241 lines
32 KiB
XML
241 lines
32 KiB
XML
<document id="52FD15D721A9DC49BD2B56F8BF66F9D8" ID-DOI="10.5281/zenodo.5753778" ID-ISSN="1638-9387" ID-Zenodo-Dep="5753778" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630331563133" checkinUser="marcus" docAuthor="Lourenço, Wilson R. & Goodman, Steven M." docDate="2008" docId="DE2D87E5FFB2FF8F7642FD9D406AF93B" docLanguage="en" docName="Zoosystema.2008.30.3.665-679.pdf" docOrigin="Zoosystema 30 (3)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Opisthacanthus milloti Lourenço & Goodman 2008, n. sp." docType="treatment" docVersion="3" lastPageNumber="673" masterDocId="2214FF9DFFB6FF877550FFA94207FF9F" masterDocTitle="Scorpions of the Réserve spéciale d’Ankarana, Madagascar, with particular reference to cave-dwelling animals and the description of two new species (Arachnida, Scorpiones)" masterLastPageNumber="679" masterPageNumber="665" pageNumber="669" updateTime="1699230852284" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="6D9CB1F58D6A979BC7D64E250EED7959">
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<mods:title id="03B107A794451BFE503D27816CB0054B">Scorpions of the Réserve spéciale d’Ankarana, Madagascar, with particular reference to cave-dwelling animals and the description of two new species (Arachnida, Scorpiones)</mods:title>
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</mods:titleInfo>
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<mods:name id="4F4EFCBB9643A8BD104C7840439EB043" type="personal">
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<mods:role id="BF134C4947EC88683F7954F3BDC7F16A">
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<mods:roleTerm id="245EE36638A47054074569556DE6C3D5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="113EB12EE2A67350820B5531F1EB1AE7">Lourenço, Wilson R.</mods:namePart>
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<mods:affiliation id="8CD2261294C9E02CFCB01E1426EB1818">Muséum national d’Histoire naturelle, Département Systématique et Évolution, USM 0602, section Arthropodes (Arachnologie), case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) arachne @ mnhn. fr</mods:affiliation>
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<mods:nameIdentifier id="A599E0D5DE5F62FAB8E47FD589FB9DA1" type="email">arachne@mnhn.fr</mods:nameIdentifier>
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</mods:name>
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<mods:name id="177EB8C61DC8F343BFD46B1B9D8B00EB" type="personal">
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<mods:role id="7190EE1592847487AF63EE68AF546EB2">
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<mods:roleTerm id="E5BBCA5DDBD971C0CE5B1B5D26E70A3F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="32FA64C51D5A07C91E3137309977F02C">Goodman, Steven M.</mods:namePart>
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<mods:affiliation id="E8BF8E3C1AEBE5DE933AAA943B680C8C">Field Museum of Natural History, 1400 South Lake Shore Drive, Chicago, Illinois 60605 (USA) and Vahatra, BP 3972, Antananarivo (101) (Madagascar) sgoodman @ fieldmuseum. org</mods:affiliation>
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<mods:typeOfResource id="98A623C67E0B468B167EB28AECB62381">text</mods:typeOfResource>
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<mods:title id="CCDE473CB3921E9ABF0C140605C2D4A5">Zoosystema</mods:title>
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</mods:titleInfo>
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<mods:part id="23225ABC1BEFF36B09257F450EF82FED">
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<mods:date id="C2DD7CF6BA58AFD08721B4DA256C90D9">2008</mods:date>
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<mods:detail id="2951604385578CD12F20145EACEC007C" type="volume">
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<mods:number id="55991D66FF7FE05C44E1CB4FA58AC125">30</mods:number>
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</mods:detail>
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<mods:detail id="1E914E95B683070337120849EE51FD9E" type="issue">
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<mods:number id="854F5576FF402E2EA7B0D8AD8E92D86D">3</mods:number>
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<mods:extent id="2FAE1E59DAFF49AFC10E8EC22520AAA5" unit="page">
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<mods:start id="64ACE4586A8432DFCB71936A594EE909">665</mods:start>
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<mods:end id="9AA5933393C4CAABD151F4062D7C7595">679</mods:end>
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</mods:extent>
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<mods:classification id="E59F65038749FB4A02B4D22C5F871AAA">journal article</mods:classification>
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<mods:identifier id="5A334313A2A1A1697239183797788C7A" type="DOI">10.5281/zenodo.5753778</mods:identifier>
