138 lines
18 KiB
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138 lines
18 KiB
XML
<document id="50366693E99F3CA00BEC0B6236C7BBD5" ID-DOI="10.1111/j.1096-3642.2011.00731.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5442095" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630452512153" checkinUser="valdenar" docAuthor="Li, Liang & Zhou, Hong-Zhang" docDate="2011" docId="E0275714D9720E0DFE9DE98EFDA3A8BA" docLanguage="en" docName="j.1096-3642.2011.00731.x.pdf" docOrigin="Zoological Journal of the Linnean Society 163 (3)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2011.00731.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Hyptiomina Casey 1906" docType="treatment" docVersion="3" lastPageNumber="694" masterDocId="1C1E2F6CD97D0E1DFFA5EC4DFFF6AE0C" masterDocTitle="Revision and phylogenetic assessment of the rove beetle genus Pseudohesperus Hayashi, with broad reference to the subtribe Philonthina (Coleoptera: Staphylinidae: Staphylinini)" masterLastPageNumber="722" masterPageNumber="679" pageNumber="694" updateTime="1699275715089" updateUser="plazi">
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<mods:title id="55F278C68D5B35766E1473B15E517164">Revision and phylogenetic assessment of the rove beetle genus Pseudohesperus Hayashi, with broad reference to the subtribe Philonthina (Coleoptera: Staphylinidae: Staphylinini)</mods:title>
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<mods:namePart id="1BB3747876FC9FD63010E5F9C6D2FE87">Li, Liang</mods:namePart>
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<mods:namePart id="62C59020E32886A6EBB35C9DD82AAC70">Zhou, Hong-Zhang</mods:namePart>
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<mods:typeOfResource id="E3341FB18AAD5236961DEDD92D981338">text</mods:typeOfResource>
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<mods:title id="90591E4E232D41F2A719936722D6BC95">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="9444D5D46604FB830C0DC8F6BE5EE29B">
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<mods:date id="83D9A7CEEFC97C1BD821BB80094743B8">2011</mods:date>
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<mods:number id="61E3B530AF109CCCCB263C1E173BB032">2011-10-25</mods:number>
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<mods:number id="845D6C9956F952A28A9B7AD835E0B71F">163</mods:number>
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<mods:identifier id="FAE30FDBB63B3CB6B26E72B65A20131E" type="DOI">10.1111/j.1096-3642.2011.00731.x</mods:identifier>
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<subSubSection id="2094B589D9720E12FE9DE98EFD81ABD4" box="[312,631,1475,1496]" pageId="15" pageNumber="694" type="nomenclature">
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<paragraph id="6831E602D9720E12FE9DE98EFD81ABD4" blockId="15.[312,631,1475,1496]" box="[312,631,1475,1496]" pageId="15" pageNumber="694">
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<heading id="3379516ED9720E12FE9DE98EFD81ABD4" box="[312,631,1475,1496]" centered="true" fontSize="8" level="2" pageId="15" pageNumber="694" reason="8">
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THE SUBTRIBE
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<taxonomicName id="AF8E9D81D9720E12FE4DE98EFD81ABD4" ID-CoL="8XJ2X" authorityName="Casey" authorityYear="1906" box="[488,631,1475,1496]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="694" phylum="Arthropoda" rank="subTribe" subTribe="Hyptiomina">HYPTIOMINA</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="2094B589D9720E0DFF01E9A7FDA3A8BA" lastPageId="16" lastPageNumber="695" pageId="15" pageNumber="694" type="description">
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<paragraph id="6831E602D9720E0DFF01E9A7FDA3A8BA" blockId="15.[164,779,1514,1903]" lastBlockId="16.[144,759,1512,1718]" lastPageId="16" lastPageNumber="695" pageId="15" pageNumber="694">
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The subtribe
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<taxonomicName id="AF8E9D81D9720E12FEF6E9A7FE28A80C" authorityName="Casey" authorityYear="1906" box="[339,478,1514,1536]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="694" phylum="Arthropoda" rank="subTribe" subTribe="Hyptiomina">Hyptiomina</taxonomicName>
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, which was established by
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<bibRefCitation id="0C1F9BF3D9720E12FF6BEA45FE9DA812" author="Casey TL" box="[206,363,1544,1566]" pageId="15" pageNumber="694" pagination="125 - 434" refId="ref20246" refString="Casey TL. 1906. Observations on the staphylinid groups Aleocharinae and Xantholinini chiefly of America. Transactions of the Academy of Science of St. Louis 16: 125 - 434." type="journal article" year="1906">Casey (1906)</bibRefCitation>
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, contains only the genus
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<taxonomicName id="AF8E9D81D9720E12FD11EA45FEB9A830" authority="Erichson, 1839" authorityName="Erichson" authorityYear="1839" class="Insecta" family="Staphylinidae" genus="Holisus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="694" phylum="Arthropoda" rank="genus">
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<emphasis id="5AFA3A10D9720E12FD11EA45FCFCA811" box="[692,778,1544,1565]" italics="true" pageId="15" pageNumber="694">Holisus</emphasis>
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<bibRefCitation id="0C1F9BF3D9720E12FF01EA6AFEB9A830" author="Erichson WF" box="[164,335,1575,1597]" pageId="15" pageNumber="694" pagination="1 - 400" refId="ref20495" refString="Erichson WF. 1839. Genera et species Staphylinorum insectorum coleopterorum familiae, Vol. 1. Berlin: F. H. Morin, 1 - 400." type="book chapter" year="1839">Erichson, 1839</bibRefCitation>
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</taxonomicName>
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distributed in the Neotropical, Nearctic, and Ethiopian regions (
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<bibRefCitation id="0C1F9BF3D9720E12FE79EA0BFD75A850" author="Herman LH" box="[476,643,1606,1628]" pageId="15" pageNumber="694" pagination="2241 - 3020" refId="ref20779" refString="Herman LH. 2001. Catalog of the Staphylinidae (Insecta: Coleoptera) 1758 to the end of the milleenium. Bulletion of the American Museum of Natural History 265: 2241 - 3020." type="journal article" year="2001">Herman, 2001</bibRefCitation>
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).
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<bibRefCitation id="0C1F9BF3D9720E12FD33EA0BFEB2A876" author="Newton AF & Thayer MK" pageId="15" pageNumber="694" pagination="1 - 92" refId="ref21403" refString="Newton AF, Thayer MK. 1992. Current classification and family-group names in Staphyliniformia (Coleoptera). Fieldiana: Zoology (N. Ser.) 67: 1 - 92." type="journal article" year="1992">Newton & Thayer (1992)</bibRefCitation>
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treated it as a subtribe of
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<taxonomicName id="AF8E9D81D9720E12FDD0EA29FCF1A876" authorityName="Latreille" authorityYear="1802" box="[629,775,1636,1658]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="694" phylum="Arthropoda" rank="tribe" tribe="Staphylinini">Staphylinini</taxonomicName>
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. However, the current systematic position of
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<taxonomicName id="AF8E9D81D9720E12FD64EACEFF1DA8BB" authorityName="Casey" authorityYear="1906" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="694" phylum="Arthropoda" rank="subTribe" subTribe="Hyptiomina">Hyptiomina</taxonomicName>
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in the tribe
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<taxonomicName id="AF8E9D81D9720E12FED9EAECFDFCA8BB" authorityName="Latreille" authorityYear="1802" box="[380,522,1697,1719]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="694" phylum="Arthropoda" rank="tribe" tribe="Staphylinini">Staphylinini</taxonomicName>
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may still be questionable (
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<bibRefCitation id="0C1F9BF3D9720E12FF55EA8DFD60A8DA" author="Solodovnikov AY & Schomann A" box="[240,662,1728,1750]" pageId="15" pageNumber="694" pagination="443 - 466" refId="ref22211" refString="Solodovnikov AY, Schomann A. 2009. Revised systematics and biogeography of ' Quediina' of sub-Saharan Africa: new phylogenetic insights into the rove beetle tribe Staphylinini (Coleoptera: Staphylinidae). Systematic Entomology 34: 443 - 466." type="journal article" year="2009">Solodovnikov & Schomann, 2009</bibRefCitation>
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). In the present phylogenetic analyses, the genus
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<taxonomicName id="AF8E9D81D9720E12FD11EA93FCFCA8FF" authorityName="Erichson" authorityYear="1839" box="[692,778,1758,1779]" class="Insecta" family="Staphylinidae" genus="Holisus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="694" phylum="Arthropoda" rank="genus">
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<emphasis id="5AFA3A10D9720E12FD11EA93FCFCA8FF" box="[692,778,1758,1779]" italics="true" pageId="15" pageNumber="694">Holisus</emphasis>
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</taxonomicName>
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nested within the core of
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<taxonomicName id="AF8E9D81D9720E12FE67EAB0FDBFA91F" baseAuthorityName="Smetana & Davies" baseAuthorityYear="2000" box="[450,585,1789,1811]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="694" phylum="Arthropoda" rank="subTribe" subTribe="Philonthina">Philonthina</taxonomicName>
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and appeared as Clade C with one branch of the genus
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<taxonomicName id="AF8E9D81D9720E12FD22EB51FCFCA93D" box="[647,778,1820,1841]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="694" phylum="Arthropoda" rank="genus">
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<emphasis id="5AFA3A10D9720E12FD22EB51FCFCA93D" box="[647,778,1820,1841]" italics="true" pageId="15" pageNumber="694">Belonuchus</emphasis>
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</taxonomicName>
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(Clade D). This result agreed well with the recent phylogenetic study (
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<bibRefCitation id="0C1F9BF3D9720E12FE22EB14FD09A963" author="Solodovnikov AY & Schomann A" box="[391,767,1881,1903]" pageId="15" pageNumber="694" pagination="443 - 466" refId="ref22211" refString="Solodovnikov AY, Schomann A. 2009. Revised systematics and biogeography of ' Quediina' of sub-Saharan Africa: new phylogenetic insights into the rove beetle tribe Staphylinini (Coleoptera: Staphylinidae). Systematic Entomology 34: 443 - 466." type="journal article" year="2009">Solodovnikov & Schomann, 2009</bibRefCitation>
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); although these two studies were based on different sampled representatives and carried out for different purposes. The genus
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<taxonomicName id="AF8E9D81D9720E12FB95E91DFB70AB69" authorityName="Erichson" authorityYear="1839" box="[1072,1158,1360,1381]" class="Insecta" family="Staphylinidae" genus="Holisus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="694" phylum="Arthropoda" rank="genus">
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<emphasis id="5AFA3A10D9720E12FB95E91DFB70AB69" box="[1072,1158,1360,1381]" italics="true" pageId="15" pageNumber="694">Holisus</emphasis>
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</taxonomicName>
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can be distinguished by a very flat body, aciculate apical segments of labial and maxillary palpi, protibia with ctenidium of several rows of stout setae, mesoscutellum with one transverse carina, and especially the presence of an additional oblique line connecting the superior and inferior lines of the pronotal hypomeron (
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<bibRefCitation id="0C1F9BF3D9720E12FA91EA45FC85A830" author="Smetana A" pageId="15" pageNumber="694" pagination="1 - 946" refId="ref21981" refString="Smetana A. 1995. Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae) classification, phylogeny and taxonomic revision. Memoirs on Entomology, International 3: 1 - 946." type="journal article" year="1995">Smetana, 1995</bibRefCitation>
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;
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<bibRefCitation id="0C1F9BF3D9720E12FCDAEA6AFB96A831" author="Newton AF & Thayer MK & Ashe JS & Chandler DS" box="[895,1120,1574,1597]" pageId="15" pageNumber="694" pagination="272 - 418" refId="ref21494" refString="Newton AF, Thayer MK, Ashe JS, Chandler DS. 2000. Staphylinidae. In: Arnett RH, Thomas MC, eds. American beetles. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia, Vol. 1. Boca Raton, Florida: CRC Press, 272 - 418." type="book chapter" year="2000">
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Newton
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<emphasis id="5AFA3A10D9720E12FC7AEA6AFBF9A837" box="[991,1039,1574,1596]" italics="true" pageId="15" pageNumber="694">et al</emphasis>
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., 2000
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</bibRefCitation>
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). However, it shares most of its character states with the members of the subtribe
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<taxonomicName id="AF8E9D81D9720E12FC9FEA29FC37A876" baseAuthorityName="Smetana & Davies" baseAuthorityYear="2000" box="[826,961,1636,1658]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="694" phylum="Arthropoda" rank="subTribe" subTribe="Philonthina">Philonthina</taxonomicName>
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including ligula entire, apical segments of labial and maxillary palpi glabrous, mentum with only one anterolateral seta on each side, neck impunctate, gular sutures fused, claws of all tarsi without empodial setae, and even the additional connecting line of the pronotal hypomeron (similar to that of the genus
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<taxonomicName id="AF8E9D81D9720E12FC2EEB56FBCDA93C" box="[907,1083,1819,1840]" class="Insecta" family="Staphylinidae" genus="Craspedomerus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="694" phylum="Arthropoda" rank="genus">
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<emphasis id="5AFA3A10D9720E12FC2EEB56FBCDA93C" box="[907,1083,1819,1840]" italics="true" pageId="15" pageNumber="694">Craspedomerus</emphasis>
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</taxonomicName>
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, but much longer). It is also easy to understand the close relationships of
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<taxonomicName id="AF8E9D81D9720E12FAEEEB77FA57A943" authorityName="Erichson" authorityYear="1839" box="[1355,1441,1850,1871]" class="Insecta" family="Staphylinidae" genus="Holisus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="694" phylum="Arthropoda" rank="genus">
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<emphasis id="5AFA3A10D9720E12FAEEEB77FA57A943" box="[1355,1441,1850,1871]" italics="true" pageId="15" pageNumber="694">Holisus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="AF8E9D81D9720E12FCCAEB15FC04A961" box="[879,1010,1880,1901]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="694" phylum="Arthropoda" rank="genus">
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<emphasis id="5AFA3A10D9720E12FCCAEB15FC04A961" box="[879,1010,1880,1901]" italics="true" pageId="15" pageNumber="694">Belonuchus</emphasis>
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</taxonomicName>
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when considering the flat body form and the similar subcortical habitats (most species of
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<taxonomicName id="AF8E9D81D96D0E0DFF35EA4AFF10A810" authorityName="Erichson" authorityYear="1839" box="[144,230,1543,1564]" class="Insecta" family="Staphylinidae" genus="Holisus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="695" phylum="Arthropoda" rank="genus">
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<emphasis id="5AFA3A10D96D0E0DFF35EA4AFF10A810" box="[144,230,1543,1564]" italics="true" pageId="16" pageNumber="695">Holisus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="AF8E9D81D96D0E0DFE86EA4AFE50A810" box="[291,422,1543,1564]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="695" phylum="Arthropoda" rank="genus">
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<emphasis id="5AFA3A10D96D0E0DFE86EA4AFE50A810" box="[291,422,1543,1564]" italics="true" pageId="16" pageNumber="695">Belonuchus</emphasis>
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</taxonomicName>
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are found under the bark of decaying trees) (
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<bibRefCitation id="0C1F9BF3D96D0E0DFEEAEA68FD26A837" author="Solodovnikov AY & Schomann A" box="[335,720,1573,1595]" pageId="16" pageNumber="695" pagination="443 - 466" refId="ref22211" refString="Solodovnikov AY, Schomann A. 2009. Revised systematics and biogeography of ' Quediina' of sub-Saharan Africa: new phylogenetic insights into the rove beetle tribe Staphylinini (Coleoptera: Staphylinidae). Systematic Entomology 34: 443 - 466." type="journal article" year="2009">Solodovnikov & Schomann, 2009</bibRefCitation>
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). If the broad outlines of this analysis are supported by future work, the subtribe
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<taxonomicName id="AF8E9D81D96D0E0DFE18EA2FFDB3A874" authorityName="Casey" authorityYear="1906" box="[445,581,1634,1656]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="695" phylum="Arthropoda" rank="subTribe" subTribe="Hyptiomina">Hyptiomina</taxonomicName>
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may suffer the same fate as
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<taxonomicName id="AF8E9D81D96D0E0DFE8EEACCFE11A89B" box="[299,487,1665,1687]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="693" phylum="Arthropoda" rank="subTribe" subTribe="Craspedomerina">Craspedomerina</taxonomicName>
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(currently treated as a synonym of
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<taxonomicName id="AF8E9D81D96D0E0DFEBBEAEDFE50A8BA" baseAuthorityName="Smetana & Davies" baseAuthorityYear="2000" box="[286,422,1696,1718]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="695" phylum="Arthropoda" rank="subTribe" subTribe="Philonthina">Philonthina</taxonomicName>
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) in the future.
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</paragraph>
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</subSubSection>
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</treatment>
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