217 lines
29 KiB
XML
217 lines
29 KiB
XML
<document id="BF340091A74A0DBCAA01441BB2535EE8" ID-CLB-Dataset="27575" ID-DOI="10.11646/zootaxa.4554.1.9" ID-GBIF-Dataset="ff7707f0-058f-4f40-8eb8-2b6c9c75a173" ID-ISSN="1175-5326" ID-Zenodo-Dep="2623565" ID-ZooBank="A804E949-109A-468D-B58B-CF7C8BCB3059" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="diego" checkinTime="1554271225461" checkinUser="plazi" docAuthor="Vandenberg, Natalia J. & Hanson, Paul E." docDate="2019" docId="DF18F971FFD70B3DFF4BFD236E61BE8C" docLanguage="en" docName="zootaxa.4554.1.9.pdf" docOrigin="Zootaxa 4554 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Moiradiomus atropos Vandenberg & Hanson, 2019, sp. nov." docType="treatment" docVersion="10" lastPageNumber="277" masterDocId="23218109FFC20B2BFFDCFFF96C54BC27" masterDocTitle="Overview of the lady beetle tribe Diomini (Coleoptera: Coccinellidae) and description of a new phytophagous, silk-spinning genus from Costa Rica that induces food bodies on leaves of Piper (Piperaceae)" masterLastPageNumber="285" masterPageNumber="255" pageNumber="276" updateTime="1728643685052" updateUser="ExternalLinkService">
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<mods:titleInfo id="94977C8EDD26A9CE46116B0F2FA87808">
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<mods:title id="B5BE7548957F31E3379E29A6CD5583B1">Overview of the lady beetle tribe Diomini (Coleoptera: Coccinellidae) and description of a new phytophagous, silk-spinning genus from Costa Rica that induces food bodies on leaves of Piper (Piperaceae)</mods:title>
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<mods:namePart id="CA663B6D41DF2E9169829A789C5D5358">Vandenberg, Natalia J.</mods:namePart>
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<mods:name id="F3E9EF3C5C2552468D90C8C314676BA2" type="personal">
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<mods:roleTerm id="ECD965E614F5E96C49954F3F06156522">Author</mods:roleTerm>
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<mods:namePart id="A3D9331F6F4FF4C9FA941B65D203387A">Hanson, Paul E.</mods:namePart>
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<mods:typeOfResource id="E28A892A99CB89F39E91C60588FEF347">text</mods:typeOfResource>
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<mods:title id="DEA984CF8569608F3170C19A3021DF3F">Zootaxa</mods:title>
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<mods:part id="43EC1FD48FDA6770EFDF031467B13FD3">
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<mods:date id="753EE4479C5D5B764A9EFEA69E7AA4D8">2019</mods:date>
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<mods:number id="5A7B4A6FAE0BCA031352FCD376AE3478">2019-02-08</mods:number>
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<mods:number id="CF99196742E01963B0B35CED7619B059">4554</mods:number>
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<mods:identifier id="4BF9BD76EA367E23E9820E7AF294317F" type="DOI">10.11646/zootaxa.4554.1.9</mods:identifier>
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<treatment id="DF18F971FFD70B3DFF4BFD236E61BE8C" ID-DOI="http://doi.org/10.5281/zenodo.5921888" ID-GBIF-Taxon="154535008" ID-Zenodo-Dep="5921888" LSID="urn:lsid:plazi:treatment:DF18F971FFD70B3DFF4BFD236E61BE8C" httpUri="http://treatment.plazi.org/id/DF18F971FFD70B3DFF4BFD236E61BE8C" lastPageId="22" lastPageNumber="277" pageId="21" pageNumber="276">
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<subSubSection id="1FAB1BECFFD70B3EFF4BFD236DABBED2" box="[151,511,730,757]" pageId="21" pageNumber="276" type="nomenclature">
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<paragraph id="570E4867FFD70B3EFF4BFD236DABBED2" blockId="21.[151,511,730,790]" box="[151,511,730,757]" pageId="21" pageNumber="276">
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<heading id="0C46FF0BFFD70B3EFF4BFD236DABBED2" bold="true" box="[151,511,730,757]" fontSize="11" level="1" pageId="21" pageNumber="276" reason="1">
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<emphasis id="65C59475FFD70B3EFF4BFD236DABBED2" bold="true" box="[151,511,730,757]" pageId="21" pageNumber="276">
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<taxonomicName id="90B133E4FFD70B3EFF4BFD236DCFBED3" authorityName="Vandenberg & Hanson" authorityYear="2019" box="[151,411,730,756]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="276" phylum="Arthropoda" rank="species" species="atropos" status="sp. nov.">
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<emphasis id="65C59475FFD70B3EFF4BFD236DCFBED3" bold="true" box="[151,411,730,756]" italics="true" pageId="21" pageNumber="276">Moiradiomus atropos</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="7EF6290EFFD70B3EFE7EFD226DABBED2" box="[418,511,731,757]" pageId="21" pageNumber="276" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="1FAB1BECFFD70B3EFF4BFD046DF8BF31" box="[151,428,765,790]" pageId="21" pageNumber="276" type="description">
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<paragraph id="570E4867FFD70B3EFF4BFD046DF8BF31" blockId="21.[151,511,730,790]" box="[151,428,765,790]" pageId="21" pageNumber="276">
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(
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<figureCitation id="CF8A54E2FFD70B3EFF43FD046D72BF31" box="[159,294,765,790]" captionStart="FIGURES 17–20" captionStartId="14.[151,264,1965,1988]" captionTargetBox="[151,1436,260,1911]" captionTargetId="figure@14.[151,1436,260,1911]" captionTargetPageId="14" captionText="FIGURES 17–20. Moiradiomus species habitus illustrations. 17, M. atropos: holotype, male; 18, M. atropos: paratype, female. 19, M. nanita: holotype, male; 20, M. nanita: paratype, female. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.2623573" httpUri="https://zenodo.org/record/2623573/files/figure.png" pageId="21" pageNumber="276">Figs. 17, 18</figureCitation>
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,
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<figureCitation id="CF8A54E2FFD70B3EFEEEFD076D05BF31" box="[306,337,766,790]" captionStart="FIGURES 25–28" captionStartId="15.[151,264,1975,1998]" captionTargetBox="[151,1436,1466,1918]" captionTargetId="figure@15.[151,1436,1461,1924]" captionTargetPageId="15" captionText="FIGURES 25–28. Diagrammatic ventral view of Moiradiomus prosterna (setae, color patterns, and surface punctation not indicated): 25, M. clotho; 26, M. lachesis; 27, M. atopos; 28, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623578" httpUri="https://zenodo.org/record/2623578/files/figure.png" pageId="21" pageNumber="276">27</figureCitation>
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,
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<figureCitation id="CF8A54E2FFD70B3EFE81FD076D2FBF31" box="[349,379,766,790]" captionStart="FIGURES 29–32" captionStartId="16.[151,264,1700,1723]" captionTargetBox="[154,1436,190,1666]" captionTargetId="figure@16.[151,1436,189,1667]" captionTargetPageId="16" captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="21" pageNumber="276">31</figureCitation>
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,
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<figureCitation id="CF8A54E2FFD70B3EFE5AFD076DF7BF31" box="[390,419,766,790]" captionStart="FIGURES 40–42" captionStartId="20.[151,264,1957,1980]" captionTargetBox="[151,1436,193,1936]" captionTargetId="figure@20.[151,1436,193,1936]" captionTargetPageId="20" captionText="FIGURES 40–42. Tents of Moiradiomus spp.: 40, M. lachesis on Piper lanosibracteum; 41, M. nanita on P. reticulatum; 42, M. atropos on P. friedrichsthalii." figureDoi="http://doi.org/10.5281/zenodo.2623584" httpUri="https://zenodo.org/record/2623584/files/figure.png" pageId="21" pageNumber="276">42</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="1FAB1BECFFD70B3EFF4BFCBC6D1FBFCA" pageId="21" pageNumber="276" type="diagnosis">
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<paragraph id="570E4867FFD70B3EFF4BFCBC6D1FBFCA" blockId="21.[151,1437,837,2020]" pageId="21" pageNumber="276">
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<emphasis id="65C59475FFD70B3EFF4BFCBC6D45BF79" bold="true" box="[151,273,837,862]" pageId="21" pageNumber="276">Diagnosis.</emphasis>
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This species is readily distinguished from its congeners by the male genitalia which have the basal lobe distinctly humped in lateral view and apex of parameres obliquely truncate. In addition, the adult form is more slender and apically tapered and the prosternal carinae are weakly convergent, framing an elongate ovoid depression with broadly rounded apex. It is also distinguished by its larval host plant:
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<taxonomicName id="90B133E4FFD70B3EFB84FC4B6D10BFC9" authority="Casimir de Condolle, 1869" authorityName="Casimir de Condolle" authorityYear="1869" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="21" pageNumber="276" phylum="Tracheophyta" rank="species" species="friedrichsthalii">
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<emphasis id="65C59475FFD70B3EFB84FC4B694FBFED" box="[1112,1307,945,970]" italics="true" pageId="21" pageNumber="276">P. friedrichsthalii</emphasis>
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Casimir de Condolle, 1869
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="1FAB1BECFFD70B3DFF1BFC006801BD88" lastPageId="22" lastPageNumber="277" pageId="21" pageNumber="276" type="description">
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<paragraph id="570E4867FFD70B3EFF1BFC006E70BB5F" blockId="21.[151,1437,837,2020]" pageId="21" pageNumber="276">
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<emphasis id="65C59475FFD70B3EFF1BFC006EE8B835" bold="true" box="[199,700,1017,1042]" pageId="21" pageNumber="276">
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Description of
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<typeStatus id="880AF6C5FFD70B3EFE5DFC006DBCB835" box="[385,488,1017,1042]" pageId="21" pageNumber="276" type="holotype">holotype</typeStatus>
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(male) (
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<figureCitation id="CF8A54E2FFD70B3EFD88FC006EF9B835" box="[596,685,1017,1042]" captionStart="FIGURES 17–20" captionStartId="14.[151,264,1965,1988]" captionTargetBox="[151,1436,260,1911]" captionTargetId="figure@14.[151,1436,260,1911]" captionTargetPageId="14" captionText="FIGURES 17–20. Moiradiomus species habitus illustrations. 17, M. atropos: holotype, male; 18, M. atropos: paratype, female. 19, M. nanita: holotype, male; 20, M. nanita: paratype, female. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.2623573" httpUri="https://zenodo.org/record/2623573/files/figure.png" pageId="21" pageNumber="276">Fig. 17</figureCitation>
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).
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</emphasis>
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Length
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<quantity id="9049E582FFD70B3EFCF8FC036FD5B836" box="[804,897,1018,1042]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="21" pageNumber="276" unit="mm" value="1.7">1.7 mm</quantity>
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, width 1.0 mm. Form elongate, oval, tapered apically, weakly convex. Metathoracic wing present. Dorsal surfaces feebly shining, distinctly punctate, pubescent with fimbriate borders. Head, pronotum, appendages including mouthparts, straw yellow, with meso- metacoxae slightly darkened; base of pronotum in front of scutellar shield with light brown transverse mark; scutellar shield light brown; elytron light brown with apex narrowly yellow brown, epipleuron yellow brown, nearly transparent; pubescence shiny, off white. Venter predominantly light brown; prosternum pale yellow; mesepimeron, mesepisternum paler than surrounding sclerites except posterior margin of mesepimeron narrowly darkened; abdomen light brown, lighter, yellower toward apex; tarsal claw, mandibular apex dark reddish amber. Dorsal punctation moderately coarse; punctures separated by approximately 1× diameter, on elytron equal in width to eye facet, finer on head, pronotum. Dorsal pubescence semi erect, moderately dense, evenly distributed, lacking distinct setal pattern; individual setae equal to about 2/3 to 1× length of scutellar shield, arcuate; elytral setae mostly directed posteriorly except directed outwardly at sides of body, posterolaterally near suture. Head large, 0.80× width of pronotum; eye large, finely facetted, with interfacetal setae, weakly notched near antennal insertion by small rounded ocular canthus; inner orbits arcuate, strongly diverging toward vertex, weakly diverging toward clypeus, with minimum separation at slightly below midlength; interocular distance 1.5× width of eye in frontal view. Antenna composed of 10 antennomeres; antennomere 3 elongate, about 2.3× length of antennomere 4; last 4 antennomeres forming gradual club expanding apically from antennomere 7–9; antennomere 10 subrhomboidal, slightly narrower, slightly longer than 9. Maxillary palp with terminal palpomere short, strongly expanded distally; with oblique apex. Pronotum transverse, width 2.0× length, weakly, evenly convex. Elytron in dorsal view with lateral margin weakly arcuate; elytral apices dehiscent. Ventral surfaces pubescent, distinctly punctate, except glabrous, impunctate in posterior 4/5ths of metacoxal plate; punctation coarser, denser on abdomen. Prosternum (
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<figureCitation id="CF8A54E2FFD70B3EFF43F9146CA6BB21" box="[159,242,1773,1798]" captionStart="FIGURES 3–12" captionStartId="12.[151,264,1784,1807]" captionTargetBox="[151,1433,212,1735]" captionTargetId="figure@12.[151,1436,193,1763]" captionTargetPageId="12" captionText="FIGURES 3–12. Genitalia of Diomini and Selvadiina (Hyperaspidini): 3–6, female genitalia, dorsal view (spermatheca shaded gray): 3, Selvadius nunenmacheri Gordon 1970 (Hyperaspidini: Selvadiina), showing distal end of sperm duct and spermatheca, accessory gland not shown (after Gordon 1985); 4, Erratodiomus brindisi Gordon 1999 (Hyperaspidini: Selvadiina), bursa through spermatheca, accessory gland not shown (after Gordon 1999); 5, Decadiomus hughesi Gordon & Hilburn 1990 (Diomini), bursa through spermatheca (note long convoluted sperm duct); 6, M. lachesis (Diomini) (A.g.= accessory gland); 7–12, male genitalia of Diomini, left lateral view of penis: 7, D. hughesi; 8, M. lachesis; 9, Diomus donatus Gordon (capsule); 10, M. lachesis (capsule); 11, Decadiomus bigemmeus (capsule); 12, Decadiomus liebecki (capsule)" figureDoi="http://doi.org/10.5281/zenodo.2623569" httpUri="https://zenodo.org/record/2623569/files/figure.png" pageId="21" pageNumber="276">Fig. 12</figureCitation>
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) with carinae of intercoxal process extending nearly to apex, weakly convergent, framing elongate ovoid depression with broadly rounded apex. Suture between abdominal ventrites 1 and 2 distinct, linear; 5
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<superScript id="A0C4E52FFFD70B3EFAD6F8E96942BB39" attach="none" box="[1290,1302,1808,1822]" fontSize="6" pageId="21" pageNumber="276">th</superScript>
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, 6
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<superScript id="A0C4E52FFFD70B3EFAEFF8E9696BBB39" attach="left" box="[1331,1343,1808,1822]" fontSize="6" pageId="21" pageNumber="276">th</superScript>
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ventrite with apex shallowly, roundly emarginate in median ½. Tarsal claw with broad scythe-like inner tooth extending from base to apical 1/3
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<superScript id="A0C4E52FFFD70B3EFE4BF8A26DF0BB4E" attach="left" box="[407,420,1883,1897]" fontSize="6" pageId="21" pageNumber="276">rd</superScript>
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or beyond.
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</paragraph>
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<paragraph id="570E4867FFD70B3DFF1BF87A6CA7BCF2" blockId="21.[151,1437,837,2020]" lastBlockId="22.[151,1436,151,683]" lastPageId="22" lastPageNumber="277" pageId="21" pageNumber="276">
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<emphasis id="65C59475FFD70B3EFF1BF87A6DB6BBBB" bold="true" box="[199,482,1923,1948]" pageId="21" pageNumber="276">
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Male genitalia (
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<figureCitation id="CF8A54E2FFD70B3EFE5CF87A6D80BBBB" box="[384,468,1923,1948]" captionStart="FIGURES 29–32" captionStartId="16.[151,264,1700,1723]" captionTargetBox="[154,1436,190,1666]" captionTargetId="figure@16.[151,1436,189,1667]" captionTargetPageId="16" captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="21" pageNumber="276">Fig. 31</figureCitation>
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).
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</emphasis>
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Basal lobe in ventral view (
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<figureCitation id="CF8A54E2FFD70B3EFCC2F87A6F29BBBB" box="[798,893,1923,1948]" captionStart="FIGURES 29–32" captionStartId="16.[151,264,1700,1723]" captionTargetBox="[154,1436,190,1666]" captionTargetId="figure@16.[151,1436,189,1667]" captionTargetPageId="16" captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="21" pageNumber="276">Fig. 31b</figureCitation>
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) weakly, unevenly tapered toward oblique apex, with small thorn-like process at distal most point, distinctly shorter than paramere; in lateral view (
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<figureCitation id="CF8A54E2FFD70B3EFB27F85E690DBB98" box="[1275,1369,1959,1984]" captionStart="FIGURES 29–32" captionStartId="16.[151,264,1700,1723]" captionTargetBox="[154,1436,190,1666]" captionTargetId="figure@16.[151,1436,189,1667]" captionTargetPageId="16" captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="21" pageNumber="276">Fig. 31a</figureCitation>
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) with strong hump, apex flat; paramere elongate, strongly expanded toward obliquely truncated apex, about 2.5× as long as wide, about 2× longer than longest setae. Penis (
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<figureCitation id="CF8A54E2FFD40B3DFD07FF6E6F6EBC97" box="[731,826,151,176]" captionStart="FIGURES 29–32" captionStartId="16.[151,264,1700,1723]" captionTargetBox="[154,1436,190,1666]" captionTargetId="figure@16.[151,1436,189,1667]" captionTargetPageId="16" captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="22" pageNumber="277">Fig. 31c</figureCitation>
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) lacking apical flagellum, slender with wedge-shaped capsule.
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</paragraph>
|
||
<paragraph id="570E4867FFD40B3DFF1BFF26682BBD64" blockId="22.[151,1436,151,683]" pageId="22" pageNumber="277">
|
||
<emphasis id="65C59475FFD40B3DFF1BFF266DC0BCDF" bold="true" box="[199,404,223,248]" pageId="22" pageNumber="277">
|
||
Female (
|
||
<figureCitation id="CF8A54E2FFD40B3DFEF3FF266DD1BCDF" box="[303,389,223,248]" captionStart="FIGURES 17–20" captionStartId="14.[151,264,1965,1988]" captionTargetBox="[151,1436,260,1911]" captionTargetId="figure@14.[151,1436,260,1911]" captionTargetPageId="14" captionText="FIGURES 17–20. Moiradiomus species habitus illustrations. 17, M. atropos: holotype, male; 18, M. atropos: paratype, female. 19, M. nanita: holotype, male; 20, M. nanita: paratype, female. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.2623573" httpUri="https://zenodo.org/record/2623573/files/figure.png" pageId="22" pageNumber="277">Fig. 18</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Similar to male except pronotum with brown blotch occupying median 2/3; pronotal base with narrow brown margin. 5
|
||
<superScript id="A0C4E52FFFD40B3DFE03FEFA6DBFBD36" attach="left" box="[479,491,259,273]" fontSize="6" pageId="22" pageNumber="277">th</superScript>
|
||
abdominal ventrite with posterior margin truncate; 6
|
||
<superScript id="A0C4E52FFFD40B3DFBE7FEFA6813BD36" attach="left" box="[1083,1095,259,273]" fontSize="6" pageId="22" pageNumber="277">th</superScript>
|
||
ventrite with posterior margin arcuate. Tarsal claw with short triangular tooth near base, not extending beyond apical ½.
|
||
</paragraph>
|
||
<paragraph id="570E4867FFD40B3DFF1BFEB76CAEBDAC" blockId="22.[151,1436,151,683]" pageId="22" pageNumber="277">
|
||
<emphasis id="65C59475FFD40B3DFF1BFEB76D68BD40" bold="true" box="[199,316,334,359]" pageId="22" pageNumber="277">Variation.</emphasis>
|
||
Length
|
||
<quantity id="9049E582FFD40B3DFE41FEB76E7CBD40" box="[413,552,334,359]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="22" pageNumber="277" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.5–1.7 mm</quantity>
|
||
. Individuals vary in the intensity of brown pigmentation on dorsal and ventral surfaces.
|
||
</paragraph>
|
||
<paragraph id="570E4867FFD40B3DFF1BFE6F6801BD88" blockId="22.[151,1436,151,683]" box="[199,1109,406,431]" pageId="22" pageNumber="277">
|
||
<emphasis id="65C59475FFD40B3DFF1BFE6F6DCEBD88" bold="true" box="[199,410,406,431]" pageId="22" pageNumber="277">Trophic relations.</emphasis>
|
||
Larvae develop on food bodies of
|
||
<taxonomicName id="90B133E4FFD40B3DFCFDFE616FB1BD88" authorityName="Casimir de Condolle" authorityYear="1869" box="[801,997,406,431]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="22" pageNumber="277" phylum="Tracheophyta" rank="species" species="friedrichsthalii">
|
||
<emphasis id="65C59475FFD40B3DFCFDFE616FB1BD88" box="[801,997,406,431]" italics="true" pageId="22" pageNumber="277">P. friedrichsthalii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="CF8A54E2FFD40B3DFC28FE6F6812BD88" box="[1012,1094,406,431]" captionStart="FIGURES 40–42" captionStartId="20.[151,264,1957,1980]" captionTargetBox="[151,1436,193,1936]" captionTargetId="figure@20.[151,1436,193,1936]" captionTargetPageId="20" captionText="FIGURES 40–42. Tents of Moiradiomus spp.: 40, M. lachesis on Piper lanosibracteum; 41, M. nanita on P. reticulatum; 42, M. atropos on P. friedrichsthalii." figureDoi="http://doi.org/10.5281/zenodo.2623584" httpUri="https://zenodo.org/record/2623584/files/figure.png" pageId="22" pageNumber="277">Fig. 42</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1FAB1BECFFD40B3DFF1BFE436F23BE3D" pageId="22" pageNumber="277" type="etymology">
|
||
<paragraph id="570E4867FFD40B3DFF1BFE436F23BE3D" blockId="22.[151,1436,151,683]" pageId="22" pageNumber="277">
|
||
<emphasis id="65C59475FFD40B3DFF1BFE436D1ABDF4" bold="true" box="[199,334,442,467]" pageId="22" pageNumber="277">Etymology.</emphasis>
|
||
Atropos, proper noun in apposition. Ancient Greek Ἄτρoπoς (Atropos), literally meaning inflexible or not turning, from the Greek a-, (not) + tropos, (to turn). In Greek mythology, one of the three Fates, or Moirae; the cutter of the thread of life, depicted as an old woman.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1FAB1BECFFD40B3DFF1BFDDF6E61BE8C" pageId="22" pageNumber="277" type="materials_examined">
|
||
<paragraph id="570E4867FFD40B3DFF1BFDDF6E61BE8C" blockId="22.[151,1436,151,683]" pageId="22" pageNumber="277">
|
||
<emphasis id="65C59475FFD40B3DFF1BFDDF6D23BE18" bold="true" box="[199,375,550,575]" pageId="22" pageNumber="277">Type material.</emphasis>
|
||
<materialsCitation id="E7D9423AFFD40B3DFE5CFDDF6E61BE8C" ID-GBIF-Occurrence="2234228025" collectingDate="2014-04-16" collectingDateMax="2017-05" collectingDateMin="2014-04-16" collectionCode="USNM, MZUCR" collectorName="Res. Biol. A. Brenes & P. Hanson & E. Chacon" country="Costa Rica" elevation="900" location="San Ramon" pageId="22" pageNumber="277" specimenCount="10" specimenCount-female="1" specimenCount-male="4" stateProvince="Alajuela" typeStatus="holotype">
|
||
<typeStatus id="880AF6C5FFD40B3DFE5CFDDF6DBCBE18" box="[384,488,550,575]" pageId="22" pageNumber="277" type="holotype">Holotype</typeStatus>
|
||
(male) with labels: “
|
||
<collectingCountry id="2FA608F7FFD40B3DFD00FDDE6FDDBE18" box="[732,905,551,575]" name="Costa Rica" pageId="22" pageNumber="277">COSTA RICA</collectingCountry>
|
||
:
|
||
<collectingRegion id="95758685FFD40B3DFC4AFDDF6FADBE18" box="[918,1017,550,575]" country="Costa Rica" name="Alajuela" pageId="22" pageNumber="277">Alajuela</collectingRegion>
|
||
:
|
||
<collectorName id="FA442DB1FFD40B3DFBDAFDD168A0BE18" box="[1030,1268,550,575]" pageId="22" pageNumber="277">Res. Biol. A. Brenes</collectorName>
|
||
(
|
||
<location id="526E1EBCFFD40B3DFADAFDDE69C4BE18" LSID="urn:lsid:plazi:treatment:DF18F971FFD70B3DFF4BFD236E61BE8C:526E1EBCFFD40B3DFADAFDDE69C4BE18" box="[1286,1424,551,575]" country="Costa Rica" name="San Ramon" pageId="22" pageNumber="277" stateProvince="Alajuela">San Ramon</location>
|
||
),
|
||
<quantity id="9049E582FFD40B3DFF4BFDB26C83BE44" box="[151,215,587,611]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="22" pageNumber="277" unit="m" value="900.0">
|
||
<elevation id="DC9CAF54FFD40B3DFF4BFDB26C83BE44" box="[151,215,587,611]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="22" pageNumber="277" unit="m" value="900.0">900m</elevation>
|
||
</quantity>
|
||
<date id="230F6EA7FFD40B3DFF01FDB36D08BE44" box="[221,348,586,611]" pageId="22" pageNumber="277" value="2014-04-16">
|
||
<collectingDate id="334B974FFFD40B3DFF01FDB36D08BE44" box="[221,348,586,611]" pageId="22" pageNumber="277" value="2014-04-16">16.IV.2014</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="FA442DB1FFD40B3DFEBAFDB26D81BE45" box="[358,469,587,610]" pageId="22" pageNumber="277">P. Hanson</collectorName>
|
||
/ ex.
|
||
<taxonomicName id="90B133E4FFD40B3DFDCEFDB56F22BE44" authority="" (USNM)" baseAuthorityName="USNM" box="[530,886,586,611]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="22" pageNumber="277" phylum="Tracheophyta" rank="species" species="friedrichsthalii">
|
||
<emphasis id="65C59475FFD40B3DFDCEFDB56EABBE44" box="[530,767,586,611]" italics="true" pageId="22" pageNumber="277">Piper friedrichsthalii</emphasis>
|
||
” (
|
||
<collectionCode id="31A0D0A2FFD40B3DFCC6FDB26F3ABE44" box="[794,878,587,611]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="22" pageNumber="277" type="Museum">USNM</collectionCode>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<specimenCount id="41B783EEFFD40B3DFC77FDB3687EBE44" box="[939,1066,586,611]" pageId="22" pageNumber="277" type="generic" typeStatus="paratypes">5 paratypes</specimenCount>
|
||
(
|
||
<specimenCount id="41B783EEFFD40B3DFBE5FDB268C7BE44" box="[1081,1171,586,611]" pageId="22" pageNumber="277" type="male">4 males</specimenCount>
|
||
,
|
||
<specimenCount id="41B783EEFFD40B3DFB42FDB26950BE44" box="[1182,1284,586,611]" pageId="22" pageNumber="277" type="female">1 female</specimenCount>
|
||
): 1 with same labels as
|
||
<typeStatus id="880AF6C5FFD40B3DFF21FD976D37BEA0" box="[253,355,622,647]" pageId="22" pageNumber="277" type="holotype">holotype</typeStatus>
|
||
; 4 with labels: “
|
||
<collectingCountry id="2FA608F7FFD40B3DFDC8FD966EEABEA0" box="[532,702,623,647]" name="Costa Rica" pageId="22" pageNumber="277">COSTA RICA</collectingCountry>
|
||
:
|
||
<collectingRegion id="95758685FFD40B3DFD15FD976F78BEA0" box="[713,812,622,647]" country="Costa Rica" name="Alajuela" pageId="22" pageNumber="277">Alajuela</collectingRegion>
|
||
:
|
||
<collectorName id="FA442DB1FFD40B3DFCEAFD89684CBEA0" box="[822,1048,622,647]" pageId="22" pageNumber="277">Res. Biol. A.Brenes</collectorName>
|
||
, ex.
|
||
<taxonomicName id="90B133E4FFD40B3DFB97FD89696CBEA0" box="[1099,1336,622,647]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="22" pageNumber="277" phylum="Tracheophyta" rank="species" species="friedrichsthalii">
|
||
<emphasis id="65C59475FFD40B3DFB97FD89696CBEA0" box="[1099,1336,622,647]" italics="true" pageId="22" pageNumber="277">Piper friedrichsthalii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<date id="230F6EA7FFD40B3DFA99FD9669C3BEA0" box="[1349,1431,623,647]" pageId="22" pageNumber="277" value="2017-05">
|
||
<collectingDate id="334B974FFFD40B3DFA99FD9669C3BEA0" box="[1349,1431,623,647]" pageId="22" pageNumber="277" value="2017-05">V.2017</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="FA442DB1FFD40B3DFF4BFD6A6D5ABE8C" box="[151,270,658,683]" pageId="22" pageNumber="277">E. Chacon</collectorName>
|
||
” (3,
|
||
<collectionCode id="31A0D0A2FFD40B3DFE98FD6A6DCDBE8C" box="[324,409,659,683]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="22" pageNumber="277" type="Museum">USNM</collectionCode>
|
||
; 2,
|
||
<collectionCode id="31A0D0A2FFD40B3DFE1CFD6A6E7DBE8C" box="[448,553,659,683]" pageId="22" pageNumber="277">MZUCR</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |