192 lines
22 KiB
XML
192 lines
22 KiB
XML
<document id="01DF368923BDC9953468712890F9FF21" ID-DOI="10.11646/phytotaxa.263.2.9" ID-ISSN="1179-3163" ID-Zenodo-Dep="13671037" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725398873111" checkinUser="felipe" docAuthor="Bartolucci, Fabrizio & Mráz, Patrik" docDate="2016" docId="E14A87C94E453038FEF526D84AAB1AEE" docLanguage="en" docName="phytotaxa.263.2.9.pdf" docOrigin="Phytotaxa 263 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.263.2.9" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Thymus thracicus Velenovsky 1892" docType="treatment" docVersion="2" lastPageNumber="163" masterDocId="1D73FFB14E47303AFE7D2637482C1E54" masterDocTitle="A new combination in the genus Thymus and clarification on the type material of Th. thracicus (Lamiaceae)" masterLastPageNumber="164" masterPageNumber="161" pageNumber="163" updateTime="1725420602202" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F0DED0F006B0FCADF56FDB66742626BA">A new combination in the genus Thymus and clarification on the type material of Th. thracicus (Lamiaceae)</mods:title>
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<mods:namePart id="96D74530A13A0F7A14AA65B902F043D4">Bartolucci, Fabrizio</mods:namePart>
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<mods:namePart id="7430398614A51DA3ABD596E6667F3E1A">Mráz, Patrik</mods:namePart>
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<mods:affiliation id="EE2E238CADEAC8BA2688868E034D03E6">Herbarium and Department of Botany, Charles University in Prague, Benátská 2, 12801 Prague, Czech Republic;</mods:affiliation>
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<mods:title id="10D2903F787395AC453C05E42A9BA9A2">Phytotaxa</mods:title>
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<mods:part id="740E2A7C6F575F5FED4C1CBCB4B92D1F">
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<mods:date id="06E70D03E4F56D249B2D6D30FDAF05CB">2016</mods:date>
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<mods:number id="A73FDF712A32B34009598C416EB16C1A">2016-06-01</mods:number>
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<mods:number id="42A8A2B4064818CA18E4255894D7D067">263</mods:number>
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<subSubSection id="21F965544E453038FEF526D84A721F5C" box="[136,606,239,264]" pageId="2" pageNumber="163" type="nomenclature">
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<paragraph id="695C36DF4E453038FEF526D84A721F5C" blockId="2.[136,606,239,264]" box="[136,606,239,264]" pageId="2" pageNumber="163">
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<heading id="321481B34E453038FEF526D84A721F5C" box="[136,606,239,264]" fontSize="11" level="2" pageId="2" pageNumber="163" reason="3">
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<treatmentCitationGroup id="49F311F14E453038FEF526D84A721F5C" box="[136,606,239,264]" pageId="2" pageNumber="163">
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<treatmentCitation id="E84210CE4E453038FEF526D84A721F5C" author="Velenovsky, J." box="[136,606,239,264]" page=")" pageId="2" pageNumber="163" year="1892">
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<taxonomicName id="AEE34D5C4E453038FEF526D84A721F5C" authority="Velenovsky (1892: 16)" authorityName="Velenovsky" authorityPageNumber="16" authorityYear="1892" box="[136,606,239,264]" class="Insecta" family="Eulophidae" genus="Thymus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="thracicus">
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<emphasis id="5B97EACD4E453038FEF526D8497B1F5C" bold="true" box="[136,343,239,264]" italics="true" pageId="2" pageNumber="163">Thymus thracicus</emphasis>
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<bibRefCitation id="0D724B2E4E453038FF2326D84A721F5C" author="Velenovsky, J." box="[350,606,239,264]" pageId="2" pageNumber="163" pagination="14 - 17" refId="ref4102" refString="Velenovsky, J. (1892) Nachtrage zur Flora bulgarica. Oesterreichische Botanische Zeitschrift 42: 14 - 17. http: // dx. doi. org / 10.1007 / BF 01680639" type="journal article" year="1892">Velenovský (1892: 16)</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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||
</treatmentCitationGroup>
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||
</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="21F965544E453038FEF527224AAB1AEE" pageId="2" pageNumber="163" type="materials_examined">
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<paragraph id="695C36DF4E453038FEF5272249891F26" blockId="2.[136,1452,277,370]" pageId="2" pageNumber="163">
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<emphasis id="5B97EACD4E453038FEF5272248961F7E" bold="true" box="[136,186,277,298]" pageId="2" pageNumber="163">Type</emphasis>
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(
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<typeStatus id="B658887D4E453038FEBA2722490D1F7E" box="[199,289,277,298]" pageId="2" pageNumber="163" type="lectotype">lectotype</typeStatus>
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designated by
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<bibRefCitation id="0D724B2E4E453038FFCE27224A7B1F7E" author="Jalas, J." box="[435,599,277,298]" pageId="2" pageNumber="163" pagination="247 - 275" refId="ref3441" refString="Jalas, J. (1971 b) Notes on Thymus L. (Labiatae) in Europe. II. Comments on species and subspecies. Botanical Journal of the Linnean Society 64: 247 - 275." type="journal article" year="1971">Jalas 1971b: 265</bibRefCitation>
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, its herbarium placement is corrected here):—
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<materialsCitation id="D98B3C824E453038FA6B2722498E1F26" collectingDate="1891-06" collectionCode="V" collectorName="W No." country="Bulgaria" location="Ad Stanimaka" pageId="2" pageNumber="163" specimenCount="1" typeStatus="lectotype">
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[
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<collectingCountry id="11F4764F4E453038FA6127224C8C1F7E" box="[1052,1184,277,298]" name="Bulgaria" pageId="2" pageNumber="163">BULGARIA</collectingCountry>
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].
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<location id="6C3C60044E453038FAD327224D151F7E" LSID="urn:lsid:plazi:treatment:E14A87C94E453038FEF526D84AAB1AEE:6C3C60044E453038FAD327224D151F7E" box="[1198,1337,277,298]" country="Bulgaria" name="Ad Stanimaka" pageId="2" pageNumber="163">Ad Stanimaka</location>
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,
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<date id="1D5D101F4E453038FB3E27224D8B1F7E" box="[1347,1447,277,298]" pageId="2" pageNumber="163" value="1891-06">
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<collectingDate id="0D19E9F74E453038FB3E27224D8B1F7E" box="[1347,1447,277,298]" pageId="2" pageNumber="163" value="1891-06">June 1891</collectingDate>
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</date>
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,
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<emphasis id="5B97EACD4E453038FEC0270E49601F1A" box="[189,332,313,334]" italics="true" pageId="2" pageNumber="163">
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Stříbrný
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<collectionCode id="0FF2AE1A4E453038FF6F270E490D1F1A" box="[274,289,313,334]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="163" type="Museum">V</collectionCode>
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. s.n.
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</emphasis>
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(
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<collectorName id="C41653094E453038FF25270E49B61F1A" box="[344,410,313,334]" pageId="2" pageNumber="163">W No.</collectorName>
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1991-0001745 [digital image!]; image of the
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<typeStatus id="B658887D4E453038FD2E270E4B871F1A" box="[851,939,313,334]" pageId="2" pageNumber="163" type="lectotype">lectotype</typeStatus>
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is available at http://herbarium.univie.ac.at/database/ detail.php?ID=159084)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="695C36DF4E453038FEF527AB4AAB1AEE" blockId="2.[136,1452,412,1210]" pageId="2" pageNumber="163">
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<emphasis id="5B97EACD4E453038FEF527AB48911FE7" bold="true" box="[136,189,412,435]" pageId="2" pageNumber="163">Note</emphasis>
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:—The name
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<taxonomicName id="AEE34D5C4E453038FF3227AB4A7A1FE7" authority="Velen." authorityName="Velenovsky" authorityYear="1892" box="[335,598,412,435]" class="Insecta" family="Eulophidae" genus="Thymus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="thracicus">
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<emphasis id="5B97EACD4E453038FF3227AB4A201FE7" box="[335,524,412,435]" italics="true" pageId="2" pageNumber="163">Thymus thracicus</emphasis>
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Velen.
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</taxonomicName>
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was lectotypified by
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<bibRefCitation id="0D724B2E4E453038FD4427AB4BE41FE7" author="Jalas, J." box="[825,968,412,435]" pageId="2" pageNumber="163" pagination="247 - 275" refId="ref3441" refString="Jalas, J. (1971 b) Notes on Thymus L. (Labiatae) in Europe. II. Comments on species and subspecies. Botanical Journal of the Linnean Society 64: 247 - 275." type="journal article" year="1971">Jalas (1971b)</bibRefCitation>
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and as a type he choose the above mentioned specimen collected by
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<collectionCode id="0FF2AE1A4E453038FF06278849A21F82" box="[379,398,447,470]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="163" type="Museum">V</collectionCode>
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. Stříbrný and determined / labeled by J. Velenovský. As a herbarium where the
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<typeStatus id="B658887D4E453038FAA327884D6C1F82" box="[1246,1344,447,470]" pageId="2" pageNumber="163" type="lectotype">lectotype</typeStatus>
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specimen had been kept, Jalas cited the herbarium of the Charles University in Prague (
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<collectingCountry id="11F4764F4E453038FDAA27D44C271FAE" box="[983,1035,483,506]" name="China" pageId="2" pageNumber="163">PRC</collectingCountry>
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)—the institution where J. Velenovský was Professor of Botany and where his herbarium has always been preserved. However, when searching for this specimen in
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<collectingCountry id="11F4764F4E453038FEDB241E48F41C14" box="[166,216,553,576]" name="China" pageId="2" pageNumber="163">PRC</collectingCountry>
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for the purposes of this contribution, we were not able to find it. Surprisingly, the specimen was retraced in the Herbarium of Natural History Museum in Wien (W). The specimen under the accession number W1991-0001745 perfectly fits the protologue data, the indications by
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<bibRefCitation id="0D724B2E4E453038FC3224584ACC1CD2" author="Jalas, J." box="[591,736,623,646]" pageId="2" pageNumber="163" pagination="247 - 275" refId="ref3441" refString="Jalas, J. (1971 b) Notes on Thymus L. (Labiatae) in Europe. II. Comments on species and subspecies. Botanical Journal of the Linnean Society 64: 247 - 275." type="journal article" year="1971">Jalas (1971b)</bibRefCitation>
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, and his revision label bearing his
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<typeStatus id="B658887D4E453038FA2D24584C9E1CD2" box="[1104,1202,623,646]" pageId="2" pageNumber="163" type="lectotype">lectotype</typeStatus>
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annotation from 1969. The explanation why the
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<typeStatus id="B658887D4E453038FFDE24A44A291CFE" box="[419,517,659,682]" pageId="2" pageNumber="163" type="lectotype">lectotype</typeStatus>
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specimen is no more in
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<collectingCountry id="11F4764F4E453038FD6124A44B621CFE" box="[796,846,659,682]" name="China" pageId="2" pageNumber="163">PRC</collectingCountry>
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but in W is as follows. In the fall 1967 a huge parcel consisting of 1976 specimens of the genus
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<taxonomicName id="AEE34D5C4E453038FC2424814A801C99" authorityName="Linnaeus" authorityYear="1753" box="[601,684,694,717]" class="Insecta" family="Eulophidae" genus="Thymus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="163" phylum="Arthropoda" rank="genus">
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<emphasis id="5B97EACD4E453038FC2424814A801C99" box="[601,684,694,717]" italics="true" pageId="2" pageNumber="163">Thymus</emphasis>
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</taxonomicName>
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from
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<collectingCountry id="11F4764F4E453038FC9224814B0D1C99" box="[751,801,694,717]" name="China" pageId="2" pageNumber="163">PRC</collectingCountry>
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was sent on loan to J. Jalas (Herbarium at the University of Helsinky, H). In the 70s of the last century the loan returned to Prague, but apparently not complete (unfortunately, there is no receipt documentation in
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<collectingCountry id="11F4764F4E453038FFC624CB49DE1D47" box="[443,498,764,787]" name="China" pageId="2" pageNumber="163">PRC</collectingCountry>
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). Importantly, in fairly the same periode (1971–1993) a huge amount of
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<taxonomicName id="AEE34D5C4E453038FA8724CB4D611D47" authorityName="Linnaeus" authorityYear="1753" box="[1274,1357,764,787]" class="Insecta" family="Eulophidae" genus="Thymus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="163" phylum="Arthropoda" rank="genus">
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<emphasis id="5B97EACD4E453038FA8724CB4D611D47" box="[1274,1357,764,787]" italics="true" pageId="2" pageNumber="163">Thymus</emphasis>
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</taxonomicName>
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material from W was on loan by J. Jalas in Helsinki for his treatment of Flora Iranica. Unfortunately, as in the case of the loan from PR, the checking of the received material in W did not work well. Thus, the number of missing / supernumerary specimens was completely unclear (E. Vitek, in e-mail). Therefore, we suppose that the most plausible explanation of the misplacement of the
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<typeStatus id="B658887D4E453038FEB225BE491D1DF4" box="[207,305,905,928]" pageId="2" pageNumber="163" type="lectotype">lectotype</typeStatus>
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of
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<taxonomicName id="AEE34D5C4E453038FF2A25BE49C91DF4" authorityName="Velenovsky" authorityYear="1892" box="[343,485,905,928]" class="Insecta" family="Eulophidae" genus="Thymus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="thracicus">
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<emphasis id="5B97EACD4E453038FF2A25BE49C91DF4" box="[343,485,905,928]" italics="true" pageId="2" pageNumber="163">Th. thracicus</emphasis>
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</taxonomicName>
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is that this specimen was accidently sent from H to W instead of
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<collectingCountry id="11F4764F4E453038FAC825BE4CCB1DF4" box="[1205,1255,905,928]" name="China" pageId="2" pageNumber="163">PRC</collectingCountry>
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and the specimen was subsequently incorporated into W. This explanation is furthermore supported by the erroneous shipment of several specimens of
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<taxonomicName id="AEE34D5C4E453038FF6225F8495E1DB2" authorityName="Linnaeus" authorityYear="1753" box="[287,370,975,998]" class="Insecta" family="Eulophidae" genus="Thymus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="163" phylum="Arthropoda" rank="genus">
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<emphasis id="5B97EACD4E453038FF6225F8495E1DB2" box="[287,370,975,998]" italics="true" pageId="2" pageNumber="163">Thymus</emphasis>
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</taxonomicName>
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(the material of Rechinger fil.) from H to
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<collectingCountry id="11F4764F4E453038FD3A25F84B501DB2" box="[839,892,975,998]" name="China" pageId="2" pageNumber="163">PRC</collectingCountry>
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, though the loan was originally from G. Moreover, all original material to the
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<taxonomicName id="AEE34D5C4E453038FFD025C44A2C1A5E" authorityName="Linnaeus" authorityYear="1753" box="[429,512,1011,1034]" class="Insecta" family="Eulophidae" genus="Thymus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="163" phylum="Arthropoda" rank="genus">
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<emphasis id="5B97EACD4E453038FFD025C44A2C1A5E" box="[429,512,1011,1034]" italics="true" pageId="2" pageNumber="163">Thymus</emphasis>
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</taxonomicName>
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taxa described by J. Velenovský could be find in
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<collectingCountry id="11F4764F4E453038FA5F25C44C781A5E" box="[1058,1108,1011,1034]" name="China" pageId="2" pageNumber="163">PRC</collectingCountry>
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(
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<bibRefCitation id="0D724B2E4E453038FA1825C44D2B1A5E" author="Stepanek, J." box="[1125,1287,1010,1034]" pageId="2" pageNumber="163" refId="ref4032" refString="Stepanek, J. (2006 - 2010) Prehled balkanskych taxonu kapradorostu a semennych rostlin popsanych prof. J. Velenovskym. msc., depon. in the Herbarium of the Charles University in Prague." type="book" year="2006">Štěpánek 2006</bibRefCitation>
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–2010), but the specimens to three names—
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<taxonomicName id="AEE34D5C4E453038FFCE22214A131A79" authorityName="Velenovsky" authorityYear="1892" box="[435,575,1046,1069]" class="Insecta" family="Eulophidae" genus="Thymus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="thracicus">
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<emphasis id="5B97EACD4E453038FFCE22214A131A79" box="[435,575,1046,1069]" italics="true" pageId="2" pageNumber="163">Th. thracicus</emphasis>
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</taxonomicName>
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,
|
||
<taxonomicName id="AEE34D5C4E453038FC3122214BE91A79" authority="Velenovsky (1902: 53)" authorityName="Velenovsky" authorityPageNumber="53" authorityYear="1902" box="[588,965,1046,1069]" class="Insecta" family="Eulophidae" genus="Thymus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="skorpilii">
|
||
<emphasis id="5B97EACD4E453038FC3122214AE21A79" box="[588,718,1046,1069]" italics="true" pageId="2" pageNumber="163">Th. skorpilii</emphasis>
|
||
<bibRefCitation id="0D724B2E4E453038FCAE22214BE91A79" author="Velenovsky, J." box="[723,965,1046,1069]" pageId="2" pageNumber="163" pagination="49 - 55" refId="ref4136" refString="Velenovsky, J. (1902) Neunter Nachtrag zur Flora von Bulgarien. Oesterreichische Botanische Zeitschrift 52: 49 - 55. http: // dx. doi. org / 10.1007 / BF 01673075" type="journal article" year="1902">Velenovský (1902: 53)</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="AEE34D5C4E453038FD85222149A61A04" authority="Velenovsky (1906: 286)" authorityName="Velenovsky" authorityPageNumber="286" authorityYear="1906" class="Insecta" family="Eulophidae" genus="Thymus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="163" phylum="Arthropoda" rank="variety" species="striatus" variety="pindicus">
|
||
<emphasis id="5B97EACD4E453038FD8522214C5D1A79" box="[1016,1137,1046,1069]" italics="true" pageId="2" pageNumber="163">Th. striatus</emphasis>
|
||
<bibRefCitation id="0D724B2E4E453038FA0A22214D321A79" author="Vahl, M." box="[1143,1310,1046,1069]" pageId="2" pageNumber="163" pagination="1 - 104" refId="ref4070" refString="Vahl, M. (1794) Symbolae Botanicae 3. Excudebant Nicolaus Moller et filius, Aulae Regiae Typographi, Hauniae [Copenhagen], pp. 1 - 104." type="book chapter" year="1794">Vahl (1794: 78)</bibRefCitation>
|
||
var.
|
||
<emphasis id="5B97EACD4E453038FB2C22214D871A79" box="[1361,1451,1046,1069]" italics="true" pageId="2" pageNumber="163">pindicus</emphasis>
|
||
<bibRefCitation id="0D724B2E4E453038FEF5220E49A61A04" author="Velenovsky, J." box="[136,394,1081,1104]" pageId="2" pageNumber="163" pagination="271 - 287" refId="ref4172" refString="Velenovsky, J. (1906) Vorstudien zu einer Monographie der Gattung Thymus L. Beihefte zum Botanischen Centralblatt, Abteilung 2: 271 - 287." type="journal article" year="1906">Velenovský (1906: 286)</bibRefCitation>
|
||
</taxonomicName>
|
||
. In addition to the
|
||
<typeStatus id="B658887D4E453038FC31220E4A821A04" box="[588,686,1081,1104]" pageId="2" pageNumber="163" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="AEE34D5C4E453038FCB2220E4B771A04" authorityName="Velenovsky" authorityYear="1892" box="[719,859,1081,1104]" class="Insecta" family="Eulophidae" genus="Thymus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="thracicus">
|
||
<emphasis id="5B97EACD4E453038FCB2220E4B771A04" box="[719,859,1081,1104]" italics="true" pageId="2" pageNumber="163">Th. thracicus</emphasis>
|
||
</taxonomicName>
|
||
, also the
|
||
<typeStatus id="B658887D4E453038FDC3220E4C0C1A04" box="[958,1056,1081,1104]" pageId="2" pageNumber="163" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="AEE34D5C4E453038FA3C220E4CEE1A04" authorityName="Velenovsky" authorityYear="1902" box="[1089,1218,1081,1104]" class="Insecta" family="Eulophidae" genus="Thymus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="skorpilii">
|
||
<emphasis id="5B97EACD4E453038FA3C220E4CEE1A04" box="[1089,1218,1081,1104]" italics="true" pageId="2" pageNumber="163">Th. skorpilii</emphasis>
|
||
</taxonomicName>
|
||
is currently kept in W (W1991-0001748, digital image seen at http://jacq.nhm-wien.ac.at/djatoka/jacq-viewer/viewer.html?rft_id=w_1991000174 8&identifiers=w_19910001748). Based on these facts the herbarium citation of the
|
||
<taxonomicName id="AEE34D5C4E453038FD8422484CA81AC2" authorityName="Velenovsky" authorityYear="1892" box="[1017,1156,1151,1174]" class="Insecta" family="Eulophidae" genus="Thymus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="thracicus">
|
||
<emphasis id="5B97EACD4E453038FD8422484CA81AC2" box="[1017,1156,1151,1174]" italics="true" pageId="2" pageNumber="163">Th. thracicus</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="B658887D4E453038FAF722484CC01AC2" box="[1162,1260,1151,1174]" pageId="2" pageNumber="163" type="lectotype">lectotype</typeStatus>
|
||
is to be corrected, and now it is housed in W (No. 1991-0001745).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |