deharveng, 1990; T 12, Troglopedetes unidentified species (Tham Phakok, Tak prov.); T 13, Troglopedetes unidentified species (Tham Mon, Kanchanaburi prov.); T 14, Troglopedetes unidentified species (Tham Erawan, Lamphun prov). Empty blue circles, caves where no species of Cyphoderopsis or Troglopedetes have been found." httpUri="https://zenodo.org/record/285106/files/figure.png" pageId="9" pageNumber="58">Fig. 1</figureCitation>
).
<typeStatus id="B662AB8A025DFFD65A4A48ECFD17FA4E" box="[592,696,1446,1471]" pageId="9" pageNumber="58" type="holotype">Holotype</typeStatus>
male and five
<typeStatus id="B662AB8A025DFFD65B6E48EDFC70FA4E" box="[884,991,1447,1471]" pageId="9" pageNumber="58" type="paratype">paratypes</typeStatus>
on slides (one female; four probably subadult or female adult, genital plate not seen clearly):
<collectingCountry id="11CE55B8025DFFD65B044881FC1CFA12" box="[798,947,1483,1507]" name="Thailand" pageId="9" pageNumber="58">THAILAND</collectingCountry>
:
<collectingCountry id="11CE55B8025DFFD65BA54881FC55FA12" box="[959,1018,1483,1507]" name="India" pageId="9" pageNumber="58">Surat</collectingCountry>
Thani Province: Tha Chana district: Tham Yai,
<date id="1D6733E8025DFFD6590648A4FE31F9F6" box="[284,414,1518,1543]" pageId="9" pageNumber="58" value="1987-07-13">13.vii.1987</date>
, in cave, Berlese extraction, L. Deharveng &amp; N. Giani leg. (sample # THA-SUT-002). Coordinates:
<geoCoordinate id="0CED73EF025DFFD659374B59FE6DF9DA" box="[301,450,1555,1579]" direction="east" orientation="longitude" pageId="9" pageNumber="58" precision="1" value="99.18912">99.189119°E</geoCoordinate>
,
<geoCoordinate id="0CED73EF025DFFD659D74B58FDF6F9DA" box="[461,601,1554,1579]" direction="north" orientation="latitude" pageId="9" pageNumber="58" precision="1" value="9.539795">9.539795°N</geoCoordinate>
, altitude about
<specimenCount id="7FDFDEA1025DFFD65B1E4B59FCEBF9DB" box="[772,836,1555,1579]" pageId="9" pageNumber="58" type="generic">
<quantity id="AE21B8CD025DFFD65B1E4B59FCEFF9DB" box="[772,832,1555,1579]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="58" unit="m" value="20.0">20 m</quantity>
.
</specimenCount>
</paragraph>
<paragraph id="69661528025DFFD658DD4B7CFB36F9BF" blockId="9.[151,1437,1446,1975]" box="[199,1177,1590,1615]" pageId="9" pageNumber="58">
<typeStatus id="B662AB8A025DFFD658DD4B7CFE80F9BE" box="[199,303,1590,1615]" pageId="9" pageNumber="58" type="holotype">Holotype</typeStatus>
and two
<typeStatus id="B662AB8A025DFFD6598C4B7DFDAEF9BE" box="[406,513,1591,1615]" pageId="9" pageNumber="58" type="paratype">paratypes</typeStatus>
in PSU; three
<typeStatus id="B662AB8A025DFFD65ABC4B7DFCBDF9BE" box="[678,786,1591,1615]" pageId="9" pageNumber="58" type="paratype">paratypes</typeStatus>
(including one female) in MNHN.
</paragraph>
</subSubSection>
<subSubSection id="21C346A3025DFFD358DD4B10FA88FE5D" lastPageId="12" lastPageNumber="61" pageId="9" pageNumber="58" type="description">
<paragraph id="69661528025DFFD658DD4B10FD50F92E" blockId="9.[151,1437,1446,1975]" pageId="9" pageNumber="58">
<emphasis id="5BADC93A025DFFD658DD4B10FEF6F982" bold="true" box="[199,345,1626,1651]" pageId="9" pageNumber="58">Description.</emphasis>
Habitus relatively stout. Body length
<quantity id="AE21B8CD025DFFD65B184B11FC26F983" box="[770,905,1627,1651]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" pageId="9" pageNumber="58" unit="mm" value="1.25" valueMax="1.4" valueMin="1.1">1.1–1.4 mm</quantity>
excluding antennae and furca. Lengths of body parts in
<tableCitation id="245B2093025DFFD658EE4B34FEE3F966" box="[244,332,1662,1687]" captionStart="TABLE 2" captionStartId="11.[151,239,152,175]" captionTargetBox="[159,1369,234,426]" captionTargetPageId="11" captionText="TABLE 2. Cyphoderopsis thachana sp. nov., measurements in µm from subadult paratype (# THA- SUT- 002 slide 01); body length includes head." httpUri="http://table.plazi.org/id/3DA645A0025FFFD4588D4DD2FE1AFF3E" pageId="9" pageNumber="58" tableUuid="3DA645A0025FFFD4588D4DD2FE1AFF3E">Table 2</tableCitation>
. Fourth abdominal segment 6–7 times as long as the third one along axis. Furca well developed, 1.8–2.0 times shorter than body length. Body colour white. Eyes absent, no ocular patch. Scales present on antennae, head, body and furca, absent on ventral tube.
