treatments-xml/data/DD/4D/BD/DD4DBDD074D5F0141F66DF0FF85B9109.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.411.6858" ID-GBIF-Dataset="7280c1cf-430f-494c-9440-8f47aaacebf4" ID-PMC="PMC4042821" ID-Pensoft-Pub="1313-2970-411-145" ID-PubMed="24899858" ID-ZBK="E80CF090FD5E4F249E88ED3B04ECD603" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-411-145" ModsDocOrigin="ZooKeys 411" ModsDocTitle="Two new species of Trichoceridae from the Middle Jurassic Jiulongshan Formation of Inner Mongolia, China" checkinTime="1451245851348" checkinUser="pensoft" docAuthor="Dong, Fei, Shih, Chungkun &amp; Ren, Dong" docDate="2014" docId="DD4DBDD074D5F0141F66DF0FF85B9109" docLanguage="en" docName="ZooKeys 411: 145-160" docOrigin="ZooKeys 411" docSource="http://dx.doi.org/10.3897/zookeys.411.6858" docTitle="Eotrichocera (Archaeotrichocera) longensis Dong, Shih &amp; Ren, 2014, sp. n." docType="treatment" docUuid="8A0D358E-7BCA-476B-A0A9-6EECA93FCBA8" docUuidSource="ZooBank" docVersion="4" lastPageNumber="150" masterDocId="FFF4FFE3FF9F7D4FFFC4FF9C2955FFA3" masterDocTitle="Two new species of Trichoceridae from the Middle Jurassic Jiulongshan Formation of Inner Mongolia, China" masterLastPageNumber="160" masterPageNumber="145" pageNumber="147" updateTime="1668158530965" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Two new species of Trichoceridae from the Middle Jurassic Jiulongshan Formation of Inner Mongolia, China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dong, Fei</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shih, Chungkun</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ren, Dong</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>411</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>145</mods:start>
<mods:end>160</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.411.6858</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.411.6858</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-411-145</mods:identifier>
<mods:identifier type="ZBK">E80CF090FD5E4F249E88ED3B04ECD603</mods:identifier>
<mods:identifier type="ZooBank">E80CF090FD5E4F249E88ED3B04ECD603</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152053140" LSID="urn:lsid:zoobank.org:act:8A0D358E-7BCA-476B-A0A9-6EECA93FCBA8" httpUri="http://treatment.plazi.org/id/DD4DBDD074D5F0141F66DF0FF85B9109" lastPageId="5" lastPageNumber="150" pageId="2" pageNumber="147">
<subSubSection pageId="2" pageNumber="147" type="nomenclature">
<paragraph pageId="2" pageNumber="147">
<taxonomicName LSID="http://zoobank.org/8A0D358E-7BCA-476B-A0A9-6EECA93FCBA8" class="Insecta" family="Trichoceridae" genus="Eotrichocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrichocera (Archaeotrichocera) longensis" order="Diptera" pageId="2" pageNumber="147" phylum="Arthropoda" rank="species" species="longensis" subGenus="Archaeotrichocera">Eotrichocera (Archaeotrichocera) longensis</taxonomicName>
<taxonomicNameLabel pageId="2" pageNumber="147">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="147" type="etymology">
<paragraph pageId="2" pageNumber="147">Etymology.</paragraph>
<paragraph pageId="2" pageNumber="147">
<normalizedToken originalValue="“longensis”">&quot;longensis&quot;</normalizedToken>
is a Latin word, referring to the long leg of this specimen.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="147" type="diagnosis">
<paragraph pageId="2" pageNumber="147">Diagnosis.</paragraph>
<paragraph pageId="2" pageNumber="147">Sc rather short about 0.65 times as long as the wing and ending at anterior margin proximad of R2; Rs forking proximad of 2/3 (at about 0.55) times of wing length; the d-cell narrow and long (about 2.5 times as long as wide); A2 short and bending sharply toward anterior margin (angle about 128°).</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="147" type="holotype">
<paragraph pageId="2" pageNumber="147">Holotype.</paragraph>
<paragraph pageId="2" pageNumber="147">An almost complete female specimen with well-preserved body, wings and head. Specimen number CNU-DIP-NN2013133. Wing length 9.0 mm, width 3.8 mm (Figs 1A, 2, 3A).</paragraph>
<caption pageId="2" pageNumber="147">
<paragraph pageId="2" pageNumber="147">
Figure 1. 
<taxonomicName class="Insecta" family="Trichoceridae" genus="Eotrichocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrichocera (Archaeotrichocera) longensis" order="Diptera" pageId="2" pageNumber="147" phylum="Arthropoda" rank="species" species="longensis" subGenus="Archaeotrichocera">Eotrichocera (Archaeotrichocera) longensis</taxonomicName>
sp. n. Holotype, specimen CNU-DIP-NN2013133 A Photograph. Paratype, specimen CNU-DIP-NN2013131 B Photograph. Scale bars = 1 mm.
</paragraph>
</caption>
<caption pageId="2" pageNumber="147">
<paragraph pageId="2" pageNumber="147">
Figure 2. 
<taxonomicName class="Insecta" family="Trichoceridae" genus="Eotrichocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrichocera (Archaeotrichocera) longensis" order="Diptera" pageId="2" pageNumber="147" phylum="Arthropoda" rank="species" species="longensis" subGenus="Archaeotrichocera">Eotrichocera (Archaeotrichocera) longensis</taxonomicName>
sp. n. Holotype, specimen CNU-DIP-NN2013133 A Line drawing B Tarsus of the mid leg. Scale bars = 1 mm; t1 = the first segment of tarsus; t2 = the second segment of tarsus; t3 = the third segment of tarsus; t4 = the fourth segment of tarsus.
</paragraph>
</caption>
<caption pageId="2" pageNumber="147">
<paragraph pageId="2" pageNumber="147">
Figure 3. 
<taxonomicName class="Insecta" family="Trichoceridae" genus="Eotrichocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrichocera (Archaeotrichocera) longensis" order="Diptera" pageId="2" pageNumber="147" phylum="Arthropoda" rank="species" species="longensis" subGenus="Archaeotrichocera">Eotrichocera (Archaeotrichocera) longensis</taxonomicName>
sp. n. Holotype, specimen CNU-DIP-NN2013133 A Line drawing of left wing. Paratype, specimen CNU-DIP-NN2013131 B Line drawing of left wing. Scale bars = 1 mm
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="3" pageNumber="148" type="paratype">
<paragraph pageId="3" pageNumber="148">
<pageBreakToken pageId="3" pageNumber="148" start="start">Paratype</pageBreakToken>
.
</paragraph>
<paragraph pageId="3" pageNumber="148">A female specimen with body and wings, specimen number CNU-DIP-NN2013131. Wing length 7.7 mm, width 3 mm (Figs 1B, 3B).</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="148" type="locality and horizon">
<paragraph pageId="3" pageNumber="148">Locality and horizon.</paragraph>
<paragraph pageId="3" pageNumber="148">Jiulongshan Formation, Late Middle Jurassic, Daohugou Village, Ningcheng County, Inner Mongolia Autonomous Region, China.</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="150" pageId="3" pageNumber="148" type="description">
<paragraph pageId="3" pageNumber="148">Description.</paragraph>
<paragraph pageId="3" pageNumber="148">Based on Holotype, different characters of the paratype CNU-DIP-NN2013131 in brackets. Medium-sized winter crane flies, body length (including head) 13 mm with well preserved wings, body and head, [paratype body length (excluding head) 10.5 mm].</paragraph>
<paragraph pageId="3" pageNumber="148">Head: Antenna very long, about 3.5 times as long as the head length, palpi about two times as long as the head length, compound eyes preserved.</paragraph>
<paragraph pageId="4" pageNumber="149">
<pageBreakToken pageId="4" pageNumber="149" start="start">Thorax</pageBreakToken>
: Much higher, in lateral view, than that of the abdomen, subcircular in shape, with robust and well-developed mesonotum. The halters spoon-type and the length of halters as long as thorax.
</paragraph>
<paragraph pageId="5" pageNumber="150">
<pageBreakToken pageId="5" pageNumber="150" start="start">Wings</pageBreakToken>
: Wing is shorter than abdomen, not covering the end of the abdomen. Length 9.0 mm [Paratype with wing length of 7.7 mm], narrow and long (about 2.5 times as long as wide); venation clear, Sc rather short about 0.68 times as long as the wing [Paratype Sc rather short, about 0.65 times as long as the wing] and ending at anterior margin proximad of R2; crossvein sc-r locating at 2/3 of Rs; Rs arising about one-fifth from the base of the wing; R2+3 about 0.8 times as long as R2+3+4; R2 about one-tenth of length of R3; R3 almost three times as long as the R2+3; dM1+2 0.6 times as long as mM1+2, while M1 2.5 times of the dM1+2; a well developed m-m crossvein about three-fourth length of bM3, closing the d-cell and nearly 0.3 length of d-cell; bM1+2 nearly 1.0 times as long as the length of the r-m and the latter at one-fifth of the d-cell; d-cell narrow and long (about 2.5 times as long as wide) and almost one-fifth length of wing; both crossveins m-m and m-cu intersecting with M4 at the same point; Cu long, curved (angle about 135°) and reaching the wing posterior margin at 0.6 from the base of the wing; the stem of A divided into A1 and A2; A1 long, slightly curving and reaching the wing posterior margin; A2 short, 0.15 (right wing) [Paratype 0.14] times as long as wing and almost 0.3 times as long as length of A1, bending sharply (angle about 128°) and reaching the wing posterior margin.
</paragraph>
<paragraph pageId="5" pageNumber="150">Legs: Legs slender and long; the hind leg nearly 1.2 times as long as the abdomen and 1.3 times as long as the wing. Tarsus with five segments; the first segment of tarsus (t1) is 1.2 times as long as t2 in mid leg.</paragraph>
<paragraph pageId="5" pageNumber="150">Abdomen: Abdomen relatively long and thin, with ten segments. Female genital discernible.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="150" type="remarks">
<paragraph pageId="5" pageNumber="150">Remarks.</paragraph>
<paragraph pageId="5" pageNumber="150">
<taxonomicName class="Insecta" family="Trichoceridae" genus="Eotrichocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrichocera (Archaeotrichocera) longensis" order="Diptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="longensis" subGenus="Archaeotrichocera">Eotrichocera (Archaeotrichocera) longensis</taxonomicName>
sp. n. is assigned to 
<taxonomicName lsidName="" pageId="5" pageNumber="150" rank="subFamily" subFamily="Thrichocerinae">Thrichocerinae</taxonomicName>
based on the following characters: d-cell medium, m-cu present; A2 short, antennae long, flagellomeres thin, much longer than two times of the head length. It belongs to 
<taxonomicName class="Insecta" family="Trichoceridae" genus="Eotrichocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrichocera (Archaeotrichocera)" order="Diptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="subGenus" subGenus="Archaeotrichocera">Eotrichocera (Archaeotrichocera)</taxonomicName>
because of wing length from 7.7 to 9.0 mm and d-cell almost one-fifth of wing length. In addition, it differs from all other known 
<taxonomicName lsidName="" pageId="5" pageNumber="150" rank="subFamily" subFamily="Thrichocerinae">Thrichocerinae</taxonomicName>
by its A2 rather short and bending sharply toward anterior margin (angle about 128°), R2 relatively long, Sc forking proximad of 2/3 (at about 0.55) times of wing length, and d-cell narrow and long. To compare the key characters among the new species and other species of 
<taxonomicName class="Insecta" family="Trichoceridae" genus="Eotrichocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrichocera (Archaeotrichocera)" order="Diptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="subGenus" subGenus="Archaeotrichocera">Eotrichocera (Archaeotrichocera)</taxonomicName>
, we set up the Table 2.
</paragraph>
<caption pageId="5" pageNumber="150">
<paragraph pageId="5" pageNumber="150">
Table 2. Comparison of key characters among the two new species and other species of 
<taxonomicName class="Insecta" family="Trichoceridae" genus="Eotrichocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrichocera (Archaeotrichocera)" order="Diptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="subGenus" subGenus="Archaeotrichocera">Eotrichocera (Archaeotrichocera)</taxonomicName>
<normalizedToken originalValue="Krzemińska">Krzeminska</normalizedToken>
, 
<normalizedToken originalValue="Krzemiński">Krzeminski</normalizedToken>
&amp; Dahl, 2009. L/W = ratio of length/width; W/L = ratio of width/length.
</paragraph>
</caption>
<paragraph pageId="5" pageNumber="150">
<table pageId="5" pageNumber="150">
<tr pageId="5" pageNumber="150">
<th colspan="1" pageId="5" pageNumber="150" rowspan="1" style="text-align:left">Key characters</th>
<th colspan="1" pageId="5" pageNumber="150" rowspan="1" style="text-align:left">
<taxonomicName class="Insecta" family="Trichoceridae" genus="Eotrichocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrichocera (Archaeotrichocera) ephemera" order="Diptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="ephemera" subGenus="Archaeotrichocera">Eotrichocera (Archaeotrichocera) ephemera</taxonomicName>
Zhang, 2006
</th>
<th colspan="1" pageId="5" pageNumber="150" rowspan="1" style="text-align:left">
<taxonomicName class="Insecta" family="Trichoceridae" genus="Eotrichocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrichocera (Archaeotrichocera) conica" order="Diptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="conica" subGenus="Archaeotrichocera">Eotrichocera (Archaeotrichocera) conica</taxonomicName>
<normalizedToken originalValue="Krzemińska">Krzeminska</normalizedToken>
, 
<normalizedToken originalValue="Krzemiński">Krzeminski</normalizedToken>
&amp; Ren, 2009
</th>
<th colspan="1" pageId="5" pageNumber="150" rowspan="1" style="text-align:left">
<taxonomicName class="Insecta" family="Trichoceridae" genus="Eotrichocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrichocera (Archaeotrichocera) rara" order="Diptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="rara" subGenus="Archaeotrichocera">Eotrichocera (Archaeotrichocera) rara</taxonomicName>
<normalizedToken originalValue="Krzemińska">Krzeminska</normalizedToken>
, 
<normalizedToken originalValue="Krzemiński">Krzeminski</normalizedToken>
&amp; Ren, 2009
</th>
<th colspan="1" pageId="5" pageNumber="150" rowspan="1" style="text-align:left">
<taxonomicName class="Insecta" family="Trichoceridae" genus="Eotrichocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrichocera (Archaeotrichocera) spatiosa" order="Diptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="spatiosa" subGenus="Archaeotrichocera">Eotrichocera (Archaeotrichocera) spatiosa</taxonomicName>
Liu, Shih &amp; Ren, 2012
</th>
<th colspan="1" pageId="5" pageNumber="150" rowspan="1" style="text-align:left">
<taxonomicName class="Insecta" family="Trichoceridae" genus="Eotrichocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrichocera (Archaeotrichocera) longensis" order="Diptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="longensis" subGenus="Archaeotrichocera">Eotrichocera (Archaeotrichocera) longensis</taxonomicName>
sp. n.
</th>
<th colspan="1" pageId="5" pageNumber="150" rowspan="1" style="text-align:left">
<taxonomicName class="Insecta" family="Trichoceridae" genus="Eotrichocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrichocera (Archaeotrichocera) amabilis" order="Diptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="amabilis" subGenus="Archaeotrichocera">Eotrichocera (Archaeotrichocera) amabilis</taxonomicName>
sp. n.
</th>
</tr>
<tr pageId="5" pageNumber="150">
<td colspan="1" pageId="5" pageNumber="150" rowspan="1">2</td>
<td colspan="1" pageId="5" pageNumber="150" rowspan="1">2</td>
<td colspan="1" pageId="5" pageNumber="150" rowspan="1">2</td>
<td colspan="1" pageId="5" pageNumber="150" rowspan="1">2</td>
<td colspan="1" pageId="5" pageNumber="150" rowspan="1">2</td>
<td colspan="1" pageId="5" pageNumber="150" rowspan="1">2</td>
</tr>
<tr pageId="5" pageNumber="150">
<td colspan="1" pageId="5" pageNumber="150" rowspan="1">2</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>