treatments-xml/data/DF/5D/B2/DF5DB2B8F5B9BF85E0E16D1CE5F78DA9.xml

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<mods:title id="E198553B68CA2ED245667F02B2337BD0">A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)</mods:title>
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<mods:namePart id="013111FD74CC150CCCA6CEFB8A744687">Caterino, Michael S.</mods:namePart>
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<mods:namePart id="A0D3C70C49D08062CD46C284B4653989">Tishechkin, Alexey K.</mods:namePart>
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<subSubSection id="BB92854201CF74AF64C5443319F30087" pageId="252" pageNumber="253" type="nomenclature">
<paragraph id="10F49BA45B47CF1105280FE916220A23" pageId="252" pageNumber="253">
<taxonomicName id="F00D297355ED7112D9B67A7FA25A8336" ID-CoL="6SNKZ" LSID="http://species-id.net/wiki/Operclipygus_subdepressus" authority="Schmidt, 1889" authorityName="Schmidt" authorityYear="1889" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus subdepressus" order="Coleoptera" pageId="252" pageNumber="253" phylum="Arthropoda" rank="species" species="subdepressus">Operclipygus subdepressus (Schmidt, 1889)</taxonomicName>
Figs 67 
<normalizedToken id="6D03C240F66DC4CB0DEAB713F71E7395" originalValue="A–B68A–EMap">A-B68A-EMap</normalizedToken>
24
</paragraph>
</subSubSection>
<subSubSection id="E9649256D97AAB5B796CE5F0046901A9" pageId="252" pageNumber="253" type="reference_group">
<paragraph id="43300779A0F6969E34B7CF7F01A0A951" pageId="252" pageNumber="253">
<taxonomicName id="A4FF3E65B2C6B2055A34EAABFDF5F1A3" class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister subdepressus" order="Coleoptera" pageId="252" pageNumber="253" phylum="Arthropoda" rank="species" species="subdepressus">Phelister subdepressus</taxonomicName>
Schmidt, 1889c: 337; 
<taxonomicName id="15326F2380FA547B0A2E4EE7B3406196" class="Insecta" family="Histeridae" genus="Pseudister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudister subdepressus" order="Coleoptera" pageId="252" pageNumber="253" phylum="Arthropoda" rank="species" species="subdepressus">Pseudister subdepressus</taxonomicName>
: 
<bibRefCitation id="1D83E6285701E57DD2F308391F56644E" pageId="252" pageNumber="253">Bickhardt (1917</bibRefCitation>
: 165); 
<taxonomicName id="9CE5DC6C473C2474A06D649870F6D064" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus subdepressus" order="Coleoptera" pageId="252" pageNumber="253" phylum="Arthropoda" rank="species" species="subdepressus">Operclipygus subdepressus</taxonomicName>
: 
<bibRefCitation id="121CD965E88B432C06F4B1B64F2C10C2" author="Wenzel, RL" journalOrPublisher="Resultats Scientifiques" pageId="377" pageNumber="378" pagination="235 - 266" title="XXVII - Histeridae from Ecuador. Mission zoologique Belge aux Iles Galapagos et en Ecuador (N. et J. Leleup 1964 - 1965)." volume="3" year="1976">Wenzel (1976</bibRefCitation>
: 260).
</paragraph>
</subSubSection>
<subSubSection id="F3FEBBCEC8C8C0561E4B280B00287C14" pageId="252" pageNumber="253" type="type locality">
<paragraph id="EC76C646B93A2F2461DD00423489A7CA" pageId="252" pageNumber="253">Type locality.</paragraph>
<paragraph id="0B324CBB59F558F64B1FEFEE7C20C3B0" pageId="252" pageNumber="253">Not specified beyond Colombia.</paragraph>
</subSubSection>
<subSubSection id="4506DFB6893B661D9B368A9CA91E4B1B" pageId="252" pageNumber="253" type="type material">
<paragraph id="D507126D9BC6FAAD88C44436A6BF98A9" pageId="252" pageNumber="253">Type material.</paragraph>
<paragraph id="545338B88306746D46F311B988632991" pageId="252" pageNumber="253">
Lectotype, here designated: 
<normalizedToken id="8ED50CC920A51C521AA3CA8423141B46" originalValue="“Columb”">&quot;Columb&quot;</normalizedToken>
/ 
<normalizedToken id="50A7232E64FCE15EE2AF814FD669F8E0" originalValue="“67882”">&quot;67882&quot;</normalizedToken>
/ 
<normalizedToken id="CD9423850FD448FA0A0F25656D8817A4" originalValue="“Type”">&quot;Type&quot;</normalizedToken>
/ &quot; 
<taxonomicName id="9D98B9C8465BB6CE9AABE541D6219D91" class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister subdepressus" order="Coleoptera" pageId="252" pageNumber="253" phylum="Arthropoda" rank="species" species="subdepressus">Phelister subdepressus</taxonomicName>
Schm. typ.&quot; / &quot;LECTOTYPE 
<taxonomicName id="D141D4C4743D3202730282645BC5C065" class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister subdepressus" order="Coleoptera" pageId="252" pageNumber="253" phylum="Arthropoda" rank="species" species="subdepressus">Phelister subdepressus</taxonomicName>
Schmidt, 1889 M.S. Caterino &amp; A.K. Tishechkin des. 2010&quot; (ZMHB). This species was described from an unspecified number of specimens, and the lectotype designation fixes primary type status on the only known original specimen.
</paragraph>
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<subSubSection id="0F63F42E70591221B9DF67E9FF4C29D9" pageId="252" pageNumber="253" type="other material">
<paragraph id="BC58ED1FC3A1C360D1CFCBCE63CB416C" pageId="252" pageNumber="253">Other material.</paragraph>
<paragraph id="6510C75CD35338607611E6A35DD4B287" pageId="252" pageNumber="253">TRINIDAD: 1: xi.1903, B.W. Indies, G.E. Bryant (BMNH)</paragraph>
</subSubSection>
<subSubSection id="9813EB36A3F39F93F1D10B357E439E4C" lastPageId="254" lastPageNumber="255" pageId="252" pageNumber="253" type="description">
<paragraph id="483DD2458D3574ADE3A14796388917F1" pageId="252" pageNumber="253">Diagnostic description.</paragraph>
<paragraph id="8E2797F703E31B3202178C71696AB8B9" lastPageId="254" lastPageNumber="255" pageId="252" pageNumber="253">
Length (nontype only measured): 2.12 mm, width: 1.68 mm; body rufescent, elongate, subdepressed, approximately parallel-sided; frons weakly depressed, frontal stria divergent at sides, almost evenly arcuate across front, weakly sinuate above antennal bases; supraorbital stria absent; epistoma flat; labrum 
<pageBreakToken id="DD4906EADEAD6F1096AC97B643235877" pageId="253" pageNumber="254" start="start">about</pageBreakToken>
twice as wide as midline length, apex asymmetrical, shorter on right side than left; left mandible slightly produced at base, right with acute basal tooth; pronotum lacking prescutellar impression, disk with fine, inconspicuous ground punctation, with 
<pageBreakToken id="01CD002BBAF8F854A3CA1F98E41837AE" pageId="254" pageNumber="255" start="start">numerous</pageBreakToken>
(&gt;20) coarser punctures along sides; marginal pronotal stria broadly interrupted behind head; submarginal pronotal stria continuous along lateral and anterior margins, but weakly angulate behind eyes; median pronotal gland openings simple, slightly less than one-half pronotal length behind anterior pronotal margin; elytron with two complete epipleural striae, outer subhumeral stria interrupted at middle, inner subhumeral stria present only in apical half, striae 1-3 complete, 4th stria present in just over apical half, 5th stria present in apical half, sutural stria present in apical two-thirds; elytral disk with irregular series of apical marginal punctures; pronotal keel projecting at base, carinal striae complete, sinuate, meeting in narrow anterior arch; anterior mesoventral margin distinctly emarginate, marginal stria complete or interrupted at middle; mesometaventral stria displaced anterad, transverse across middle of mesoventrite, bend posterolaterad, continued by lateral metaventral stria toward middle of metacoxa; 1st abdominal ventrite with inner lateral stria complete, outer abbreviated, without postmetacoxal fovea; propygidium with ground punctation fine, sparse, densely covered with small coarse punctures, separated by much less than their diameters; pygidium with dense, fine ground punctation, with numerous coarser punctures interspersed, more densely toward apex; marginal pygidial sulcus complete, moderately deep, weakly crenulate. Male genitalia (Figs 68 
<normalizedToken id="38A7F5FC82E64CB1EAF615061A27BA9F" originalValue="A–E">A-E</normalizedToken>
): accessory sclerites present, very small, not basal; T8 broad and short, sides curving to subtruncate apex; apical emargination narrow, moderately deep, basal emargination broadly arcuate, basal membrane attachment line distad basal emargination by about one-fourth its depth, ventrolateral apodemes most strongly developed basally, narrowed to apex; S8 more or less parallel-sided, apical guides narrow at base, broadening to rounded apices, with distinctly membraneous inner edges, halves narrowly fused at extreme ventral base, divergent to apex; T9 with sides rounded, convergent to subacute, weakly convergent apices; T10 cordate, fused along midline in basal half; S9 [of only available male broken] with stem narrowed near apex, with narrow apical emargination, apical flanges separate; tegmen short and broad, widest about one-third from apex, with extremely large and projecting medioventral tooth, in addition to distinct ventrolateral cusp on body of tegmen; median lobe with wide gonopore, long proximal apodemes, showing weak differentiation; basal piece about one-third tegmen length.
</paragraph>
</subSubSection>
<subSubSection id="7C1A59346697AEECEB0FFAE01CE5283F" pageId="254" pageNumber="255" type="remarks">
<paragraph id="0F0D8DD2737D7B8848C143932AEB81AA" pageId="254" pageNumber="255">Remarks.</paragraph>
<paragraph id="95EBE2CF5A64E27F1979B0EF6C4EEDEC" pageId="254" pageNumber="255">
The aedeagus (Fig. 68E) of this species unequivocally indicates relationships with the 
<taxonomicName id="7706D67BD16585D69F03A5B8526C5BE4" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus fossipygus" order="Coleoptera" pageId="254" pageNumber="255" phylum="Arthropoda" rank="species" species="fossipygus">Operclipygus fossipygus</taxonomicName>
group. However, it has a number of unusual characters for that group, including the elongate, subdepressed body form (Fig. 67A), the submarginal pronotal stria showing a postocular angulation, and the lack of basal pygidial foveae (Fig. 67B).
</paragraph>
<caption id="4814FC033B876B55996EAFDB17BFA459" pageId="254" pageNumber="255">
<paragraph id="793C9E9F563D33B189557666FE3EAE89" pageId="254" pageNumber="255">
Figure 67. 
<taxonomicName id="1BD4C99089DEAFC9505552D21B272155" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus fossipygus" order="Coleoptera" pageId="254" pageNumber="255" phylum="Arthropoda" rank="species" species="fossipygus">Operclipygus fossipygus</taxonomicName>
group. A Dorsal habitus of 
<taxonomicName id="19DC91F9820E80F824C1129FF546EDF7" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus subdepressus" order="Coleoptera" pageId="254" pageNumber="255" phylum="Arthropoda" rank="species" species="subdepressus">Operclipygus subdepressus</taxonomicName>
B Pygidia of 
<taxonomicName id="731DBF84F31AB17B0D819E4E7AAB4159" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus subdepressus" order="Coleoptera" pageId="254" pageNumber="255" phylum="Arthropoda" rank="species" species="subdepressus">Operclipygus subdepressus</taxonomicName>
C Dorsal habitus of 
<taxonomicName id="550D30E3B14C71EB5B2682A012D6A8F4" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus simplicipygus" order="Coleoptera" pageId="254" pageNumber="255" phylum="Arthropoda" rank="species" species="simplicipygus">Operclipygus simplicipygus</taxonomicName>
D Pygidia of 
<taxonomicName id="225A991B7F2D68CBD33A9936D2547847" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus simplicipygus" order="Coleoptera" pageId="254" pageNumber="255" phylum="Arthropoda" rank="species" species="simplicipygus">Operclipygus simplicipygus</taxonomicName>
E Pygidia of 
<taxonomicName id="BF3472AE37C0412782BA4D7C14D4BBE9" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus therondi" order="Coleoptera" pageId="254" pageNumber="255" phylum="Arthropoda" rank="species" species="therondi">Operclipygus therondi</taxonomicName>
F Pygidia of 
<taxonomicName id="B73E459339AF65516CCA3AF002B24720" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus disconnectus" order="Coleoptera" pageId="254" pageNumber="255" phylum="Arthropoda" rank="species" species="disconnectus">Operclipygus disconnectus</taxonomicName>
G Dorsal habitus of 
<taxonomicName id="12419CFE386E12A18A9A21B5C6E9A91C" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus olivensis" order="Coleoptera" pageId="254" pageNumber="255" phylum="Arthropoda" rank="species" species="olivensis">Operclipygus olivensis</taxonomicName>
H Pygidia of 
<taxonomicName id="EC8A4A09D84E335BD1826E827A8D16BE" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus olivensis" order="Coleoptera" pageId="254" pageNumber="255" phylum="Arthropoda" rank="species" species="olivensis">Operclipygus olivensis</taxonomicName>
.
</paragraph>
</caption>
<caption id="27FAD020BA85AB9D3451D2DA0AA695EE" pageId="254" pageNumber="255">
<paragraph id="CAD81853DE310665D4CBAAA89D8CF791" pageId="254" pageNumber="255">
Figure 68. Male genitalia of 
<taxonomicName id="E142C8697B43AFABDA642C1C3CB06D0C" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus fossipygus" order="Coleoptera" pageId="254" pageNumber="255" phylum="Arthropoda" rank="species" species="fossipygus">Operclipygus fossipygus</taxonomicName>
group. A T8 of 
<taxonomicName id="F666DADABC692890A3289EEFC7DEC658" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus subdepressus" order="Coleoptera" pageId="254" pageNumber="255" phylum="Arthropoda" rank="species" species="subdepressus">Operclipygus subdepressus</taxonomicName>
B S8 of 
<taxonomicName id="7A64548DFCE3E29F43D917EB76EDA5B2" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus subdepressus" order="Coleoptera" pageId="254" pageNumber="255" phylum="Arthropoda" rank="species" species="subdepressus">Operclipygus subdepressus</taxonomicName>
C T9 &amp; T10 of 
<taxonomicName id="A9E135736F55AF238C4575D06EE835D8" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus subdepressus" order="Coleoptera" pageId="254" pageNumber="255" phylum="Arthropoda" rank="species" species="subdepressus">Operclipygus subdepressus</taxonomicName>
D S9 of 
<taxonomicName id="CC5E00F75285EF8E04C69C8D2FA82FDF" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus subdepressus" order="Coleoptera" pageId="254" pageNumber="255" phylum="Arthropoda" rank="species" species="subdepressus">Operclipygus subdepressus</taxonomicName>
, (stem and head both broken in only available male; presumed reconstruction shown in gray) E Aedeagus, dorsal and lateral views, of 
<taxonomicName id="DE2C7B3A1911F3FD9A022BC8DEE83D9A" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus subdepressus" order="Coleoptera" pageId="254" pageNumber="255" phylum="Arthropoda" rank="species" species="subdepressus">Operclipygus subdepressus</taxonomicName>
F Aedeagus, dorsal and lateral views, of 
<taxonomicName id="FB2B7829291934C3EBD62DBDB5C10594" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus simplicipygus" order="Coleoptera" pageId="254" pageNumber="255" phylum="Arthropoda" rank="species" species="simplicipygus">Operclipygus simplicipygus</taxonomicName>
G Aedeagus, dorsal and lateral views, of 
<taxonomicName id="1F2BD96A18DC541474797B7315A66DA3" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus disconnectus" order="Coleoptera" pageId="254" pageNumber="255" phylum="Arthropoda" rank="species" species="disconnectus">Operclipygus disconnectus</taxonomicName>
H T8 of 
<taxonomicName id="110C18128D4914BDFCD4D34109D1E10A" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus disconnectus" order="Coleoptera" pageId="254" pageNumber="255" phylum="Arthropoda" rank="species" species="disconnectus">Operclipygus disconnectus</taxonomicName>
I S8 of 
<taxonomicName id="8D8DF73B1F85B2F0130BDEA73A1CA72F" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus disconnectus" order="Coleoptera" pageId="254" pageNumber="255" phylum="Arthropoda" rank="species" species="disconnectus">Operclipygus disconnectus</taxonomicName>
J T9 &amp; T10 of 
<taxonomicName id="06D23C67EE634C297839A483945A97EB" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus disconnectus" order="Coleoptera" pageId="254" pageNumber="255" phylum="Arthropoda" rank="species" species="disconnectus">Operclipygus disconnectus</taxonomicName>
K S9 of 
<taxonomicName id="8D998AB9C0D34320BA67248D311CBA68" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus disconnectus" order="Coleoptera" pageId="254" pageNumber="255" phylum="Arthropoda" rank="species" species="disconnectus">Operclipygus disconnectus</taxonomicName>
.
</paragraph>
</caption>
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</treatment>
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