</paragraph>
<paragraph id="69661528025DFFD658DD4BA0FA85F846" blockId="9.[151,1437,1446,1975]" pageId="9" pageNumber="58">
<emphasis id="5BADC93A025DFFD658DD4BA0FEF9F8F2" bold="true" box="[199,342,1770,1795]" pageId="9" pageNumber="58">Mouthparts</emphasis>
. Mandible head with 4 (left) and 5 (right) teeth. Maxilla head stocky, with tridentate claw and 6 ciliate lamellae. Labial basis chaetotaxy: M1M2ReL1l2, all mesochaetae, except l2 which is laterally displaced and mic-size, with R shorter than M2 and e smooth (
<figureCitation id="F1E209AD025DFFD65AAA4A78FD5DF8BA" box="[688,754,1842,1867]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[229,1357,256,1867]" captionTargetId="figure@10.[211,1376,173,1874]" captionTargetPageId="10" captionText="FIGURE 5. Cyphoderopsis thachana sp. nov. (A) Chaetotaxy of basal area of labium; (B) Outer maxillary lobe, right side; (C) Labrum, distal part in ventral view with two lateral combs and two central curved rods; (D) Different types of antennal chaetae (see text for chaeta type numbering); (E) Tergite chaetotaxy, left side (mes number not complete); (F) Two types of S-chaetae on Th. II; (G) Foot complex and distal part of tibiotarsus III; (H) Claw and distal part of tibiotarsus III; (I) Anterior side of ventral tube; (J) Dens and mucro; (K) Mucro." httpUri="https://zenodo.org/record/285110/files/figure.png" pageId="9" pageNumber="58">Fig. 5</figureCitation>
A). Outer maxillary lobe with one basal chaeta, a simple palp and two sublobal hairs (
<figureCitation id="F1E209AD025DFFD659B64A1CFE5DF89E" box="[428,498,1878,1903]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[229,1357,256,1867]" captionTargetId="figure@10.[211,1376,173,1874]" captionTargetPageId="10" captionText="FIGURE 5. Cyphoderopsis thachana sp. nov. (A) Chaetotaxy of basal area of labium; (B) Outer maxillary lobe, right side; (C) Labrum, distal part in ventral view with two lateral combs and two central curved rods; (D) Different types of antennal chaetae (see text for chaeta type numbering); (E) Tergite chaetotaxy, left side (mes number not complete); (F) Two types of S-chaetae on Th. II; (G) Foot complex and distal part of tibiotarsus III; (H) Claw and distal part of tibiotarsus III; (I) Anterior side of ventral tube; (J) Dens and mucro; (K) Mucro." httpUri="https://zenodo.org/record/285110/files/figure.png" pageId="9" pageNumber="58">Fig. 5</figureCitation>
B). Labral formula 4/554, with 4 prelabral chaetae bent and ciliate, labral chaetae smooth and acuminate; distal part of labrum adorned with two strong central pointed spines dorsally and a complex structure constituted of two asymmetrical combs and two small adjacent curved rods ventrally (
<figureCitation id="F1E209AD025DFFD65CDF4AD4FAA8F846" box="[1221,1287,1950,1975]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[229,1357,256,1867]" captionTargetId="figure@10.[211,1376,173,1874]" captionTargetPageId="10" captionText="FIGURE 5. Cyphoderopsis thachana sp. nov. (A) Chaetotaxy of basal area of labium; (B) Outer maxillary lobe, right side; (C) Labrum, distal part in ventral view with two lateral combs and two central curved rods; (D) Different types of antennal chaetae (see text for chaeta type numbering); (E) Tergite chaetotaxy, left side (mes number not complete); (F) Two types of S-chaetae on Th. II; (G) Foot complex and distal part of tibiotarsus III; (H) Claw and distal part of tibiotarsus III; (I) Anterior side of ventral tube; (J) Dens and mucro; (K) Mucro." httpUri="https://zenodo.org/record/285110/files/figure.png" pageId="9" pageNumber="58">Fig. 5</figureCitation>
C).
</paragraph>
<caption id="3DA645A0025EFFD5588D4A1FFDCAF817" httpUri="https://zenodo.org/record/285110/files/figure.png" pageId="10" pageNumber="59" targetBox="[229,1357,256,1867]" targetPageId="10">
<paragraph id="69661528025EFFD5588D4A1FFDCAF817" blockId="10.[151,1436,1877,2022]" pageId="10" pageNumber="59">
<emphasis id="5BADC93A025EFFD5588D4A1FFEA4F89A" bold="true" box="[151,267,1877,1899]" pageId="10" pageNumber="59">FIGURE 5</emphasis>
.
<taxonomicName id="AED96EAB025EFFD559024A1FFDBCF89A" box="[280,531,1877,1899]" class="Entognatha" family="Paronellidae" genus="Cyphoderopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="59" phylum="Arthropoda" rank="species" species="thachana" status="sp. nov.">
<emphasis id="5BADC93A025EFFD559024A1FFDBCF89A" box="[280,531,1877,1899]" italics="true" pageId="10" pageNumber="59">Cyphoderopsis thachana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="409E7441025EFFD55A004A1CFDC6F89A" box="[538,617,1878,1899]" pageId="10" pageNumber="59" rank="species">
<emphasis id="5BADC93A025EFFD55A004A1CFDC6F89A" bold="true" box="[538,617,1878,1899]" pageId="10" pageNumber="59">sp. nov.</emphasis>
</taxonomicNameLabel>
(A) Chaetotaxy of basal area of labium; (B) Outer maxillary lobe, right side; (C) Labrum, distal part in ventral view with two lateral combs and two central curved rods; (D) Different types of antennal chaetae (see text for chaeta type numbering); (E) Tergite chaetotaxy, left side (mes number not complete); (F) Two types of S-chaetae on Th.II; (G) Foot complex and distal part of tibiotarsus III; (H) Claw and distal part of tibiotarsus III; (I) Anterior side of ventral tube; (J) Dens and mucro; (K) Mucro.
</paragraph>
</caption>
<caption id="3DA645A0025FFFD4588D4DD2FE1AFF3E" ID-Table-UUID="3DA645A0025FFFD4588D4DD2FE1AFF3E" httpUri="http://table.plazi.org/id/3DA645A0025FFFD4588D4DD2FE1AFF3E" pageId="11" pageNumber="60" targetBox="[159,1369,234,426]" targetIsTable="true" targetPageId="11">
<paragraph id="69661528025FFFD4588D4DD2FE1AFF3E" blockId="11.[151,1436,151,207]" pageId="11" pageNumber="60">
<emphasis id="5BADC93A025FFFD4588D4DD2FDD3FF5E" bold="true" box="[151,636,151,175]" pageId="11" pageNumber="60">
TABLE 2.
<taxonomicName id="AED96EAB025FFFD4590B4DDDFD8EFF5E" box="[273,545,151,175]" class="Entognatha" family="Paronellidae" genus="Cyphoderopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="thachana" status="sp. nov.">
<emphasis id="5BADC93A025FFFD4590B4DDDFD8EFF5E" bold="true" box="[273,545,151,175]" italics="true" pageId="11" pageNumber="60">Cyphoderopsis thachana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="409E7441025FFFD45A3D4DD2FDD3FF5E" box="[551,636,152,175]" pageId="11" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, measurements in µm from subadult paratype (# THA- SUT-002 slide 01); body length includes head.
</paragraph>
</caption>
<paragraph id="69661528025FFFD458854DA0FA9FFE5B" pageId="11" pageNumber="60">
<table id="1BD9E788025F002058854DA0FAF6FE5B" box="[159,1369,234,426]" gridcols="9" gridrows="4" pageId="11" pageNumber="60">
<tr id="D7E9176A025F002058854DA0FAF6FEF1" box="[159,1369,234,256]" gridrow="0" pageId="11" pageNumber="60">
<th id="94387E16025F002058854DA0FF77FEF1" box="[159,216,234,256]" gridcol="0" gridrow="0" pageId="11" pageNumber="60">Body</th>
<th id="94387E16025F002059374DA0FEDFFEF1" box="[301,368,234,256]" gridcol="1" gridrow="0" pageId="11" pageNumber="60">Head</th>
<th id="94387E16025F002059A64DA0FDA8FEF1" box="[444,519,234,256]" gridcol="2" gridrow="0" pageId="11" pageNumber="60">Ant</th>
<th id="94387E16025F00205A514DA0FD37FEF1" box="[587,664,234,256]" gridcol="3" gridrow="0" pageId="11" pageNumber="60">Ant.I</th>
<th id="94387E16025F00205AC04DA0FCB0FEF1" box="[730,799,234,256]" gridcol="4" gridrow="0" pageId="11" pageNumber="60">Ant.II</th>
<th id="94387E16025F00205B734DA0FC1AFEF1" box="[873,949,234,256]" gridcol="5" gridrow="0" pageId="11" pageNumber="60">Ant.III</th>
<th id="94387E16025F00205BE24DA0FBEFFEF1" box="[1016,1088,234,256]" gridcol="6" gridrow="0" pageId="11" pageNumber="60">Ant.IV</th>
<th id="94387E16025F00205C9D4DA0FB14FEF1" box="[1159,1211,234,256]" gridcol="7" gridrow="0" pageId="11" pageNumber="60">Th.II</th>
<th id="94387E16025F00205D0C4DA0FAF6FEF1" box="[1302,1369,234,256]" gridcol="8" gridrow="0" pageId="11" pageNumber="60">Th.III</th>
</tr>
<tr id="D7E9176A025F002058854C5FFAF6FEDA" box="[159,1369,277,299]" gridrow="1" pageId="11" pageNumber="60">
<th id="94387E16025F002058854C5FFF77FEDA" box="[159,216,277,299]" gridcol="0" gridrow="1" pageId="11" pageNumber="60">1356</th>
<td id="94387E16025F002059374C5FFEDFFEDA" box="[301,368,277,299]" gridcol="1" gridrow="1" pageId="11" pageNumber="60">371</td>
<td id="94387E16025F002059A64C5FFDA8FEDA" box="[444,519,277,299]" gridcol="2" gridrow="1" pageId="11" pageNumber="60">536</td>
<td id="94387E16025F00205A514C5FFD37FEDA" box="[587,664,277,299]" gridcol="3" gridrow="1" pageId="11" pageNumber="60">57</td>
<td id="94387E16025F00205AC04C5FFCB0FEDA" box="[730,799,277,299]" gridcol="4" gridrow="1" pageId="11" pageNumber="60">143</td>
<td id="94387E16025F00205B734C5FFC1AFEDA" box="[873,949,277,299]" gridcol="5" gridrow="1" pageId="11" pageNumber="60">86</td>
<td id="94387E16025F00205BE24C5FFBEFFEDA" box="[1016,1088,277,299]" gridcol="6" gridrow="1" pageId="11" pageNumber="60">250</td>
<td id="94387E16025F00205C9D4C5FFB14FEDA" box="[1159,1211,277,299]" gridcol="7" gridrow="1" pageId="11" pageNumber="60">136</td>
<td id="94387E16025F00205D0C4C5FFAF6FEDA" box="[1302,1369,277,299]" gridcol="8" gridrow="1" pageId="11" pageNumber="60">100</td>
</tr>
<tr id="D7E9176A025F002058854C23FAF6FE8E" box="[159,1369,361,383]" gridrow="2" pageId="11" pageNumber="60">
<th id="94387E16025F002058854C23FF77FE8E" box="[159,216,361,383]" gridcol="0" gridrow="2" pageId="11" pageNumber="60">Abd.I</th>
<td id="94387E16025F002059374C23FEDFFE8E" box="[301,368,361,383]" gridcol="1" gridrow="2" pageId="11" pageNumber="60">Abd.II</td>
<td id="94387E16025F002059A64C23FDA8FE8E" box="[444,519,361,383]" gridcol="2" gridrow="2" pageId="11" pageNumber="60">Abd.III</td>
<td id="94387E16025F00205A514C23FD37FE8E" box="[587,664,361,383]" gridcol="3" gridrow="2" pageId="11" pageNumber="60">Abd.IV</td>
<td id="94387E16025F00205AC04C23FCB0FE8E" box="[730,799,361,383]" gridcol="4" gridrow="2" pageId="11" pageNumber="60">Abd.V</td>
<td id="94387E16025F00205B734C23FC1AFE8E" box="[873,949,361,383]" gridcol="5" gridrow="2" pageId="11" pageNumber="60">Abd.VI</td>
<td id="94387E16025F00205BE24C23FBEFFE8E" box="[1016,1088,361,383]" gridcol="6" gridrow="2" pageId="11" pageNumber="60">Man</td>
<td id="94387E16025F00205C9D4C23FB14FE8E" box="[1159,1211,361,383]" gridcol="7" gridrow="2" pageId="11" pageNumber="60">Dens</td>
<td id="94387E16025F00205D0C4C23FAF6FE8E" box="[1302,1369,361,383]" gridcol="8" gridrow="2" pageId="11" pageNumber="60">Mucro</td>
</tr>
<tr id="D7E9176A025F002058854CDEFAF6FE5B" box="[159,1369,404,426]" gridrow="3" pageId="11" pageNumber="60">
<th id="94387E16025F002058854CDEFF77FE5B" box="[159,216,404,426]" gridcol="0" gridrow="3" pageId="11" pageNumber="60">43</th>
<td id="94387E16025F002059374CDEFEDFFE5B" box="[301,368,404,426]" gridcol="1" gridrow="3" pageId="11" pageNumber="60">36</td>
<td id="94387E16025F002059A64CDEFDA8FE5B" box="[444,519,404,426]" gridcol="2" gridrow="3" pageId="11" pageNumber="60">86</td>
<td id="94387E16025F00205A514CDEFD37FE5B" box="[587,664,404,426]" gridcol="3" gridrow="3" pageId="11" pageNumber="60">457</td>
<td id="94387E16025F00205AC04CDEFCB0FE5B" box="[730,799,404,426]" gridcol="4" gridrow="3" pageId="11" pageNumber="60">79</td>
<td id="94387E16025F00205B734CDEFC1AFE5B" box="[873,949,404,426]" gridcol="5" gridrow="3" pageId="11" pageNumber="60">50</td>
<td id="94387E16025F00205BE24CDEFBEFFE5B" box="[1016,1088,404,426]" gridcol="6" gridrow="3" pageId="11" pageNumber="60">357</td>
<td id="94387E16025F00205C9D4CDEFB14FE5B" box="[1159,1211,404,426]" gridcol="7" gridrow="3" pageId="11" pageNumber="60">278</td>
<td id="94387E16025F00205D0C4CDEFAF6FE5B" box="[1302,1369,404,426]" gridcol="8" gridrow="3" pageId="11" pageNumber="60">57</td>
</tr>
</table>
</paragraph>
<paragraph id="69661528025FFFD458DD4CA0FECCFD46" blockId="11.[151,1436,490,2027]" pageId="11" pageNumber="60">
<emphasis id="5BADC93A025FFFD458DD4CA0FE95FDF2" bold="true" box="[199,314,490,515]" pageId="11" pageNumber="60">Antennae</emphasis>
. Antennae about 1.5 times as long as head. Scales present dorsally on Ant.I and II. Ordinary chaetae as ciliate mes, numerous on all antennal segment (
<typeStatus id="B662AB8A025FFFD45ADF4F45FCADFDD6" box="[709,770,527,551]" pageId="11" pageNumber="60">types</typeStatus>
1, 2, 4 on
<figureCitation id="F1E209AD025FFFD45B6D4F44FC17FDD6" box="[887,952,526,551]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[229,1357,256,1867]" captionTargetId="figure@10.[211,1376,173,1874]" captionTargetPageId="10" captionText="FIGURE 5. Cyphoderopsis thachana sp. nov. (A) Chaetotaxy of basal area of labium; (B) Outer maxillary lobe, right side; (C) Labrum, distal part in ventral view with two lateral combs and two central curved rods; (D) Different types of antennal chaetae (see text for chaeta type numbering); (E) Tergite chaetotaxy, left side (mes number not complete); (F) Two types of S-chaetae on Th. II; (G) Foot complex and distal part of tibiotarsus III; (H) Claw and distal part of tibiotarsus III; (I) Anterior side of ventral tube; (J) Dens and mucro; (K) Mucro." httpUri="https://zenodo.org/record/285110/files/figure.png" pageId="11" pageNumber="60">Fig. 5</figureCitation>