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<mods:identifier id="C7D74290C559B81CB8BB7AE1012C9467" type="ISSN">1638-9387</mods:identifier>
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</mods:mods>
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<treatment id="DE2D87E5FFB2FF8F7642FD9D406AF93B" LSID="urn:lsid:plazi:treatment:DE2D87E5FFB2FF8F7642FD9D406AF93B" httpUri="http://treatment.plazi.org/id/DE2D87E5FFB2FF8F7642FD9D406AF93B" lastPageId="8" lastPageNumber="673" pageId="4" pageNumber="669">
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<subSubSection id="1E9E6578FFB2FF837642FD9D464BFDD0" box="[786,1100,564,591]" pageId="4" pageNumber="669" type="nomenclature">
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<paragraph id="563B36F3FFB2FF837642FD9D464BFDD0" blockId="4.[786,1100,564,623]" box="[786,1100,564,591]" pageId="4" pageNumber="669">
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<heading id="0D73819FFFB2FF837642FD9D464BFDD0" box="[786,1100,564,591]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="669" reason="2">
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<taxonomicName id="91844D70FFB2FF837642FD9D460CFDD0" ID-CoL="6T2TH" authority="Lourenço & Goodman, 2008" authorityName="Lourenço & Goodman" authorityYear="2008" box="[786,1035,564,591]" class="Arachnida" family="Hormuridae" genus="Opisthacanthus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="669" phylum="Arthropoda" rank="species" species="milloti" status="sp. nov.">
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<emphasis id="64F0EAE1FFB2FF837642FD9D460CFDD0" bold="true" box="[786,1035,564,591]" italics="true" pageId="4" pageNumber="669">Opisthacanthus milloti</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="7FC3579AFFB2FF837142FD9D464BFDD0" box="[1042,1100,564,591]" pageId="4" pageNumber="669" rank="species">n. sp.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="1E9E6578FFB2FF83762BFDFD41E4FDF0" box="[891,995,596,623]" pageId="4" pageNumber="669" type="description">
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<paragraph id="563B36F3FFB2FF83762BFDFD41E4FDF0" blockId="4.[786,1100,564,623]" box="[891,995,596,623]" pageId="4" pageNumber="669">
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(
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<figureCitation id="CEBF2A76FFB2FF8376D2FDFD41DCFDF0" box="[898,987,596,623]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="5.[113,124,1658,1675]" captionStartId-1="6.[677,688,1448,1465]" captionStartId-2="7.[113,124,1635,1652]" captionTargetBox-0="[215,1097,215,1567]" captionTargetBox-1="[680,1208,236,1372]" captionTargetBox-2="[214,1099,218,1592]" captionTargetId-0="figure-21@5.[188,757,727,1277]" captionTargetId-1="figure-16@6.[675,1212,213,1384]" captionTargetId-2="figure-17@7.[205,1118,141,1655]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIG. 2. — Opisthacanthus milloti n. sp., ♂ holotype and ♀ paratype:A, carapace; B, chelicera; C, hemispermatophore external aspect (♂); D, E, metasomal segment V and telson, lateral aspect (♂ and ♀). Scale bars: 2 mm." captionText-1="FIG. 3. — Opisthacanthus milloti n. sp., ventral aspect, showing coxapophysis,sternum,genital operculum,pectines and sternite III: A, ♂ holotype; B, ♀ paratype. Scale bar: 2 mm." captionText-2="FIG. 4. — Opisthacanthus milloti n. sp., ♂ holotype, trichobothrial pattern: A-C, chela dorso-external, external and ventral aspects; D, femur, dorsal aspect; E, fixed finger, internal aspect; F-H, patella, dorsal, external and ventral aspects; I, cutting edge of movable finger with rows of granules. Scale bars: 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5753780" figureDoi-1="http://doi.org/10.5281/zenodo.5753782" figureDoi-2="http://doi.org/10.5281/zenodo.5753784" httpUri-0="https://zenodo.org/record/5753780/files/figure.png" httpUri-1="https://zenodo.org/record/5753782/files/figure.png" httpUri-2="https://zenodo.org/record/5753784/files/figure.png" pageId="4" pageNumber="669">Figs 2-4</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="1E9E6578FFB2FF8377F5FD244176FC90" pageId="4" pageNumber="669" type="materials_examined">
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<paragraph id="563B36F3FFB2FF8377F5FD244176FC90" blockId="4.[677,1211,653,783]" pageId="4" pageNumber="669">
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<collectionCode id="3095AE36FFB2FF8377F5FD244130FD3B" box="[677,823,653,676]" pageId="4" pageNumber="669">TYPE MATERIAL</collectionCode>
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. —
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<emphasis id="64F0EAE1FFB2FF837633FD2441D9FD3A" bold="true" box="[867,990,653,677]" pageId="4" pageNumber="669">
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<collectingCountry id="2E937663FFB2FF837633FD2441DDFD3A" box="[867,986,653,677]" name="Madagascar" pageId="4" pageNumber="669">Madagascar</collectingCountry>
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.
