treatments-xml/data/DE/03/F4/DE03F47E9E400407FF19FCA1FDB4FA09.xml

<document id="C9EF0B84E4888CFAA614A64DDE6D9A3A" ID-DOI="10.11646/zootaxa.4032.2.1" ID-GBIF-Dataset="0f892064-2fb2-4422-a234-0ac163f4e1a7" ID-ISSN="1175-5326" ID-Zenodo-Dep="253740" ID-ZooBank="26E6B554-D7F1-4125-AC40-E40D838354B9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461293095294" checkinUser="plazi" docAuthor="Serrano, Artur R. M., Capela, Rúben A. &amp; Oesterle, Andreas" docDate="2015" docId="DE03F47E9E400407FF19FCA1FDB4FA09" docLanguage="en" docName="zt04032p178.pdf" docOrigin="Zootaxa 4032 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trichotaenia pepetela Serrano &amp; Capela, sp. n." docType="treatment" docVersion="8" lastPageNumber="164" masterDocId="223A8C069E47040AFF8EFFC9FFC1FE3C" masterDocTitle="Three new species of tiger beetles and new data on Cicindelina species from Angola (Coleoptera: Carabidae: Cicindelinae)" masterLastPageNumber="178" masterPageNumber="151" pageNumber="158" updateTime="1698616006471" updateUser="plazi">
<mods:mods id="951EF9DCA431E73F204E5FF28D7284C5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D0AAB040ABF6C8AF4974B5917D44E3A0">
<mods:title id="41AC6307404D139677B8FBC79F05FE0C">Three new species of tiger beetles and new data on Cicindelina species from Angola (Coleoptera: Carabidae: Cicindelinae)</mods:title>
</mods:titleInfo>
<mods:name id="A547F37B9992DAE87DA2F070D743072F" type="personal">
<mods:role id="819C0B3925B4BF68894411AC89E2162E">
<mods:roleTerm id="17803E27B6A56FA81E5DEE97714296E4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BF1C82CCACD3E2A1AE309DB34989542E">Serrano, Artur R. M.</mods:namePart>
</mods:name>
<mods:name id="54E2AB401A2268C51F1D43505A1FC4B8" type="personal">
<mods:role id="DF7CEB0E2F1F7BC90D45D86F50B593BD">
<mods:roleTerm id="42A6E0A2016B4994ACC530DB327E7732">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A6190B35DF8F5306FD49958761614548">Capela, Rúben A.</mods:namePart>
</mods:name>
<mods:name id="32F21049F3B91B8E6E13266AD451E33A" type="personal">
<mods:role id="ED3FB628EC23D2B028A79D5FAB1B17D4">
<mods:roleTerm id="BA9902F1BD091507F7C0F9C6373C8432">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="19FF823355835E5E274612F16CB30593">Oesterle, Andreas</mods:namePart>
</mods:name>
<mods:typeOfResource id="AC4B4D1A4DA58ADDC9540D946EC88B51">text</mods:typeOfResource>
<mods:relatedItem id="6FFA13A329B1D17D839C36C4963AFC48" type="host">
<mods:titleInfo id="7E320F73CB44F62F67ADD540DF6A1F48">
<mods:title id="2A1B7B380ECE0E6B5FEC180728509A63">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9F11E3A760B6171C6A3F678707650521">
<mods:date id="7D48698E4AF6C71430C104D24DD0B9F0">2015</mods:date>
<mods:detail id="7FCBD68B99267D808FFF10B078768B6B" type="volume">
<mods:number id="EA0C3857B629E8104191CA886CC7C239">4032</mods:number>
</mods:detail>
<mods:detail id="566390B853F900B9310C155CDF5BD0A6" type="issue">
<mods:number id="822D0A8B43534F4FF8A41763ABC6F93B">2</mods:number>
</mods:detail>
<mods:extent id="DF700E7F9E33EF6BA2041702362DBDF4" unit="page">
<mods:start id="0736846A0D5DB73D0F208E1DA8834DD6">151</mods:start>
<mods:end id="A302884BFE7800525E14CB0E7E874F0B">178</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="0702DC72155AFF7132535CBF5CD053C2">journal article</mods:classification>
<mods:identifier id="82016C5C00A24D0B0CB4F5A3A0900071" type="DOI">10.11646/zootaxa.4032.2.1</mods:identifier>
<mods:identifier id="5B26462E1EC03BE2C329679B6EE56A4C" type="GBIF-Dataset">0f892064-2fb2-4422-a234-0ac163f4e1a7</mods:identifier>
<mods:identifier id="90FB8AF20DEADFEDB81FD9004BC87CF7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B062F7F156C23398A8E0595C6A0146D5" type="Zenodo-Dep">253740</mods:identifier>
<mods:identifier id="8564D5D5DE9E9DDDC9778A8AF6B4E379" type="ZooBank">26E6B554-D7F1-4125-AC40-E40D838354B9</mods:identifier>
</mods:mods>
<treatment id="DE03F47E9E400407FF19FCA1FDB4FA09" ID-DOI="http://doi.org/10.5281/zenodo.5658899" ID-GBIF-Taxon="119642059" ID-Zenodo-Dep="5658899" LSID="urn:lsid:plazi:treatment:DE03F47E9E400407FF19FCA1FDB4FA09" httpUri="http://treatment.plazi.org/id/DE03F47E9E400407FF19FCA1FDB4FA09" lastPageId="13" lastPageNumber="164" pageId="7" pageNumber="158">
<subSubSection id="1EB016E39E40040DFF19FCA1FEB8FD9F" pageId="7" pageNumber="158" type="nomenclature">
<paragraph id="561545689E40040DFF19FCA1FD10FDBE" blockId="7.[151,721,872,931]" box="[151,721,872,898]" pageId="7" pageNumber="158">
<heading id="0D5DF2049E40040DFF19FCA1FD10FDBE" bold="true" box="[151,721,872,898]" fontSize="11" level="1" pageId="7" pageNumber="158" reason="1">
<emphasis id="64DE997A9E40040DFF19FCA1FD10FDBE" bold="true" box="[151,721,872,898]" pageId="7" pageNumber="158">
<taxonomicName id="91AA3EEB9E40040DFF19FCA1FD46FDBE" authority="Serrano &amp; Capela" authorityName="Serrano &amp; Capela" box="[151,647,872,898]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="pepetela" status="sp. nov.">
<emphasis id="64DE997A9E40040DFF19FCA1FE5DFDBE" bold="true" box="[151,412,872,898]" italics="true" pageId="7" pageNumber="158">Trichotaenia pepetela</emphasis>
Serrano &amp; Capela
</taxonomicName>
<taxonomicNameLabel id="7FED24019E40040DFD01FCA1FD10FDBE" box="[655,721,872,898]" pageId="7" pageNumber="158" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="561545689E40040DFF19FC43FEB8FD9F" blockId="7.[151,721,872,931]" box="[151,377,906,931]" pageId="7" pageNumber="158">
(
<figureCitation id="CE9159ED9E40040DFF11FC43FF26FD9F" box="[159,231,906,931]" captionStart="FIGURE 4" captionStartId="8.[151,250,1885,1907]" captionTargetBox="[168,1422,570,1847]" captionTargetId="figure@8.[151,1436,553,1864]" captionTargetPageId="8" captionText="FIGURE 4. Facies of Trichotaenia pepetela sp. n.: a) male holotype, b) female allotype." httpUri="https://zenodo.org/record/253744/files/figure.png" pageId="7" pageNumber="158">Figs 4</figureCitation>
, 
<figureCitation id="CE9159ED9E40040DFF7DFC43FEC2FD9F" box="[243,259,906,931]" captionStart="FIGURE 5" captionStartId="9.[151,250,1276,1298]" captionTargetBox="[164,1424,208,1244]" captionTargetId="figure@9.[151,1435,193,1255]" captionTargetPageId="9" captionText="FIGURE 5. Trichotaenia pepetela sp. n., head, mandibles and labrum, pronotum (dorsal view): a, c) male; b, d) female." httpUri="https://zenodo.org/record/253745/files/figure.png" pageId="7" pageNumber="158">5</figureCitation>
, 
<figureCitation id="CE9159ED9E40040DFE81FC43FEDFFD9F" box="[271,286,906,931]" captionStart="FIGURE 6" captionStartId="10.[151,250,1973,1995]" captionTargetBox="[212,1369,576,1946]" captionTargetId="figure@10.[201,1386,567,1962]" captionTargetPageId="10" captionText="FIGURE 6. Aedeagus (lateral view): a) Trichotaenia nzingae sp. n., paratype, Kakande, Bié, Angola, b) Trichotaenia pepetela sp. n., holotype, Catota, Bié, Angola." httpUri="https://zenodo.org/record/253746/files/figure.png" pageId="7" pageNumber="158">6</figureCitation>
b, 7a, b)
</paragraph>
</subSubSection>
<subSubSection id="1EB016E39E40040DFF19FC1BFC9DFA0F" pageId="7" pageNumber="158" type="materials_examined">
<paragraph id="561545689E40040DFF19FC1BFC9DFA0F" blockId="7.[151,1437,977,2011]" pageId="7" pageNumber="158">
<emphasis id="64DE997A9E40040DFF19FC1BFEDFFDD7" bold="true" box="[151,286,978,1003]" pageId="7" pageNumber="158">
<typeStatus id="8911FBCA9E40040DFF19FC1BFF13FDD7" box="[151,210,978,1003]" pageId="7" pageNumber="158">Type</typeStatus>
series
</emphasis>
. 
<typeStatus id="8911FBCA9E40040DFEA3FC1AFE59FDD7" box="[301,408,979,1003]" pageId="7" pageNumber="158" type="holotype">Holotype</typeStatus>
, ♂; 
<collectingCountry id="2EBD05F89E40040DFE45FC1AFDDFFDD7" box="[459,542,979,1003]" name="Angola" pageId="7" pageNumber="158">Angola</collectingCountry>
(
<collectionCode id="30BBDDAD9E40040DFDA1FC18FDA0FDD7" box="[559,609,977,1003]" httpUri="http://grbio.org/cool/7k2m-q4ze" name="Instituto di Entomologia" pageId="7" pageNumber="158">BIÉ</collectionCode>
), Catota (coord.: 
<geoCoordinate id="339E23AF9E40040DFCA7FC1AFC29FDD7" box="[809,1000,979,1003]" direction="south" orientation="latitude" pageId="7" pageNumber="158" precision="1" value="-14.010325">14º 00´37.17´´ S</geoCoordinate>
, 
<geoCoordinate id="339E23AF9E40040DFC79FC1AFB00FDD6" box="[1015,1217,979,1003]" direction="east" orientation="longitude" pageId="7" pageNumber="158" precision="1" value="17.400091">17º 24´00.33´´ E</geoCoordinate>
, 
<quantity id="9152E88D9E40040DFB40FC1BFAE4FDD6" box="[1230,1317,978,1002]" metricMagnitude="3" metricUnit="m" metricValue="1.5319999999999998" pageId="7" pageNumber="158" unit="m" value="1532.0">1532 m</quantity>
alt., 323), (
<collectionCode id="30BBDDAD9E40040DFF11FC3FFF10FA33" box="[159,209,1014,1039]" httpUri="http://grbio.org/cool/7k2m-q4ze" name="Instituto di Entomologia" pageId="7" pageNumber="158">BIÉ</collectionCode>
), 
<date id="221463A89E40040DFF6AFC3EFE99FA33" box="[228,344,1015,1039]" pageId="7" pageNumber="158" value="2014-11-02">2.XI.2014</date>
, DO, A. Serrano leg. Allotype ♀ and 
<typeStatus id="8911FBCA9E40040DFD71FC3EFCAAFA33" box="[767,875,1015,1039]" pageId="7" pageNumber="158" type="paratype">paratypes</typeStatus>
same locality, 
<date id="221463A89E40040DFB9CFC3EFB44FA33" box="[1042,1157,1015,1039]" pageId="7" pageNumber="158" value="2014-11-02">2.XI.2014</date>
, 
<specimenCount id="40AC8EE19E40040DFB1FFC3EFB74FA33" box="[1169,1205,1015,1039]" pageId="7" pageNumber="158" type="generic">1♂</specimenCount>
, 1♀, 
<date id="221463A89E40040DFB7DFC3EFAA6FA33" box="[1267,1383,1015,1039]" pageId="7" pageNumber="158" value="2014-11-03">3.XI.2014</date>
, 
<specimenCount id="40AC8EE19E40040DFAFCFC3EFA56FA33" box="[1394,1431,1015,1039]" pageId="7" pageNumber="158" type="generic">1♂</specimenCount>
, DO, A. Serrano leg. 
<typeStatus id="8911FBCA9E40040DFEF0FBD3FE27FA0F" box="[382,486,1050,1075]" pageId="7" pageNumber="158" type="holotype">Holotype</typeStatus>
and 
<typeStatus id="8911FBCA9E40040DFD93FBD2FD49FA0F" box="[541,648,1051,1075]" pageId="7" pageNumber="158" type="paratype">paratypes</typeStatus>
deposited in ASC.
</paragraph>
</subSubSection>
<subSubSection id="1EB016E39E40040DFF49FBF7FAB1FAA3" pageId="7" pageNumber="158" type="etymology">
<paragraph id="561545689E40040DFF49FBF7FAB1FAA3" blockId="7.[151,1437,977,2011]" pageId="7" pageNumber="158">
<emphasis id="64DE997A9E40040DFF49FBF7FE60FA6B" bold="true" box="[199,417,1086,1111]" pageId="7" pageNumber="158">
<emphasis id="64DE997A9E40040DFF49FBF7FE5BFA6A" bold="true" box="[199,410,1086,1110]" italics="true" pageId="7" pageNumber="158">Derivatio nominis</emphasis>
.
</emphasis>
The specific epithet, 
<taxonomicName id="91AA3EEB9E40040DFD2DFBF7FCC3FA6B" box="[675,770,1086,1111]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="pepetela">
<emphasis id="64DE997A9E40040DFD2DFBF7FCC3FA6B" box="[675,770,1086,1111]" italics="true" pageId="7" pageNumber="158">pepetela</emphasis>
</taxonomicName>
, is used as a noun in apposition based on the pen name “ 
<taxonomicName id="91AA3EEB9E40040DFF2FFBABFEC4FA47" box="[161,261,1122,1147]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="pepetela" status="sp. nov.">Pepetela</taxonomicName>
” of Artur Carlos Pestana dos Santos, a great Angolan writer descent of Portuguese colones of Benguela, which fought together with MPLA (Popular Liberation Movement of 
<collectingCountry id="2EBD05F89E40040DFC2AFB4FFC39FAA3" box="[932,1016,1158,1183]" name="Angola" pageId="7" pageNumber="158">Angola</collectingCountry>
) for liberation of their homeland.
</paragraph>
</subSubSection>
<subSubSection id="1EB016E39E40040DFF49FB63FE81F8E2" pageId="7" pageNumber="158" type="diagnosis">
<paragraph id="561545689E40040DFF49FB63FE81F8E2" blockId="7.[151,1437,977,2011]" pageId="7" pageNumber="158">
<emphasis id="64DE997A9E40040DFF49FB63FEFBFAFF" bold="true" box="[199,314,1194,1219]" pageId="7" pageNumber="158">Diagnosis</emphasis>
. A winged 
<taxonomicName id="91AA3EEB9E40040DFE4EFB63FD8EFAFF" box="[448,591,1194,1219]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="64DE997A9E40040DFE4EFB63FD8EFAFF" box="[448,591,1194,1219]" italics="true" pageId="7" pageNumber="158">Trichotaenia</emphasis>
</taxonomicName>
species, black coloured with some slight punctual purpurish and bluish or violet reflections on head, pronotum and elytra. Clypeus mostly setose, blackish with some purpurish blue-greenish reflections. Labrum large, rounded and 5-toothed in front, very slightly setose on half posteriad surface with just four whitish marginal sensorial setae, only triangularly doubled blackened in the first basal third (males) or triangularly blackened in the two basal thirds (females). A stripe of dense white recumbent pubescence on genae, continuing behind on the ventral half of proepisterna up to the lateral sides of first five abdominal sternites, where it is much narrower. Sternum and abdominal sternites otherwise glabrous. Antennae barely attaining nearly the half of the elytral length in male, shorter in female; antennomeres 1-4 dark metallic black with bluish or violaceous reflections, antennomeres 5-10 strongly foliated. Elytral shoulders and parts of the disk glabrous. Elytral decumbent pubescence (
<figureCitation id="CE9159ED9E40040DFE27FA27FE31F83A" box="[425,496,1518,1543]" captionStart="FIGURE 7" captionStartId="11.[151,250,1499,1521]" captionTargetBox="[171,1423,206,1462]" captionTargetId="figure@11.[151,1437,193,1477]" captionTargetPageId="11" captionText="FIGURE 7. Elytra (dorsal view): a and b) Trichotaenia pepetela sp. n., male and female, respectively, c) Trichotaenia tereticollis (W. Horn), d) Trichotaenia pseudotereticollis (W. Horn), e) Trichotaenia suturata (W. Horn)." httpUri="https://zenodo.org/record/253747/files/figure.png" pageId="7" pageNumber="158">Figs 7</figureCitation>
a‒7b) covering the lateral sides, from the middle of basal third to the apex where it becomes slightly wider, emitting two rami on disc (one slightly arcuate above the apex and one almost straight and large near the middle). A small tuft of closely and diagonally appressed setae on each side of the base of scutellum and a subsutural elongate tuft of closely appressed setae below the scutellum. Absence of decumbent setae along the suture. Elytral decumbent pubescence leaving three naked areoles in between, one large above the middle stripe, the second one immediately below the middle stripe and a third one smaller between the subapical stripe and the apical area.