D), and a few smooth basal mic on Ant.I, II and III (
<typeStatus id="B662AB8A025FFFD458E94F79FE8AFDBA" box="[243,293,563,587]" pageId="11" pageNumber="60">type</typeStatus>
11 on
<figureCitation id="F1E209AD025FFFD4596F4F78FE18FDBA" box="[373,439,562,587]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[229,1357,256,1867]" captionTargetId="figure@10.[211,1376,173,1874]" captionTargetPageId="10" captionText="FIGURE 5. Cyphoderopsis thachana sp. nov. (A) Chaetotaxy of basal area of labium; (B) Outer maxillary lobe, right side; (C) Labrum, distal part in ventral view with two lateral combs and two central curved rods; (D) Different types of antennal chaetae (see text for chaeta type numbering); (E) Tergite chaetotaxy, left side (mes number not complete); (F) Two types of S-chaetae on Th. II; (G) Foot complex and distal part of tibiotarsus III; (H) Claw and distal part of tibiotarsus III; (I) Anterior side of ventral tube; (J) Dens and mucro; (K) Mucro." httpUri="https://zenodo.org/record/285110/files/figure.png" pageId="11" pageNumber="60">Fig. 5</figureCitation>
D); S-chaetae of various morphology, present on all antennal segments (
<typeStatus id="B662AB8A025FFFD45CF14F79FA88FDBA" box="[1259,1319,563,587]" pageId="11" pageNumber="60">types</typeStatus>
3, 7, 8, 9, 10, 12, 13 on
<figureCitation id="F1E209AD025FFFD4592D4F1CFED5FD9E" box="[311,378,598,623]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[229,1357,256,1867]" captionTargetId="figure@10.[211,1376,173,1874]" captionTargetPageId="10" captionText="FIGURE 5. Cyphoderopsis thachana sp. nov. (A) Chaetotaxy of basal area of labium; (B) Outer maxillary lobe, right side; (C) Labrum, distal part in ventral view with two lateral combs and two central curved rods; (D) Different types of antennal chaetae (see text for chaeta type numbering); (E) Tergite chaetotaxy, left side (mes number not complete); (F) Two types of S-chaetae on Th. II; (G) Foot complex and distal part of tibiotarsus III; (H) Claw and distal part of tibiotarsus III; (I) Anterior side of ventral tube; (J) Dens and mucro; (K) Mucro." httpUri="https://zenodo.org/record/285110/files/figure.png" pageId="11" pageNumber="60">Fig. 5</figureCitation>
D).
<typeStatus id="B662AB8A025FFFD459BC4F1DFE46FD9E" box="[422,489,599,623]" pageId="11" pageNumber="60">Types</typeStatus>
<date id="1D6733E8025FFFD459EB4F1CFDD0FD9E" box="[497,639,598,623]" pageId="11" pageNumber="60">5 and 6 may</date>
correspond to smooth and to very slightly serrated variants of a same
<typeStatus id="B662AB8A025FFFD4588D4F31FF68FD62" box="[151,199,635,659]" pageId="11" pageNumber="60">type</typeStatus>
of chaeta. Ant.IV not subdivided or annulated, apical bulb absent, subapical organ rather long, thick, slightly thickened distally.
</paragraph>
<paragraph id="69661528025FFFD458DD4F88FB5FFCD2" blockId="11.[151,1436,490,2027]" pageId="11" pageNumber="60">
<emphasis id="5BADC93A025FFFD458DD4F88FE34FD2A" bold="true" box="[199,411,706,731]" pageId="11" pageNumber="60">Dorsal chaetotaxy</emphasis>
. Dorsal chaetae of 6
<typeStatus id="B662AB8A025FFFD45A9D4F89FD69FD2A" box="[647,710,707,731]" pageId="11" pageNumber="60">types</typeStatus>
: ciliate, very long and thin tric; serrated or ciliate mac, of various length and thickness; smooth and rather thin mes; modified mes around tric (enlarged with special morphology); sens of 2
<typeStatus id="B662AB8A025FFFD4591B4E41FEEFFCD2" box="[257,320,779,803]" pageId="11" pageNumber="60">types</typeStatus>
: dark, short, straight, pointed (
<typeStatus id="B662AB8A025FFFD45A8F4E41FD68FCD2" box="[661,711,779,803]" pageId="11" pageNumber="60">type</typeStatus>
1) and longer, thinner, hyaline (
<typeStatus id="B662AB8A025FFFD45C344E41FBCFFCD2" box="[1070,1120,779,803]" pageId="11" pageNumber="60">type</typeStatus>
2) (
<figureCitation id="F1E209AD025FFFD45C974E40FB7FFCD2" box="[1165,1232,778,803]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[229,1357,256,1867]" captionTargetId="figure@10.[211,1376,173,1874]" captionTargetPageId="10" captionText="FIGURE 5. Cyphoderopsis thachana sp. nov. (A) Chaetotaxy of basal area of labium; (B) Outer maxillary lobe, right side; (C) Labrum, distal part in ventral view with two lateral combs and two central curved rods; (D) Different types of antennal chaetae (see text for chaeta type numbering); (E) Tergite chaetotaxy, left side (mes number not complete); (F) Two types of S-chaetae on Th. II; (G) Foot complex and distal part of tibiotarsus III; (H) Claw and distal part of tibiotarsus III; (I) Anterior side of ventral tube; (J) Dens and mucro; (K) Mucro." httpUri="https://zenodo.org/record/285110/files/figure.png" pageId="11" pageNumber="60">Fig. 5</figureCitation>
F).
</paragraph>
<paragraph id="69661528025FFFD458DD4E64FE9CFC7E" blockId="11.[151,1436,490,2027]" pageId="11" pageNumber="60">
Dorsal macrochaeta and pseudopore arrangement illustrated on
<figureCitation id="F1E209AD025FFFD45B834E64FC74FCB6" box="[921,987,814,839]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[229,1357,256,1867]" captionTargetId="figure@10.[211,1376,173,1874]" captionTargetPageId="10" captionText="FIGURE 5. Cyphoderopsis thachana sp. nov. (A) Chaetotaxy of basal area of labium; (B) Outer maxillary lobe, right side; (C) Labrum, distal part in ventral view with two lateral combs and two central curved rods; (D) Different types of antennal chaetae (see text for chaeta type numbering); (E) Tergite chaetotaxy, left side (mes number not complete); (F) Two types of S-chaetae on Th. II; (G) Foot complex and distal part of tibiotarsus III; (H) Claw and distal part of tibiotarsus III; (I) Anterior side of ventral tube; (J) Dens and mucro; (K) Mucro." httpUri="https://zenodo.org/record/285110/files/figure.png" pageId="11" pageNumber="60">Fig. 5</figureCitation>
E. Trichobothria formula 0, 0/0, 2, 3, 3. Macrochaeta formula 0/4, 0/0, 2, 4, 4 (chaeta posterior to T3 not developed in mac). Dorsal pseudopore formula 1, 1/1, 1, 1, 1+4.
