283 lines
41 KiB
XML
283 lines
41 KiB
XML
<document id="AF27FF8DFFE9A6ACD4325F05977C59CD" ID-CLB-Dataset="105312" ID-DOI="10.11646/zootaxa.5168.3.2" ID-GBIF-Dataset="53574cb3-f6fb-436c-bf7e-8cc59f163982" ID-ISSN="1175-5326" ID-Zenodo-Dep="6882742" ID-ZooBank="8B7C6BF7-0B3D-420F-BDAF-EB4F15E211AC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658485582440" checkinUser="plazi" docAuthor="Kataev, Boris M." docDate="2022" docId="E04B4F00FFE2FFC3FF4FF9C77F4F576A" docLanguage="en" docName="zootaxa.5168.3.2.pdf" docOrigin="Zootaxa 5168 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coleolissus perlucens" docType="treatment" docVersion="7" lastPageNumber="289" masterDocId="1C723778FFE3FFC7FFD8FFA37C5B5029" masterDocTitle="Notes on carabid beetles of the genera Coleolissus and Siopelus (Coleoptera: Carabidae: Harpalini), with description of four new species with unusual aedeagi from India and China" masterLastPageNumber="305" masterPageNumber="285" pageNumber="286" updateTime="1699361575247" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="78C8B6FFD29A4723FDF24EE1AB6C1D79">Notes on carabid beetles of the genera Coleolissus and Siopelus (Coleoptera: Carabidae: Harpalini), with description of four new species with unusual aedeagi from India and China</mods:title>
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<mods:namePart id="03D42CD4D3830BDE0256FC87FA1F344C">Kataev, Boris M.</mods:namePart>
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<mods:title id="6239B077AF424D83E1925F3A4222FC35">Zootaxa</mods:title>
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<mods:date id="FB8EDE8A7EE4F80C38F2ABE071F42CF3">2022</mods:date>
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<treatment id="E04B4F00FFE2FFC3FF4FF9C77F4F576A" ID-DOI="http://doi.org/10.5281/zenodo.6882730" ID-GBIF-Taxon="197852161" ID-Zenodo-Dep="6882730" LSID="urn:lsid:plazi:treatment:E04B4F00FFE2FFC3FF4FF9C77F4F576A" httpUri="http://treatment.plazi.org/id/E04B4F00FFE2FFC3FF4FF9C77F4F576A" lastPageId="4" lastPageNumber="289" pageId="1" pageNumber="286">
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<subSubSection id="20F8AD9DFFE2FFC6FF4FF9C77E1E5657" box="[151,581,1636,1663]" pageId="1" pageNumber="286" type="nomenclature">
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<heading id="3315497AFFE2FFC6FF4FF9C77E1E5657" bold="true" box="[151,581,1636,1663]" fontSize="11" level="1" pageId="1" pageNumber="286" reason="1">
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<taxonomicName id="AFE28595FFE2FFC6FF4FF9C77E1E5657" ID-CoL="WXX9" authority="(Bates, 1878)" baseAuthorityName="Bates" baseAuthorityYear="1878" box="[151,581,1636,1663]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="286" phylum="Arthropoda" rank="species" species="perlucens">
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<emphasis id="5A962204FFE2FFC6FF4FF9C77E1E5657" bold="true" box="[151,581,1636,1663]" pageId="1" pageNumber="286">
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<emphasis id="5A962204FFE2FFC6FF4FF9C77DC75656" bold="true" box="[151,412,1636,1663]" italics="true" pageId="1" pageNumber="286">Coleolissus perlucens</emphasis>
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(
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<bibRefCitation id="0C7383E7FFE2FFC6FE74F9C77E665657" author="Bates, H. W." box="[428,573,1636,1662]" pageId="1" pageNumber="286" pagination="713 - 721" refId="ref11963" refString="Bates, H. W. (1878) On new species of coleopterous insects (Geodephaga and Longicornia) taken by Dr. Stoliczka during the Forsyth Expedition to Kashgar in 1873 - 74. Proceedings of the Scientific Meetings of the Zoological Society of London, 1878, 713 - 721. https: // doi. org / 10.1111 / j. 1469 - 7998.1878. tb 08010. x" type="book chapter" year="1878">Bates, 1878</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="20F8AD9DFFE2FFC6FF4FF92B7D49568A" box="[151,274,1672,1699]" pageId="1" pageNumber="286" type="description">
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<paragraph id="685DFE16FFE2FFC6FF4FF92B7D49568A" blockId="1.[151,581,1636,1699]" box="[151,274,1672,1699]" pageId="1" pageNumber="286">
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(
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<figureCitation id="F0D9E293FFE2FFC6FF47F92B7D52568A" box="[159,265,1672,1699]" captionStart-0="FIGURES 1, 2" captionStart-1="FIGURES 3–8" captionStartId-0="2.[151,264,1359,1383]" captionStartId-1="4.[151,264,1294,1318]" captionTargetBox-0="[204,1394,191,1324]" captionTargetBox-1="[163,1417,192,1260]" captionTargetId-0="figure-30@2.[188,1398,181,1336]" captionTargetId-1="figure-30@4.[151,1436,181,1271]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURES 1, 2. Coleolissus perlucens (Bates, 1878), habitus (1, male, Islamabad; 2, female, holotype). Scale bar = 1.0 mm." captionText-1="FIGURES 3–8. Coleolissus perlucens (Bates, 1878). 3, laterotergite, gonosubcoxite, and gonocoxite; 4, gonocoxite; 5, 6, median lobe of aedeagus; 7, left paramere; 8, right paramere. Ventral (3), left lateral (4, 5, 8), dorsal (6) and right lateral (7) views. Scale bar = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6882746" figureDoi-1="http://doi.org/10.5281/zenodo.6882750" httpUri-0="https://zenodo.org/record/6882746/files/figure.png" httpUri-1="https://zenodo.org/record/6882750/files/figure.png" pageId="1" pageNumber="286">Figs. 1–8</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="20F8AD9DFFE2FFC6FF4FF96B7E7356F6" box="[151,552,1735,1760]" pageId="1" pageNumber="286" type="reference_group">
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<paragraph id="685DFE16FFE2FFC6FF4FF96B7E7356F6" blockId="1.[151,552,1735,1760]" box="[151,552,1735,1760]" pageId="1" pageNumber="286">
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<treatmentCitation id="E943D807FFE2FFC6FF4FF96B7E7F56F6" author="Bates, H. W." box="[151,548,1735,1760]" page="715" pageId="1" pageNumber="286" year="1878">
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<taxonomicName id="AFE28595FFE2FFC6FF4FF96B7E7F56F6" authority="Bates, 1878: 715" authorityName="Bates" authorityPageNumber="715" authorityYear="1878" box="[151,548,1735,1760]" class="Insecta" family="Elateridae" genus="Hypolithus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="286" phylum="Arthropoda" rank="species" species="perlucens">
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<emphasis id="5A962204FFE2FFC6FF4FF96B7D3456C9" box="[151,367,1736,1760]" italics="true" pageId="1" pageNumber="286">Hypolithus perlucens</emphasis>
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<bibRefCitation id="0C7383E7FFE2FFC6FEAEF96B7E7F56F6" author="Bates, H. W." box="[374,548,1735,1760]" pageId="1" pageNumber="286" pagination="713 - 721" refId="ref11963" refString="Bates, H. W. (1878) On new species of coleopterous insects (Geodephaga and Longicornia) taken by Dr. Stoliczka during the Forsyth Expedition to Kashgar in 1873 - 74. Proceedings of the Scientific Meetings of the Zoological Society of London, 1878, 713 - 721. https: // doi. org / 10.1111 / j. 1469 - 7998.1878. tb 08010. x" type="book chapter" year="1878">Bates, 1878: 715</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<emphasis id="5A962204FFE2FFC6FF4FF8AA7D1D570B" bold="true" box="[151,326,1800,1827]" pageId="1" pageNumber="286">Type material.</emphasis>
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<materialsCitation id="D88AF44BFFE2FFC6FE96F8AB7E29576E" ID-GBIF-Occurrence="3865867306" collectionCode="MNHN" collectorName="Ex Musaeo H. W. Bates" pageId="1" pageNumber="286" specimenCount="1" typeStatus="holotype">
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<typeStatus id="B75940B4FFE2FFC6FE96F8AB7DE0570B" box="[334,443,1800,1826]" pageId="1" pageNumber="286" type="holotype">Holotype</typeStatus>
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: female, “Jhelum Vall., Cashmere”, “
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<taxonomicName id="AFE28595FFE2FFC6FCBAF8AB78C0570A" authority="Bates" authorityName="Bates" authorityYear="1878" box="[866,1179,1800,1827]" class="Insecta" family="Elateridae" genus="Hypolithus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="286" phylum="Arthropoda" rank="species" species="perlucens">
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<emphasis id="5A962204FFE2FFC6FCBAF8AB7808570B" box="[866,1107,1800,1826]" italics="true" pageId="1" pageNumber="286">Hypolithus perlucens</emphasis>
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Bates
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</taxonomicName>
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” [Bates’ handwriting], “
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<collectorName id="C5179BC0FFE2FFC6FF7AF88E7DF0576E" box="[162,427,1837,1863]" pageId="1" pageNumber="286">Ex Musaeo H.W. Bates</collectorName>
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, 1892” (
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).
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</paragraph>
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<paragraph id="685DFE16FFE2FFC6FF1FF8F37954579A" blockId="1.[151,1437,1800,2043]" pageId="1" pageNumber="286">
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<emphasis id="5A962204FFE2FFC6FF1FF8F37E745743" bold="true" box="[199,559,1872,1898]" pageId="1" pageNumber="286">Additional material examined.</emphasis>
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<materialsCitation id="D88AF44BFFE2FFC6FDEBF8F379C257A6" ID-GBIF-Occurrence="3865867322" collectingDate="2003-07-12" collectingDateMax="2003-07-24" collectingDateMin="2003-07-12" collectionCode="ZIN, SMNS" collectorName="W. Heinz" country="Pakistan" county="Islamabad" elevation="1000" location="The National Forest" municipality="The National Forest Park" pageId="1" pageNumber="286" specimenCount="4" specimenCount-female="3" specimenCount-male="1">
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.
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,
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,
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Park
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,
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<collectingDate id="0C18213EFFE2FFC6FAD4F8F279CC5742" box="[1292,1431,1873,1899]" pageId="1" pageNumber="286" value="2003-07-12">12.VII.2003</collectingDate>
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,
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<collectorName id="C5179BC0FFE2FFC6FF4FF8D67D1357A7" box="[151,328,1908,1935]" pageId="1" pageNumber="286">S. Ovchinnikov</collectorName>
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leg. (
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<collectionCode id="0EF366D3FFE2FFC6FE51F8D67DE657A6" box="[393,445,1909,1935]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="286" type="Museum">ZIN</collectionCode>
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);
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,
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<location id="6D3DA8CDFFE2FFC6FD9BF8D77EB057A7" LSID="urn:lsid:plazi:treatment:E04B4F00FFE2FFC3FF4FF9C77F4F576A:6D3DA8CDFFE2FFC6FD9BF8D77EB057A7" box="[579,747,1908,1934]" country="Pakistan" county="Islamabad" municipality="The National Forest Park" name="Kaghan Valley" pageId="1" pageNumber="286">Kaghan Valley</location>
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,
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<collectorName id="C5179BC0FFE2FFC6FD2EF8D77F0B57A7" box="[758,848,1908,1934]" pageId="1" pageNumber="286">Balakot</collectorName>
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, ~
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</quantity>
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, 21/
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<date id="1C5CD8D6FFE2FFC6FC25F8D678D357A6" box="[1021,1160,1909,1935]" pageId="1" pageNumber="286" value="2003-07-24">
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<collectingDate id="0C18213EFFE2FFC6FC25F8D678D357A6" box="[1021,1160,1909,1935]" pageId="1" pageNumber="286" value="2003-07-24">24.VII.2003</collectingDate>
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,
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<collectorName id="C5179BC0FFE2FFC6FB4BF8D678A657A7" box="[1171,1277,1908,1935]" pageId="1" pageNumber="286">W. Heinz</collectorName>
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leg. (
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<collectionCode id="0EF366D3FFE2FFC6FAE6F8D679CB57A6" box="[1342,1424,1909,1935]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="1" pageNumber="286">SMNS</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="D88AF44BFFE2FFC6FF4FF83B7957579A" ID-GBIF-Occurrence="3865867308" box="[151,1292,1944,1971]" collectingDate="1994-07-20" country="India" location="Karnaprayas" pageId="1" pageNumber="286" specimenCount="1" specimenCount-female="1" stateProvince="Uttarakhand">
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<collectingCountry id="10F5BE86FFE2FFC6FF4FF83B7C82579B" box="[151,217,1944,1970]" name="India" pageId="1" pageNumber="286">India</collectingCountry>
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.
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</emphasis>
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<collectingRegion id="AA2630F4FFE2FFC6FF3DF83B7D2F579B" box="[229,372,1944,1970]" country="India" name="Uttarakhand" pageId="1" pageNumber="286">Uttarakhand</collectingRegion>
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:
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,
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<collectingCountry id="10F5BE86FFE2FFC6FE35F83B7E7C579B" box="[493,551,1944,1970]" name="India" pageId="1" pageNumber="286">India</collectingCountry>
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bor.,
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<collectingRegion id="AA2630F4FFE2FFC6FDBFF83A7F5B579B" box="[615,768,1944,1971]" country="India" name="Uttar Pradesh" pageId="1" pageNumber="286">Uttar Pradesh</collectingRegion>
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bor., env.
|
||
<location id="6D3DA8CDFFE2FFC6FCACF83A785F579A" LSID="urn:lsid:plazi:treatment:E04B4F00FFE2FFC3FF4FF9C77F4F576A:6D3DA8CDFFE2FFC6FCACF83A785F579A" box="[884,1028,1945,1971]" country="India" name="Karnaprayas" pageId="1" pageNumber="286" stateProvince="Uttarakhand">Karnaprayas</location>
|
||
,
|
||
<date id="1C5CD8D6FFE2FFC6FBC8F83A78C3579A" box="[1040,1176,1945,1971]" pageId="1" pageNumber="286" value="1994-07-20">
|
||
<collectingDate id="0C18213EFFE2FFC6FBC8F83A78C3579A" box="[1040,1176,1945,1971]" pageId="1" pageNumber="286" value="1994-07-20">20.VII.1994</collectingDate>
|
||
</date>
|
||
(cFCCH)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="685DFE16FFE2FFC6FF1FF81F7E6F57D2" blockId="1.[151,1437,1800,2043]" pageId="1" pageNumber="286">
|
||
<emphasis id="5A962204FFE2FFC6FF1FF81F7D2C57FF" bold="true" box="[199,375,1980,2006]" pageId="1" pageNumber="286">Re-description</emphasis>
|
||
(
|
||
<specimenCount id="7EE4359FFFE2FFC6FE50F81E7D8B57FF" box="[392,464,1980,2007]" pageId="1" pageNumber="286" type="male">1 male</specimenCount>
|
||
and
|
||
<specimenCount id="7EE4359FFFE2FFC6FDDEF81F7E2A57FF" box="[518,625,1980,2007]" pageId="1" pageNumber="286" type="female">5 females</specimenCount>
|
||
measured). Body medium-sized for the genus, length
|
||
<quantity id="AF1A53F3FFE2FFC6FB12F81F790957FE" box="[1226,1362,1980,2007]" metricMagnitude="-3" metricUnit="m" metricValue="8.85" metricValueMax="9.1" metricValueMin="8.6" pageId="1" pageNumber="286" unit="mm" value="8.85" valueMax="9.1" valueMin="8.6">8.6–9.1 mm</quantity>
|
||
, width
|
||
<quantity id="AF1A53F3FFE2FFC6FF4FF8427D7A57D2" box="[151,289,2017,2043]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" metricValueMax="3.9" metricValueMin="3.7" pageId="1" pageNumber="286" unit="mm" value="3.8" valueMax="3.9" valueMin="3.7">3.7–3.9 mm</quantity>
|
||
. Habitus as in
|
||
<figureCitation id="F0D9E293FFE2FFC6FE1DF8437E6B57D2" box="[453,560,2016,2043]" captionStart="FIGURES 1, 2" captionStartId="2.[151,264,1359,1383]" captionTargetBox="[204,1394,191,1324]" captionTargetId="figure-30@2.[188,1398,181,1336]" captionTargetPageId="2" captionText="FIGURES 1, 2. Coleolissus perlucens (Bates, 1878), habitus (1, male, Islamabad; 2, female, holotype). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6882746" httpUri="https://zenodo.org/record/6882746/files/figure.png" pageId="1" pageNumber="286">Figs. 1, 2</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="3C9DAE9EFFE1FFC5FF4FFAEC79DE554E" ID-DOI="http://doi.org/10.5281/zenodo.6882746" ID-Zenodo-Dep="6882746" box="[151,1413,1359,1384]" httpUri="https://zenodo.org/record/6882746/files/figure.png" pageId="2" pageNumber="287" startId="2.[151,264,1359,1383]" targetBox="[204,1394,191,1324]" targetPageId="2">
|
||
<paragraph id="685DFE16FFE1FFC5FF4FFAEC79DE554E" blockId="2.[151,1413,1359,1384]" box="[151,1413,1359,1384]" pageId="2" pageNumber="287">
|
||
<emphasis id="5A962204FFE1FFC5FF4FFAEC7D61554E" bold="true" box="[151,314,1359,1383]" pageId="2" pageNumber="287">FIGURES 1, 2.</emphasis>
|
||
<taxonomicName id="AFE28595FFE1FFC5FE99FAF37EEB5541" authority="(Bates, 1878)" baseAuthorityName="Bates" baseAuthorityYear="1878" box="[321,688,1360,1384]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="287" phylum="Arthropoda" rank="species" species="perlucens">
|
||
<emphasis id="5A962204FFE1FFC5FE99FAF37E455541" box="[321,542,1360,1384]" italics="true" pageId="2" pageNumber="287">Coleolissus perlucens</emphasis>
|
||
(
|
||
<bibRefCitation id="0C7383E7FFE1FFC5FDF4FAF37EF35541" author="Bates, H. W." box="[556,680,1360,1384]" pageId="2" pageNumber="287" pagination="713 - 721" refId="ref11963" refString="Bates, H. W. (1878) On new species of coleopterous insects (Geodephaga and Longicornia) taken by Dr. Stoliczka during the Forsyth Expedition to Kashgar in 1873 - 74. Proceedings of the Scientific Meetings of the Zoological Society of London, 1878, 713 - 721. https: // doi. org / 10.1111 / j. 1469 - 7998.1878. tb 08010. x" type="book chapter" year="1878">Bates, 1878</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, habitus (1, male, Islamabad; 2, female, holotype). Scale bar = 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="685DFE16FFE1FFC5FF1FFA3578C355D1" blockId="2.[151,1437,1430,2032]" pageId="2" pageNumber="287">Body dark brown to black, shiny and iridescent, with labrum externally, mandibles basally and very narrow lateral margins of pronotum reddish brown; abdominal sternites reddish black; palps, antennae and legs yellowish brown; femora slightly paler than tibiae and tarsi; coxae blackish brown or blackish yellow.</paragraph>
|
||
<paragraph id="685DFE16FFE1FFC4FF1FF9A17D1F514A" blockId="2.[151,1437,1430,2032]" lastBlockId="3.[151,1437,151,2023]" lastPageId="3" lastPageNumber="288" pageId="2" pageNumber="287">
|
||
Head medium-sized (HWmax/PWmax 0.61–0.64, HWmin/PWmax 0.50–0.52), impunctate. Eyes large, moderately convex (HWmax/HWmin 1.20–1.27), almost touching ventrally margins of buccal fissure. Genae glabrous. Tempora moderately long, slightly convex, somewhat steeply sloping to neck. Labrum flat, with almost straight anterior margin. Clypeus very slightly convex, very shallowly emarginated along anterior margin, with one setigerous pore at each outer angle and with very fine and short longitudinal wrinkles adjacent medially to the pore. Fronto-clypeal suture very fine, superficial, almost straight. Frontal foveae oval, more or less deepened, without prolongation on clypeus. Fronto-ocular furrows fine and short, not reaching supraorbital furrows. Supraorbital furrows very narrow, touching upper margin of eyes. Supraorbital setigerous pores small, situated slightly before level of posterior margin of eyes and removed from supraorbital furrows by distance less than width of antennomere 2 basally.