</paragraph>
</subSubSection>
<subSubSection id="1EB016E39E400401FF49F923FC35F8E3" lastPageId="11" lastPageNumber="162" pageId="7" pageNumber="158" type="description">
<paragraph id="561545689E40040DFF49F923FA50F93F" blockId="7.[151,1437,977,2011]" box="[199,1425,1770,1795]" pageId="7" pageNumber="158">
<emphasis id="64DE997A9E40040DFF49F923FE98F93F" bold="true" box="[199,345,1770,1795]" pageId="7" pageNumber="158">Description.</emphasis>
Length of 
<typeStatus id="8911FBCA9E40040DFE5AF923FD80F93F" box="[468,577,1770,1795]" pageId="7" pageNumber="158" type="holotype">Holotype</typeStatus>
: 
<quantity id="9152E88D9E40040DFDC5F922FD73F93E" box="[587,690,1771,1795]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="7" pageNumber="158" unit="mm" value="10.2">10.2 mm</quantity>
. Length of 
<typeStatus id="8911FBCA9E40040DFCBFF922FC60F93F" box="[817,929,1771,1795]" pageId="7" pageNumber="158" type="paratype">paratypes</typeStatus>
: 
<quantity id="9152E88D9E40040DFC25F922FBCEF93E" box="[939,1039,1771,1795]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="7" pageNumber="158" unit="mm" value="10.1">10.1 mm</quantity>
(males), 
<quantity id="9152E88D9E40040DFBF6F922FAD9F93E" box="[1144,1304,1771,1795]" metricMagnitude="-2" metricUnit="m" metricValue="1.1300000000000001" metricValueMax="1.1400000000000001" metricValueMin="1.1199999999999999" pageId="7" pageNumber="158" unit="mm" value="11.3" valueMax="11.4" valueMin="11.2">11.2–11.4 mm</quantity>
(females).
</paragraph>
<paragraph id="561545689E400402FF49F8C7FAE1FFC9" blockId="7.[151,1437,977,2011]" lastBlockId="8.[151,1437,151,501]" lastPageId="8" lastPageNumber="159" pageId="7" pageNumber="158">
<emphasis id="64DE997A9E40040DFF49F8C7FEC5F91B" box="[199,260,1806,1831]" italics="true" pageId="7" pageNumber="158">Head</emphasis>
(
<figureCitation id="CE9159ED9E40040DFE9CF8C7FE98F91B" box="[274,345,1806,1831]" captionStart="FIGURE 5" captionStartId="9.[151,250,1276,1298]" captionTargetBox="[164,1424,208,1244]" captionTargetId="figure@9.[151,1435,193,1255]" captionTargetPageId="9" captionText="FIGURE 5. Trichotaenia pepetela sp. n., head, mandibles and labrum, pronotum (dorsal view): a, c) male; b, d) female." httpUri="https://zenodo.org/record/253745/files/figure.png" pageId="7" pageNumber="158">Figs 5</figureCitation>
a‒5b). Wider than long [length: 
<quantity id="9152E88D9E40040DFD4EF8C7FCA2F91A" box="[704,867,1806,1831]" metricMagnitude="-3" metricUnit="m" metricValue="1.825" metricValueMax="2.02" metricValueMin="1.63" pageId="7" pageNumber="158" unit="mm" value="1.825" valueMax="2.02" valueMin="1.63">1.63–2.02 mm</quantity>
(males), 
<quantity id="9152E88D9E40040DFC45F8C7FBA6F91B" box="[971,1127,1806,1831]" metricMagnitude="-3" metricUnit="m" metricValue="2.065" metricValueMax="2.08" metricValueMin="2.05" pageId="7" pageNumber="158" unit="mm" value="2.065" valueMax="2.08" valueMin="2.05">2.05–2.08mm</quantity>
(females), width 
<quantity id="9152E88D9E40040DFAA2F8C7FF02F976" metricMagnitude="-3" metricUnit="m" metricValue="2.7649999999999997" metricValueMax="2.78" metricValueMin="2.75" pageId="7" pageNumber="158" unit="mm" value="2.7649999999999997" valueMax="2.78" valueMin="2.75">2.75–2.78 mm</quantity>
(males) and 
<quantity id="9152E88D9E40040DFEDCF8FAFE2FF977" box="[338,494,1843,1867]" metricMagnitude="-3" metricUnit="m" metricValue="3.07" metricValueMax="3.1" metricValueMin="3.04" pageId="7" pageNumber="158" unit="mm" value="3.07" valueMax="3.1" valueMin="3.04">3.04–3.10mm</quantity>
(females)], with large eyes, slightly narrower than body (
<figureCitation id="CE9159ED9E40040DFBE0F8FBFB73F977" box="[1134,1202,1842,1867]" captionStart="FIGURE 4" captionStartId="8.[151,250,1885,1907]" captionTargetBox="[168,1422,570,1847]" captionTargetId="figure@8.[151,1436,553,1864]" captionTargetPageId="8" captionText="FIGURE 4. Facies of Trichotaenia pepetela sp. n.: a) male holotype, b) female allotype." httpUri="https://zenodo.org/record/253744/files/figure.png" pageId="7" pageNumber="158">Fig. 4</figureCitation>
), dorsal colour black with coppery reflections; surface sculpture rugulose, rugae forming irregular longitudinal meshes on vertex and transverse meshes on occiput; rugae straight, more or less parallel on orbital plates and frons; occiput, vertex and frons covered with dense and short piceous decumbent setae, clypeus with sparsely decumbent setae, temples (lateral area of head behind eyes) indistinctly striaterugulose, glabrous, glossy; genae covered with dense white decumbent pubescence; orbital plates with two sublateral setigerous punctures; labrum large, shape nearly semicircular in both sexes (
<figureCitation id="CE9159ED9E4F0402FE46FF75FDCFFEE9" box="[456,526,188,213]" captionStart="FIGURE 5" captionStartId="9.[151,250,1276,1298]" captionTargetBox="[164,1424,208,1244]" captionTargetId="figure@9.[151,1435,193,1255]" captionTargetPageId="9" captionText="FIGURE 5. Trichotaenia pepetela sp. n., head, mandibles and labrum, pronotum (dorsal view): a, c) male; b, d) female." httpUri="https://zenodo.org/record/253745/files/figure.png" pageId="8" pageNumber="159">Figs 5</figureCitation>
a‒5b), transverse, wider than long [length: 
<quantity id="9152E88D9E4F0402FC61FF75FB53FEE8" box="[1007,1170,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.09" metricValueMax="1.12" metricValueMin="1.06" pageId="8" pageNumber="159" unit="mm" value="1.09" valueMax="1.12" valueMin="1.06">1.06–1.12 mm</quantity>
(males), 
<quantity id="9152E88D9E4F0402FB74FF75FA5CFEE8" box="[1274,1437,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.36" metricValueMin="1.34" pageId="8" pageNumber="159" unit="mm" value="1.35" valueMax="1.36" valueMin="1.34">1.34–1.36 mm</quantity>
(females), width 
<quantity id="9152E88D9E4F0402FED6FF16FE3DFECB" box="[344,508,223,248]" metricMagnitude="-3" metricUnit="m" metricValue="1.505" metricValueMax="1.54" metricValueMin="1.47" pageId="8" pageNumber="159" unit="mm" value="1.505" valueMax="1.54" valueMin="1.47">1.47–1.54 mm</quantity>
(males) and 
<quantity id="9152E88D9E4F0402FD1EFF16FCF5FECB" box="[656,820,223,248]" metricMagnitude="-3" metricUnit="m" metricValue="1.7349999999999999" metricValueMax="1.76" metricValueMin="1.71" pageId="8" pageNumber="159" unit="mm" value="1.7349999999999999" valueMax="1.76" valueMin="1.71">1.71–1.76 mm</quantity>
(females)], anterior margin five-toothed, middle basal half slightly protruding, four sublateral sensorial setae, basal portions covered with few sparse, white decumbent setae (males: 1–10, females: 4–8); colour yellowish with outer margin black, males triangularly doubled blackened in the first basal third, females triangularly blackened almost in the two basal thirds; antennae reaching nearly the half of the elytral length in male, shorter in female; antennomeres 1–4 dark metallic black with bluish or violaceous reflections, antennomeres 5–10 strongly dilated and foliated, more pronounced in females; mandibles four-toothed, black with a large yellow patch basally on outer edge; maxillary and labial palpi light yellow, except terminal palpomeres black with metallic reflections; penultimate palpomere of labial palpus moderately inflated.
</paragraph>
<caption id="02D515E09E4F0402FF19F894FBDAF94F" box="[151,1051,1885,1907]" httpUri="https://zenodo.org/record/253744/files/figure.png" pageId="8" pageNumber="159" targetBox="[168,1422,570,1847]" targetPageId="8">
<paragraph id="561545689E4F0402FF19F894FBDAF94F" blockId="8.[151,1051,1885,1907]" box="[151,1051,1885,1907]" pageId="8" pageNumber="159">
<emphasis id="64DE997A9E4F0402FF19F894FED3F94F" bold="true" box="[151,274,1885,1907]" pageId="8" pageNumber="159">FIGURE 4.</emphasis>
Facies of 
<taxonomicName id="91AA3EEB9E4F0402FEF5F894FD98F94F" box="[379,601,1885,1907]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="pepetela" status="sp. nov.">
<emphasis id="64DE997A9E4F0402FEF5F894FD98F94F" box="[379,601,1885,1907]" italics="true" pageId="8" pageNumber="159">Trichotaenia pepetela</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7FED24019E4F0402FDD1F897FD59F94F" box="[607,664,1886,1907]" pageId="8" pageNumber="159" rank="species">
<emphasis id="64DE997A9E4F0402FDD1F897FDB6F94F" bold="true" box="[607,631,1886,1907]" pageId="8" pageNumber="159">sp</emphasis>
. 
<emphasis id="64DE997A9E4F0402FD0DF897FD59F94F" bold="true" box="[643,664,1886,1907]" pageId="8" pageNumber="159">n.</emphasis>
</taxonomicNameLabel>
: a) male holotype, b) female allotype.
</paragraph>
</caption>
<caption id="02D515E09E4E0403FF19FB35FA9BFB2E" box="[151,1370,1276,1298]" httpUri="https://zenodo.org/record/253745/files/figure.png" pageId="9" pageNumber="160" targetBox="[164,1424,208,1244]" targetPageId="9">
<paragraph id="561545689E4E0403FF19FB35FA9BFB2E" blockId="9.[151,1370,1276,1298]" box="[151,1370,1276,1298]" pageId="9" pageNumber="160">
<emphasis id="64DE997A9E4E0403FF19FB35FED3FB2E" bold="true" box="[151,274,1276,1298]" pageId="9" pageNumber="160">FIGURE 5.</emphasis>
<taxonomicName id="91AA3EEB9E4E0403FE97FB35FE36FB2E" box="[281,503,1276,1298]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="pepetela" status="sp. nov.">
<emphasis id="64DE997A9E4E0403FE97FB35FE36FB2E" box="[281,503,1276,1298]" italics="true" pageId="9" pageNumber="160">Trichotaenia pepetela</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7FED24019E4E0403FE72FB34FDF7FB2E" box="[508,566,1277,1298]" pageId="9" pageNumber="160" rank="species">
<emphasis id="64DE997A9E4E0403FE72FB34FDD2FB2E" bold="true" box="[508,531,1277,1298]" pageId="9" pageNumber="160">sp</emphasis>
. 
<emphasis id="64DE997A9E4E0403FDAFFB34FDF7FB2E" bold="true" box="[545,566,1277,1298]" pageId="9" pageNumber="160">n.</emphasis>
</taxonomicNameLabel>
, head, mandibles and labrum, pronotum (dorsal view): a, c) male; b, d) female.
</paragraph>
</caption>
<paragraph id="561545689E4E0403FF49FAF3FE21F8E2" blockId="9.[151,1437,1338,2011]" pageId="9" pageNumber="160">
<emphasis id="64DE997A9E4E0403FF49FAF3FEE1FB6F" box="[199,288,1338,1363]" italics="true" pageId="9" pageNumber="160">Thorax.</emphasis>
Pronotum (
<figureCitation id="CE9159ED9E4E0403FE29FAF3FE2FFB6F" box="[423,494,1338,1363]" captionStart="FIGURE 5" captionStartId="9.[151,250,1276,1298]" captionTargetBox="[164,1424,208,1244]" captionTargetId="figure@9.[151,1435,193,1255]" captionTargetPageId="9" captionText="FIGURE 5. Trichotaenia pepetela sp. n., head, mandibles and labrum, pronotum (dorsal view): a, c) male; b, d) female." httpUri="https://zenodo.org/record/253745/files/figure.png" pageId="9" pageNumber="160">Figs 5</figureCitation>
c‒5d) more or less as long as wide (1.0–1.1 times) [length: 
<quantity id="9152E88D9E4E0403FB15FAF3FA81FB6E" box="[1179,1344,1338,1362]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="1.98" metricValueMin="1.82" pageId="9" pageNumber="160" unit="mm" value="1.9" valueMax="1.98" valueMin="1.82">1.82–1.98 mm</quantity>
(males) and 
<quantity id="9152E88D9E4E0403FF47FA97FEAEFB4A" box="[201,367,1374,1398]" metricMagnitude="-3" metricUnit="m" metricValue="2.06" metricValueMax="2.1" metricValueMin="2.02" pageId="9" pageNumber="160" unit="mm" value="2.06" valueMax="2.1" valueMin="2.02">2.02–2.10 mm</quantity>
(females); width: 
<quantity id="9152E88D9E4E0403FDC8FA94FD2DFB4A" box="[582,748,1373,1398]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="1.94" metricValueMin="1.86" pageId="9" pageNumber="160" unit="mm" value="1.9" valueMax="1.94" valueMin="1.86">1.86–1.94 mm</quantity>
(males) and 
<quantity id="9152E88D9E4E0403FC08FA97FC2CFB4A" box="[902,1005,1374,1398]" metricMagnitude="-3" metricUnit="m" metricValue="2.08" pageId="9" pageNumber="160" unit="mm" value="2.08">2.08 mm</quantity>
(females)], subcordiform-shaped, the anterior margin slightly larger [width: 
<quantity id="9152E88D9E4E0403FDD0FA4BFCC7FBA6" box="[606,774,1410,1434]" metricMagnitude="-3" metricUnit="m" metricValue="1.78" metricValueMax="1.82" metricValueMin="1.74" pageId="9" pageNumber="160" unit="mm" value="1.78" valueMax="1.82" valueMin="1.74">1.74–1.82 mm</quantity>
(males) and 
<quantity id="9152E88D9E4E0403FC28FA4BFBCEFBA6" box="[934,1039,1410,1435]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" pageId="9" pageNumber="160" unit="mm" value="2.05">2.05 mm</quantity>
(females)] than the posterior one [width: 
<quantity id="9152E88D9E4E0403FF7CFA6FFE58FB82" box="[242,409,1446,1470]" metricMagnitude="-3" metricUnit="m" metricValue="1.7149999999999999" metricValueMax="1.73" metricValueMin="1.7" pageId="9" pageNumber="160" unit="mm" value="1.7149999999999999" valueMax="1.73" valueMin="1.7">1.70–1.73 mm</quantity>
(males) and 
<quantity id="9152E88D9E4E0403FDBAFA6CFD5AFB82" box="[564,667,1445,1470]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" pageId="9" pageNumber="160" unit="mm" value="1.95">1.95 mm</quantity>
(females)]; black, on each side with a broad sublateral band with cupric reflections, this bands are covered with dense piceous decumbent pubescence, which is transversaly directed on median lobe, oblique on posterior and anterior lobes; along each side of median lobe and on central portions of anterior and posterior lobes a few additional thin piceous setae; surface of pronotum rugulose, rugae forming irregular meshes on latero-dorsal and dorsal portions; dorsal half of proepisterna glabrous, ventral half densely covered with white decumbent pubescence which continues that of genae and extends to mesepisterna, metepisterna, lateral portions of metasternum and metacoxae to lateral portions of the first five abdominal sterni; proepisterna slightly visible from above; coupling sulcus: a deeply pitted, funnel-formed impression centrally in upper half of mesepisternum.
</paragraph>
<paragraph id="561545689E4E0400FF49F923FB06FF03" blockId="9.[151,1437,1338,2011]" lastBlockId="10.[151,1437,151,536]" lastPageId="10" lastPageNumber="161" pageId="9" pageNumber="160">
<emphasis id="64DE997A9E4E0403FF49F923FECCF93F" box="[199,269,1770,1795]" italics="true" pageId="9" pageNumber="160">Elytra</emphasis>
(
<figureCitation id="CE9159ED9E4E0403FEAEF923FEABF93F" box="[288,362,1770,1795]" captionStart="FIGURE 7" captionStartId="11.[151,250,1499,1521]" captionTargetBox="[171,1423,206,1462]" captionTargetId="figure@11.[151,1437,193,1477]" captionTargetPageId="11" captionText="FIGURE 7. Elytra (dorsal view): a and b) Trichotaenia pepetela sp. n., male and female, respectively, c) Trichotaenia tereticollis (W. Horn), d) Trichotaenia pseudotereticollis (W. Horn), e) Trichotaenia suturata (W. Horn)." httpUri="https://zenodo.org/record/253747/files/figure.png" pageId="9" pageNumber="160">Figs 7</figureCitation>
a‒7b) longer than wide (1.8–1.9 times) [(length: 
<quantity id="9152E88D9E4E0403FC21F923FB97F93E" box="[943,1110,1770,1795]" metricMagnitude="-3" metricUnit="m" metricValue="6.369999999999999" metricValueMax="6.47" metricValueMin="6.27" pageId="9" pageNumber="160" unit="mm" value="6.369999999999999" valueMax="6.47" valueMin="6.27">6.27–6.47 mm</quantity>
(males) and 
<quantity id="9152E88D9E4E0403FB7BF923FA5CF93E" box="[1269,1437,1770,1795]" metricMagnitude="-3" metricUnit="m" metricValue="7.195" metricValueMax="7.26" metricValueMin="7.13" pageId="9" pageNumber="160" unit="mm" value="7.195" valueMax="7.26" valueMin="7.13">7.13–7.26 mm</quantity>
(females); width: 
<quantity id="9152E88D9E4E0403FEEEF8C4FDC4F91A" box="[352,517,1805,1830]" metricMagnitude="-3" metricUnit="m" metricValue="3.395" metricValueMax="3.46" metricValueMin="3.33" pageId="9" pageNumber="160" unit="mm" value="3.395" valueMax="3.46" valueMin="3.33">3.33–3.46 mm</quantity>
(males) and 
<quantity id="9152E88D9E4E0403FD14F8C4FCFFF91A" box="[666,830,1805,1830]" metricMagnitude="-3" metricUnit="m" metricValue="3.9299999999999997" metricValueMax="3.96" metricValueMin="3.9" pageId="9" pageNumber="160" unit="mm" value="3.9299999999999997" valueMax="3.96" valueMin="3.9">3.90–3.96 mm</quantity>
(females)], subrectangular-shaped, elongated (males) (
<figureCitation id="CE9159ED9E4E0403FF11F8FBFF23F976" box="[159,226,1842,1867]" captionStart="FIGURE 7" captionStartId="11.[151,250,1499,1521]" captionTargetBox="[171,1423,206,1462]" captionTargetId="figure@11.[151,1437,193,1477]" captionTargetPageId="11" captionText="FIGURE 7. Elytra (dorsal view): a and b) Trichotaenia pepetela sp. n., male and female, respectively, c) Trichotaenia tereticollis (W. Horn), d) Trichotaenia pseudotereticollis (W. Horn), e) Trichotaenia suturata (W. Horn)." httpUri="https://zenodo.org/record/253747/files/figure.png" pageId="9" pageNumber="160">Fig. 7</figureCitation>
a), more convex and enlarged (females) (
<figureCitation id="CE9159ED9E4E0403FD22F8FBFD2EF977" box="[684,751,1842,1867]" captionStart="FIGURE 7" captionStartId="11.[151,250,1499,1521]" captionTargetBox="[171,1423,206,1462]" captionTargetId="figure@11.[151,1437,193,1477]" captionTargetPageId="11" captionText="FIGURE 7. Elytra (dorsal view): a and b) Trichotaenia pepetela sp. n., male and female, respectively, c) Trichotaenia tereticollis (W. Horn), d) Trichotaenia pseudotereticollis (W. Horn), e) Trichotaenia suturata (W. Horn)." httpUri="https://zenodo.org/record/253747/files/figure.png" pageId="9" pageNumber="160">Fig. 7</figureCitation>
b), shoulders smooth and glossy very marked, apex ending in a short but acute sutural spine in both sexes, disc surface sculptured by coarse, but very densely arranged polygonal (quadrangular to hexagonal) alveoli most slightly flattened walled in the middle and posteriad half, apical margin with distinct microserrulation; posteriad disc near suture not (males) or slightly protruding in relief in its uppermost region (females); colour of elytra black with more or less coppery lustre only in the pubescent areas, otherwise with some bluish or violaceous reflections; Elytra with white-piceous decumbent pubescence (
<figureCitation id="CE9159ED9E4D0400FB02FF5EFB15FE93" box="[1164,1236,151,176]" captionStart="FIGURE 7" captionStartId="11.[151,250,1499,1521]" captionTargetBox="[171,1423,206,1462]" captionTargetId="figure@11.[151,1437,193,1477]" captionTargetPageId="11" captionText="FIGURE 7. Elytra (dorsal view): a and b) Trichotaenia pepetela sp. n., male and female, respectively, c) Trichotaenia tereticollis (W. Horn), d) Trichotaenia pseudotereticollis (W. Horn), e) Trichotaenia suturata (W. Horn)." httpUri="https://zenodo.org/record/253747/files/figure.png" pageId="10" pageNumber="161">Figs 7</figureCitation>
a‒7b), consisting of a marginal band, from the middle of basal third to the apex where it becomes slightly wider, emitting two rami on disc, one almost straight and large near the middle and another subapical slightly arcuate; a small tuft of closely and diagonally appressed setae on each side of the base of scutellum and a subsutural elongate tuft of closely appressed setae below the scutellum. Absence of of sparce or decumbent setae along the suture.
</paragraph>
<paragraph id="561545689E4D0400FF49FE85FDC3FF91" blockId="10.[151,1437,151,536]" pageId="10" pageNumber="161">Ventral surface. Black, with stronger coppery-golden reflections in head and thorax parts, violet lustre in abdominal sternites; sides of these sternites, except the ultimate, with decumbent pubescence, becoming sparser towards apex; trochanters black.</paragraph>
<paragraph id="561545689E4D0400FF49FE70FDF8FFC9" blockId="10.[151,1437,151,536]" pageId="10" pageNumber="161">
<emphasis id="64DE997A9E4D0400FF49FE70FEC2FFEC" box="[199,259,441,464]" italics="true" pageId="10" pageNumber="161">Legs.</emphasis>
Metallic black with greenish reflections in tarsi and violaceous-bluish in femora and tibiae, a few rows of spiniform setae on femora and tibiae;
</paragraph>
<paragraph id="561545689E4D0400FF49FE36FAAEFC2B" blockId="10.[151,1437,151,536]" box="[199,1391,511,536]" pageId="10" pageNumber="161">
<emphasis id="64DE997A9E4D0400FF49FE36FEF5FC24" box="[199,308,511,536]" italics="true" pageId="10" pageNumber="161">Aedeagus</emphasis>
(
<figureCitation id="CE9159ED9E4D0400FECDFE36FE44FC24" box="[323,389,511,536]" captionStart="FIGURE 6" captionStartId="10.[151,250,1973,1995]" captionTargetBox="[212,1369,576,1946]" captionTargetId="figure@10.[201,1386,567,1962]" captionTargetPageId="10" captionText="FIGURE 6. Aedeagus (lateral view): a) Trichotaenia nzingae sp. n., paratype, Kakande, Bié, Angola, b) Trichotaenia pepetela sp. n., holotype, Catota, Bié, Angola." httpUri="https://zenodo.org/record/253746/files/figure.png" pageId="10" pageNumber="161">Fig. 6</figureCitation>
b) relatively small (length: 
<quantity id="9152E88D9E4D0400FD38FE36FC9CFC24" box="[694,861,511,536]" metricMagnitude="-3" metricUnit="m" metricValue="2.71" metricValueMax="2.78" metricValueMin="2.64" pageId="10" pageNumber="161" unit="mm" value="2.71" valueMax="2.78" valueMin="2.64">2.64–2.78 mm</quantity>
), arched, tapering, with a straight, simple apex.
</paragraph>
<caption id="02D515E09E4D0400FF19F87CFDD3F9D6" httpUri="https://zenodo.org/record/253746/files/figure.png" pageId="10" pageNumber="161" targetBox="[212,1369,576,1946]" targetPageId="10">
<paragraph id="561545689E4D0400FF19F87CFDD3F9D6" blockId="10.[151,1436,1973,2026]" pageId="10" pageNumber="161">
<emphasis id="64DE997A9E4D0400FF19F87CFED3F9F6" bold="true" box="[151,274,1973,1995]" pageId="10" pageNumber="161">FIGURE 6.</emphasis>
Aedeagus (lateral view): a) 
<taxonomicName id="91AA3EEB9E4D0400FDBFF87CFCC6F9F7" box="[561,775,1973,1995]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="nzingae" status="sp. nov.">
<emphasis id="64DE997A9E4D0400FDBFF87CFCC6F9F7" box="[561,775,1973,1995]" italics="true" pageId="10" pageNumber="161">Trichotaenia nzingae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7FED24019E4D0400FC83F87CFC87F9F6" box="[781,838,1973,1995]" pageId="10" pageNumber="161" rank="species">
<emphasis id="64DE997A9E4D0400FC83F87CFCE4F9F6" bold="true" box="[781,805,1973,1994]" pageId="10" pageNumber="161">sp</emphasis>
. 
<emphasis id="64DE997A9E4D0400FCBCF87CFC87F9F6" bold="true" box="[818,838,1973,1994]" pageId="10" pageNumber="161">n.</emphasis>
</taxonomicNameLabel>
, paratype, Kakande, Bié, Angola, b) 
<taxonomicName id="91AA3EEB9E4D0400FB31F87CFA5DF9F7" box="[1215,1436,1973,1995]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="pepetela" status="sp. nov.">
<emphasis id="64DE997A9E4D0400FB31F87CFA5DF9F7" box="[1215,1436,1973,1995]" italics="true" pageId="10" pageNumber="161">Trichotaenia pepetela</emphasis>
</taxonomicName>
<emphasis id="64DE997A9E4D0400FF19F81DFF11F9D5" bold="true" box="[151,208,2004,2025]" pageId="10" pageNumber="161">
<taxonomicNameLabel id="7FED24019E4D0400FF19F81DFF11F9D5" box="[151,208,2004,2025]" pageId="10" pageNumber="161" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype, Catota, Bié, Angola.
</paragraph>
</caption>
<caption id="02D515E09E4C0401FF19FA12FB76F833" httpUri="https://zenodo.org/record/253747/files/figure.png" pageId="11" pageNumber="162" targetBox="[171,1423,206,1462]" targetPageId="11">
<paragraph id="561545689E4C0401FF19FA12FB76F833" blockId="11.[151,1436,1499,1551]" pageId="11" pageNumber="162">
<emphasis id="64DE997A9E4C0401FF19FA12FED9FBCC" bold="true" box="[151,280,1499,1521]" pageId="11" pageNumber="162">FIGURE 7.</emphasis>
Elytra (dorsal view): a and b) 
<taxonomicName id="91AA3EEB9E4C0401FDFFFA12FC92FBCD" box="[625,851,1499,1521]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="pepetela" status="sp. nov.">
<emphasis id="64DE997A9E4C0401FDFFFA12FC92FBCD" box="[625,851,1499,1521]" italics="true" pageId="11" pageNumber="162">Trichotaenia pepetela</emphasis>
</taxonomicName>
<emphasis id="64DE997A9E4C0401FCD0FA12FC5DFBCC" bold="true" box="[862,924,1499,1520]" pageId="11" pageNumber="162">
<taxonomicNameLabel id="7FED24019E4C0401FCD0FA12FC5DFBCC" box="[862,924,1499,1520]" pageId="11" pageNumber="162" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male and female, respectively, c) 
<taxonomicName id="91AA3EEB9E4C0401FA95FA12FEB3F833" authority="W. Horn" authorityName="W. Horn" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="tereticollis">
<emphasis id="64DE997A9E4C0401FA95FA12FEC2F833" italics="true" pageId="11" pageNumber="162">Trichotaenia tereticollis</emphasis>
(W. Horn)
</taxonomicName>
, d) 
<taxonomicName id="91AA3EEB9E4C0401FE16FA30FC83F833" authority="W. Horn" authorityName="W. Horn" box="[408,834,1529,1551]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="pseudotereticollis">
<emphasis id="64DE997A9E4C0401FE16FA30FD13F833" box="[408,722,1529,1551]" italics="true" pageId="11" pageNumber="162">Trichotaenia pseudotereticollis</emphasis>
(W. Horn)
</taxonomicName>
, e) 
<taxonomicName id="91AA3EEB9E4C0401FCE8FA30FB72F833" authority="W. Horn" authorityName="W. Horn" box="[870,1203,1529,1551]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="suturata">
<emphasis id="64DE997A9E4C0401FCE8FA30FB82F833" box="[870,1091,1529,1551]" italics="true" pageId="11" pageNumber="162">Trichotaenia suturata</emphasis>
(W. Horn)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="561545689E4C0401FF49F9FFFC35F8E3" blockId="11.[151,1437,1590,2012]" pageId="11" pageNumber="162">
<emphasis id="64DE997A9E4C0401FF49F9FFFE13F873" bold="true" box="[199,466,1590,1615]" pageId="11" pageNumber="162">Intraspecific variation</emphasis>
. The range of variability observed in 
<taxonomicName id="91AA3EEB9E4C0401FC0DF9F1FC3EF873" box="[899,1023,1590,1615]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="pepetela" status="sp. nov.">
<emphasis id="64DE997A9E4C0401FC0DF9F1FC3EF873" box="[899,1023,1590,1615]" italics="true" pageId="11" pageNumber="162">T. pepetela</emphasis>
</taxonomicName>
<emphasis id="64DE997A9E4C0401FB89F9FFFB86F873" bold="true" box="[1031,1095,1590,1615]" pageId="11" pageNumber="162">
<taxonomicNameLabel id="7FED24019E4C0401FB89F9FFFB86F873" box="[1031,1095,1590,1615]" pageId="11" pageNumber="162" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<specimenCount id="40AC8EE19E4C0401FBD6F9FFFB27F873" box="[1112,1254,1590,1615]" pageId="11" pageNumber="162" type="generic">5 specimens</specimenCount>
) affects slightly the number of labral decumbent setae (see description), but not the elytral decumbent pubescence. 
<collectingCountry id="2EBD05F89E4C0401FB6DF992FAC3F84F" box="[1251,1282,1627,1651]" name="American Samoa" pageId="11" pageNumber="162">As</collectingCountry>
happens with 
<taxonomicName id="91AA3EEB9E4C0401FF19F949FECBF8AB" box="[151,266,1664,1687]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="nzingae" status="sp. nov.">
<emphasis id="64DE997A9E4C0401FF19F949FECBF8AB" box="[151,266,1664,1687]" italics="true" pageId="11" pageNumber="162">T. nzingae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7FED24019E4C0401FE9CF9B7FE92F8AB" box="[274,339,1662,1687]" pageId="11" pageNumber="162" rank="species">
<emphasis id="64DE997A9E4C0401FE9CF9B7FE92F8AB" bold="true" box="[274,339,1662,1687]" pageId="11" pageNumber="162">sp. n.</emphasis>
</taxonomicNameLabel>
the shapes of pronotum and elytra are very conservative, but variation in the length of the apical denticle of elytra is very common, with individuals having longer denticles than others. Asymmetries in the length of left and right elytron denticles within the same specimen are common too.
</paragraph>
</subSubSection>
<subSubSection id="1EB016E39E4C0401FF49F922FAB9F953" pageId="11" pageNumber="162" type="discussion">
<paragraph id="561545689E4C0401FF49F922FAB9F953" blockId="11.[151,1437,1590,2012]" pageId="11" pageNumber="162">
<emphasis id="64DE997A9E4C0401FF49F922FEFAF938" bold="true" box="[199,315,1771,1796]" pageId="11" pageNumber="162">Remarks.</emphasis>
A total of fifteen species, including the two new ones, are known so far within the genus 
<taxonomicName id="91AA3EEB9E4C0401FF19F8C7FEE7F91B" box="[151,294,1806,1831]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="genus">
<emphasis id="64DE997A9E4C0401FF19F8C7FEE7F91B" box="[151,294,1806,1831]" italics="true" pageId="11" pageNumber="162">Trichotaenia</emphasis>
</taxonomicName>
. 
<bibRefCitation id="323B38999E4C0401FEB8F8C7FE20F91B" author="Cassola" box="[310,481,1806,1831]" pageId="11" pageNumber="162" refString="Cassola, F. (1983) Studi sui Cicindelidi. XXXVIII. Revisione del genere Trichotaenia Rivalier, com descrizione di due nuove specie (Coleoptera, Cicindelidae). Revue de Zoologie africaine, 97 (4), 689 - 704." type="journal article" year="1983">Cassola (1983)</bibRefCitation>
revised the eight then known species of this Afrotropical genus and 
<bibRefCitation id="323B38999E4C0401FB73F8C7FA5DF91B" author="Schule" box="[1277,1436,1806,1831]" pageId="11" pageNumber="162" refString="Schule, P. (2011) A new species of Trichotaenia Rivalier, 1957 from Zambia (Coleoptera: Cicindelidae). Entomologische Zeitschrift, 121 (5), 211 - 214." type="journal article" year="2011">Schüle (2011)</bibRefCitation>
provided an identification key for the thirtheen then known species. Until now the genus was represented in 
<collectingCountry id="2EBD05F89E4C0401FAC4F8FAFA5DF970" box="[1354,1436,1843,1868]" name="Angola" pageId="11" pageNumber="162">Angola</collectingCountry>
by two species [
<taxonomicName id="91AA3EEB9E4C0401FEC2F891FD19F953" authority="W. Horn, 1914" authorityName="W. Horn" authorityYear="1914" box="[332,728,1878,1903]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="pseudosuturalis">
<emphasis id="64DE997A9E4C0401FEC2F891FDD9F953" box="[332,536,1878,1903]" italics="true" pageId="11" pageNumber="162">T. pseudosuturalis</emphasis>
(W. Horn, 1914)
</taxonomicName>
and 
<taxonomicName id="91AA3EEB9E4C0401FC80F891FB89F953" authority="W. Horn, 1915" authorityName="W. Horn" authorityYear="1915" box="[782,1096,1878,1903]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="suturata">
<emphasis id="64DE997A9E4C0401FC80F891FC47F953" box="[782,902,1880,1903]" italics="true" pageId="11" pageNumber="162">T. suturata</emphasis>
(W. Horn, 1915)
</taxonomicName>
] (
<bibRefCitation id="323B38999E4C0401FBD0F89EFAADF953" author="Serrano" box="[1118,1388,1878,1903]" pageId="11" pageNumber="162" refString="Serrano, A. R. M. &amp; Capela, R. (2013) The tiger beetles (Coleoptera: Carabidae, Cicindelinae) of Angola: a descriptive catalogue and designation of neotypes. Zootaxa, 3731 (4), 401 - 444." type="journal article" year="2013">Serrano &amp; Capela 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="1EB016E39E4C0407FF49F8B2FDB4FA09" lastPageId="13" lastPageNumber="164" pageId="11" pageNumber="162" type="reference_group">
<paragraph id="561545689E4C0406FF49F8B2FC39FFC8" blockId="11.[151,1437,1590,2012]" lastBlockId="12.[151,1437,151,501]" lastPageId="12" lastPageNumber="163" pageId="11" pageNumber="162">
The 
<taxonomicName id="91AA3EEB9E4C0401FF74F8B2FE48F9A8" box="[250,393,1915,1940]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="genus">
<emphasis id="64DE997A9E4C0401FF74F8B2FE48F9A8" box="[250,393,1915,1940]" italics="true" pageId="11" pageNumber="162">Trichotaenia</emphasis>
</taxonomicName>
species can be included in some artificial groups based on two sets of characteristics (
<tableCitation id="1B2870D39E4C0401FAD2F8B2FF65F98B" captionStart="TABLE 1" captionStartId="12.[151,239,548,571]" captionTargetBox="[208,1371,777,1327]" captionTargetPageId="12" captionText="TABLE 1. Artificial groups of Trichotaenia species based on two sets of characters: presence or not of labral pubescence and elytral humeri distinct or effaced. Within each set of characters, groups are joined simultaneously by the same background color and the same small letters." httpUri="http://table.plazi.org/id/02D515E09E4B0406FF19FDEDFDB2FC47" pageId="11" pageNumber="162" tableUuid="02D515E09E4B0406FF19FDEDFDB2FC47">Table 1</tableCitation>
). Two groups are easily discriminated based on the presence / absence of full wings, a condition that is correlated with the evidence of distinct elytral humeri against effaced elytral humeri and three groups based on the presence / absence of decumbent setae on the labrum. So far, it has not been proved to be any phylogenetic mean of these features, but they are practical to discriminate those groups of species. A lot of eight species seems to share both the elytral humeri distinct plus the labrum setose [
<taxonomicName id="91AA3EEB9E4B0406FD53FF28FB8BFEC4" authority="W. Horn, 1929" authorityName="W. Horn" authorityYear="1929" box="[733,1098,223,248]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="duplosetosa">
<emphasis id="64DE997A9E4B0406FD53FF28FC40FEC4" box="[733,897,223,248]" italics="true" pageId="12" pageNumber="163">T. duplosetosa</emphasis>
(W. Horn, 1929)
</taxonomicName>
, 
<taxonomicName id="91AA3EEB9E4B0406FBD6FF28FABBFEC4" authority="Cassola, 2008" authorityName="Cassola" authorityYear="2008" box="[1112,1402,223,248]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="kudrnai">
<emphasis id="64DE997A9E4B0406FBD6FF28FB0CFEC4" box="[1112,1229,223,248]" italics="true" pageId="12" pageNumber="163">T. kudrnai</emphasis>
<bibRefCitation id="323B38999E4B0406FB59FF16FABBFEC4" author="Cassola" box="[1239,1402,223,248]" pageId="12" pageNumber="163" refString="Cassola, F. (2008) A new Trichotaenia species from Tanzania (Coleoptera, Cicindelidae). Fragmenta entomologica, 40 (2), 243 - 248. http: // dx. doi. org / 10.4081 / fe. 2008.96" type="journal article" year="2008">Cassola, 2008</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="91AA3EEB9E4B0406FA09FF28FF2FFF20" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="nzingae" status="sp. nov.">
<emphasis id="64DE997A9E4B0406FA09FF28FF2FFF20" italics="true" pageId="12" pageNumber="163">T. nzingae</emphasis>
</taxonomicName>
<emphasis id="64DE997A9E4B0406FF7BFECDFE8FFF21" bold="true" box="[245,334,260,285]" pageId="12" pageNumber="163">
<taxonomicNameLabel id="7FED24019E4B0406FF7BFECDFE8FFF21" box="[245,334,260,285]" pageId="12" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="91AA3EEB9E4B0406FED2FECCFE16FF21" box="[348,471,260,285]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="pepetela" status="sp. nov.">
<emphasis id="64DE997A9E4B0406FED2FECCFE16FF21" box="[348,471,260,285]" italics="true" pageId="12" pageNumber="163">T. pepetela</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7FED24019E4B0406FE50FECDFDF7FF20" box="[478,566,260,285]" pageId="12" pageNumber="163" rank="species">
<emphasis id="64DE997A9E4B0406FE50FECDFDF1FF21" bold="true" box="[478,560,260,285]" pageId="12" pageNumber="163">sp. nov</emphasis>
.
</taxonomicNameLabel>
, 
<taxonomicName id="91AA3EEB9E4B0406FDCBFECCFC0CFF20" authority="W. Horn 1914" authorityName="W. Horn" authorityYear="1914" box="[581,973,260,285]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="pseudosuturalis">
<emphasis id="64DE997A9E4B0406FDCBFECCFCD3FF21" box="[581,786,260,285]" italics="true" pageId="12" pageNumber="163">T. pseudosuturalis</emphasis>
(W. Horn 1914)
</taxonomicName>
, 
<taxonomicName id="91AA3EEB9E4B0406FC56FECCFABAFF20" authority="W. Horn, 1929" authorityName="W. Horn" authorityYear="1929" box="[984,1403,260,285]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="pseudotereticollis">
<emphasis id="64DE997A9E4B0406FC56FECCFB7BFF21" box="[984,1210,260,285]" italics="true" pageId="12" pageNumber="163">T. pseudotereticollis</emphasis>
(W. Horn, 1929)
</taxonomicName>
, 
<taxonomicName id="91AA3EEB9E4B0406FA09FECCFE04FF7C" authority="Basilewsky, 1958" authorityName="Basilewsky" authorityYear="1958" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="rivalieri">
<emphasis id="64DE997A9E4B0406FA09FECCFF32FF7C" italics="true" pageId="12" pageNumber="163">T. rivalieri</emphasis>
Basilewsky, 1958
</taxonomicName>
and 
<taxonomicName id="91AA3EEB9E4B0406FD8FFEE0FC87FF7C" authority="W. Horn, 1915" authorityName="W. Horn" authorityYear="1915" box="[513,838,295,320]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="suturata">
<emphasis id="64DE997A9E4B0406FD8FFEE0FDBDFF7C" box="[513,636,297,320]" italics="true" pageId="12" pageNumber="163">T. suturata</emphasis>
(W. Horn, 1915)
</taxonomicName>
] and other three the elytral humeri effaced plus the labrum glabrous (
<taxonomicName id="91AA3EEB9E4B0406FEEFFE84FDA8FF58" authority="Werner, 2003" authorityName="Werner" authorityYear="2003" box="[353,617,332,357]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="mireki">
<emphasis id="64DE997A9E4B0406FEEFFE84FE03FF59" box="[353,450,332,357]" italics="true" pageId="12" pageNumber="163">T. mireki</emphasis>
<bibRefCitation id="323B38999E4B0406FE40FE84FDA8FF58" author="Werner" box="[462,617,332,356]" pageId="12" pageNumber="163" refString="Werner, K. (2003) Description of four new tiger beetle species from Tanzania and Zambia (Coleoptera: Cicindelidae). Lambillionea, 103 (4), 559 - 584." type="journal article" year="2003">Werner, 2003</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="91AA3EEB9E4B0406FDF8FE84FB75FF58" authority="Cassola, Werner &amp; Schule, 2009" authorityName="Cassola, Werner &amp; Schule" authorityYear="2009" box="[630,1204,332,357]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="mufumbweana">
<emphasis id="64DE997A9E4B0406FDF8FE84FCF9FF59" box="[630,824,332,357]" italics="true" pageId="12" pageNumber="163">T. mufumbweana</emphasis>
<bibRefCitation id="323B38999E4B0406FCCEFE85FB75FF58" author="Cassola" box="[832,1204,332,357]" pageId="12" pageNumber="163" refString="Cassola, F., Werner, K. &amp; Schule, P. (2009) Two new Trichotaenia species from Zambia (Coleoptera: Cicindelidae). Mitteilungen der Munchner Entomologischen Gesellschaft, 99, 11 - 16." type="journal article" year="2009">Cassola, Werner &amp; Schüle, 2009</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="91AA3EEB9E4B0406FB60FE84FDCCFFB4" authority="Cassola, Werner &amp; Schule, 2009" authorityName="Cassola, Werner &amp; Schule" authorityYear="2009" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="mwinilungae">
<emphasis id="64DE997A9E4B0406FB60FE84FA5DFF59" box="[1262,1436,332,357]" italics="true" pageId="12" pageNumber="163">T. mwinilungae</emphasis>
<bibRefCitation id="323B38999E4B0406FF19FEA6FDCCFFB4" author="Cassola" box="[151,525,367,392]" pageId="12" pageNumber="163" refString="Cassola, F., Werner, K. &amp; Schule, P. (2009) Two new Trichotaenia species from Zambia (Coleoptera: Cicindelidae). Mitteilungen der Munchner Entomologischen Gesellschaft, 99, 11 - 16." type="journal article" year="2009">Cassola, Werner &amp; Schüle, 2009</bibRefCitation>
</taxonomicName>
). The remaining four species, by presenting either the labrum setose or glabrous [
<taxonomicName id="91AA3EEB9E4B0406FF2EFE5CFE07FF90" authority="Cassola, 1983" authorityName="Cassola" authorityYear="1983" box="[160,454,404,429]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="64DE997A9E4B0406FF2EFE5CFEDDFF90" box="[160,284,404,428]" italics="true" pageId="12" pageNumber="163">T. africana</emphasis>
<bibRefCitation id="323B38999E4B0406FEA8FE5DFE07FF90" author="Cassola" box="[294,454,404,429]" pageId="12" pageNumber="163" refString="Cassola, F. (1983) Studi sui Cicindelidi. XXXVIII. Revisione del genere Trichotaenia Rivalier, com descrizione di due nuove specie (Coleoptera, Cicindelidae). Revue de Zoologie africaine, 97 (4), 689 - 704." type="journal article" year="1983">Cassola, 1983</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="91AA3EEB9E4B0406FD8FFE5CFCA9FF91" authority="Boheman, 1860" authorityName="Boheman" authorityYear="1860" box="[513,872,404,429]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="tereticollis">
<emphasis id="64DE997A9E4B0406FD8FFE5CFD57FF91" box="[513,662,404,429]" italics="true" pageId="12" pageNumber="163">T. tereticollis</emphasis>
(Boheman, 1860)
</taxonomicName>
] or by presenting a distinct elytral humeri plus a glabrous labrum (
<taxonomicName id="91AA3EEB9E4B0406FEE9FE70FD40FFEC" authority="Cassola, 1983" authorityName="Cassola" authorityYear="1983" box="[359,641,439,464]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="allardi">
<emphasis id="64DE997A9E4B0406FEE9FE70FE11FFEC" box="[359,464,439,464]" italics="true" pageId="12" pageNumber="163">T. allardi</emphasis>
<bibRefCitation id="323B38999E4B0406FE50FE7EFD40FFEC" author="Cassola" box="[478,641,439,464]" pageId="12" pageNumber="163" refString="Cassola, F. (1983) Studi sui Cicindelidi. XXXVIII. Revisione del genere Trichotaenia Rivalier, com descrizione di due nuove specie (Coleoptera, Cicindelidae). Revue de Zoologie africaine, 97 (4), 689 - 704." type="journal article" year="1983">Cassola, 1983</bibRefCitation>
</taxonomicName>
) or even exhibiting elytral humeri effaced plus a setose labrum (
<taxonomicName id="91AA3EEB9E4B0406FA09FE70FE47FFC8" authority="Schule, 2011" authorityName="Schule" authorityYear="2011" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="minettii">
<emphasis id="64DE997A9E4B0406FA09FE70FF2DFFC8" italics="true" pageId="12" pageNumber="163">T. minettii</emphasis>
<bibRefCitation id="323B38999E4B0406FF7DFE15FE47FFC8" author="Schule" box="[243,390,476,501]" pageId="12" pageNumber="163" refString="Schule, P. (2011) A new species of Trichotaenia Rivalier, 1957 from Zambia (Coleoptera: Cicindelidae). Entomologische Zeitschrift, 121 (5), 211 - 214." type="journal article" year="2011">Schüle, 2011</bibRefCitation>
</taxonomicName>
) do not enter wholy in the two former groups (
<tableCitation id="1B2870D39E4B0406FC1DFE15FC26FFC8" box="[915,999,476,501]" captionStart="TABLE 1" captionStartId="12.[151,239,548,571]" captionTargetBox="[208,1371,777,1327]" captionTargetPageId="12" captionText="TABLE 1. Artificial groups of Trichotaenia species based on two sets of characters: presence or not of labral pubescence and elytral humeri distinct or effaced. Within each set of characters, groups are joined simultaneously by the same background color and the same small letters." httpUri="http://table.plazi.org/id/02D515E09E4B0406FF19FDEDFDB2FC47" pageId="12" pageNumber="163" tableUuid="02D515E09E4B0406FF19FDEDFDB2FC47">Table 1</tableCitation>
).
</paragraph>
<caption id="02D515E09E4B0406FF19FDEDFDB2FC47" ID-Table-UUID="02D515E09E4B0406FF19FDEDFDB2FC47" httpUri="http://table.plazi.org/id/02D515E09E4B0406FF19FDEDFDB2FC47" pageId="12" pageNumber="163" targetBox="[208,1371,777,1327]" targetIsTable="true" targetPageId="12">
<paragraph id="561545689E4B0406FF19FDEDFDB2FC47" blockId="12.[151,1436,548,635]" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FF19FDEDFEC9FC07" bold="true" box="[151,264,548,571]" pageId="12" pageNumber="163">TABLE 1.</emphasis>
Artificial groups of 
<taxonomicName id="91AA3EEB9E4B0406FE6CFDEDFDABFC07" box="[482,618,548,571]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="64DE997A9E4B0406FE6CFDEDFDABFC07" box="[482,618,548,571]" italics="true" pageId="12" pageNumber="163">Trichotaenia</emphasis>
</taxonomicName>
species based on two sets of characters: presence or not of labral pubescence and elytral humeri distinct or effaced. Within each set of characters, groups are joined simultaneously by the same background color and the same small letters.
</paragraph>
</caption>
<paragraph id="561545689E4B0407FF5EFD05FF21FF20" blockId="12.[208,1310,716,739]" lastBlockId="13.[151,1436,151,1077]" lastPageId="13" lastPageNumber="164" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FF5EFD05FADFFCDF" bold="true" box="[208,1310,716,739]" pageId="12" pageNumber="163">Species Labrum Elytral humeri</emphasis>
The two new species belong to the first group presenting well distinct elytral shoulders as well as a labrum more (
<taxonomicName id="91AA3EEB9E4B0406FF6DFA6CFE98FB80" box="[227,345,1445,1468]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="nzingae" status="sp. nov.">
<emphasis id="64DE997A9E4B0406FF6DFA6CFE98FB80" box="[227,345,1445,1468]" italics="true" pageId="12" pageNumber="163">T. nzingae</emphasis>
</taxonomicName>
<emphasis id="64DE997A9E4B0406FEECFA6AFE6FFB80" bold="true" box="[354,430,1443,1468]" pageId="12" pageNumber="163">
<taxonomicNameLabel id="7FED24019E4B0406FEECFA6AFE64FB80" box="[354,421,1443,1468]" pageId="12" pageNumber="163" rank="species">sp. n.</taxonomicNameLabel>
)
</emphasis>
or less (
<taxonomicName id="91AA3EEB9E4B0406FD9BFA6CFD53FB80" box="[533,658,1443,1468]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="pepetela" status="sp. nov.">
<emphasis id="64DE997A9E4B0406FD9BFA6CFD53FB80" box="[533,658,1443,1468]" italics="true" pageId="12" pageNumber="163">T. pepetela</emphasis>
</taxonomicName>
<emphasis id="64DE997A9E4B0406FD12FA6AFD26FB80" bold="true" box="[668,743,1443,1468]" pageId="12" pageNumber="163">
<taxonomicNameLabel id="7FED24019E4B0406FD12FA6AFD1FFB80" box="[668,734,1443,1468]" pageId="12" pageNumber="163" rank="species">sp. n.</taxonomicNameLabel>
)
</emphasis>
setose. Within this group both new species, and taking into account the morphological characters, are more akin with 
<taxonomicName id="91AA3EEB9E4B0406FCBCFA01FBC0FBDC" box="[818,1025,1479,1504]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="pseudosuturalis">
<emphasis id="64DE997A9E4B0406FCBCFA01FBC0FBDC" box="[818,1025,1479,1504]" italics="true" pageId="12" pageNumber="163">T. pseudosuturalis</emphasis>
</taxonomicName>
and 
<taxonomicName id="91AA3EEB9E4B0406FBB2FA01FADEFBDC" box="[1084,1311,1479,1504]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="pseudotereticollis">
<emphasis id="64DE997A9E4B0406FBB2FA01FADEFBDC" box="[1084,1311,1479,1504]" italics="true" pageId="12" pageNumber="163">T. pseudotereticollis</emphasis>
</taxonomicName>
. However, they can be segregated from the former species by the position of the elytral median transverse decumbent setae stripe located around or slightly after the middle (located before the middle in 
<taxonomicName id="91AA3EEB9E4B0406FB86F9D9FB14F814" box="[1032,1237,1551,1576]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="pseudosuturalis">
<emphasis id="64DE997A9E4B0406FB86F9D9FB14F814" box="[1032,1237,1551,1576]" italics="true" pageId="12" pageNumber="163">T. pseudosuturalis</emphasis>
</taxonomicName>
), by the presence of a subapical arcuate decumbent stripe of setae (absent in 
<taxonomicName id="91AA3EEB9E4B0406FCB6F9FCFBC7F870" box="[824,1030,1587,1612]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="pseudosuturalis">
<emphasis id="64DE997A9E4B0406FCB6F9FCFBC7F870" box="[824,1030,1587,1612]" italics="true" pageId="12" pageNumber="163">T. pseudosuturalis</emphasis>
</taxonomicName>
) and even by the absence of sparse pubescence on shoulders (present in 
<taxonomicName id="91AA3EEB9E4B0406FDCDF991FCD2F84C" box="[579,787,1623,1648]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="pseudosuturalis">
<emphasis id="64DE997A9E4B0406FDCDF991FCD2F84C" box="[579,787,1623,1648]" italics="true" pageId="12" pageNumber="163">T. pseudosuturalis</emphasis>
</taxonomicName>
). Both new species can be discriminated from the later species also by the shoulders slightly more distinct and the shape of pronotum which is slightly cordiform (subrectangular in 
<taxonomicName id="91AA3EEB9E4B0406FEFAF969FD9AF884" box="[372,603,1695,1720]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="pseudotereticollis">
<emphasis id="64DE997A9E4B0406FEFAF969FD9AF884" box="[372,603,1695,1720]" italics="true" pageId="12" pageNumber="163">T. pseudotereticollis</emphasis>
</taxonomicName>
). Moreover 
<taxonomicName id="91AA3EEB9E4B0406FD7AF956FC27F88B" box="[756,998,1695,1720]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="nzingae" status="sp. nov.">
<emphasis id="64DE997A9E4B0406FD7AF956FC27F88B" box="[756,998,1695,1720]" italics="true" pageId="12" pageNumber="163">Trichotaenia nzingae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7FED24019E4B0406FC7DF956FBF9F88B" box="[1011,1080,1695,1720]" pageId="12" pageNumber="163" rank="species">
<emphasis id="64DE997A9E4B0406FC7DF956FBF1F884" bold="true" box="[1011,1072,1695,1720]" pageId="12" pageNumber="163">sp. n</emphasis>
.
</taxonomicNameLabel>
can be discriminated from 
<taxonomicName id="91AA3EEB9E4B0406FA09F969FE9CF8E0" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="pseudotereticollis">
<emphasis id="64DE997A9E4B0406FA09F969FA55F88B" box="[1415,1428,1696,1719]" italics="true" pageId="12" pageNumber="163">T</emphasis>
. 
<emphasis id="64DE997A9E4B0406FF19F90AFE9CF8E0" box="[151,349,1731,1756]" italics="true" pageId="12" pageNumber="163">pseudotereticollis</emphasis>
</taxonomicName>
by a pronotum slightly wider than long, a different elytral decumbent setae distribution pattern (cf. 
<figureCitation id="CE9159ED9E4B0406FF45F92EFED6F93C" box="[203,279,1767,1792]" captionStart="FIGURE 3" captionStartId="6.[151,250,1973,1995]" captionTargetBox="[256,1345,213,1925]" captionTargetId="figure@6.[238,1359,193,1948]" captionTargetPageId="6" captionText="FIGURE 3: Trichotaenia nzingae sp. n., elytra (dorsal view): a) typical decumbent setae pattern, b ‒ d) examples of variability among the decumbent setae pattern." httpUri="https://zenodo.org/record/253743/files/figure.png" pageId="12" pageNumber="163">Figs 3</figureCitation>
and 
<figureCitation id="CE9159ED9E4B0406FED5F92EFEA8F93C" box="[347,361,1767,1792]" captionStart="FIGURE 7" captionStartId="11.[151,250,1499,1521]" captionTargetBox="[171,1423,206,1462]" captionTargetId="figure@11.[151,1437,193,1477]" captionTargetPageId="11" captionText="FIGURE 7. Elytra (dorsal view): a and b) Trichotaenia pepetela sp. n., male and female, respectively, c) Trichotaenia tereticollis (W. Horn), d) Trichotaenia pseudotereticollis (W. Horn), e) Trichotaenia suturata (W. Horn)." httpUri="https://zenodo.org/record/253747/files/figure.png" pageId="12" pageNumber="163">7</figureCitation>
d) and the alveoli of the elytral disk slightly sharply walled (most flattened walled in 
<taxonomicName id="91AA3EEB9E4B0406FA09F921FE9CF918" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="pseudotereticollis">
<emphasis id="64DE997A9E4B0406FA09F921FE9CF918" italics="true" pageId="12" pageNumber="163">T. pseudotereticollis</emphasis>
</taxonomicName>
, conferring to this species a coppery lustre on whole disk, but not in the new species). On the other hand, 
<taxonomicName id="91AA3EEB9E4B0406FE92F8E6FDD3F974" box="[284,530,1839,1864]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="pepetela" status="sp. nov.">
<emphasis id="64DE997A9E4B0406FE92F8E6FDD3F974" box="[284,530,1839,1864]" italics="true" pageId="12" pageNumber="163">Trichotaenia pepetela</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7FED24019E4B0406FD97F8E6FD98F97B" box="[537,601,1839,1864]" pageId="12" pageNumber="163" rank="species">
<emphasis id="64DE997A9E4B0406FD97F8E6FD90F974" bold="true" box="[537,593,1839,1864]" pageId="12" pageNumber="163">sp. n</emphasis>
.
</taxonomicNameLabel>
can be separated also from 
<taxonomicName id="91AA3EEB9E4B0406FC1AF8F9FBB4F974" box="[916,1141,1839,1864]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="pseudotereticollis">
<emphasis id="64DE997A9E4B0406FC1AF8F9FBB4F974" box="[916,1141,1839,1864]" italics="true" pageId="12" pageNumber="163">T. pseudotereticollis</emphasis>
</taxonomicName>
by a very different elytral decumbent setae distribution pattern (cf. 
<figureCitation id="CE9159ED9E4B0406FDE6F89AFD6CF950" box="[616,685,1875,1900]" captionStart="FIGURE 7" captionStartId="11.[151,250,1499,1521]" captionTargetBox="[171,1423,206,1462]" captionTargetId="figure@11.[151,1437,193,1477]" captionTargetPageId="11" captionText="FIGURE 7. Elytra (dorsal view): a and b) Trichotaenia pepetela sp. n., male and female, respectively, c) Trichotaenia tereticollis (W. Horn), d) Trichotaenia pseudotereticollis (W. Horn), e) Trichotaenia suturata (W. Horn)." httpUri="https://zenodo.org/record/253747/files/figure.png" pageId="12" pageNumber="163">Figs 7</figureCitation>
a‒7b and 7d) and labral blackish colour pattern. Both new species can be discriminated easily from 
<taxonomicName id="91AA3EEB9E4B0406FD9CF8B1FDB9F9AC" box="[530,632,1911,1936]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="allardi">
<emphasis id="64DE997A9E4B0406FD9CF8B1FDB9F9AC" box="[530,632,1911,1936]" italics="true" pageId="12" pageNumber="163">T. allardi</emphasis>
</taxonomicName>
by the 5–9 antennomeres enlarged and foliated and the setose labrum, from 
<taxonomicName id="91AA3EEB9E4B0406FF5AF854FEB7F988" box="[212,374,1947,1972]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="duplosetosa">
<emphasis id="64DE997A9E4B0406FF5AF854FEB7F988" box="[212,374,1947,1972]" italics="true" pageId="12" pageNumber="163">T. duplosetosa</emphasis>
</taxonomicName>
by the absence of depigmented areas in the elytrae, from 
<taxonomicName id="91AA3EEB9E4B0406FC70F854FBB7F988" box="[1022,1142,1949,1972]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="suturata">
<emphasis id="64DE997A9E4B0406FC70F854FBB7F988" box="[1022,1142,1949,1972]" italics="true" pageId="12" pageNumber="163">T. suturata</emphasis>
</taxonomicName>
by a very different elytral pubescence pattern (cf. 
<figureCitation id="CE9159ED9E4B0406FE11F876FE24F9E4" box="[415,485,1983,2008]" captionStart="FIGURE 3" captionStartId="6.[151,250,1973,1995]" captionTargetBox="[256,1345,213,1925]" captionTargetId="figure@6.[238,1359,193,1948]" captionTargetPageId="6" captionText="FIGURE 3: Trichotaenia nzingae sp. n., elytra (dorsal view): a) typical decumbent setae pattern, b ‒ d) examples of variability among the decumbent setae pattern." httpUri="https://zenodo.org/record/253743/files/figure.png" pageId="12" pageNumber="163">Figs 3</figureCitation>
, 
<figureCitation id="CE9159ED9E4B0406FE7FF876FE3FF9E4" box="[497,510,1983,2008]" captionStart="FIGURE 7" captionStartId="11.[151,250,1499,1521]" captionTargetBox="[171,1423,206,1462]" captionTargetId="figure@11.[151,1437,193,1477]" captionTargetPageId="11" captionText="FIGURE 7. Elytra (dorsal view): a and b) Trichotaenia pepetela sp. n., male and female, respectively, c) Trichotaenia tereticollis (W. Horn), d) Trichotaenia pseudotereticollis (W. Horn), e) Trichotaenia suturata (W. Horn)." httpUri="https://zenodo.org/record/253747/files/figure.png" pageId="12" pageNumber="163">7</figureCitation>
a‒7b and 7e, see also Plate 46, 
<figureCitation id="CE9159ED9E4B0406FCD6F876FC67F9E4" box="[856,934,1983,2008]" captionStart="FIGURE 12" captionStartId="22.[151,250,1978,2000]" captionTargetBox="[166,1427,1004,1953]" captionTargetId="figure@22.[151,1436,995,1965]" captionTargetPageId="22" captionText="FIGURE 12. Aedeagus (lateral view) and median lobe apex (ventral view): a) Cylindera agualusai n. sp., paratype, Cachingues ‒ Chipica, Angola; b) Cylindera lutaria (Guérin-Méneville), Canchungo ‒ Calequisse, Guinea-Bissau." httpUri="https://zenodo.org/record/253751/files/figure.png" pageId="12" pageNumber="163">
Fig. 
<quantity id="9152E88D9E4B0406FC04F809FC67F9E4" box="[906,934,1984,2008]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="12" pageNumber="163" unit="in" value="12.0">12</quantity>
</figureCitation>
in 
<bibRefCitation id="323B38999E4B0406FC47F809FB85F9E4" author="Horn" box="[969,1092,1984,2008]" pageId="12" pageNumber="163" refString="Horn, W. (1938) 2000 Zeichnungen von Cicindelinae. Entomologische Beihefte aus Berlin-Dahlem, 5, 1 - 71." type="journal article" year="1938">Horn 1938</bibRefCitation>
), from 
<taxonomicName id="91AA3EEB9E4B0406FB1AF809FAC8F9E4" box="[1172,1289,1983,2008]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="rivalieri">
<emphasis id="64DE997A9E4B0406FB1AF809FAC8F9E4" box="[1172,1289,1983,2008]" italics="true" pageId="12" pageNumber="163">T. rivalieri</emphasis>
</taxonomicName>
by the shape of pronotum (cordiform or subcordiform 
<emphasis id="64DE997A9E4A0407FDF7FF50FD51FE8C" box="[633,656,153,176]" italics="true" pageId="13" pageNumber="164">vs</emphasis>
. subquadrate) and the very distinct notopleural suture and from 
<taxonomicName id="91AA3EEB9E4A0407FA09FF50FF2FFEE9" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="kudrnai">
<emphasis id="64DE997A9E4A0407FA09FF50FF2FFEE9" italics="true" pageId="13" pageNumber="164">T. kudrnai</emphasis>
</taxonomicName>
by the position of the elytral median transverse decumbent stripe of setae located around or only slightly after the middle and the presence of a transverse and arcuate subapical band (cf. 
<figureCitation id="CE9159ED9E4A0407FB98FF16FB9DFEC4" box="[1046,1116,223,248]" captionStart="FIGURE 3" captionStartId="6.[151,250,1973,1995]" captionTargetBox="[256,1345,213,1925]" captionTargetId="figure@6.[238,1359,193,1948]" captionTargetPageId="6" captionText="FIGURE 3: Trichotaenia nzingae sp. n., elytra (dorsal view): a) typical decumbent setae pattern, b ‒ d) examples of variability among the decumbent setae pattern." httpUri="https://zenodo.org/record/253743/files/figure.png" pageId="13" pageNumber="164">Figs 3</figureCitation>
, 
<figureCitation id="CE9159ED9E4A0407FBE6FF16FBB4FEC4" box="[1128,1141,223,248]" captionStart="FIGURE 7" captionStartId="11.[151,250,1499,1521]" captionTargetBox="[171,1423,206,1462]" captionTargetId="figure@11.[151,1437,193,1477]" captionTargetPageId="11" captionText="FIGURE 7. Elytra (dorsal view): a and b) Trichotaenia pepetela sp. n., male and female, respectively, c) Trichotaenia tereticollis (W. Horn), d) Trichotaenia pseudotereticollis (W. Horn), e) Trichotaenia suturata (W. Horn)." httpUri="https://zenodo.org/record/253747/files/figure.png" pageId="13" pageNumber="164">7</figureCitation>
a‒7b and 
<figureCitation id="CE9159ED9E4A0407FB6FFF16FAE3FEC4" box="[1249,1314,223,248]" captionStart="FIGURE 1" captionStartId="4.[151,255,1856,1879]" captionTargetBox="[166,1418,465,1810]" captionTargetId="figure@4.[151,1437,447,1831]" captionTargetPageId="4" captionText="FIGURE 1. Facies of Trichotaenia nzingae sp. n.: a) male holotype, b) female allotype." httpUri="https://zenodo.org/record/253741/files/figure.png" pageId="13" pageNumber="164">
Fig. 
<quantity id="9152E88D9E4A0407FA9AFF29FAE3FEC4" box="[1300,1314,224,248]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="13" pageNumber="164" unit="in" value="1.0">1</quantity>
</figureCitation>
in 
<bibRefCitation id="323B38999E4A0407FACBFF16FF15FF20" author="Cassola" pageId="13" pageNumber="164" refString="Cassola, F. (2008) A new Trichotaenia species from Tanzania (Coleoptera, Cicindelidae). Fragmenta entomologica, 40 (2), 243 - 248. http: // dx. doi. org / 10.4081 / fe. 2008.96" type="journal article" year="2008">Cassola 2008</bibRefCitation>
).
</paragraph>
<paragraph id="561545689E4B0406FD44FCC0FA82FB13" pageId="12" pageNumber="163">
<table id="24AAB7C89E4BFBF5FF5EFCC0FA9AFB13" box="[208,1371,777,1327]" gridcols="5" gridrows="16" pageId="12" pageNumber="163">
<tr id="E89A472A9E4BFBF5FF5EFCC0FA9AFD1C" box="[208,1371,777,800]" gridrow="0" pageId="12" pageNumber="163" rowspan-0="1">
<th id="AB4B2E569E4BFBF5FD44FCC0FC00FD1C" box="[714,961,777,800]" gridcol="1" gridrow="0" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FD44FCC0FC00FD1C" bold="true" box="[714,961,777,800]" pageId="12" pageNumber="163">With decumbent setae</emphasis>
</th>
<th id="AB4B2E569E4BFBF5FC6FFCC0FB88FD1C" box="[993,1097,777,800]" gridcol="2" gridrow="0" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FC6FFCC0FB88FD1C" bold="true" box="[993,1097,777,800]" pageId="12" pageNumber="163">Glabrous</emphasis>
</th>
<th id="AB4B2E569E4BFBF5FB0CFCC0FB18FD1C" box="[1154,1241,777,800]" gridcol="3" gridrow="0" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB0CFCC0FB18FD1C" bold="true" box="[1154,1241,777,800]" pageId="12" pageNumber="163">Distinct</emphasis>
</th>
<th id="AB4B2E569E4BFBF5FA88FCC0FA9AFD1C" box="[1286,1371,777,800]" gridcol="4" gridrow="0" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FA88FCC0FA9AFD1C" bold="true" box="[1286,1371,777,800]" pageId="12" pageNumber="163">Effaced</emphasis>
</th>
</tr>
<tr id="E89A472A9E4BFBF5FF5EFCEEFA9AFD03" box="[208,1371,807,831]" gridrow="1" pageId="12" pageNumber="163">
<th id="AB4B2E569E4BFBF5FF5EFCEEFD65FD03" box="[208,676,807,831]" gridcol="0" gridrow="1" pageId="12" pageNumber="163">
<taxonomicName id="91AA3EEB9E4B0406FF5EFCEEFE1DFD02" authority="Cassola, 1983" authorityName="Cassola" authorityYear="1983" box="[208,476,807,830]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="64DE997A9E4B0406FF5EFCEEFE82FD02" box="[208,323,807,830]" italics="true" pageId="12" pageNumber="163">T. africana</emphasis>
Cassola, 1983
</taxonomicName>
</th>
<td id="AB4B2E569E4BFBF5FD44FCEEFC00FD03" box="[714,961,807,831]" gridcol="1" gridrow="1" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FCA1FCE1FC9DFD03" bold="true" box="[815,860,808,831]" pageId="12" pageNumber="163">+(a)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FC6FFCEEFB88FD03" box="[993,1097,807,831]" gridcol="2" gridrow="1" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB86FCE1FBF7FD03" bold="true" box="[1032,1078,808,831]" pageId="12" pageNumber="163">+(a)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FB0CFCEEFB18FD03" box="[1154,1241,807,831]" gridcol="3" gridrow="1" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB17FCE1FB00FD03" bold="true" box="[1177,1217,808,831]" pageId="12" pageNumber="163">-(d)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FA88FCEEFA9AFD03" box="[1286,1371,807,831]" gridcol="4" gridrow="1" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FA97FCE1FA89FD03" bold="true" box="[1305,1352,808,831]" pageId="12" pageNumber="163">+(d)</emphasis>
</td>
</tr>
<tr id="E89A472A9E4BFBF5FF5EFC8FFA9AFD62" box="[208,1371,838,862]" gridrow="2" pageId="12" pageNumber="163">
<th id="AB4B2E569E4BFBF5FF5EFC8FFD65FD62" box="[208,676,838,862]" gridcol="0" gridrow="2" pageId="12" pageNumber="163">
<taxonomicName id="91AA3EEB9E4B0406FF5EFC8FFE0AFD61" authority="Cassola, 1983" authorityName="Cassola" authorityYear="1983" box="[208,459,838,861]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="allardi">
<emphasis id="64DE997A9E4B0406FF5EFC8FFEF3FD61" box="[208,306,838,861]" italics="true" pageId="12" pageNumber="163">T. allardi</emphasis>
Cassola, 1983
</taxonomicName>
</th>
<td id="AB4B2E569E4BFBF5FD44FC8FFC00FD62" box="[714,961,838,862]" gridcol="1" gridrow="2" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FCBFFC8EFC98FD62" bold="true" box="[817,857,839,862]" pageId="12" pageNumber="163">-(b)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FC6FFC8FFB88FD62" box="[993,1097,838,862]" gridcol="2" gridrow="2" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB86FC8EFBF7FD62" bold="true" box="[1032,1078,839,862]" pageId="12" pageNumber="163">+(b)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FB0CFC8FFB18FD62" box="[1154,1241,838,862]" gridcol="3" gridrow="2" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB16FC8EFB02FD62" bold="true" box="[1176,1219,839,862]" pageId="12" pageNumber="163">+(e)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FA88FC8FFA9AFD62" box="[1286,1371,838,862]" gridcol="4" gridrow="2" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FA90FC8EFA82FD62" bold="true" box="[1310,1347,839,862]" pageId="12" pageNumber="163">-(e)</emphasis>
</td>
</tr>
<tr id="E89A472A9E4BFBF5FF5EFCACFA9AFD41" box="[208,1371,869,893]" gridrow="3" pageId="12" pageNumber="163">
<th id="AB4B2E569E4BFBF5FF5EFCACFD65FD41" box="[208,676,869,893]" gridcol="0" gridrow="3" pageId="12" pageNumber="163">
<taxonomicName id="91AA3EEB9E4B0406FF5EFCAFFDDAFD40" authority="W. Horn, 1929" authorityName="W. Horn" authorityYear="1929" box="[208,539,869,893]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="duplosetosa">
<emphasis id="64DE997A9E4B0406FF5EFCAFFEA9FD41" box="[208,360,870,893]" italics="true" pageId="12" pageNumber="163">T. duplosetosa</emphasis>
(W. Horn, 1929)
</taxonomicName>
</th>
<td id="AB4B2E569E4BFBF5FD44FCACFC00FD41" box="[714,961,869,893]" gridcol="1" gridrow="3" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FCBEFCAFFC9DFD41" bold="true" box="[816,860,870,893]" pageId="12" pageNumber="163">+(c)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FC6FFCACFB88FD41" box="[993,1097,869,893]" gridcol="2" gridrow="3" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB82FCAFFBF0FD41" bold="true" box="[1036,1073,870,893]" pageId="12" pageNumber="163">-(c)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FB0CFCACFB18FD41" box="[1154,1241,869,893]" gridcol="3" gridrow="3" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB16FCAFFB02FD41" bold="true" box="[1176,1219,870,893]" pageId="12" pageNumber="163">+(e)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FA88FCACFA9AFD41" box="[1286,1371,869,893]" gridcol="4" gridrow="3" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FA90FCAFFA82FD41" bold="true" box="[1310,1347,870,893]" pageId="12" pageNumber="163">-(e)</emphasis>
</td>
</tr>
<tr id="E89A472A9E4BFBF5FF5EFC4DFA9AFDA0" box="[208,1371,900,924]" gridrow="4" pageId="12" pageNumber="163">
<th id="AB4B2E569E4BFBF5FF5EFC4DFD65FDA0" box="[208,676,900,924]" gridcol="0" gridrow="4" pageId="12" pageNumber="163">
<taxonomicName id="91AA3EEB9E4B0406FF5EFC4CFE14FDA7" authority="Cassola, 2008" authorityName="Cassola" authorityYear="2008" box="[208,469,900,924]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="kudrnai">
<emphasis id="64DE997A9E4B0406FF5EFC4CFEFDFDA0" box="[208,316,901,924]" italics="true" pageId="12" pageNumber="163">T. kudrnai</emphasis>
Cassola, 2008
</taxonomicName>
</th>
<td id="AB4B2E569E4BFBF5FD44FC4DFC00FDA0" box="[714,961,900,924]" gridcol="1" gridrow="4" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FCBEFC4CFC9DFDA0" bold="true" box="[816,860,901,924]" pageId="12" pageNumber="163">+(c)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FC6FFC4DFB88FDA0" box="[993,1097,900,924]" gridcol="2" gridrow="4" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB82FC4CFBF0FDA0" bold="true" box="[1036,1073,901,924]" pageId="12" pageNumber="163">-(c)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FB0CFC4DFB18FDA0" box="[1154,1241,900,924]" gridcol="3" gridrow="4" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB16FC4CFB02FDA0" bold="true" box="[1176,1219,901,924]" pageId="12" pageNumber="163">+(e)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FA88FC4DFA9AFDA0" box="[1286,1371,900,924]" gridcol="4" gridrow="4" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FA90FC4CFA82FDA0" bold="true" box="[1310,1347,901,924]" pageId="12" pageNumber="163">-(e)</emphasis>
</td>
</tr>
<tr id="E89A472A9E4BFBF5FF5EFC6AFA9AFD87" box="[208,1371,931,955]" gridrow="5" pageId="12" pageNumber="163">
<th id="AB4B2E569E4BFBF5FF5EFC6AFD65FD87" box="[208,676,931,955]" gridcol="0" gridrow="5" pageId="12" pageNumber="163">
<taxonomicName id="91AA3EEB9E4B0406FF5EFC6DFE08FD86" authority="Schule, 2011" authorityName="Schule" authorityYear="2011" box="[208,457,931,955]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="minettii">
<emphasis id="64DE997A9E4B0406FF5EFC6DFEFAFD87" box="[208,315,932,955]" italics="true" pageId="12" pageNumber="163">T. minettii</emphasis>
Schüle, 2011
</taxonomicName>
</th>
<td id="AB4B2E569E4BFBF5FD44FC6AFC00FD87" box="[714,961,931,955]" gridcol="1" gridrow="5" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FCBEFC6DFC9DFD87" bold="true" box="[816,860,932,955]" pageId="12" pageNumber="163">+(c)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FC6FFC6AFB88FD87" box="[993,1097,931,955]" gridcol="2" gridrow="5" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB82FC6DFBF0FD87" bold="true" box="[1036,1073,932,955]" pageId="12" pageNumber="163">-(c)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FB0CFC6AFB18FD87" box="[1154,1241,931,955]" gridcol="3" gridrow="5" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB17FC6DFB00FD87" bold="true" box="[1177,1217,932,955]" pageId="12" pageNumber="163">-(d)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FA88FC6AFA9AFD87" box="[1286,1371,931,955]" gridcol="4" gridrow="5" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FA97FC6DFA89FD87" bold="true" box="[1305,1352,932,955]" pageId="12" pageNumber="163">+(d)</emphasis>
</td>
</tr>
<tr id="E89A472A9E4BFBF5FF5EFC0BFA9AFDE7" box="[208,1371,962,987]" gridrow="6" pageId="12" pageNumber="163">
<th id="AB4B2E569E4BFBF5FF5EFC0BFD65FDE7" box="[208,676,962,987]" gridcol="0" gridrow="6" pageId="12" pageNumber="163">
<taxonomicName id="91AA3EEB9E4B0406FF5EFC0AFE02FDE5" authority="Werner, 2003" authorityName="Werner" authorityYear="2003" box="[208,451,962,986]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="mireki">
<emphasis id="64DE997A9E4B0406FF5EFC0AFEEFFDE6" box="[208,302,963,986]" italics="true" pageId="12" pageNumber="163">T. mireki</emphasis>
Werner, 2003
</taxonomicName>
</th>
<td id="AB4B2E569E4BFBF5FD44FC0BFC00FDE7" box="[714,961,962,987]" gridcol="1" gridrow="6" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FCBFFC0DFC98FDE7" bold="true" box="[817,857,964,987]" pageId="12" pageNumber="163">-(b)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FC6FFC0BFB88FDE7" box="[993,1097,962,987]" gridcol="2" gridrow="6" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB86FC0DFBF7FDE7" bold="true" box="[1032,1078,964,987]" pageId="12" pageNumber="163">+(b)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FB0CFC0BFB18FDE7" box="[1154,1241,962,987]" gridcol="3" gridrow="6" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB17FC0DFB00FDE7" bold="true" box="[1177,1217,964,987]" pageId="12" pageNumber="163">-(d)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FA88FC0BFA9AFDE7" box="[1286,1371,962,987]" gridcol="4" gridrow="6" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FA97FC0DFA89FDE7" bold="true" box="[1305,1352,964,987]" pageId="12" pageNumber="163">+(d)</emphasis>
</td>
</tr>
<tr id="E89A472A9E4BFBF5FF5EFC28FA9AFA2A" box="[208,1371,993,1046]" gridrow="7" pageId="12" pageNumber="163">
<th id="AB4B2E569E4BFBF5FF5EFC28FD65FA2A" box="[208,676,993,1046]" gridcol="0" gridrow="7" pageId="12" pageNumber="163">
<taxonomicName id="91AA3EEB9E4B0406FF5EFC2BFEC5FA2A" authority="Cassola, Werner &amp; Schule, 2009" authorityName="Cassola, Werner &amp; Schule" authorityYear="2009" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="mufumbweana">
<emphasis id="64DE997A9E4B0406FF5EFC2BFE42FDC5" box="[208,387,994,1017]" italics="true" pageId="12" pageNumber="163">T. mufumbweana</emphasis>
Cassola, Werner &amp; Schüle, 2009
</taxonomicName>
</th>
<td id="AB4B2E569E4BFBF5FD44FC28FC00FA2A" box="[714,961,993,1046]" gridcol="1" gridrow="7" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FCBFFC2AFC98FDC6" bold="true" box="[817,857,995,1018]" pageId="12" pageNumber="163">-(b)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FC6FFC28FB88FA2A" box="[993,1097,993,1046]" gridcol="2" gridrow="7" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB86FC2AFBF7FDC6" bold="true" box="[1032,1078,995,1018]" pageId="12" pageNumber="163">+(b)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FB0CFC28FB18FA2A" box="[1154,1241,993,1046]" gridcol="3" gridrow="7" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB17FC2AFB00FDC6" bold="true" box="[1177,1217,995,1018]" pageId="12" pageNumber="163">-(d)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FA88FC28FA9AFA2A" box="[1286,1371,993,1046]" gridcol="4" gridrow="7" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FA97FC2AFA89FDC6" bold="true" box="[1305,1352,995,1018]" pageId="12" pageNumber="163">+(d)</emphasis>
</td>
</tr>
<tr id="E89A472A9E4BFBF5FF5EFBD6FA9AFA6F" box="[208,1371,1055,1107]" gridrow="8" pageId="12" pageNumber="163">
<th id="AB4B2E569E4BFBF5FF5EFBD6FD65FA6F" box="[208,676,1055,1107]" gridcol="0" gridrow="8" pageId="12" pageNumber="163">
<taxonomicName id="91AA3EEB9E4B0406FF5EFBD6FEC5FA6F" authority="Cassola, Werner &amp; Schule, 2009" authorityName="Cassola, Werner &amp; Schule" authorityYear="2009" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="mwinilungae">
<emphasis id="64DE997A9E4B0406FF5EFBD6FEB3FA0A" box="[208,370,1055,1078]" italics="true" pageId="12" pageNumber="163">T. mwinilungae</emphasis>
Cassola, Werner &amp; Schüle, 2009
</taxonomicName>
</th>
<td id="AB4B2E569E4BFBF5FD44FBD6FC00FA6F" box="[714,961,1055,1107]" gridcol="1" gridrow="8" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FCBFFBE9FC98FA0B" bold="true" box="[817,857,1056,1079]" pageId="12" pageNumber="163">-(b)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FC6FFBD6FB88FA6F" box="[993,1097,1055,1107]" gridcol="2" gridrow="8" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB86FBE9FBF7FA0B" bold="true" box="[1032,1078,1056,1079]" pageId="12" pageNumber="163">+(b)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FB0CFBD6FB18FA6F" box="[1154,1241,1055,1107]" gridcol="3" gridrow="8" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB17FBE9FB00FA0B" bold="true" box="[1177,1217,1056,1079]" pageId="12" pageNumber="163">-(d)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FA88FBD6FA9AFA6F" box="[1286,1371,1055,1107]" gridcol="4" gridrow="8" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FA97FBE9FA89FA0B" bold="true" box="[1305,1352,1056,1079]" pageId="12" pageNumber="163">+(d)</emphasis>
</td>
</tr>
<tr id="E89A472A9E4BFBF5FF5EFB95FA9AFA48" box="[208,1371,1116,1140]" gridrow="9" pageId="12" pageNumber="163">
<th id="AB4B2E569E4BFBF5FF5EFB95FD65FA48" box="[208,676,1116,1140]" gridcol="0" gridrow="9" pageId="12" pageNumber="163">
<taxonomicName id="91AA3EEB9E4B0406FF5EFB95FEFDFA4F" box="[208,316,1116,1139]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="nzingae" status="sp. nov.">
<emphasis id="64DE997A9E4B0406FF5EFB95FEFDFA4F" box="[208,316,1116,1139]" italics="true" pageId="12" pageNumber="163">T. nzingae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7FED24019E4B0406FECDFB95FEBCFA4F" box="[323,381,1116,1139]" pageId="12" pageNumber="163" rank="species">
<emphasis id="64DE997A9E4B0406FECDFB95FEBCFA4F" bold="true" box="[323,381,1116,1139]" pageId="12" pageNumber="163">sp. n.</emphasis>
</taxonomicNameLabel>
</th>
<td id="AB4B2E569E4BFBF5FD44FB95FC00FA48" box="[714,961,1116,1140]" gridcol="1" gridrow="9" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FCBEFB94FC9DFA48" bold="true" box="[816,860,1117,1140]" pageId="12" pageNumber="163">+(c)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FC6FFB95FB88FA48" box="[993,1097,1116,1140]" gridcol="2" gridrow="9" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB82FB94FBF0FA48" bold="true" box="[1036,1073,1117,1140]" pageId="12" pageNumber="163">-(c)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FB0CFB95FB18FA48" box="[1154,1241,1116,1140]" gridcol="3" gridrow="9" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB16FB94FB02FA48" bold="true" box="[1176,1219,1117,1140]" pageId="12" pageNumber="163">+(e)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FA88FB95FA9AFA48" box="[1286,1371,1116,1140]" gridcol="4" gridrow="9" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FA90FB94FA82FA48" bold="true" box="[1310,1347,1117,1140]" pageId="12" pageNumber="163">-(e)</emphasis>
</td>
</tr>
<tr id="E89A472A9E4BFBF5FF5EFBB2FA9AFAAF" box="[208,1371,1147,1171]" gridrow="10" pageId="12" pageNumber="163">
<th id="AB4B2E569E4BFBF5FF5EFBB2FD65FAAF" box="[208,676,1147,1171]" gridcol="0" gridrow="10" pageId="12" pageNumber="163">
<taxonomicName id="91AA3EEB9E4B0406FF5EFBB2FE82FAAE" box="[208,323,1147,1170]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="pepetela" status="sp. nov.">
<emphasis id="64DE997A9E4B0406FF5EFBB2FE82FAAE" box="[208,323,1147,1170]" italics="true" pageId="12" pageNumber="163">T. pepetela</emphasis>
</taxonomicName>
<emphasis id="64DE997A9E4B0406FEC4FBB2FE44FAAE" bold="true" box="[330,389,1147,1170]" pageId="12" pageNumber="163">
<taxonomicNameLabel id="7FED24019E4B0406FEC4FBB2FE44FAAE" box="[330,389,1147,1170]" pageId="12" pageNumber="163" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</th>
<td id="AB4B2E569E4BFBF5FD44FBB2FC00FAAF" box="[714,961,1147,1171]" gridcol="1" gridrow="10" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FCBEFBB5FC9DFAAF" bold="true" box="[816,860,1148,1171]" pageId="12" pageNumber="163">+(c)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FC6FFBB2FB88FAAF" box="[993,1097,1147,1171]" gridcol="2" gridrow="10" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB82FBB5FBF0FAAF" bold="true" box="[1036,1073,1148,1171]" pageId="12" pageNumber="163">-(c)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FB0CFBB2FB18FAAF" box="[1154,1241,1147,1171]" gridcol="3" gridrow="10" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB16FBB5FB02FAAF" bold="true" box="[1176,1219,1148,1171]" pageId="12" pageNumber="163">+(e)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FA88FBB2FA9AFAAF" box="[1286,1371,1147,1171]" gridcol="4" gridrow="10" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FA90FBB5FA82FAAF" bold="true" box="[1310,1347,1148,1171]" pageId="12" pageNumber="163">-(e)</emphasis>
</td>
</tr>
<tr id="E89A472A9E4BFBF5FF5EFB53FA9AFA8E" box="[208,1371,1178,1202]" gridrow="11" pageId="12" pageNumber="163">
<th id="AB4B2E569E4BFBF5FF5EFB53FD65FA8E" box="[208,676,1178,1202]" gridcol="0" gridrow="11" pageId="12" pageNumber="163">
<taxonomicName id="91AA3EEB9E4B0406FF5EFB52FD82FA8D" authority="W. Horn, 1914" authorityName="W. Horn" authorityYear="1914" box="[208,579,1178,1202]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="pseudosuturalis">
<emphasis id="64DE997A9E4B0406FF5EFB52FE51FA8E" box="[208,400,1179,1202]" italics="true" pageId="12" pageNumber="163">T. pseudosuturalis</emphasis>
(W. Horn, 1914)
</taxonomicName>
</th>
<td id="AB4B2E569E4BFBF5FD44FB53FC00FA8E" box="[714,961,1178,1202]" gridcol="1" gridrow="11" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FCBEFB52FC9DFA8E" bold="true" box="[816,860,1179,1202]" pageId="12" pageNumber="163">+(c)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FC6FFB53FB88FA8E" box="[993,1097,1178,1202]" gridcol="2" gridrow="11" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB82FB52FBF0FA8E" bold="true" box="[1036,1073,1179,1202]" pageId="12" pageNumber="163">-(c)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FB0CFB53FB18FA8E" box="[1154,1241,1178,1202]" gridcol="3" gridrow="11" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB16FB52FB02FA8E" bold="true" box="[1176,1219,1179,1202]" pageId="12" pageNumber="163">+(e)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FA88FB53FA9AFA8E" box="[1286,1371,1178,1202]" gridcol="4" gridrow="11" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FA90FB52FA82FA8E" bold="true" box="[1310,1347,1179,1202]" pageId="12" pageNumber="163">-(e)</emphasis>
</td>
</tr>
<tr id="E89A472A9E4BFBF5FF5EFB70FA9AFAEE" box="[208,1371,1209,1234]" gridrow="12" pageId="12" pageNumber="163">
<th id="AB4B2E569E4BFBF5FF5EFB70FD65FAEE" box="[208,676,1209,1234]" gridcol="0" gridrow="12" pageId="12" pageNumber="163">
<taxonomicName id="91AA3EEB9E4B0406FF5EFB73FD96FAEC" authority="W. Horn, 1929" authorityName="W. Horn" authorityYear="1929" box="[208,599,1209,1233]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="pseudotereticollis">
<emphasis id="64DE997A9E4B0406FF5EFB73FE65FAED" box="[208,420,1210,1233]" italics="true" pageId="12" pageNumber="163">T. pseudotereticollis</emphasis>
(W. Horn, 1929)
</taxonomicName>
</th>
<td id="AB4B2E569E4BFBF5FD44FB70FC00FAEE" box="[714,961,1209,1234]" gridcol="1" gridrow="12" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FCBEFB72FC9DFAEE" bold="true" box="[816,860,1211,1234]" pageId="12" pageNumber="163">+(c)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FC6FFB70FB88FAEE" box="[993,1097,1209,1234]" gridcol="2" gridrow="12" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB82FB72FBF0FAEE" bold="true" box="[1036,1073,1211,1234]" pageId="12" pageNumber="163">-(c)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FB0CFB70FB18FAEE" box="[1154,1241,1209,1234]" gridcol="3" gridrow="12" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB16FB72FB02FAEE" bold="true" box="[1176,1219,1211,1234]" pageId="12" pageNumber="163">+(e)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FA88FB70FA9AFAEE" box="[1286,1371,1209,1234]" gridcol="4" gridrow="12" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FA90FB72FA82FAEE" bold="true" box="[1310,1347,1211,1234]" pageId="12" pageNumber="163">-(e)</emphasis>
</td>
</tr>
<tr id="E89A472A9E4BFBF5FF5EFB11FA9AFACD" box="[208,1371,1240,1265]" gridrow="13" pageId="12" pageNumber="163">
<th id="AB4B2E569E4BFBF5FF5EFB11FD65FACD" box="[208,676,1240,1265]" gridcol="0" gridrow="13" pageId="12" pageNumber="163">
<taxonomicName id="91AA3EEB9E4B0406FF5EFB10FDC1FAD3" authority="Basilewsky, 1958" authorityName="Basilewsky" authorityYear="1958" box="[208,512,1240,1264]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="rivalieri">
<emphasis id="64DE997A9E4B0406FF5EFB10FE81FACC" box="[208,320,1241,1264]" italics="true" pageId="12" pageNumber="163">T. rivalieri</emphasis>
Basilewsky, 1958
</taxonomicName>
</th>
<td id="AB4B2E569E4BFBF5FD44FB11FC00FACD" box="[714,961,1240,1265]" gridcol="1" gridrow="13" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FCBEFB13FC9DFACD" bold="true" box="[816,860,1242,1265]" pageId="12" pageNumber="163">+(c)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FC6FFB11FB88FACD" box="[993,1097,1240,1265]" gridcol="2" gridrow="13" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB82FB13FBF0FACD" bold="true" box="[1036,1073,1242,1265]" pageId="12" pageNumber="163">-(c)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FB0CFB11FB18FACD" box="[1154,1241,1240,1265]" gridcol="3" gridrow="13" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB16FB13FB02FACD" bold="true" box="[1176,1219,1242,1265]" pageId="12" pageNumber="163">+(e)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FA88FB11FA9AFACD" box="[1286,1371,1240,1265]" gridcol="4" gridrow="13" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FA90FB13FA82FACD" bold="true" box="[1310,1347,1242,1265]" pageId="12" pageNumber="163">-(e)</emphasis>
</td>
</tr>
<tr id="E89A472A9E4BFBF5FF5EFB3EFA9AFB2C" box="[208,1371,1271,1296]" gridrow="14" pageId="12" pageNumber="163">
<th id="AB4B2E569E4BFBF5FF5EFB3EFD65FB2C" box="[208,676,1271,1296]" gridcol="0" gridrow="14" pageId="12" pageNumber="163">
<taxonomicName id="91AA3EEB9E4B0406FF5EFB31FE34FB32" authority="W. Horn, 1915" authorityName="W. Horn" authorityYear="1915" box="[208,501,1271,1295]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="suturata">
<emphasis id="64DE997A9E4B0406FF5EFB31FE83FB33" box="[208,322,1272,1295]" italics="true" pageId="12" pageNumber="163">T. suturata</emphasis>
(W. Horn, 1915)
</taxonomicName>
</th>
<td id="AB4B2E569E4BFBF5FD44FB3EFC00FB2C" box="[714,961,1271,1296]" gridcol="1" gridrow="14" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FCBEFB30FC9DFB2C" bold="true" box="[816,860,1273,1296]" pageId="12" pageNumber="163">+(c)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FC6FFB3EFB88FB2C" box="[993,1097,1271,1296]" gridcol="2" gridrow="14" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB82FB30FBF0FB2C" bold="true" box="[1036,1073,1273,1296]" pageId="12" pageNumber="163">-(c)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FB0CFB3EFB18FB2C" box="[1154,1241,1271,1296]" gridcol="3" gridrow="14" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FB16FB30FB02FB2C" bold="true" box="[1176,1219,1273,1296]" pageId="12" pageNumber="163">+(e)</emphasis>
</td>
<td id="AB4B2E569E4BFBF5FA88FB3EFA9AFB2C" box="[1286,1371,1271,1296]" gridcol="4" gridrow="14" pageId="12" pageNumber="163">
<emphasis id="64DE997A9E4B0406FA90FB30FA82FB2C" bold="true" box="[1310,1347,1273,1296]" pageId="12" pageNumber="163">-(e)</emphasis>
</td>
</tr>
<tr id="E89A472A9E4BFBF5FF5EFADEFA9AFB13" box="[208,1371,1303,1327]" gridrow="15" pageId="12" pageNumber="163">
<th id="AB4B2E569E4BFBF5FF5EFADEFD65FB13" box="[208,676,1303,1327]" gridcol="0" gridrow="15" pageId="12" pageNumber="163">
<taxonomicName id="91AA3EEB9E4B0406FF5EFADEFDD6FB12" authority="Boheman, 1860" authorityName="Boheman" authorityYear="1860" box="[208,535,1303,1326]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="tereticollis">
<emphasis id="64DE997A9E4B0406FF5EFADEFE9BFB12" box="[208,346,1303,1326]" italics="true" pageId="12" pageNumber="163">T. tereticollis</emphasis>
(Boheman, 1860)
</taxonomicName>
</th>
<td id="AB4B2E569E4BFBF5FD44FADEFC00FB13" box="[714,961,1303,1327]" gridcol="1" gridrow="15" pageId="12" pageNumber="163">+(a)</td>
<td id="AB4B2E569E4BFBF5FC6FFADEFB88FB13" box="[993,1097,1303,1327]" gridcol="2" gridrow="15" pageId="12" pageNumber="163">+(a)</td>
<td id="AB4B2E569E4BFBF5FB0CFADEFB18FB13" box="[1154,1241,1303,1327]" gridcol="3" gridrow="15" pageId="12" pageNumber="163">+(e)</td>
<td id="AB4B2E569E4BFBF5FA88FADEFA9AFB13" box="[1286,1371,1303,1327]" gridcol="4" gridrow="15" pageId="12" pageNumber="163">-(e)</td>
</tr>
</table>
</paragraph>
<paragraph id="561545689E4A0407FF49FEEEFC49FFBB" blockId="13.[151,1436,151,1077]" pageId="13" pageNumber="164">
The two new species are very close also with 
<taxonomicName id="91AA3EEB9E4A0407FD42FEE0FCA1FF7C" box="[716,864,295,320]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="tereticollis">
<emphasis id="64DE997A9E4A0407FD42FEE0FCA1FF7C" box="[716,864,295,320]" italics="true" pageId="13" pageNumber="164">T. tereticollis</emphasis>
</taxonomicName>
, a species with distinct shoulders, but with labrum either setose or glabrous. Both new species can be segregated from 
<taxonomicName id="91AA3EEB9E4A0407FC1CFE84FBE4FF59" box="[914,1061,332,357]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="tereticollis">
<emphasis id="64DE997A9E4A0407FC1CFE84FBE4FF59" box="[914,1061,332,357]" italics="true" pageId="13" pageNumber="164">T. tereticollis</emphasis>
</taxonomicName>
by a different elytral pubescence pattern (cf. 
<figureCitation id="CE9159ED9E4A0407FE97FEA6FE9EFFB4" box="[281,351,367,392]" captionStart="FIGURE 3" captionStartId="6.[151,250,1973,1995]" captionTargetBox="[256,1345,213,1925]" captionTargetId="figure@6.[238,1359,193,1948]" captionTargetPageId="6" captionText="FIGURE 3: Trichotaenia nzingae sp. n., elytra (dorsal view): a) typical decumbent setae pattern, b ‒ d) examples of variability among the decumbent setae pattern." httpUri="https://zenodo.org/record/253743/files/figure.png" pageId="13" pageNumber="164">Figs 3</figureCitation>
, 
<figureCitation id="CE9159ED9E4A0407FEE2FEA6FEB8FFB4" box="[364,377,367,392]" captionStart="FIGURE 7" captionStartId="11.[151,250,1499,1521]" captionTargetBox="[171,1423,206,1462]" captionTargetId="figure@11.[151,1437,193,1477]" captionTargetPageId="11" captionText="FIGURE 7. Elytra (dorsal view): a and b) Trichotaenia pepetela sp. n., male and female, respectively, c) Trichotaenia tereticollis (W. Horn), d) Trichotaenia pseudotereticollis (W. Horn), e) Trichotaenia suturata (W. Horn)." httpUri="https://zenodo.org/record/253747/files/figure.png" pageId="13" pageNumber="164">7</figureCitation>
a‒7b and 7c) as well as the shape of pronotum.
</paragraph>
<paragraph id="561545689E4A0407FF49FE5DFD9BFC5C" blockId="13.[151,1436,151,1077]" pageId="13" pageNumber="164">
The two new species are easily discriminated between them by some features such as the elytral decumbent setae distribution pattern (cf. 
<figureCitation id="CE9159ED9E4A0407FE51FE7EFDE5FFEC" box="[479,548,439,464]" captionStart="FIGURE 3" captionStartId="6.[151,250,1973,1995]" captionTargetBox="[256,1345,213,1925]" captionTargetId="figure@6.[238,1359,193,1948]" captionTargetPageId="6" captionText="FIGURE 3: Trichotaenia nzingae sp. n., elytra (dorsal view): a) typical decumbent setae pattern, b ‒ d) examples of variability among the decumbent setae pattern." httpUri="https://zenodo.org/record/253743/files/figure.png" pageId="13" pageNumber="164">Figs 3</figureCitation>
and 
<figureCitation id="CE9159ED9E4A0407FDD5FE7EFDA8FFEC" box="[603,617,439,464]" captionStart="FIGURE 7" captionStartId="11.[151,250,1499,1521]" captionTargetBox="[171,1423,206,1462]" captionTargetId="figure@11.[151,1437,193,1477]" captionTargetPageId="11" captionText="FIGURE 7. Elytra (dorsal view): a and b) Trichotaenia pepetela sp. n., male and female, respectively, c) Trichotaenia tereticollis (W. Horn), d) Trichotaenia pseudotereticollis (W. Horn), e) Trichotaenia suturata (W. Horn)." httpUri="https://zenodo.org/record/253747/files/figure.png" pageId="13" pageNumber="164">7</figureCitation>
a‒7b), prothoracic epipleura visible (
<taxonomicName id="91AA3EEB9E4A0407FB89FE70FB40FFEC" box="[1031,1153,439,464]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="pepetela" status="sp. nov.">
<emphasis id="64DE997A9E4A0407FB89FE70FB40FFEC" box="[1031,1153,439,464]" italics="true" pageId="13" pageNumber="164">T. pepetela</emphasis>
</taxonomicName>
<emphasis id="64DE997A9E4A0407FB06FE7EFB11FFEC" bold="true" box="[1160,1232,439,464]" pageId="13" pageNumber="164">
<taxonomicNameLabel id="7FED24019E4A0407FB06FE7EFB09FFEC" box="[1160,1224,439,464]" pageId="13" pageNumber="164" rank="species">sp. n.</taxonomicNameLabel>
)
</emphasis>
or not (
<taxonomicName id="91AA3EEB9E4A0407FAA7FE70FA5DFFEC" box="[1321,1436,441,464]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="nzingae" status="sp. nov.">
<emphasis id="64DE997A9E4A0407FAA7FE70FA5DFFEC" box="[1321,1436,441,464]" italics="true" pageId="13" pageNumber="164">T. nzingae</emphasis>
</taxonomicName>
<emphasis id="64DE997A9E4A0407FF19FE15FF1EFFC9" bold="true" box="[151,223,476,501]" pageId="13" pageNumber="164">
<taxonomicNameLabel id="7FED24019E4A0407FF19FE15FF17FFC9" box="[151,214,476,501]" pageId="13" pageNumber="164" rank="species">sp. n.</taxonomicNameLabel>
)
</emphasis>
from above and elytral punctures in apical half not anastomosing to longitudinal chains (
<taxonomicName id="91AA3EEB9E4A0407FB30FE14FAF1FFC8" box="[1214,1328,477,500]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="nzingae" status="sp. nov.">
<emphasis id="64DE997A9E4A0407FB30FE14FAF1FFC8" box="[1214,1328,477,500]" italics="true" pageId="13" pageNumber="164">T. nzingae</emphasis>
</taxonomicName>
<emphasis id="64DE997A9E4A0407FAB8FE15FABFFFC9" bold="true" box="[1334,1406,476,501]" pageId="13" pageNumber="164">
<taxonomicNameLabel id="7FED24019E4A0407FAB8FE15FAB7FFC9" box="[1334,1398,476,501]" pageId="13" pageNumber="164" rank="species">sp. n.</taxonomicNameLabel>
)
</emphasis>
or conversely anastomosing to longitudinal chains (
<taxonomicName id="91AA3EEB9E4A0407FD37FDC8FCF2FC24" box="[697,819,511,536]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="pepetela" status="sp. nov.">
<emphasis id="64DE997A9E4A0407FD37FDC8FCF2FC24" box="[697,819,511,536]" italics="true" pageId="13" pageNumber="164">T. pepetela</emphasis>
</taxonomicName>
<emphasis id="64DE997A9E4A0407FCB7FE36FC49FC24" bold="true" box="[825,904,511,536]" pageId="13" pageNumber="164">
<taxonomicNameLabel id="7FED24019E4A0407FCB7FE36FCB8FC24" box="[825,889,511,536]" pageId="13" pageNumber="164" rank="species">sp. n.</taxonomicNameLabel>
).
</emphasis>
Finally both new species, by the distinct elytral shoulders, are easily separated from the remaining species of 
<taxonomicName id="91AA3EEB9E4A0407FCC6FDEDFC16FC01" box="[840,983,548,573]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="64DE997A9E4A0407FCC6FDEDFC16FC01" box="[840,983,548,573]" italics="true" pageId="13" pageNumber="164">Trichotaenia</emphasis>
</taxonomicName>
which are characterized by more or less effaced humeri, among other characters.
</paragraph>
<paragraph id="561545689E4A0407FF49FDA5FDB4FA09" blockId="13.[151,1436,151,1077]" pageId="13" pageNumber="164">
<emphasis id="64DE997A9E4A0407FF49FDA5FE75FCB9" bold="true" box="[199,436,620,645]" pageId="13" pageNumber="164">Ecological remarks.</emphasis>
Both 
<taxonomicName id="91AA3EEB9E4A0407FE76FDA5FD46FCB9" box="[504,647,620,645]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="64DE997A9E4A0407FE76FDA5FD46FCB9" box="[504,647,620,645]" italics="true" pageId="13" pageNumber="164">Trichotaenia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7FED24019E4A0407FD00FDA4FCD4FCB9" box="[654,789,620,645]" pageId="13" pageNumber="164" rank="species">new species</taxonomicNameLabel>
are until now restricted to 
<collectingCountry id="2EBD05F89E4A0407FBCFFDA5FB56FCB9" box="[1089,1175,620,645]" name="Angola" pageId="13" pageNumber="164">Angola</collectingCountry>
. The adults were found together with some adults of other tiger beetles (see also 
<taxonomicName id="91AA3EEB9E4A0407FCACFD58FC0CFC94" ID-CoL="4N49J" box="[802,973,655,680]" class="Insecta" family="Carabidae" genus="Prothymidia" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="angusticollis">
<emphasis id="64DE997A9E4A0407FCACFD58FC0CFC94" box="[802,973,655,680]" italics="true" pageId="13" pageNumber="164">P. angusticollis</emphasis>
</taxonomicName>
ecological remarks concerning Kakande and Catota localities) in countryside roads of whitish sandy soil. At such roads and paths with sand-clayish soil (or even small fields) in secondary open forest, 
<taxonomicName id="91AA3EEB9E4A0407FD07FD10FD3DFCCC" box="[649,764,729,752]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="nzingae" status="sp. nov.">
<emphasis id="64DE997A9E4A0407FD07FD10FD3DFCCC" box="[649,764,729,752]" italics="true" pageId="13" pageNumber="164">T. nzingae</emphasis>
</taxonomicName>
<emphasis id="64DE997A9E4A0407FC8DFD1EFC82FCCC" bold="true" box="[771,835,727,752]" pageId="13" pageNumber="164">
<taxonomicNameLabel id="7FED24019E4A0407FC8DFD1EFC82FCCC" box="[771,835,727,752]" pageId="13" pageNumber="164" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
could be observed at the Cachingues localities rather at the margins with darker soil, litter and herbaceous plants. Other sympatric tiger beetles (with preferences of their micro-habitats) have been 
<taxonomicName id="91AA3EEB9E4A0407FE4AFCE8FDAEFD04" ID-CoL="4N49J" box="[452,623,799,824]" class="Insecta" family="Carabidae" genus="Prothymidia" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="angusticollis">
<emphasis id="64DE997A9E4A0407FE4AFCE8FDAEFD04" box="[452,623,799,824]" italics="true" pageId="13" pageNumber="164">P. angusticollis</emphasis>
</taxonomicName>
(litter), 
<taxonomicName id="91AA3EEB9E4A0407FD40FCE8FC66FD04" ID-CoL="37PSG" box="[718,935,799,824]" class="Insecta" family="Carabidae" genus="Dromica" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="serietuberculata">
<emphasis id="64DE997A9E4A0407FD40FCE8FC66FD04" box="[718,935,799,824]" italics="true" pageId="13" pageNumber="164">D. serietuberculata</emphasis>
</taxonomicName>
f. 
<emphasis id="64DE997A9E4A0407FC47FCD6FBE4FD04" box="[969,1061,799,824]" italics="true" pageId="13" pageNumber="164">lundana</emphasis>
(litter), 
<taxonomicName id="91AA3EEB9E4A0407FB0DFCE9FA5DFD04" box="[1155,1436,800,824]" class="Insecta" family="Carabidae" genus="Ophryodera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="subSpecies" species="rofomarginata" subSpecies="poggei">
<emphasis id="64DE997A9E4A0407FB0DFCE9FA5DFD04" box="[1155,1436,800,824]" italics="true" pageId="13" pageNumber="164">
<collectingCountry id="2EBD05F89E4A0407FB0DFCE9FB5BFD0B" box="[1155,1178,800,823]" name="Iceland" pageId="13" pageNumber="164">O</collectingCountry>
. rofomarginata poggei
</emphasis>
</taxonomicName>
(white sandy soil), 
<taxonomicName id="91AA3EEB9E4A0407FEFDFC8CFDCEFD61" ID-CoL="3W4YK" box="[371,527,836,861]" class="Insecta" family="Carabidae" genus="Lophyra" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="infuscatula">
<emphasis id="64DE997A9E4A0407FEFDFC8CFDCEFD61" box="[371,527,836,861]" italics="true" pageId="13" pageNumber="164">L. infuscatula</emphasis>
</taxonomicName>
(white sandy soil), 
<taxonomicName id="91AA3EEB9E4A0407FD7DFC8CFC9BFD60" box="[755,858,837,860]" class="Insecta" family="Carabidae" genus="Elliptica" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="muata">
<emphasis id="64DE997A9E4A0407FD7DFC8CFC9BFD60" box="[755,858,837,860]" italics="true" pageId="13" pageNumber="164">E. muata</emphasis>
</taxonomicName>
f. 
<emphasis id="64DE997A9E4A0407FCF2FC8DFBEBFD61" box="[892,1066,836,861]" italics="true" pageId="13" pageNumber="164">parallelestriata</emphasis>
, (dark soil), 
<taxonomicName id="91AA3EEB9E4A0407FB31FC8CFA99FD61" ID-CoL="3W4YR" box="[1215,1368,836,861]" class="Insecta" family="Carabidae" genus="Lophyra" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="saraliensis">
<emphasis id="64DE997A9E4A0407FB31FC8CFA99FD61" box="[1215,1368,836,861]" italics="true" pageId="13" pageNumber="164">L. saraliensis</emphasis>
</taxonomicName>
(dark soil), 
<taxonomicName id="91AA3EEB9E4A0407FF57FCA0FE87FDBC" ID-CoL="37PSL" box="[217,326,871,896]" class="Insecta" family="Carabidae" genus="Dromica" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="64DE997A9E4A0407FF57FCA0FE87FDBC" box="[217,326,871,896]" italics="true" pageId="13" pageNumber="164">D. similis</emphasis>
</taxonomicName>
and 
<taxonomicName id="91AA3EEB9E4A0407FE0FFCA1FDD6FDBC" box="[385,535,871,896]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="agualusae" status="sp. nov.">
<emphasis id="64DE997A9E4A0407FE0FFCA1FDD6FDBC" box="[385,535,871,896]" italics="true" pageId="13" pageNumber="164">C. agualusae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7FED24019E4A0407FDAFFCAEFDA3FD43" box="[545,610,871,896]" pageId="13" pageNumber="164" rank="species">
<emphasis id="64DE997A9E4A0407FDAFFCAEFD9AFDBC" bold="true" box="[545,603,871,896]" pageId="13" pageNumber="164">sp. n</emphasis>
.
</taxonomicNameLabel>
Unlike to what was said by 
<bibRefCitation id="323B38999E4A0407FC3EFCAEFB5EFDBC" box="[944,1183,871,896]" pageId="13" pageNumber="164" refString="Cassola, F., Werner, K. &amp; Schule, P. (2009) Two new Trichotaenia species from Zambia (Coleoptera: Cicindelidae). Mitteilungen der Munchner Entomologischen Gesellschaft, 99, 11 - 16." type="journal article">
Cassola 
<emphasis id="64DE997A9E4A0407FB9EFCA0FB85FDBC" box="[1040,1092,871,896]" italics="true" pageId="13" pageNumber="164">et al</emphasis>
. (2009)
</bibRefCitation>
that “all 
<taxonomicName id="91AA3EEB9E4A0407FA83FCAEFA5DFDBC" box="[1293,1436,871,896]" class="Insecta" family="Carabidae" genus="Trichotaenia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="64DE997A9E4A0407FA83FCAEFA5DFDBC" box="[1293,1436,871,896]" italics="true" pageId="13" pageNumber="164">Trichotaenia</emphasis>
</taxonomicName>
species, even if normally winged, never fly and prefer to rapidly run on the ground through the grasses”, most of the adults of both new species were captured when attempted to fly rapidly into the neighbouring open secondary forest. Some specimens were captured also on the ground when attempted to run into the edges of the roads, but this behaviour seems more connected with the lower environmental temperature conditions in early morning or later near dusk or even under overcast sky.
</paragraph>
</subSubSection>
</treatment>
</document>