</paragraph>
<paragraph id="69661528025FFFD458DD4ED0FDA6FA92" blockId="11.[151,1436,490,2027]" pageId="11" pageNumber="60">
Dorsal chaetotaxy of head similar to that of
<taxonomicName id="AED96EAB025FFFD45AA64ED1FCE1FC42" box="[700,846,922,947]" class="Entognatha" family="Paronellidae" genus="Cyphoderopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="phangnga" status="sp. nov.">
<emphasis id="5BADC93A025FFFD45AA64ED1FCE1FC42" box="[700,846,922,947]" italics="true" pageId="11" pageNumber="60">C. phangnga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="409E7441025FFFD45B4F4ED0FC1FFC42" box="[853,944,922,947]" pageId="11" pageNumber="60" rank="species">
<emphasis id="5BADC93A025FFFD45B4F4ED0FC1FFC42" bold="true" box="[853,944,922,947]" pageId="11" pageNumber="60">sp. nov.</emphasis>
</taxonomicNameLabel>
No dorsal mac on head except an antennobasal row of 6–8 mac on each side; cephalic mes short, feebly serrated, equal, symmetrically arranged. A pair of thin trichobothria-like chaetae present antero-laterally on head (as on
<figureCitation id="F1E209AD025FFFD45BB34EA8FC40FC0A" box="[937,1007,994,1019]" captionStart="FIGURE 3" captionStartId="7.[151,250,1889,1911]" captionTargetBox="[271,1329,238,1873]" captionTargetId="figure@7.[238,1354,193,1882]" captionTargetPageId="7" captionText="FIGURE 3. Cyphoderopsis phangnga sp. nov., continued, (A) Two types of S-chaetae on tergites; (B) Dorsal chaetotaxy of head; some fallen mes may have been unnoticed; (C) Antero-lateral trichobothrium-like chaeta on head (X); (D) Chaetotaxy of tergites, left side (M, macrochaeta; lateral group of macrochaetae encircled by dotted line; mes number not complete); (E) Trichobothrial complexes of Abd. II – III, left side; (F) Trichobothrial complex of Abd. IV (T 1 and T 2), left side. Filled black circles on Abd. II-Abd. IV trichobothrial areas: sockets of modified mesochaetae." httpUri="https://zenodo.org/record/285108/files/figure.png" pageId="11" pageNumber="60">Figs 3</figureCitation>
B, C). Suture zone visible on head (as on
<figureCitation id="F1E209AD025FFFD458A6494CFEAEFBEE" box="[188,257,1030,1055]" captionStart="FIGURE 3" captionStartId="7.[151,250,1889,1911]" captionTargetBox="[271,1329,238,1873]" captionTargetId="figure@7.[238,1354,193,1882]" captionTargetPageId="7" captionText="FIGURE 3. Cyphoderopsis phangnga sp. nov., continued, (A) Two types of S-chaetae on tergites; (B) Dorsal chaetotaxy of head; some fallen mes may have been unnoticed; (C) Antero-lateral trichobothrium-like chaeta on head (X); (D) Chaetotaxy of tergites, left side (M, macrochaeta; lateral group of macrochaetae encircled by dotted line; mes number not complete); (E) Trichobothrial complexes of Abd. II – III, left side; (F) Trichobothrial complex of Abd. IV (T 1 and T 2), left side. Filled black circles on Abd. II-Abd. IV trichobothrial areas: sockets of modified mesochaetae." httpUri="https://zenodo.org/record/285108/files/figure.png" pageId="11" pageNumber="60">Fig. 3</figureCitation>
B). Thin serrated mes present on all tergites in uncertain number. Th.II with 4+4 central mac and 2+2 antero-lateral sens (1+1 sens1 and 1+1 sens2) (
<figureCitation id="F1E209AD025FFFD45ABB4960FD4CFBB2" box="[673,739,1066,1091]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[229,1357,256,1867]" captionTargetId="figure@10.[211,1376,173,1874]" captionTargetPageId="10" captionText="FIGURE 5. Cyphoderopsis thachana sp. nov. (A) Chaetotaxy of basal area of labium; (B) Outer maxillary lobe, right side; (C) Labrum, distal part in ventral view with two lateral combs and two central curved rods; (D) Different types of antennal chaetae (see text for chaeta type numbering); (E) Tergite chaetotaxy, left side (mes number not complete); (F) Two types of S-chaetae on Th. II; (G) Foot complex and distal part of tibiotarsus III; (H) Claw and distal part of tibiotarsus III; (I) Anterior side of ventral tube; (J) Dens and mucro; (K) Mucro." httpUri="https://zenodo.org/record/285110/files/figure.png" pageId="11" pageNumber="60">Fig. 5</figureCitation>
F). Th.III without mac, sens not seen. Abd.I without mac, with 1+1 sens1. Abd.II with 2+2 tric, 1+1 sens2 near internal tric, 1+1 mac external to internal tric, 1+1 mac external to external tric, and several modified mes around tric (
<figureCitation id="F1E209AD025FFFD45AC24938FCB6FB7A" box="[728,793,1138,1163]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[229,1357,256,1867]" captionTargetId="figure@10.[211,1376,173,1874]" captionTargetPageId="10" captionText="FIGURE 5. Cyphoderopsis thachana sp. nov. (A) Chaetotaxy of basal area of labium; (B) Outer maxillary lobe, right side; (C) Labrum, distal part in ventral view with two lateral combs and two central curved rods; (D) Different types of antennal chaetae (see text for chaeta type numbering); (E) Tergite chaetotaxy, left side (mes number not complete); (F) Two types of S-chaetae on Th. II; (G) Foot complex and distal part of tibiotarsus III; (H) Claw and distal part of tibiotarsus III; (I) Anterior side of ventral tube; (J) Dens and mucro; (K) Mucro." httpUri="https://zenodo.org/record/285110/files/figure.png" pageId="11" pageNumber="60">Fig. 5</figureCitation>
E). Abd.III with 3+3 tric, 4+4 mac (1+1 near internal tric, 3+3 near postero-external tric), several modified mes around tric, and 2+2 sens located like in
<taxonomicName id="AED96EAB025FFFD45CC449DDFADEFB5E" box="[1246,1393,1174,1199]" class="Entognatha" family="Paronellidae" genus="Cyphoderopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="phangnga" status="sp. nov.">
<emphasis id="5BADC93A025FFFD45CC449DDFADEFB5E" box="[1246,1393,1174,1199]" italics="true" pageId="11" pageNumber="60">C. phangnga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="409E7441025FFFD45D6149DCFF67FB22" pageId="11" pageNumber="60" rank="species">
<emphasis id="5BADC93A025FFFD45D6149DCFF67FB22" bold="true" pageId="11" pageNumber="60">sp. nov.</emphasis>
</taxonomicNameLabel>
(1+1 sens2 anterior to internal tric and 1+1 sens1 posterior to external tric). Abd.IV with 3+3 tric (2+2 anterior, 1+1 postero-lateral), 4+4 mac (1+1 anterior to pseudopores, 1+1 near T2, 1+1 near T3, 1+1 posterior), and several mac external to the anterior trichobothrial complex T1-T2; several modified mes around T1 and T2; presence of short to medium-size chaetae, not easily assignable to mes or S-chaetae. Abd.V with 2+2 sens2. Abd.VI chaetotaxy not analyzed.
</paragraph>
<paragraph id="69661528025FFFD458DD4824FD19F9AE" blockId="11.[151,1436,490,2027]" pageId="11" pageNumber="60">
<emphasis id="5BADC93A025FFFD458DD4824FF50FA76" bold="true" box="[199,255,1390,1415]" pageId="11" pageNumber="60">Legs</emphasis>
. Tibiotarsus chaetotaxy mostly composed of large ciliate mesochaetae, finely ciliate chaetae internally, 2– 3 shorter dorsal ciliate mes (1/5 of the long ones), one thick smooth ventro-subapical chaeta on hind tita, a dorsoapical tenent hair rather thick, smooth and acuminate on all tita, and 2–3 smooth pointed dorsal mic, the distal one close to the tenent hair; distal row of 10 serrated chaetae arranged obliquely on all legs (
<figureCitation id="F1E209AD025FFFD45C9B4890FB67FA02" box="[1153,1224,1498,1523]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[229,1357,256,1867]" captionTargetId="figure@10.[211,1376,173,1874]" captionTargetPageId="10" captionText="FIGURE 5. Cyphoderopsis thachana sp. nov. (A) Chaetotaxy of basal area of labium; (B) Outer maxillary lobe, right side; (C) Labrum, distal part in ventral view with two lateral combs and two central curved rods; (D) Different types of antennal chaetae (see text for chaeta type numbering); (E) Tergite chaetotaxy, left side (mes number not complete); (F) Two types of S-chaetae on Th. II; (G) Foot complex and distal part of tibiotarsus III; (H) Claw and distal part of tibiotarsus III; (I) Anterior side of ventral tube; (J) Dens and mucro; (K) Mucro." httpUri="https://zenodo.org/record/285110/files/figure.png" pageId="11" pageNumber="60">Figs 5</figureCitation>
G, H). Claw small, 7.5–8.5 times shorter than tita; with 2 teeth at 55% and 75% of inner edge, a small dorsal tooth basally and a pair of inner basal teeth of unequal size (
<figureCitation id="F1E209AD025FFFD45A154B68FDFDF9CA" box="[527,594,1570,1595]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[229,1357,256,1867]" captionTargetId="figure@10.[211,1376,173,1874]" captionTargetPageId="10" captionText="FIGURE 5. Cyphoderopsis thachana sp. nov. (A) Chaetotaxy of basal area of labium; (B) Outer maxillary lobe, right side; (C) Labrum, distal part in ventral view with two lateral combs and two central curved rods; (D) Different types of antennal chaetae (see text for chaeta type numbering); (E) Tergite chaetotaxy, left side (mes number not complete); (F) Two types of S-chaetae on Th. II; (G) Foot complex and distal part of tibiotarsus III; (H) Claw and distal part of tibiotarsus III; (I) Anterior side of ventral tube; (J) Dens and mucro; (K) Mucro." httpUri="https://zenodo.org/record/285110/files/figure.png" pageId="11" pageNumber="60">Fig. 5</figureCitation>
H). Unguiculus pointed, narrow, lanceolate and elongate, more than half as long as claw, its external edge smooth (
<figureCitation id="F1E209AD025FFFD45A554B0CFD3DF9AE" box="[591,658,1606,1631]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[229,1357,256,1867]" captionTargetId="figure@10.[211,1376,173,1874]" captionTargetPageId="10" captionText="FIGURE 5. Cyphoderopsis thachana sp. nov. (A) Chaetotaxy of basal area of labium; (B) Outer maxillary lobe, right side; (C) Labrum, distal part in ventral view with two lateral combs and two central curved rods; (D) Different types of antennal chaetae (see text for chaeta type numbering); (E) Tergite chaetotaxy, left side (mes number not complete); (F) Two types of S-chaetae on Th. II; (G) Foot complex and distal part of tibiotarsus III; (H) Claw and distal part of tibiotarsus III; (I) Anterior side of ventral tube; (J) Dens and mucro; (K) Mucro." httpUri="https://zenodo.org/record/285110/files/figure.png" pageId="11" pageNumber="60">Fig. 5</figureCitation>
H).
</paragraph>
<paragraph id="69661528025FFFD458DD4B20FB5AF956" blockId="11.[151,1436,490,2027]" pageId="11" pageNumber="60">
<emphasis id="5BADC93A025FFFD458DD4B20FEF3F972" bold="true" box="[199,348,1642,1667]" pageId="11" pageNumber="60">Ventral tube</emphasis>
. With 4+4 long serrated mac anteriorly, at least 5 smooth mes on each lateral flap (
<figureCitation id="F1E209AD025FFFD45CE74B20FAEEF972" box="[1277,1345,1642,1667]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[229,1357,256,1867]" captionTargetId="figure@10.[211,1376,173,1874]" captionTargetPageId="10" captionText="FIGURE 5. Cyphoderopsis thachana sp. nov. (A) Chaetotaxy of basal area of labium; (B) Outer maxillary lobe, right side; (C) Labrum, distal part in ventral view with two lateral combs and two central curved rods; (D) Different types of antennal chaetae (see text for chaeta type numbering); (E) Tergite chaetotaxy, left side (mes number not complete); (F) Two types of S-chaetae on Th. II; (G) Foot complex and distal part of tibiotarsus III; (H) Claw and distal part of tibiotarsus III; (I) Anterior side of ventral tube; (J) Dens and mucro; (K) Mucro." httpUri="https://zenodo.org/record/285110/files/figure.png" pageId="11" pageNumber="60">
Fig.
<date id="1D6733E8025FFFD45D284B20FAEEF972" box="[1330,1345,1642,1667]" pageId="11" pageNumber="60">5</date>
</figureCitation>
I), and a combination of long and short ciliate mes posteriorly (exact number uncertain from our specimens).
</paragraph>
<paragraph id="69661528025FFFD358DD4BF8FB3EFF24" blockId="11.[151,1436,490,2027]" lastBlockId="12.[151,1436,151,717]" lastPageId="12" lastPageNumber="61" pageId="11" pageNumber="60">
<emphasis id="5BADC93A025FFFD458DD4BF8FEA1F93A" bold="true" box="[199,270,1714,1739]" pageId="11" pageNumber="60">Furca</emphasis>
. Tenaculum with 4 teeth on each branch and a strong, densely serrated, bent distally chaeta. Manubrium about as long as or slightly longer than mucrodens. Dens 4.5–5.0 times as long as mucro (
<figureCitation id="F1E209AD025FFFD45C664B9CFB10F91E" box="[1148,1215,1750,1775]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[229,1357,256,1867]" captionTargetId="figure@10.[211,1376,173,1874]" captionTargetPageId="10" captionText="FIGURE 5. Cyphoderopsis thachana sp. nov. (A) Chaetotaxy of basal area of labium; (B) Outer maxillary lobe, right side; (C) Labrum, distal part in ventral view with two lateral combs and two central curved rods; (D) Different types of antennal chaetae (see text for chaeta type numbering); (E) Tergite chaetotaxy, left side (mes number not complete); (F) Two types of S-chaetae on Th. II; (G) Foot complex and distal part of tibiotarsus III; (H) Claw and distal part of tibiotarsus III; (I) Anterior side of ventral tube; (J) Dens and mucro; (K) Mucro." httpUri="https://zenodo.org/record/285110/files/figure.png" pageId="11" pageNumber="60">Fig. 5</figureCitation>
J). Manubrium with dorsal ciliate mes (none smooth) arranged in two longitudinal stripes laterally separated by a glabrous axial stripe, distal area with 4+4 mes and 2+2 pseudopores; ventrally with oval scales and a distal group of 2–3+2–3 ciliate mes. Dens elongate, hairy, with 2 dorso-lateral rows of spines, sometimes asymmetrical; the external row with 14–15 thick serrated spiny chaetae, blunt apically; the internal row with 23 to 25 spines, subequal, pointed, smooth and shorter than the external ones in adult (
<figureCitation id="F1E209AD025FFFD45A614AC0FD6AF852" box="[635,709,1930,1955]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[229,1357,256,1867]" captionTargetId="figure@10.[211,1376,173,1874]" captionTargetPageId="10" captionText="FIGURE 5. Cyphoderopsis thachana sp. nov. (A) Chaetotaxy of basal area of labium; (B) Outer maxillary lobe, right side; (C) Labrum, distal part in ventral view with two lateral combs and two central curved rods; (D) Different types of antennal chaetae (see text for chaeta type numbering); (E) Tergite chaetotaxy, left side (mes number not complete); (F) Two types of S-chaetae on Th. II; (G) Foot complex and distal part of tibiotarsus III; (H) Claw and distal part of tibiotarsus III; (I) Anterior side of ventral tube; (J) Dens and mucro; (K) Mucro." httpUri="https://zenodo.org/record/285110/files/figure.png" pageId="11" pageNumber="60">Fig. 5</figureCitation>
J). Dorsally between the two spine rows, one (distally) to 3 (proximally) irregular lines of ciliate chaetae of various length. Scales on ventral and ventro-lateral sides of dentes, arranged in multiplets of 3–4 scales, more visible distally. Scales elongate, enlarged distally, of various morphology (see
<figureCitation id="F1E209AD0258FFD358D04DDDFEA3FF41" box="[202,268,151,176]" captionStart="FIGURE 4" captionStartId="8.[151,250,1918,1940]" captionTargetBox="[250,1345,289,1877]" captionTargetId="figure@8.[233,1354,193,1897]" captionTargetPageId="8" captionText="FIGURE 4. Cyphoderopsis phangnga sp. nov., continued, (A) Tibiotarsus II showing three smooth pointed dorsal mic and tenent hair (ordinary chaetae not drawn); (B) Foot complexand distal part of tibiotarsus III; (C) Ventral tube in antero-lateral view; (D) Posterior side of ventral tube; (E) Dens and mucro (subadult); (F) Different types of dental chaetae: ordinary chaetae (1 – 4) and spines (5 – 7); (G) Different types of scales on dens (1 – 3) and on ventral side of manubrium (4)." httpUri="https://zenodo.org/record/285109/files/figure.png" pageId="12" pageNumber="61">Fig. 4</figureCitation>
G). Mucro rather stout and rather long, straight, with 3 main teeth, the apical one blunt, the subapical one small and acute, the dorso-basal one acute and longer, with 1–4 toothlets basally (
<figureCitation id="F1E209AD0258FFD35C334DF6FBC3FF24" box="[1065,1132,188,213]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[229,1357,256,1867]" captionTargetId="figure@10.[211,1376,173,1874]" captionTargetPageId="10" captionText="FIGURE 5. Cyphoderopsis thachana sp. nov. (A) Chaetotaxy of basal area of labium; (B) Outer maxillary lobe, right side; (C) Labrum, distal part in ventral view with two lateral combs and two central curved rods; (D) Different types of antennal chaetae (see text for chaeta type numbering); (E) Tergite chaetotaxy, left side (mes number not complete); (F) Two types of S-chaetae on Th. II; (G) Foot complex and distal part of tibiotarsus III; (H) Claw and distal part of tibiotarsus III; (I) Anterior side of ventral tube; (J) Dens and mucro; (K) Mucro." httpUri="https://zenodo.org/record/285110/files/figure.png" pageId="12" pageNumber="61">Fig. 5</figureCitation>
K).
</paragraph>
<paragraph id="696615280258FFD358DD4D95FD6EFF09" blockId="12.[151,1436,151,717]" box="[199,705,223,248]" pageId="12" pageNumber="61">
<emphasis id="5BADC93A0258FFD358DD4D95FEFDFF09" bold="true" box="[199,338,223,248]" pageId="12" pageNumber="61">Occurrence</emphasis>
. Only known from
<typeStatus id="B662AB8A0258FFD35A2A4DAAFDCFFF09" box="[560,608,224,248]" pageId="12" pageNumber="61">type</typeStatus>
locality.
</paragraph>
<paragraph id="696615280258FFD358DD4C4EFA88FE5D" blockId="12.[151,1436,151,717]" pageId="12" pageNumber="61">
<emphasis id="5BADC93A0258FFD358DD4C4EFE8CFEEC" bold="true" box="[199,291,260,285]" pageId="12" pageNumber="61">Habitat</emphasis>
. Cave. The new species was formerly recorded as
<taxonomicName id="AED96EAB0258FFD35B754C4EFBA2FEEC" box="[879,1037,260,285]" class="Entognatha" family="Paronellidae" genus="Troglopedetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="5BADC93A0258FFD35B754C4EFBA2FEEC" box="[879,1037,260,285]" italics="true" pageId="12" pageNumber="61">Troglopedetes</emphasis>
</taxonomicName>
sp. from Tham Yai (Deharveng &amp; Bedos 1988). This small cave has
<quantity id="AE21B8CD0258FFD35A064C6DFDCCFECE" box="[540,611,295,320]" metricMagnitude="2" metricUnit="m" metricValue="1.96" pageId="12" pageNumber="61" unit="m" value="196.0">196 m</quantity>
of passages in a 6 km-long isolated limestone outcrop about
<quantity id="AE21B8CD0258FFD35D004C62FAFBFEB1" box="[1306,1364,295,320]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="61" unit="km" value="4.0">4 km</quantity>
SE of Tha Chana. The specimens were found in dark humid oligotrophic micro-habitats of the westernmost part of the cave, together with one specimen of another slightly troglomorphic species of
<taxonomicName id="AED96EAB0258FFD35C1C4C25FB02FE79" box="[1030,1197,367,392]" class="Entognatha" family="Paronellidae" genus="Cyphoderopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="5BADC93A0258FFD35C1C4C25FB02FE79" box="[1030,1197,367,392]" italics="true" pageId="12" pageNumber="61">Cyphoderopsis</emphasis>
</taxonomicName>
(thinner claw devoid of distal inner tooth). A species of
<taxonomicName id="AED96EAB0258FFD35A064CDEFC69FE5C" authority="Carl &amp; Lebedinsky 1905" authorityName="Carl &amp; Lebedinsky" authorityYear="1905" box="[540,966,404,429]" class="Entognatha" family="Oncopoduridae" genus="Oncopodura" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="5BADC93A0258FFD35A064CDEFD05FE5C" box="[540,682,404,429]" italics="true" pageId="12" pageNumber="61">Oncopodura</emphasis>
Carl &amp; Lebedinsky 1905
</taxonomicName>
was also collected in this cave.
</paragraph>
</subSubSection>
<subSubSection id="21C346A30258FFD358DD4CFDFCD8FE21" box="[199,887,439,464]" pageId="12" pageNumber="61" type="etymology">
<paragraph id="696615280258FFD358DD4CFDFCD8FE21" blockId="12.[151,1436,151,717]" box="[199,887,439,464]" pageId="12" pageNumber="61">
<emphasis id="5BADC93A0258FFD358DD4CFDFEE9FE21" bold="true" box="[199,326,439,464]" pageId="12" pageNumber="61">Etymology</emphasis>
. The species name is taken from the
<typeStatus id="B662AB8A0258FFD35AFC4CF2FCB9FE21" box="[742,790,440,464]" pageId="12" pageNumber="61">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="21C346A30258FFD358DD4C96FC8AFD3C" pageId="12" pageNumber="61" type="discussion">
<paragraph id="696615280258FFD358DD4C96FC8AFD3C" blockId="12.[151,1436,151,717]" pageId="12" pageNumber="61">
<emphasis id="5BADC93A0258FFD358DD4C96FE9BFE04" bold="true" box="[199,308,476,501]" pageId="12" pageNumber="61">Remarks</emphasis>
.
<taxonomicName id="AED96EAB0258FFD3595F4C96FDF0FE04" box="[325,607,476,501]" class="Entognatha" family="Paronellidae" genus="Cyphoderopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="thachana" status="sp. nov.">
<emphasis id="5BADC93A0258FFD3595F4C96FDF0FE04" box="[325,607,476,501]" italics="true" pageId="12" pageNumber="61">Cyphoderopsis thachana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="409E74410258FFD35A704C96FD67FE04" box="[618,712,476,501]" pageId="12" pageNumber="61" rank="species">
<emphasis id="5BADC93A0258FFD35A704C96FD67FE04" bold="true" box="[618,712,476,501]" pageId="12" pageNumber="61">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="AED96EAB0258FFD35B7D4C97FC53FE04" box="[871,1020,476,501]" class="Entognatha" family="Paronellidae" genus="Cyphoderopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="phangnga" status="sp. nov.">
<emphasis id="5BADC93A0258FFD35B7D4C97FC53FE04" box="[871,1020,476,501]" italics="true" pageId="12" pageNumber="61">C. phangnga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="409E74410258FFD35C1C4C96FBCBFE04" box="[1030,1124,476,501]" pageId="12" pageNumber="61" rank="species">
<emphasis id="5BADC93A0258FFD35C1C4C96FBCBFE04" bold="true" box="[1030,1124,476,501]" pageId="12" pageNumber="61">sp. nov.</emphasis>
</taxonomicNameLabel>
by the characters listed in Table 5; in particular, this new species usually has a longer body (1.1–1.4
<emphasis id="5BADC93A0258FFD35BFC4F4BFB81FDE9" box="[998,1070,513,536]" italics="true" pageId="12" pageNumber="61">versus</emphasis>
0.9–1.2) and two inner teeth on unguis (
<emphasis id="5BADC93A0258FFD358E94F6FFE95FDCD" box="[243,314,549,572]" italics="true" pageId="12" pageNumber="61">versus</emphasis>
one in
<taxonomicName id="AED96EAB0258FFD3598B4F6FFD8DFDCC" box="[401,546,548,573]" class="Entognatha" family="Paronellidae" genus="Cyphoderopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="phangnga" status="sp. nov.">
<emphasis id="5BADC93A0258FFD3598B4F6FFD8DFDCC" box="[401,546,548,573]" italics="true" pageId="12" pageNumber="61">C. phangnga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="409E74410258FFD35A304F6EFD2AFDCC" box="[554,645,548,573]" pageId="12" pageNumber="61" rank="species">
<emphasis id="5BADC93A0258FFD35A304F6EFD2AFDCC" bold="true" box="[554,645,548,573]" pageId="12" pageNumber="61">sp. nov.</emphasis>
</taxonomicNameLabel>
); tenent hairs of both species are acuminate but they are thicker in
<taxonomicName id="AED96EAB0258FFD35D984F6FFF50FD91" class="Entognatha" family="Paronellidae" genus="Cyphoderopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="thachana" status="sp. nov.">
<emphasis id="5BADC93A0258FFD35D984F6FFF50FD91" italics="true" pageId="12" pageNumber="61">C. thachana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="409E74410258FFD3591F4F0DFEF0FD91" box="[261,351,583,608]" pageId="12" pageNumber="61" rank="species">
<emphasis id="5BADC93A0258FFD3591F4F0DFEF0FD91" bold="true" box="[261,351,583,608]" pageId="12" pageNumber="61">sp. nov.</emphasis>
</taxonomicNameLabel>
; ventral tube has 4+4 anterior mac (
<emphasis id="5BADC93A0258FFD35AE94F03FC94FD91" box="[755,827,585,608]" italics="true" pageId="12" pageNumber="61">versus</emphasis>
3+
<quantity id="AE21B8CD0258FFD35B454F02FC26FD91" box="[863,905,583,608]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="12" pageNumber="61" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="AED96EAB0258FFD35B954F02FB8FFD91" box="[911,1056,583,608]" class="Entognatha" family="Paronellidae" genus="Cyphoderopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="phangnga" status="sp. nov.">
<emphasis id="5BADC93A0258FFD35B954F02FB8FFD91" box="[911,1056,583,608]" italics="true" pageId="12" pageNumber="61">C. phangnga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="409E74410258FFD35C3C4F0DFBD2FD91" box="[1062,1149,583,608]" pageId="12" pageNumber="61" rank="species">
<emphasis id="5BADC93A0258FFD35C3C4F0DFBD7FD91" bold="true" box="[1062,1144,583,608]" pageId="12" pageNumber="61">sp. nov</emphasis>
.
</taxonomicNameLabel>
); the ratio dens: mucro is higher than in
<taxonomicName id="AED96EAB0258FFD359224F27FE67FD74" box="[312,456,620,645]" class="Entognatha" family="Paronellidae" genus="Cyphoderopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="phangnga" status="sp. nov.">
<emphasis id="5BADC93A0258FFD359224F27FE67FD74" box="[312,456,620,645]" italics="true" pageId="12" pageNumber="61">C. phangnga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="409E74410258FFD359D44F26FD8BFD74" box="[462,548,620,645]" pageId="12" pageNumber="61" rank="species">
<emphasis id="5BADC93A0258FFD359D44F26FD8FFD74" bold="true" box="[462,544,620,645]" pageId="12" pageNumber="61">sp. nov</emphasis>
.
</taxonomicNameLabel>
(5
<emphasis id="5BADC93A0258FFD35A5F4F27FD23FD75" box="[581,652,621,644]" italics="true" pageId="12" pageNumber="61">versus</emphasis>
3–4) and there are more spines in the dorso-lateral rows of the dens in this new species (internal: 14–15
<emphasis id="5BADC93A0258FFD35A094FDBFDF4FD59" box="[531,603,657,680]" italics="true" pageId="12" pageNumber="61">versus</emphasis>
10–12; external: 23–25
<emphasis id="5BADC93A0258FFD35B754FDBFC19FD59" box="[879,950,657,680]" italics="true" pageId="12" pageNumber="61">versus</emphasis>
19). It should be stressed however that we lack information on variability in number of dental spines.
</paragraph>
</subSubSection>
</treatment>
</document>
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