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</emphasis>
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<collectingRegion id="9440F811FFB2FF8376B4FD2446B1FD3A" box="[996,1206,653,677]" country="Madagascar" name="Antsiranana" pageId="4" pageNumber="669">Antsiranana Province</collectingRegion>
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, Réserve spéciale d’Ankarana, in the entry of the grotte des Chauve-souris, found resting on soil under stone,
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<date id="223A1033FFB2FF8377F6FD7540FCFD6B" box="[678,763,732,756]" pageId="4" pageNumber="669" value="2001-09">IX.2001</date>
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, W.
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<collectionCode id="3095AE36FFB2FF837675FD75413FFD6B" box="[805,824,732,756]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="669">R</collectionCode>
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. Lourenço coll.,
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<specimenCount id="4082FD7AFFB2FF837688FD7541F8FD6A" box="[984,1023,732,757]" pageId="4" pageNumber="669" type="male">1 ♂</specimenCount>
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<typeStatus id="893F8851FFB2FF837154FD75465BFD6B" box="[1028,1116,732,756]" pageId="4" pageNumber="669" type="holotype">holotype</typeStatus>
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,
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<specimenCount id="4082FD7AFFB2FF837136FD75468BFD6A" box="[1126,1164,732,757]" pageId="4" pageNumber="669" type="male">1 ♂</specimenCount>
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,
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<specimenCount id="4082FD7AFFB2FF8371C7FD7546BCFD6A" box="[1175,1211,732,757]" pageId="4" pageNumber="669" type="female">1 ♀</specimenCount>
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<typeStatus id="893F8851FFB2FF8377F5FD5E4106FC90" box="[677,769,759,783]" pageId="4" pageNumber="669" type="paratype">paratypes</typeStatus>
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(
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<collectionCode id="3095AE36FFB2FF83765EFD5E4161FC90" box="[782,870,759,783]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="669">MNHN</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="1E9E6578FFB2FF8377F5FC8E4130FB54" pageId="4" pageNumber="669" type="diagnosis">
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<paragraph id="563B36F3FFB2FF8377F5FC8E469DFB8D" blockId="4.[675,1211,807,1227]" pageId="4" pageNumber="669">
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DIAGNOSIS. — Small size scorpions: males
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||
<quantity id="917C9B16FFB2FF837117FC8E4688FCA0" box="[1095,1167,807,831]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="4" pageNumber="669" unit="mm" value="36.0">36 mm</quantity>
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and female
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<quantity id="917C9B16FFB2FF8377B8FCE8412AFCC6" box="[744,813,833,857]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="4" pageNumber="669" unit="mm" value="32.0">32 mm</quantity>
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||
in total length. Coloration reddish-yellow to reddish-brown with some blackish areas over pedipalp carinae. Pectines with 6-6 teeth in males and female. Hemispermatophore small, with the distal lamina weakly enlarged and longer than that observed for
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<taxonomicName id="91844D70FFB2FF837165FC0246BFFC5C" authorityName="Lourenco & Goodman" authorityYear="2006" box="[1077,1208,939,963]" class="Arachnida" family="Hormuridae" genus="Opisthacanthus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="669" phylum="Arthropoda" rank="species" species="darainensis">
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<emphasis id="64F0EAE1FFB2FF837165FC0246BFFC5C" box="[1077,1208,939,963]" italics="true" pageId="4" pageNumber="669">O. darainensis</emphasis>
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</taxonomicName>
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(
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<figureCitation id="CEBF2A76FFB2FF8377FBFC6C40F2FC42" box="[683,757,965,989]" captionStart="FIG" captionStartId="5.[113,124,1658,1675]" captionTargetBox="[215,1097,215,1567]" captionTargetId="figure-21@5.[188,757,727,1277]" captionTargetPageId="5" captionText="FIG. 2. — Opisthacanthus milloti n. sp., ♂ holotype and ♀ paratype:A, carapace; B, chelicera; C, hemispermatophore external aspect (♂); D, E, metasomal segment V and telson, lateral aspect (♂ and ♀). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5753780" httpUri="https://zenodo.org/record/5753780/files/figure.png" pageId="4" pageNumber="669">Fig. 2C</figureCitation>
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). Female genital operculum large, with an almost oval-shape, and lacking incision in the base (
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<figureCitation id="CEBF2A76FFB2FF83710FFC4946ABFC67" box="[1119,1196,992,1016]" captionStart="FIG" captionStartId="6.[677,688,1448,1465]" captionTargetBox="[680,1208,236,1372]" captionTargetId="figure-16@6.[675,1212,213,1384]" captionTargetPageId="6" captionText="FIG. 3. — Opisthacanthus milloti n. sp., ventral aspect, showing coxapophysis,sternum,genital operculum,pectines and sternite III: A, ♂ holotype; B, ♀ paratype. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5753782" httpUri="https://zenodo.org/record/5753782/files/figure.png" pageId="4" pageNumber="669">Fig. 3B</figureCitation>
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). Trichobothrial pattern of
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<typeStatus id="893F8851FFB2FF8376CFFC5341CEFB8D" box="[927,969,1018,1042]" pageId="4" pageNumber="669">type</typeStatus>
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C, orthobothriotaxy.
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</paragraph>
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<paragraph id="563B36F3FFB2FF8377EBFBBC4130FB54" blockId="4.[675,1211,807,1227]" pageId="4" pageNumber="669">
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The new species can be distinguished from
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<taxonomicName id="91844D70FFB2FF837115FBBC40BEFBD8" authorityName="Lourenco & Goodman" authorityYear="2006" class="Arachnida" family="Hormuridae" genus="Opisthacanthus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="669" phylum="Arthropoda" rank="species" species="darainensis">
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<emphasis id="64F0EAE1FFB2FF837115FBBC40BEFBD8" italics="true" pageId="4" pageNumber="669">O. darainensis</emphasis>
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</taxonomicName>
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, described from the more southerly Loky-Manambato (= Daraina) region, by the following characters: 1) smaller global size; 2) distinct morphometric values (see Table 1); 3) presence of four moderate carinae on sternite VII; and 4) distal lamina of hemispermatophore narrower and elongated.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="1E9E6578FFB2FF8377F5FB4A41E3FAAF" pageId="4" pageNumber="669" type="etymology">
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<paragraph id="563B36F3FFB2FF8377F5FB4A41E3FAAF" blockId="4.[677,1211,1251,1328]" pageId="4" pageNumber="669">ETYMOLOGY. — Patronym in honour of the late Prof. Jacques Millot, who devoted many years of his life to the study of the Malagasy fauna.</paragraph>
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</subSubSection>
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||
<subSubSection id="1E9E6578FFB2FF8F77F5FAC4406AF93B" lastPageId="8" lastPageNumber="673" pageId="4" pageNumber="669" type="description">
|
||
<paragraph id="563B36F3FFB2FF8377F5FAC44132FA1A" blockId="4.[674,1212,1389,1700]" box="[677,821,1389,1415]" pageId="4" pageNumber="669">DESCRIPTION</paragraph>
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<paragraph id="563B36F3FFB2FF8377F5FA244642FA39" blockId="4.[674,1212,1389,1700]" box="[677,1093,1420,1447]" pageId="4" pageNumber="669">
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(based on
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<specimenCount id="4082FD7AFFB2FF837643FA26412EFA3A" box="[787,809,1423,1445]" pageId="4" pageNumber="669" type="male">♂</specimenCount>
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||
<typeStatus id="893F8851FFB2FF837661FA244197FA38" box="[817,912,1421,1447]" pageId="4" pageNumber="669" type="holotype">holotype</typeStatus>
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||
and
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<specimenCount id="4082FD7AFFB2FF837696FA2641DEFA3A" box="[966,985,1423,1445]" pageId="4" pageNumber="669" type="female">♀</specimenCount>
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<typeStatus id="893F8851FFB2FF8376B0FA254639FA39" box="[992,1086,1420,1446]" pageId="4" pageNumber="669" type="paratype">paratype</typeStatus>
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||
)
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||
</paragraph>
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||
<paragraph id="563B36F3FFB2FF8377F5FA024114FA5A" blockId="4.[674,1212,1389,1700]" box="[677,787,1451,1477]" pageId="4" pageNumber="669">
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||
<emphasis id="64F0EAE1FFB2FF8377F5FA024114FA5A" box="[677,787,1451,1477]" italics="true" pageId="4" pageNumber="669">Coloration</emphasis>
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||
</paragraph>
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||
<paragraph id="563B36F3FFB2FF8177F5FA624341FD8F" blockId="4.[674,1212,1389,1700]" lastBlockId="6.[111,649,216,528]" lastPageId="6" lastPageNumber="671" pageId="4" pageNumber="669">Basically reddish-yellow to reddish-brown with some blackish areas on the pedipalp carinae. Carapace reddish-brown with a paler section on the posterior edge; median and lateral eyes surrounded with black pigment.Tergites reddish-brown with two vestigial longitudinal pale yellow spots. Metasomal segments reddish-brown, with some vestigial yellowish variegated areas; vesicle yellowish,with lateral reddish bands on males, but absent on female; aculeus dark reddish. Chelicerae reddish-brown; base of fingers blackish; the whole surface with diffuse variegated dark spots; fingers blackish with reddish teeth. Pedipalps reddish-brown; most carinae blackish. Venter: sternites brownish-yellow; coxapophysis and sternum reddish-brown; pectines and genital operculum yellowish; legs reddish-yellow with diffused dark spots.</paragraph>
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<caption id="02FB667BFFB3FF827521F9D3414CF93E" ID-DOI="http://doi.org/10.5281/zenodo.5753780" ID-Zenodo-Dep="5753780" httpUri="https://zenodo.org/record/5753780/files/figure.png" pageId="5" pageNumber="670" startId="5.[113,124,1658,1675]" targetBox="[215,1097,215,1567]" targetPageId="5">
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||
<paragraph id="563B36F3FFB3FF827521F9D3414CF93E" blockId="5.[113,1209,1657,1698]" pageId="5" pageNumber="670">
|
||
FIG. 2. —
|
||
<taxonomicName id="91844D70FFB3FF827590F9D04370F915" authority="Lourenço & Goodman, 2008" authorityName="Lourenço & Goodman" authorityYear="2008" box="[192,375,1657,1674]" class="Arachnida" family="Hormuridae" genus="Opisthacanthus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="670" phylum="Arthropoda" rank="species" species="milloti" status="sp. nov.">
|
||
<emphasis id="64F0EAE1FFB3FF827590F9D04370F915" box="[192,375,1657,1674]" italics="true" pageId="5" pageNumber="670">Opisthacanthus milloti</emphasis>
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||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FC3579AFFB3FF82742CF9D343AAF914" box="[380,429,1658,1675]" pageId="5" pageNumber="670" rank="species">n. sp.</taxonomicNameLabel>
|
||
, ♂ holotype and ♀ paratype:
|
||
<emphasis id="64F0EAE1FFB3FF8277CCF9D040AEF915" bold="true" box="[668,681,1657,1674]" pageId="5" pageNumber="670">A</emphasis>
|
||
, carapace;
|
||
<emphasis id="64F0EAE1FFB3FF82765AF9D04110F915" bold="true" box="[778,791,1657,1674]" pageId="5" pageNumber="670">B</emphasis>
|
||
, chelicera;
|
||
<emphasis id="64F0EAE1FFB3FF827626F9D04183F915" bold="true" box="[886,900,1657,1674]" pageId="5" pageNumber="670">C</emphasis>
|
||
, hemispermatophore external aspect (♂);
|
||
<emphasis id="64F0EAE1FFB3FF8275C4F93942A5F93E" bold="true" box="[148,162,1680,1697]" pageId="5" pageNumber="670">D</emphasis>
|
||
,
|
||
<emphasis id="64F0EAE1FFB3FF8275FCF93942BFF93E" bold="true" box="[172,184,1680,1697]" pageId="5" pageNumber="670">E</emphasis>
|
||
, metasomal segment V and telson, lateral aspect (♂ and ♀). Scale bars: 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="563B36F3FFB0FF817521FD9D42EBFDD1" blockId="6.[111,649,564,1700]" box="[113,236,564,590]" pageId="6" pageNumber="671">
|
||
<emphasis id="64F0EAE1FFB0FF817521FD9D42EBFDD1" box="[113,236,564,590]" italics="true" pageId="6" pageNumber="671">Morphology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="563B36F3FFB0FF817521FDFD4083F93B" blockId="6.[111,649,564,1700]" pageId="6" pageNumber="671">
|
||
Carapace smooth with intense punctation; lateral edges, in males, with some minute granulations; furrows shallow. Anterior margin with a strong concavity reaching as far as the level of the second lateral eye (
|
||
<figureCitation id="CEBF2A76FFB0FF817427FD7A43D7FD71" box="[375,464,723,750]" captionStart="FIG" captionStartId="5.[113,124,1658,1675]" captionTargetBox="[215,1097,215,1567]" captionTargetId="figure-21@5.[188,757,727,1277]" captionTargetPageId="5" captionText="FIG. 2. — Opisthacanthus milloti n. sp., ♂ holotype and ♀ paratype:A, carapace; B, chelicera; C, hemispermatophore external aspect (♂); D, E, metasomal segment V and telson, lateral aspect (♂ and ♀). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5753780" httpUri="https://zenodo.org/record/5753780/files/figure.png" pageId="6" pageNumber="671">Fig. 2A</figureCitation>
|
||
). Median ocular tubercle flattened and almost in the center of the carapace; median eyes moderate to small, separated by less than one ocular diameter; three pairs of lateral eyes. Sternum pentagonal, wider than long. Genital operculum formed by two semi-oval plates in males, and one single large, almost oval-like shaped plate in females (
|
||
<figureCitation id="CEBF2A76FFB0FF817767FC19407DFC54" box="[567,634,944,971]" captionStart="FIG" captionStartId="6.[677,688,1448,1465]" captionTargetBox="[680,1208,236,1372]" captionTargetId="figure-16@6.[675,1212,213,1384]" captionTargetPageId="6" captionText="FIG. 3. — Opisthacanthus milloti n. sp., ventral aspect, showing coxapophysis,sternum,genital operculum,pectines and sternite III: A, ♂ holotype; B, ♀ paratype. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5753782" httpUri="https://zenodo.org/record/5753782/files/figure.png" pageId="6" pageNumber="671">Fig. 3</figureCitation>
|
||
). Tergites with one vestigial median carina, and with intense punctation. Pectinal tooth count 6-
|
||
<quantity id="917C9B16FFB0FF817706FC594083FB94" box="[598,644,1008,1035]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="6" pageNumber="671" unit="in" value="6.0">6 in</quantity>
|
||
male
|
||
<typeStatus id="893F8851FFB0FF8175E1FBB94316FBB4" box="[177,273,1040,1067]" pageId="6" pageNumber="671" type="holotype">holotype</typeStatus>
|
||
and in female
|
||
<typeStatus id="893F8851FFB0FF8174ECFBB94019FBB4" box="[444,542,1040,1067]" pageId="6" pageNumber="671" type="paratype">paratype</typeStatus>
|
||
. Sternites smooth and shiny, with punctations laterally; VII with four moderate carinae. Metasomal segments I to V longer than wide with sparse granulations. All carinae weakly developed in segments I-IV of male, moderately evident in female; segment V slightly rounded with spinoids granules on ventro-lateral and ventro-median carinae. All segments with moderate setation, more pronounced in males. Telson with a pear-like shape; smooth and covered with strong setation (
|
||
<figureCitation id="CEBF2A76FFB0FF81741EFAE543C6FAF8" box="[334,449,1356,1383]" captionStart="FIG" captionStartId="5.[113,124,1658,1675]" captionTargetBox="[215,1097,215,1567]" captionTargetId="figure-21@5.[188,757,727,1277]" captionTargetPageId="5" captionText="FIG. 2. — Opisthacanthus milloti n. sp., ♂ holotype and ♀ paratype:A, carapace; B, chelicera; C, hemispermatophore external aspect (♂); D, E, metasomal segment V and telson, lateral aspect (♂ and ♀). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5753780" httpUri="https://zenodo.org/record/5753780/files/figure.png" pageId="6" pageNumber="671">Fig. 2D, E</figureCitation>
|
||
). Pedipalps: femur with dorsal internal, dorsal external, ventral internal, and ventral external carinae strong, tuberculate; dorsal face with moderately marked granulation; ventral face with a thin granulation; internal face moderately granulose. Patella with internal and external faces weakly granulated; dorsal and ventral faces smooth and lustrous, with punctations; dorsal internal, dorsal external, ventral internal, ventral external, and external carinae moderate to strong; other carinae less well-marked. Chela strongly
|
||
</paragraph>
|
||
<paragraph id="563B36F3FFB0FF8F7791FF74435FF999" blockId="6.[705,722,221,246]" lastBlockId="8.[111,648,1293,1542]" lastPageId="8" lastPageNumber="673" pageId="6" pageNumber="671">
|
||
A granular on dorso-internal and external faces; other faces punctated; dorsal marginal, external secondary, ventro-internal and ventral median carina moderate to strong; other carinae less well-marked. Chelicerae typical of Scorpionoidea (
|
||
<bibRefCitation id="32154B02FFBEFF8F74BAFAA4407DFAB7" author="VACHON M." box="[490,634,1293,1320]" pageId="8" pageNumber="673" pagination="161 - 166" refId="ref7003" refString="VACHON M. 1963. - De l'utilite, en systematique, d'une nomenclature des dents des cheliceres chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, Paris, 2 e ser., 35 (2): 161 - 166." type="journal article" year="1963">Vachon 1963</bibRefCitation>
|
||
); teeth sharp (
|
||
<figureCitation id="CEBF2A76FFBEFF8F75ACFA844354FAD7" box="[252,339,1325,1352]" captionStart="FIG" captionStartId="5.[113,124,1658,1675]" captionTargetBox="[215,1097,215,1567]" captionTargetId="figure-21@5.[188,757,727,1277]" captionTargetPageId="5" captionText="FIG. 2. — Opisthacanthus milloti n. sp., ♂ holotype and ♀ paratype:A, carapace; B, chelicera; C, hemispermatophore external aspect (♂); D, E, metasomal segment V and telson, lateral aspect (♂ and ♀). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5753780" httpUri="https://zenodo.org/record/5753780/files/figure.png" pageId="8" pageNumber="673">Fig. 2B</figureCitation>
|
||
). Trichobothriotaxy
|
||
<typeStatus id="893F8851FFBEFF8F7769FA84406FFAD7" box="[569,616,1325,1352]" pageId="8" pageNumber="673">type</typeStatus>
|
||
C (
|
||
<figureCitation id="CEBF2A76FFBEFF8F752BFAE542B9FAF8" box="[123,190,1356,1383]" captionStart="FIG" captionStartId="7.[113,124,1635,1652]" captionTargetBox="[214,1099,218,1592]" captionTargetId="figure-17@7.[205,1118,141,1655]" captionTargetPageId="7" captionText="FIG. 4. — Opisthacanthus milloti n. sp., ♂ holotype, trichobothrial pattern: A-C, chela dorso-external, external and ventral aspects; D, femur, dorsal aspect; E, fixed finger, internal aspect; F-H, patella, dorsal, external and ventral aspects; I, cutting edge of movable finger with rows of granules. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5753784" httpUri="https://zenodo.org/record/5753784/files/figure.png" pageId="8" pageNumber="673">Fig. 4</figureCitation>
|
||
A-H); orthobothriotaxic (
|
||
<bibRefCitation id="32154B02FFBEFF8F748FFAE5407BFAF8" author="VACHON M." box="[479,636,1356,1383]" pageId="8" pageNumber="673" pagination="857 - 958" refId="ref7084" refString="VACHON M. 1974. - Etude des caracteres utilises pour classer les familles et les genres de Scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, Paris 3 e ser., nº 140, Zoologie 104: 857 - 958 (date 1973, publie en 1974)." type="journal article" year="1974">Vachon 1974</bibRefCitation>
|
||
). Legs: tarsi of legs III and IV with two prolateral and two retrolateral spines, surrounded by a few long setae. Spurs moderate. Hemispermatophore as in
|
||
<figureCitation id="CEBF2A76FFBEFF8F75FCFA62431CFA79" box="[172,283,1483,1510]" captionStart="FIG" captionStartId="5.[113,124,1658,1675]" captionTargetBox="[215,1097,215,1567]" captionTargetId="figure-21@5.[188,757,727,1277]" captionTargetPageId="5" captionText="FIG. 2. — Opisthacanthus milloti n. sp., ♂ holotype and ♀ paratype:A, carapace; B, chelicera; C, hemispermatophore external aspect (♂); D, E, metasomal segment V and telson, lateral aspect (♂ and ♀). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5753780" httpUri="https://zenodo.org/record/5753780/files/figure.png" pageId="8" pageNumber="673">Figure 2C</figureCitation>
|
||
with the distal lamina elongated and weakly enlarged.
|
||
</paragraph>
|
||
<caption id="02FB667BFFB0FF8177F5FA014629FA78" ID-DOI="http://doi.org/10.5281/zenodo.5753782" ID-Zenodo-Dep="5753782" httpUri="https://zenodo.org/record/5753782/files/figure.png" pageId="6" pageNumber="671" startId="6.[677,688,1448,1465]" targetBox="[680,1208,236,1372]" targetPageId="6">
|
||
<paragraph id="563B36F3FFB0FF8177F5FA014629FA78" blockId="6.[675,1212,1448,1511]" pageId="6" pageNumber="671">
|
||
FIG. 3. —
|
||
<taxonomicName id="91844D70FFB0FF8177A8FA0141B6FA26" authority="Lourenço & Goodman, 2008" authorityName="Lourenço & Goodman" authorityYear="2008" box="[760,945,1448,1465]" class="Arachnida" family="Hormuridae" genus="Opisthacanthus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="671" phylum="Arthropoda" rank="species" species="milloti" status="sp. nov.">
|
||
<emphasis id="64F0EAE1FFB0FF8177A8FA0141B6FA26" box="[760,945,1448,1465]" italics="true" pageId="6" pageNumber="671">Opisthacanthus milloti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FC3579AFFB0FF8176E7FA0141EDFA26" box="[951,1002,1448,1465]" pageId="6" pageNumber="671" rank="species">n. sp.</taxonomicNameLabel>
|
||
, ventral aspect, showing coxapophysis,sternum,genital operculum,pectines and sternite III:
|
||
<emphasis id="64F0EAE1FFB0FF8177F3FA7C40B7FA79" bold="true" box="[675,688,1493,1510]" pageId="6" pageNumber="671">A</emphasis>
|
||
, ♂ holotype;
|
||
<emphasis id="64F0EAE1FFB0FF817671FA7C4129FA79" bold="true" box="[801,814,1493,1510]" pageId="6" pageNumber="671">B</emphasis>
|
||
, ♀ paratype. Scale bar: 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="02FB667BFFB1FF807521F9CA43FDF93E" ID-DOI="http://doi.org/10.5281/zenodo.5753784" ID-Zenodo-Dep="5753784" httpUri="https://zenodo.org/record/5753784/files/figure.png" pageId="7" pageNumber="672" startId="7.[113,124,1635,1652]" targetBox="[214,1099,218,1592]" targetPageId="7">
|
||
<paragraph id="563B36F3FFB1FF807521F9CA43FDF93E" blockId="7.[113,1211,1635,1697]" pageId="7" pageNumber="672">
|
||
FIG. 4. —
|
||
<taxonomicName id="91844D70FFB1FF807594F9CA437AF9EB" authority="Lourenço & Goodman, 2008" authorityName="Lourenço & Goodman" authorityYear="2008" box="[196,381,1635,1652]" class="Arachnida" family="Hormuridae" genus="Opisthacanthus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="672" phylum="Arthropoda" rank="species" species="milloti" status="sp. nov.">
|
||
<emphasis id="64F0EAE1FFB1FF807594F9CA437AF9EB" box="[196,381,1635,1652]" italics="true" pageId="7" pageNumber="672">Opisthacanthus milloti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FC3579AFFB1FF8074D3F9CA43B1F9EB" box="[387,438,1635,1652]" pageId="7" pageNumber="672" rank="species">n. sp.</taxonomicNameLabel>
|
||
, ♂ holotype, trichobothrial pattern:
|
||
<emphasis id="64F0EAE1FFB1FF8077B1F9CA4104F9EB" bold="true" box="[737,771,1635,1652]" pageId="7" pageNumber="672">A -C</emphasis>
|
||
, chela dorso-external, external and ventral aspects;
|
||
<emphasis id="64F0EAE1FFB1FF807521F9D04278F915" bold="true" box="[113,127,1657,1674]" pageId="7" pageNumber="672">D</emphasis>
|
||
, femur, dorsal aspect;
|
||
<emphasis id="64F0EAE1FFB1FF80746EF9D0434DF915" bold="true" box="[318,330,1657,1674]" pageId="7" pageNumber="672">E</emphasis>
|
||
, fixed finger, internal aspect;
|
||
<emphasis id="64F0EAE1FFB1FF80776FF9D04058F915" bold="true" box="[575,607,1657,1675]" pageId="7" pageNumber="672">F -H</emphasis>
|
||
, patella, dorsal, external and ventral aspects;
|
||
<emphasis id="64F0EAE1FFB1FF80768FF9D041E2F915" bold="true" box="[991,997,1657,1674]" pageId="7" pageNumber="672">I</emphasis>
|
||
, cutting edge of movable finger with rows of granules. Scale bars: 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="02FB667BFFBEFF8F741DFB6C41D8FB49" box="[333,991,1220,1238]" pageId="8" pageNumber="673" startId="8.[333,344,1221,1238]">
|
||
<paragraph id="563B36F3FFBEFF8F741DFB6C41D8FB49" blockId="8.[333,991,1220,1238]" box="[333,991,1220,1238]" pageId="8" pageNumber="673">
|
||
FIG. 5. —
|
||
<taxonomicName id="91844D70FFBEFF8F74CDFB6D4063FB4A" authority="Lourenço & Goodman, 2008" authorityName="Lourenço & Goodman" authorityYear="2008" box="[413,612,1220,1237]" class="Arachnida" family="Hormuridae" genus="Opisthacanthus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="673" phylum="Arthropoda" rank="species" species="pauliani" status="sp. nov.">
|
||
<emphasis id="64F0EAE1FFBEFF8F74CDFB6D4063FB4A" box="[413,612,1220,1237]" italics="true" pageId="8" pageNumber="673">Opisthacanthus pauliani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FC3579AFFBEFF8F7739FB6C409CFB49" box="[617,667,1221,1238]" pageId="8" pageNumber="673" rank="species">n. sp.</taxonomicNameLabel>
|
||
, ♂ holotype, habitus. Scale bar: 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="563B36F3FFBEFF8F7521F9834369F9DB" blockId="8.[111,648,1578,1700]" box="[113,366,1578,1604]" pageId="8" pageNumber="673">
|
||
<emphasis id="64F0EAE1FFBEFF8F7521F9834369F9DB" box="[113,366,1578,1604]" italics="true" pageId="8" pageNumber="673">Ecology and distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="563B36F3FFBEFF8F7521F9E3406AF93B" blockId="8.[111,648,1578,1700]" pageId="8" pageNumber="673">
|
||
This new species is only known from its
|
||
<typeStatus id="893F8851FFBEFF8F7756F9E34034F9FA" box="[518,563,1610,1637]" pageId="8" pageNumber="673">type</typeStatus>
|
||
locality. Because it was collected at the cave entrance only, it is not to be considered as a troglobitic element.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |