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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1142.90097" ID-Pensoft-Pub="1313-2970-1142-1" ID-Pensoft-UUID="1D4BDF04F72B59B9984E00D0B4FB338D" ID-ZooBank="A1129EE50F9941CFB73AE771B66E2486" ModsDocID="1313-2970-1142-1" checkinTime="1674655519517" checkinUser="pensoft" docAuthor="Tongkerd, Piyoros, Tumpeesuwan, Sakboworn, Inkhavilay, Khamla, Prasankok, Pongpun, Jeratthitikul, Ekgachai, Panha, Somsak &amp; Sutcharit, Chirasak" docDate="2023" docId="DD8864CB7857556E809760FA2168DC7B" docLanguage="en" docName="ZooKeys 1142: 1-144" docOrigin="ZooKeys 1142" docPubDate="2023-01-24" docSource="http://dx.doi.org/10.3897/zookeys.1142.90097" docTitle="Rhiostoma marioni" docType="treatment" docVersion="1" id="1D4BDF04F72B59B9984E00D0B4FB338D" lastPageNumber="1" masterDocId="1D4BDF04F72B59B9984E00D0B4FB338D" masterDocTitle="Systematic revision of the snorkel snail genus Rhiostoma Benson, 1860 (Gastropoda, Caenogastropoda, Cyclophoridae) with descriptions of new species" masterLastPageNumber="144" masterPageNumber="1" pageNumber="1" updateTime="1674655519517" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Systematic revision of the snorkel snail genus Rhiostoma Benson, 1860 (Gastropoda, Caenogastropoda, Cyclophoridae) with descriptions of new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tongkerd, Piyoros</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tumpeesuwan, Sakboworn</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand &amp; Present address: Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai, Maha Sarakham 44150 Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Inkhavilay, Khamla</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Natural Science, National University of Laos, P. O. Box 7322, Dongdok, Vientiane, Laos</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Prasankok, Pongpun</mods:namePart>
<mods:affiliation>School of Biology, Institute of Science, Suranaree University of Technology, Nakhon Ratchasima 30000, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jeratthitikul, Ekgachai</mods:namePart>
<mods:affiliation>Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Panha, Somsak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">jirasak4@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-24</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1142</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>144</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1142.90097</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1142.90097</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1142-1</mods:identifier>
<mods:identifier type="ZooBank">A1129EE50F9941CFB73AE771B66E2486</mods:identifier>
<mods:identifier type="Pensoft-UUID">1D4BDF04F72B59B9984E00D0B4FB338D</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:DD8864CB7857556E809760FA2168DC7B" httpUri="http://treatment.plazi.org/id/DD8864CB7857556E809760FA2168DC7B" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
7.
<taxonomicName ID-CoL="7SZX9" LSID="DD8864CB-7857-556E-8097-60FA2168DC7B" authority="(Ancey, 1898)" baseAuthorityName="Ancey" baseAuthorityYear="1898" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma marioni" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="marioni">Rhiostoma marioni (Ancey, 1898)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution map of Rhiostoma housei - species group." figureDoi="10.3897/zookeys.1142.90097.figure18" httpUri="https://binary.pensoft.net/fig/799876" pageId="0" pageNumber="1">Figs 18</figureCitation>
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A specimen mentioned as being the syntype of &quot; Pterocyclos marioni Ancey, 1898 &quot; NMW 1955.158.24090. B-F shell of Rhiostoma marioni B published figure of the type specimen (after Ancey 1898: pl. 9, fig. f.) and C-F specimens CUMZ 10210 from Feuang, Vientiane, Laos." figureDoi="10.3897/zookeys.1142.90097.figure24" httpUri="https://binary.pensoft.net/fig/799882" pageId="0" pageNumber="1">, 24B-F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Ancey" authorityYear="1898" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pterocyclos marioni" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="marioni">Pterocyclos marioni</taxonomicName>
(in part) Ancey, 1898: 137, pl. 9, fig. f. Type locality: Luang-prabang, Laos and Mont Hou, Tonkin.
<bibRefCitation author="Fischer, H" editor="Pavie, A" journalOrPublisher="Ernest Leroux, Editeur, Paris" pageId="0" pageNumber="1" pagination="390 - 450" refId="B42" refString="Fischer, H, Dautzenberg, P, 1904. Catalogue des mollusques terrestres et fluviatites de ľIndo-Chine orientale cites jusqu a ce jour. In: Pavie, A, Ed., Mission Pavie, Indo-Chine 1879-1895, Etudes Diverse Recherches sur Ľhistoire Naturelle de Ľhistoire Indo-Chine Orientale, Tome III. Ernest Leroux, Editeur, Paris: 390 - 450" title="Catalogue des mollusques terrestres et fluviatites de ľIndo-Chine orientale cites jusqu a ce jour." volumeTitle="Mission Pavie, Indo-Chine 1879 - 1895, Etudes Diverse Recherches sur Ľhistoire Naturelle de Ľhistoire Indo-Chine Orientale, Tome III." year="1904">Fischer and Dautzenberg 1904</bibRefCitation>
: 430.
<bibRefCitation author="Wood, H" journalOrPublisher="Biotir Report" pageId="0" pageNumber="1" pagination="1 - 162" refId="B186" refString="Wood, H, Gallichan, J, 2008. The new molluscan names of Cesar-Marie-Felix Ancey including illustrated type material from the National Museum of Wales. Studies in Biodiversity and Systematics of Terrestrial Organism from the National Museum of Wales. Biotir Report 3: 1 - 162" title="The new molluscan names of Cesar-Marie-Felix Ancey including illustrated type material from the National Museum of Wales. Studies in Biodiversity and Systematics of Terrestrial Organism from the National Museum of Wales." volume="3" year="2008">Wood and Gallichan 2008</bibRefCitation>
: 64, pl. 25, fig. 3 (syntype).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Pterocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterocyclus marioni" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="marioni">Pterocyclus marioni</taxonomicName>
-Kobelt 1911: 757.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
The figured specimens in
<bibRefCitation author="Ancey, C-F" journalOrPublisher="Bulletin du Musee ďHistoire Naturelle de Marseille" pageId="0" pageNumber="1" pagination="125 - 150" refId="B3" refString="Ancey, C-F, 1898. Observations sur les mollusques terrestres et fluviatiles recueillies dans I'Indo-Chine et particulaierement au Laos, par M. Henri Counillon, accompagnees de remarques sur ďautres mollusques de cette region. Bulletin du Musee ďHistoire Naturelle de Marseille 1: 125 - 150" title="Observations sur les mollusques terrestres et fluviatiles recueillies dans I'Indo-Chine et particulaierement au Laos, par M. Henri Counillon, accompagnees de remarques sur ďautres mollusques de cette region." volume="1" year="1898">Ancey (1898</bibRefCitation>
: pl. 9, fig. f) could not be located.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingCountry name="Laos">Laos</collectingCountry>
</emphasis>
: Wat Non Hinhare, Feuang District,
<collectingRegion country="Laos" name="Viangchan">Vientiane Province</collectingRegion>
: CUMZ 10210 (7 shells; Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A specimen mentioned as being the syntype of &quot; Pterocyclos marioni Ancey, 1898 &quot; NMW 1955.158.24090. B-F shell of Rhiostoma marioni B published figure of the type specimen (after Ancey 1898: pl. 9, fig. f.) and C-F specimens CUMZ 10210 from Feuang, Vientiane, Laos." figureDoi="10.3897/zookeys.1142.90097.figure24" httpUri="https://binary.pensoft.net/fig/799882" pageId="0" pageNumber="1">24C-F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Shell medium, thick and depressed shell. Detached whorl long, ~2
<normalizedToken originalValue="×">x</normalizedToken>
longer than aperture width, curved, slightly descending. Breathing device tubular and straight. Shell colour uniformly brownish or with irregular blotches.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Ancey" baseAuthorityYear="1898" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma marioni" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="marioni">
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma marioni</emphasis>
</taxonomicName>
has been considered a junior synonym of
<taxonomicName lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
<emphasis italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
</taxonomicName>
from Thailand (Kobelt 1911). However, this species differs from
<taxonomicName lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
<emphasis italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
</taxonomicName>
in having a monochrome brownish shell colour, long detached whorl, and long straight tubular breathing device. In comparison,
<taxonomicName lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
<emphasis italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
</taxonomicName>
tends to have a brownish zigzag pattern, medium detached-whorl length, and a curved breathing tube.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Shell</emphasis>
.
</emphasis>
Shell medium, cW 17.1-20.5 mm, cH 9.1-10.6 mm, thick, and sub-discoidal to discoidal shape; detached-whorl length 14.0-20.5 mm. Apex acute; spire slightly elevated. Whorls 5 to 6, convex, increasing regularly; suture wide and deep; last whorl rounded and stout. Shell surface with fine growth lines. Periostracum thin corneous, and transparent. Shell colour uniformly brownish to dark brown (rarely with brownish blotches); ventral shell surface with paler colour; with narrow and dark spiral band on periphery. Detached whorl long, ~ 2
<normalizedToken originalValue="×">x</normalizedToken>
longer than apertural width, curved and descending. Peristome circular and double; lip thickened, expanded and multi-layered. Aperture opened sub-ventrally. Breathing device tubular and its tip often attached to preceding whorl; outer lip forming a long, straight, closed tube, located just behind apertural lip; inner lip with small hole inside aperture. Umbilicus widely opened and deep. Operculum calcareous, cup-shaped, and multispiral (Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A specimen mentioned as being the syntype of &quot; Pterocyclos marioni Ancey, 1898 &quot; NMW 1955.158.24090. B-F shell of Rhiostoma marioni B published figure of the type specimen (after Ancey 1898: pl. 9, fig. f.) and C-F specimens CUMZ 10210 from Feuang, Vientiane, Laos." figureDoi="10.3897/zookeys.1142.90097.figure24" httpUri="https://binary.pensoft.net/fig/799882" pageId="0" pageNumber="1">24B-F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is currently known only from Vientiane Province, Laos (Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution map of Rhiostoma housei - species group." figureDoi="10.3897/zookeys.1142.90097.figure18" httpUri="https://binary.pensoft.net/fig/799876" pageId="0" pageNumber="1">18</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species was described based on two specimen lots from &quot;Luang-prabang, Laos&quot; and &quot;Mont Hou, Tonkin&quot; [probably in the area of Muang Khua and Yot Ou Districts, Phongsaly Province, Laos (see
<bibRefCitation author="Pavie, A" journalOrPublisher="Augustin Challamel, Editeur, Paris, France" pageId="0" pageNumber="1" refId="B126" refString="Pavie, A, 1903. Mission Pavie Indo-Chine, 1879-1895, Atlas, Notices et Cartes. Augustin Challamel, Editeur, Paris, France" title="Mission Pavie Indo-Chine, 1879 - 1895, Atlas, Notices et Cartes." year="1903">Pavie 1903</bibRefCitation>
: pl. 1)]. The original description includes an illustration of a single specimen and one set of measurements (
<bibRefCitation author="Ancey, C-F" journalOrPublisher="Bulletin du Musee ďHistoire Naturelle de Marseille" pageId="0" pageNumber="1" pagination="125 - 150" refId="B3" refString="Ancey, C-F, 1898. Observations sur les mollusques terrestres et fluviatiles recueillies dans I'Indo-Chine et particulaierement au Laos, par M. Henri Counillon, accompagnees de remarques sur ďautres mollusques de cette region. Bulletin du Musee ďHistoire Naturelle de Marseille 1: 125 - 150" title="Observations sur les mollusques terrestres et fluviatiles recueillies dans I'Indo-Chine et particulaierement au Laos, par M. Henri Counillon, accompagnees de remarques sur ďautres mollusques de cette region." volume="1" year="1898">Ancey 1898</bibRefCitation>
). Unfortunately, this figured specimen could not be located from the RBINS or NMW collection. The specimen lot NMW 1955.158.24090 consisting of a single specimen (with operculum) that has
<normalizedToken originalValue="Ancey’s">Ancey's</normalizedToken>
type label and collection locality
<normalizedToken originalValue="“Luang-prabang”">&quot;Luang-prabang&quot;</normalizedToken>
was considered the possible holotype by
<bibRefCitation author="Wood, H" journalOrPublisher="Biotir Report" pageId="0" pageNumber="1" pagination="1 - 162" refId="B186" refString="Wood, H, Gallichan, J, 2008. The new molluscan names of Cesar-Marie-Felix Ancey including illustrated type material from the National Museum of Wales. Studies in Biodiversity and Systematics of Terrestrial Organism from the National Museum of Wales. Biotir Report 3: 1 - 162" title="The new molluscan names of Cesar-Marie-Felix Ancey including illustrated type material from the National Museum of Wales. Studies in Biodiversity and Systematics of Terrestrial Organism from the National Museum of Wales." volume="3" year="2008">Wood and Gallichan (2008</bibRefCitation>
: pl. 25, figs 3, iv). However, this surviving specimen looks different from the actual illustrated specimen in the original publication. This specimen has a flattened shell, short detached whorl, and notch-shaped breathing device projecting anteriorly (Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A specimen mentioned as being the syntype of &quot; Pterocyclos marioni Ancey, 1898 &quot; NMW 1955.158.24090. B-F shell of Rhiostoma marioni B published figure of the type specimen (after Ancey 1898: pl. 9, fig. f.) and C-F specimens CUMZ 10210 from Feuang, Vientiane, Laos." figureDoi="10.3897/zookeys.1142.90097.figure24" httpUri="https://binary.pensoft.net/fig/799882" pageId="0" pageNumber="1">24A</figureCitation>
). Although the operculum is not clearly mentioned in the original description, the operculum accompanied by this specimen lot (NMW 1955.158.24090) is dark corneous, plate-shaped and multispiral with elevated lamellae (Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A specimen mentioned as being the syntype of &quot; Pterocyclos marioni Ancey, 1898 &quot; NMW 1955.158.24090. B-F shell of Rhiostoma marioni B published figure of the type specimen (after Ancey 1898: pl. 9, fig. f.) and C-F specimens CUMZ 10210 from Feuang, Vientiane, Laos." figureDoi="10.3897/zookeys.1142.90097.figure24" httpUri="https://binary.pensoft.net/fig/799882" pageId="0" pageNumber="1">24A</figureCitation>
). It is distinct from the typical operculum of
<taxonomicName authorityName="Tongkerd &amp; Tumpeesuwan &amp; Inkhavilay &amp; Prasankok &amp; Jeratthitikul &amp; Panha &amp; Sutcharit" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Schematic of shell morphology and terminology of the Rhiostoma A apical view with shell terminology and measurements of coiled-whorl width and detached-whorl length B operculum showing side and cross-section views C apertural views showing various shell shapes as recognised in this revision: flattened, discoidal, sub-discoidal and heliciform." figureDoi="10.3897/zookeys.1142.90097.figure2" httpUri="https://binary.pensoft.net/fig/799860" pageId="0" pageNumber="1">2</figureCitation>
), and more closely resembling the
<taxonomicName authorityName="Mollendorff" authorityYear="1885" class="Gastropoda" family="Cyclophoridae" genus="Ptychopoma" kingdom="Animalia" lsidName="Ptychopoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Ptychopoma</emphasis>
</taxonomicName>
or
<taxonomicName authorityName="W. T. Blanford" authorityYear="1863" class="Gastropoda" family="Cyclophoridae" genus="Scabrina." kingdom="Animalia" lsidName="Scabrina." order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Scabrina.</emphasis>
</taxonomicName>
This specimen is contradicting to
<normalizedToken originalValue="Ancey’s">Ancey's</normalizedToken>
concept of &quot;
<taxonomicName authorityName="Ancey" authorityYear="1898" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos marioni" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="marioni">
<emphasis italics="true" pageId="0" pageNumber="1">Pterocyclos marioni</emphasis>
</taxonomicName>
&quot;, whereas the species description agrees well with the specimen illustration (
<bibRefCitation author="Ancey, C-F" journalOrPublisher="Bulletin du Musee ďHistoire Naturelle de Marseille" pageId="0" pageNumber="1" pagination="125 - 150" refId="B3" refString="Ancey, C-F, 1898. Observations sur les mollusques terrestres et fluviatiles recueillies dans I'Indo-Chine et particulaierement au Laos, par M. Henri Counillon, accompagnees de remarques sur ďautres mollusques de cette region. Bulletin du Musee ďHistoire Naturelle de Marseille 1: 125 - 150" title="Observations sur les mollusques terrestres et fluviatiles recueillies dans I'Indo-Chine et particulaierement au Laos, par M. Henri Counillon, accompagnees de remarques sur ďautres mollusques de cette region." volume="1" year="1898">Ancey 1898</bibRefCitation>
: pl. 9, fig. f). Therefore, we consider specimen NMW 1955.158.24090 (Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A specimen mentioned as being the syntype of &quot; Pterocyclos marioni Ancey, 1898 &quot; NMW 1955.158.24090. B-F shell of Rhiostoma marioni B published figure of the type specimen (after Ancey 1898: pl. 9, fig. f.) and C-F specimens CUMZ 10210 from Feuang, Vientiane, Laos." figureDoi="10.3897/zookeys.1142.90097.figure24" httpUri="https://binary.pensoft.net/fig/799882" pageId="0" pageNumber="1">24A</figureCitation>
;
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.834.28800" author="Inkhavilay, K" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 166" refId="B65" refString="Inkhavilay, K, Sutcharit, C, Bantaowong, U, Chanabun, R, Siriwut, W, Srisonchai, R, Pholyotha, A, Jirapatrasilp, P, Panha, S, 2019. Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys 834: 1 - 166, DOI: https://doi.org/10.3897/zookeys.834.28800" title="Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda)." url="https://doi.org/10.3897/zookeys.834.28800" volume="834" year="2019">Inkhavilay et al. 2019</bibRefCitation>
: fig. 11a;
<bibRefCitation DOI="https://doi.org/10.35885/ruthenica.2021.30(3).4" author="Do, DS" journalOrPublisher="Ruthenica" pageId="0" pageNumber="1" pagination="165 - 172" refId="B36" refString="Do, DS, Nguyen, TS, Do, HL, 2020b. The land snail genus Rhiostoma Benson, 1860 from Vietnam (Gastropoda: Caenogastropoda: Cyclophoridae), with description of a new species. Ruthenica 30 (3): 165 - 172, DOI: https://doi.org/10.35885/ruthenica.2021.30(3).4" title="The land snail genus Rhiostoma Benson, 1860 from Vietnam (Gastropoda: Caenogastropoda: Cyclophoridae), with description of a new species." url="https://doi.org/10.35885/ruthenica.2021.30(3).4" volume="30" year="2020 b">Do et al. 2020b</bibRefCitation>
: fig. 1d) as a distinct species. We have excluded that specimen from our attention in the type series.
</paragraph>
<paragraph pageId="0" pageNumber="1">
This redescription is based on the recently collected specimens from Feuang District, Vientiane Province, Laos. Before establishing the French Protectorate of Laos in 1893 (
<bibRefCitation author="Pavie, A" journalOrPublisher="Augustin Challamel, Editeur, Paris, France" pageId="0" pageNumber="1" refId="B126" refString="Pavie, A, 1903. Mission Pavie Indo-Chine, 1879-1895, Atlas, Notices et Cartes. Augustin Challamel, Editeur, Paris, France" title="Mission Pavie Indo-Chine, 1879 - 1895, Atlas, Notices et Cartes." year="1903">Pavie 1903</bibRefCitation>
: pl. 6;
<bibRefCitation author="Tips, WJE" journalOrPublisher="White Lotus, Bangkok, Thailand: Bangkok" pageId="0" pageNumber="1" refId="B171" refString="Tips, WJE, 1999. Atlas of the Pavie Mission: Laos, Cambodia, Siam, Yunnan, and Vietnam. The Pavie Mission Indochina Papers, 1879-1895; Volume 2. White Lotus, Bangkok, Thailand: Bangkok" title="Atlas of the Pavie Mission: Laos, Cambodia, Siam, Yunnan, and Vietnam. The Pavie Mission Indochina Papers, 1879 - 1895; Volume 2." year="1999">Tips 1999</bibRefCitation>
: 131), the historical name &quot;Luang Prabang&quot; generally referred to the city or the Kingdom Luang Prabang (during the 18th to 19th century). This name encompasses a wide geographical area, mainly present-day northern Laos: Bokeo, Houa Phanh, Luang Namtha, Luang Prabang, Oudomxay, Phongsaly and Vientiane provinces. Therefore, this newly collected specimen from Vientiane Province is considered the topotypic specimen of this species. This specimen lot possesses a long detached whorl and a long tubular-shaped breathing device identical to the unique characteristic of
<taxonomicName lsidName="R. marioni" pageId="0" pageNumber="1" rank="species" species="marioni">
<emphasis italics="true" pageId="0" pageNumber="1">R. marioni</emphasis>
</taxonomicName>
; it agrees well with the illustration (Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A specimen mentioned as being the syntype of &quot; Pterocyclos marioni Ancey, 1898 &quot; NMW 1955.158.24090. B-F shell of Rhiostoma marioni B published figure of the type specimen (after Ancey 1898: pl. 9, fig. f.) and C-F specimens CUMZ 10210 from Feuang, Vientiane, Laos." figureDoi="10.3897/zookeys.1142.90097.figure24" httpUri="https://binary.pensoft.net/fig/799882" pageId="0" pageNumber="1">24B</figureCitation>
) and the species description.
</paragraph>
<paragraph pageId="0" pageNumber="1">
The previous records under the name
<taxonomicName authorityName="Tongkerd &amp; Tumpeesuwan &amp; Inkhavilay &amp; Prasankok &amp; Jeratthitikul &amp; Panha &amp; Sutcharit" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma</emphasis>
</taxonomicName>
&quot;
<taxonomicName baseAuthorityName="Ancey" baseAuthorityYear="1898" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma marioni" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="marioni">
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma marioni</emphasis>
</taxonomicName>
&quot; from Lai Chu, Son La, Dien Bien and Lau Cai provinces, North Vietnam, as mentioned by
<bibRefCitation DOI="https://doi.org/10.35885/ruthenica.2021.30(3).4" author="Do, DS" journalOrPublisher="Ruthenica" pageId="0" pageNumber="1" pagination="165 - 172" refId="B36" refString="Do, DS, Nguyen, TS, Do, HL, 2020b. The land snail genus Rhiostoma Benson, 1860 from Vietnam (Gastropoda: Caenogastropoda: Cyclophoridae), with description of a new species. Ruthenica 30 (3): 165 - 172, DOI: https://doi.org/10.35885/ruthenica.2021.30(3).4" title="The land snail genus Rhiostoma Benson, 1860 from Vietnam (Gastropoda: Caenogastropoda: Cyclophoridae), with description of a new species." url="https://doi.org/10.35885/ruthenica.2021.30(3).4" volume="30" year="2020 b">Do et al. (2020b)</bibRefCitation>
, should be referred to another species (see under
<taxonomicName lsidName="R. abletti" pageId="0" pageNumber="1" rank="species" species="abletti">
<emphasis italics="true" pageId="0" pageNumber="1">R. abletti</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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