|
||
<collectingRegion id="AA2630F4FFE1FFC5FECBF8E57D295749" box="[275,370,1862,1888]" country="Myanmar" name="Chin" pageId="2" pageNumber="287">Mentum</collectingRegion>
|
||
separated from submentum by complete transverse suture, with a prominent, rather wide median tooth; epilobes markedly widened apically; submentum with one pair of long lateral setae. Ligular sclerite moderately widened distally, almost truncate at apex, with one pair of ventroapical setae. Paraglossae glabrous, narrow, markedly projecting beyond ligular sclerite and separated from it apically. Basal labial palpomere not carinate; penultimate labial palpomere as long as apical one. Mandibles elongate, somewhat narrow, evenly curved apically, with very fine oblique wrinkles in apical half dorsally; left mandible acute, at most only slightly blunted at tip. Dorsal microsculpture distinct at anterior margin of clypeus, obliterated on remaining surface, consisting of mixture of isodiametric and slightly transverse meshes. Antennae slender, surpassing pronotal basal edge by approximately one and a half apical antennomeres (in female slightly shorter than in male), pubescent from middle of antennomere 3, with antennomeres 4–8 about 2.5 times as long as wide and basal antennomere about as long as antennomere 3.
|
||
</paragraph>
|
||
<paragraph id="685DFE16FFE0FFC4FF1FFECE78E85319" blockId="3.[151,1437,151,2023]" pageId="3" pageNumber="288">Pronotum wider than long (PWmax/PL 1.38–1.45), widest just before the middle, slightly more strongly narrowed apically than basally (PWmax/PWmin-ap 1.49–1.53, PWmax/PWmin-bas 1.24–1.32), with one lateral seta inserted slightly before widest point. Sides evenly rounded along entire length; lateral bead complete, very narrow throughout. Apical margin markedly emarginated, almost straight medially, very narrowly bordered along entire length. Apical angles protruding anteriorly, narrowly rounded at apex. Basal margin almost straight or slightly concave in middle portion, slightly rounded laterally, bordered along entire length, slightly longer than apical margin (PWmin-bas/PWmin-ap 1.14–1.21) and slightly shorter than base of elytra between humeral angles; basal edge without fringe of short setae. Basal angles widely rounded. Disc convex. Lateral depressions beginning from apical angles as grooves; these depressions markedly widened from lateral setae and fused basally with basal foveae forming deep oblique laterobasal depressions isolated from each other by a convex area. Median line fine, superficial, not reaching apical margin and basal margins. Anterior transverse depression very shallow or indistinct. Disc densely punctate, more coarsely along margins and very finely in middle portion, with coarsest and confluent punctures in laterobasal depressions. Microsculpture not recognized under 56x magnification.</paragraph>
|
||
<paragraph id="685DFE16FFE0FFC4FF1FFC9A787654F3" blockId="3.[151,1437,151,2023]" pageId="3" pageNumber="288">Elytra convex, oval (EL/EW 1.48–1.50, EL/PL 2.62–2.71, EW/PWmax 1.22–1.29), widest before the apical third; sides rounded apically and basally, almost straightly diverging before the middle; subapical sinuation very shallow, almost indistinct. Humerus angularly rounded, smooth or with indistinct denticle at apex recognizable from behind. Sutural angle slightly blunted at tip, not extended posteriorly. Basal edge markedly arched, forming an obtuse angle with lateral margin. Striae with granulate microsculpture on bottom, impressed along entire length, reaching anteriorly basal elytral edge and widened apically. Intervals glabrous, strongly narrowed apically, slightly convex on disc and rather strongly convex before apex; two or three lateral intervals finely punctate, more distinctly in apical half. Parascutellar (abbreviate) striole long, with a large setigerous pore basally isolated from basal elytral edge. Interval 3 with a series of 7–10 small discal setigerous pores adjoining stria 2 along its entire length; intervals 5 and 7 without discal pores. Marginal umbilicate series without distinct gap at middle, consisting of 20–23 setigerous pores. Lateral groove narrow and flat, slightly widened apically, with granulate meshes. Microsculpture on intervals highly obliterated, at most with indistinct transverse lines. Wings fully developed.</paragraph>
|
||
<paragraph id="685DFE16FFE0FFC4FF1FFB477DDC5508" blockId="3.[151,1437,151,2023]" pageId="3" pageNumber="288">Pro- and metasternum with very fine and short setae. Metepisternum markedly longer than wide, strongly narrowed posteriorly.</paragraph>
|
||
<paragraph id="685DFE16FFE0FFC4FF1FFA88794F5649" blockId="3.[151,1437,151,2023]" pageId="3" pageNumber="288">Metacoxa with two obligatory setigerous pores, without additional pores and setae. Profemur with deep longitudinal excavation on inner side. Metafemur ventrally with two long setae at posterior margin and with several very short setae at anterior margin. Protibia with longitudinal sulcus on dorsal side, with one preapical spine on outer margin in male and with three such spines in females. Tarsi almost glabrous dorsally, with a few very short and fine, faintly visible setae on male pro- and mesotarsi. In male, protarsomeres 1–4 and mesotarsomeres 2–4 markedly widened, with biseriate adhesive scales ventrally; mesotarsomere 1 weakly widened, with scales only apically, shorter (in female longer) than mesotarsomeres 2 and 3 combined. Metatarsus slender, markedly longer than HWmax, with metatarsomeres 2–4 slightly widened distally; metatarsomere 1 elongate, about as long as metatarsomeres 2–3 combined. Tarsomere 5 with two (occasionally three) pairs of ventro-lateral setae.</paragraph>
|
||
<paragraph id="685DFE16FFE0FFC4FF1FF9CA79D2568E" blockId="3.[151,1437,151,2023]" pageId="3" pageNumber="288">
|
||
Abdominal sternites finely pubescent; apex of last visible sternite (
|
||
<collectionCode id="0EF366D3FFE0FFC4FC18F9CA7FB156AA" box="[960,1002,1641,1667]" pageId="3" pageNumber="288">VII</collectionCode>
|
||
) in both sexes rounded (in male more widely than in female, almost subtruncate), with two pairs of marginal setae; these setae not distant from margin.
|
||
</paragraph>
|
||
<paragraph id="685DFE16FFE0FFC4FF1FF9137818571C" blockId="3.[151,1437,151,2023]" pageId="3" pageNumber="288">
|
||
Female genitalia (
|
||
<figureCitation id="F0D9E293FFE0FFC4FE43F9137E4956E3" box="[411,530,1712,1738]" captionStart="FIGURES 3–8" captionStartId="4.[151,264,1294,1318]" captionTargetBox="[163,1417,192,1260]" captionTargetId="figure-30@4.[151,1436,181,1271]" captionTargetPageId="4" captionText="FIGURES 3–8. Coleolissus perlucens (Bates, 1878). 3, laterotergite, gonosubcoxite, and gonocoxite; 4, gonocoxite; 5, 6, median lobe of aedeagus; 7, left paramere; 8, right paramere. Ventral (3), left lateral (4, 5, 8), dorsal (6) and right lateral (7) views. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6882750" httpUri="https://zenodo.org/record/6882750/files/figure.png" pageId="3" pageNumber="288">Figs. 3, 4</figureCitation>
|
||
). Laterotergite symmetrical, longer than wide, with two thick setae apically. Gonosubcoxite shorter than laterotergite, markedly widened posteriorly, with one preapical spine on outer margin. Gonocoxite narrow, about 0.7 times as long as gonosubcoxite, moderately curved, with relatively narrow base and with a tiny spine (short thick seta) on both ventral and dorsal edges of outer margin.
|
||
</paragraph>
|
||
<paragraph id="685DFE16FFE0FFC4FF1FF89D78E057CF" blockId="3.[151,1437,151,2023]" pageId="3" pageNumber="288">
|
||
Aedeagus (
|
||
<figureCitation id="F0D9E293FFE0FFC4FE9EF89D7DE95770" box="[326,434,1854,1881]" captionStart="FIGURES 3–8" captionStartId="4.[151,264,1294,1318]" captionTargetBox="[163,1417,192,1260]" captionTargetId="figure-30@4.[151,1436,181,1271]" captionTargetPageId="4" captionText="FIGURES 3–8. Coleolissus perlucens (Bates, 1878). 3, laterotergite, gonosubcoxite, and gonocoxite; 4, gonocoxite; 5, 6, median lobe of aedeagus; 7, left paramere; 8, right paramere. Ventral (3), left lateral (4, 5, 8), dorsal (6) and right lateral (7) views. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6882750" httpUri="https://zenodo.org/record/6882750/files/figure.png" pageId="3" pageNumber="288">Figs. 5–8</figureCitation>
|
||
) robust, with basal bulb and parameres comparatively large (
|
||
<figureCitation id="F0D9E293FFE0FFC4FBB2F89D78835771" box="[1130,1240,1854,1881]" captionStart="FIGURES 3–8" captionStartId="4.[151,264,1294,1318]" captionTargetBox="[163,1417,192,1260]" captionTargetId="figure-30@4.[151,1436,181,1271]" captionTargetPageId="4" captionText="FIGURES 3–8. Coleolissus perlucens (Bates, 1878). 3, laterotergite, gonosubcoxite, and gonocoxite; 4, gonocoxite; 5, 6, median lobe of aedeagus; 7, left paramere; 8, right paramere. Ventral (3), left lateral (4, 5, 8), dorsal (6) and right lateral (7) views. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6882750" httpUri="https://zenodo.org/record/6882750/files/figure.png" pageId="3" pageNumber="288">Figs. 7, 8</figureCitation>
|
||
). Median lobe of aedeagus in lateral view (
|
||
<figureCitation id="F0D9E293FFE0FFC4FE60F8C27DA75755" box="[440,508,1889,1916]" captionStart="FIGURES 3–8" captionStartId="4.[151,264,1294,1318]" captionTargetBox="[163,1417,192,1260]" captionTargetId="figure-30@4.[151,1436,181,1271]" captionTargetPageId="4" captionText="FIGURES 3–8. Coleolissus perlucens (Bates, 1878). 3, laterotergite, gonosubcoxite, and gonocoxite; 4, gonocoxite; 5, 6, median lobe of aedeagus; 7, left paramere; 8, right paramere. Ventral (3), left lateral (4, 5, 8), dorsal (6) and right lateral (7) views. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6882750" httpUri="https://zenodo.org/record/6882750/files/figure.png" pageId="3" pageNumber="288">Fig. 5</figureCitation>
|
||
) arcuate on dorsal side, with ventral margin almost straight in middle portion and slightly convex in the apical quarter; oblique apical capitulum slightly hooked ventrally; in dorsal view median lobe (
|
||
<figureCitation id="F0D9E293FFE0FFC4FF47F80B7CB957EB" box="[159,226,1960,1986]" captionStart="FIGURES 3–8" captionStartId="4.[151,264,1294,1318]" captionTargetBox="[163,1417,192,1260]" captionTargetId="figure-30@4.[151,1436,181,1271]" captionTargetPageId="4" captionText="FIGURES 3–8. Coleolissus perlucens (Bates, 1878). 3, laterotergite, gonosubcoxite, and gonocoxite; 4, gonocoxite; 5, 6, median lobe of aedeagus; 7, left paramere; 8, right paramere. Ventral (3), left lateral (4, 5, 8), dorsal (6) and right lateral (7) views. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6882750" httpUri="https://zenodo.org/record/6882750/files/figure.png" pageId="3" pageNumber="288">Fig. 6</figureCitation>
|
||
) almost straight, with terminal lamella comparatively narrow, about as long as wide, rounded apically. Apical orifice in dorsal position, wide apically. Internal sac with three groups of small spines apically.
|
||
</paragraph>
|
||
<caption id="3C9DAE9EFFE7FFC3FF4FFAAD7DF75547" ID-DOI="http://doi.org/10.5281/zenodo.6882750" ID-Zenodo-Dep="6882750" httpUri="https://zenodo.org/record/6882750/files/figure.png" pageId="4" pageNumber="289" startId="4.[151,264,1294,1318]" targetBox="[163,1417,192,1260]" targetPageId="4">
|
||
<paragraph id="685DFE16FFE7FFC3FF4FFAAD7DF75547" blockId="4.[151,1437,1294,1391]" pageId="4" pageNumber="289">
|
||
<emphasis id="5A962204FFE7FFC3FF4FFAAD7D64550E" bold="true" box="[151,319,1294,1319]" pageId="4" pageNumber="289">FIGURES 3–8.</emphasis>
|
||
<taxonomicName id="AFE28595FFE7FFC3FE90FAAC7E9F550E" authority="(Bates, 1878)" baseAuthorityName="Bates" baseAuthorityYear="1878" box="[328,708,1295,1319]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="289" phylum="Arthropoda" rank="species" species="perlucens">
|
||
<emphasis id="5A962204FFE7FFC3FE90FAAC7E72550E" box="[328,553,1295,1319]" italics="true" pageId="4" pageNumber="289">Coleolissus perlucens</emphasis>
|
||
(
|
||
<bibRefCitation id="0C7383E7FFE7FFC3FDE3FAAC7EE7550E" author="Bates, H. W." box="[571,700,1295,1319]" pageId="4" pageNumber="289" pagination="713 - 721" refId="ref11963" refString="Bates, H. W. (1878) On new species of coleopterous insects (Geodephaga and Longicornia) taken by Dr. Stoliczka during the Forsyth Expedition to Kashgar in 1873 - 74. Proceedings of the Scientific Meetings of the Zoological Society of London, 1878, 713 - 721. https: // doi. org / 10.1111 / j. 1469 - 7998.1878. tb 08010. x" type="book chapter" year="1878">Bates, 1878</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. 3, laterotergite, gonosubcoxite, and gonocoxite; 4, gonocoxite; 5, 6, median lobe of aedeagus; 7, left paramere; 8, right paramere. Ventral (3), left lateral (4, 5, 8), dorsal (6) and right lateral (7) views. Scale bar = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="685DFE16FFE7FFC3FF1FFA3E78F4569A" blockId="4.[151,1437,1437,1860]" pageId="4" pageNumber="289">
|
||
<emphasis id="5A962204FFE7FFC3FF1FFA3E7D39559E" bold="true" box="[199,354,1437,1463]" pageId="4" pageNumber="289">Comparison.</emphasis>
|
||
<taxonomicName id="AFE28595FFE7FFC3FEAAFA3E7E34559E" baseAuthorityName="Bates" baseAuthorityYear="1878" box="[370,623,1437,1463]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="289" phylum="Arthropoda" rank="species" species="perlucens">
|
||
<emphasis id="5A962204FFE7FFC3FEAAFA3E7E34559E" box="[370,623,1437,1463]" italics="true" pageId="4" pageNumber="289">Coleolissus perlucens</emphasis>
|
||
</taxonomicName>
|
||
, originally described as
|
||
<taxonomicName id="AFE28595FFE7FFC3FC7DFA3E78C7559E" authorityName="Bates" authorityYear="1878" box="[933,1180,1437,1463]" class="Insecta" family="Elateridae" genus="Hypolithus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="289" phylum="Arthropoda" rank="species" species="perlucens">
|
||
<emphasis id="5A962204FFE7FFC3FC7DFA3E78C7559E" box="[933,1180,1437,1463]" italics="true" pageId="4" pageNumber="289">Hypolithus perlucens</emphasis>
|
||
</taxonomicName>
|
||
, the
|
||
<typeStatus id="B75940B4FFE7FFC3FB3CFA3D794F5591" box="[1252,1300,1438,1464]" pageId="4" pageNumber="289">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="AFE28595FFE7FFC3FF4FFA627D4355F2" authorityName="Bates" authorityYear="1892" box="[151,280,1473,1499]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="289" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5A962204FFE7FFC3FF4FFA627D4355F2" box="[151,280,1473,1499]" italics="true" pageId="4" pageNumber="289">Coleolissus</emphasis>
|
||
</taxonomicName>
|
||
, is recognizable by the following combination of characters: pronotal basal angles widely rounded, protibia sulcate on dorsal side, elytra clearly punctate on lateral intervals and at apex, and abdominal sternites finely pubescent. In general habitus with widely rounded basal angles of pronotum and oval elytra, this species is somewhat similar to
|
||
<taxonomicName id="AFE28595FFE7FFC3FE5BF98E7EDA566E" authority="Ito, 2016" authorityName="Ito" authorityYear="2016" box="[387,641,1581,1608]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="289" phylum="Arthropoda" rank="species" species="nakajimai">
|
||
<emphasis id="5A962204FFE7FFC3FE5BF98E7E4F566E" box="[387,532,1581,1607]" italics="true" pageId="4" pageNumber="289">C. nakajimai</emphasis>
|
||
<bibRefCitation id="0C7383E7FFE7FFC3FDC3F98D7EDA566E" author="Ito, N." box="[539,641,1581,1608]" pageId="4" pageNumber="289" pagination="359 - 372" refId="ref12220" refString="Ito, N. (2016) Six new species of the Selenophori carabids from India and adjacent regions (Coleoptera, Carabidae, Hatrpalini). Elytra, New Series, 6 (2), 359 - 372." type="journal article" year="2016">Ito, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
described from
|
||
<specimenCount id="7EE4359FFFE7FFC3FCE1F98D7FEF566E" box="[825,948,1581,1608]" pageId="4" pageNumber="289" type="female">one female</specimenCount>
|
||
from the “Animalai” (= Anamalai) Hills in the Indian state of
|
||
<collectingRegion id="AA2630F4FFE7FFC3FEBAF9F27DB35642" box="[354,488,1617,1643]" country="India" name="Tamil Nadu" pageId="4" pageNumber="289">Tamil Nadu</collectingRegion>
|
||
, but the latter species, according to the original description (
|
||
<bibRefCitation id="0C7383E7FFE7FFC3FB59F9F278B15642" author="Ito, N." box="[1153,1258,1617,1643]" pageId="4" pageNumber="289" pagination="359 - 372" refId="ref12220" refString="Ito, N. (2016) Six new species of the Selenophori carabids from India and adjacent regions (Coleoptera, Carabidae, Hatrpalini). Elytra, New Series, 6 (2), 359 - 372." type="journal article" year="2016">Ito, 2016</bibRefCitation>
|
||
), is larger (body length
|
||
<quantity id="AF1A53F3FFE7FFC3FF3CF9D67D1056A6" box="[228,331,1653,1680]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="4" pageNumber="289" unit="mm" value="12.5">12.5 mm</quantity>
|
||
), with vestigial wings, elytra without distinct punctation on lateral intervals (only very sparsely and microscopically punctate on disc), protibia not sulcate and abdominal sternites not pubescent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="20F8AD9DFFE7FFC3FF1FF91E7F4F576A" pageId="4" pageNumber="289" type="distribution">
|
||
<paragraph id="685DFE16FFE7FFC3FF1FF91E7F4F576A" blockId="4.[151,1437,1437,1860]" pageId="4" pageNumber="289">
|
||
<emphasis id="5A962204FFE7FFC3FF1FF91E7D3A56FE" bold="true" box="[199,353,1725,1751]" pageId="4" pageNumber="289">Distribution.</emphasis>
|
||
This species has been described from
|
||
<specimenCount id="7EE4359FFFE7FFC3FCDDF91D7F2456FE" box="[773,895,1725,1752]" pageId="4" pageNumber="289" type="female">one female</specimenCount>
|
||
collected in Jhelum Valley (
|
||
<collectingRegion id="AA2630F4FFE7FFC3FB6DF91E7C9656D5" country="India" name="Jammu and Kashmir" pageId="4" pageNumber="289">Jammu and Kashmir state</collectingRegion>
|
||
,
|
||
<collectingCountry id="10F5BE86FFE7FFC3FF01F9427D4F56D2" box="[217,276,1761,1787]" name="India" pageId="4" pageNumber="289">India</collectingCountry>
|
||
) and has been known for a long time only from this female
|
||
<typeStatus id="B75940B4FFE7FFC3FC60F942784556D2" box="[952,1054,1761,1787]" pageId="4" pageNumber="289" type="holotype">holotype</typeStatus>
|
||
. According to recent data (
|
||
<bibRefCitation id="0C7383E7FFE7FFC3FA93F9427D1F5736" author="Kataev, B. M. & Wrase, D. W." pageId="4" pageNumber="289" pagination="1810" refId="ref12531" refString="Kataev, B. M. & Wrase, D. W. (2017) Subtribe Harpalina Bonelli, 1810. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga - Adephaga. Revised and Updated Edition. Brill, Leiden, pp. 515 - 562." type="book chapter" year="2017">Kataev & Wrase, 2017</bibRefCitation>
|
||
; examined material listed above), this species also occurs in the Indian state of
|
||
<collectingRegion id="AA2630F4FFE7FFC3FB1CF8A679155736" box="[1220,1358,1797,1823]" country="India" name="Uttarakhand" pageId="4" pageNumber="289">Uttarakhand</collectingRegion>
|
||
and in
|
||
<collectingCountry id="10F5BE86FFE7FFC3FF4FF88A7CAD576A" box="[151,246,1833,1859]" name="Pakistan" pageId="4" pageNumber="289">Pakistan</collectingCountry>
|
||
and seems to be endemic to the West Himalaya.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |