treatments-xml/data/DD/05/DA/DD05DA243464E963A895654D5885F9D7.xml

<document id="9F0449C2305C3EEF7378182266324405" ID-DOI="10.1649/0010-065X-69.4.531" ID-Zenodo-Dep="5334774" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630175991725" checkinUser="diego" docAuthor="Barr, Cheryl B., Gibson, J. Randy &amp; Diaz, Peter H." docDate="2015" docId="DD05DA243464E963A895654D5885F9D7" docLanguage="en" docName="0010-065X-69.4.531.pdf" docOrigin="The Coleopterists Bulletin 69 (4)" docSource="http://dx.doi.org/10.1649/0010-065x-69.4.531" docStyle="DocumentStyle:A0A70CFE30726088A421CCB226FF86C7.4:TheColeopBull.2010-2016.journal_article.1cover" docStyleId="A0A70CFE30726088A421CCB226FF86C7" docStyleName="TheColeopBull.2010-2016.journal_article.1cover" docStyleVersion="4" docTitle="Typhloelmis finegan Barr &amp; Gibson &amp; Diaz 2015, new species" docType="treatment" docVersion="3" lastPageNumber="558" masterDocId="213CA25C3476E970A852670D5926FFEB" masterDocTitle="TyphloelmisBarr (Coleoptera: Elmidae: Elminae), a New Stygobiontic Riffle Beetle Genus with Three New Species from Texas, USA" masterLastPageNumber="558" masterPageNumber="531" pageNumber="531" updateTime="1699175497231" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="B29771E026473589BD53253EABB64282">TyphloelmisBarr (Coleoptera: Elmidae: Elminae), a New Stygobiontic Riffle Beetle Genus with Three New Species from Texas, USA</mods:title>
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<mods:namePart id="6E6C542C95E677D6C3DCBA25AB4D9845">Barr, Cheryl B.</mods:namePart>
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<mods:namePart id="207EA58F9D42078A7D94953A35945EA9">Gibson, J. Randy</mods:namePart>
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<mods:namePart id="93762EDE8BE519E353A06513B93E259B">Diaz, Peter H.</mods:namePart>
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<mods:title id="E6C3BD9296C1A6893C453DCD76C81136">The Coleopterists Bulletin</mods:title>
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<mods:date id="761D7AD7F0FB6A876B2E4F52C6A8D2D7">2015</mods:date>
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<subSubSection id="1DB638B93464E962A895654D58DCFD9B" pageId="18" type="nomenclature">
<paragraph id="55136B323464E962A895654D58DCFD9B" blockId="18.[199,589,576,624]" pageId="18">
<emphasis id="67D8B7203464E962A895654D5B6BFDBD" bold="true" box="[199,589,576,598]" pageId="18">
<taxonomicName id="92AC10B13464E962A895654D58EDFDBD" ID-CoL="59WR3" authority="Barr" authorityName="Barr &amp; Gibson &amp; Diaz" authorityYear="2015" box="[199,459,576,598]" class="Insecta" family="Elmidae" genus="Typhloelmis" kingdom="Animalia" order="Coleoptera" pageId="18" phylum="Arthropoda" rank="species" species="finegan" status="sp. nov.">
<emphasis id="67D8B7203464E962A895654D58B7FDBE" bold="true" box="[199,401,576,597]" italics="true" pageId="18">Typhloelmis finegan</emphasis>
Barr
</taxonomicName>
, 
<taxonomicNameLabel id="7CEB0A5B3464E962A98A654C5B6BFDBD" box="[472,589,577,598]" pageId="18" rank="species">new species</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="CD9777B73464E962A97065565841FD9B" box="[290,359,603,624]" captionStart="Figs" captionStartId="3.[197,237,1662,1681]" captionTargetBox="[250,1154,192,1639]" captionTargetId="figure-1@3.[187,1178,191,1640]" captionTargetPageId="3" captionText="Figs. 1–2. 1) Map of Terrell and Val Verde counties, Texas, showing the type localities of Typhoelmis caroline at Caroline Springs, Typhoelmis sanfelipe at San Felipe Springs, and Typhoelmis finegan at Finegan Springs; 2) Map of the Edwards and Trinity Aquifer regions of Texas. GWD = groundwater divide; black dots = type locality springs." figureDoi="http://doi.org/10.5281/zenodo.5334776" httpUri="https://zenodo.org/record/5334776/files/figure.png" pageId="18">Figs. 1</figureCitation>
, 
<figureCitation id="CD9777B73464E962A927655658A4FD9B" box="[373,386,603,624]" captionStart="Figs" captionStartId="7.[197,237,1661,1680]" captionTargetBox="[173,1193,194,1638]" captionTargetId="figure-1@7.[172,1194,193,1638]" captionTargetPageId="7" captionText="Figs. 6–8. Typhloelmis species, adult dry specimens, dorsal view (length = pronotum + elytra). 6) T. caroline, ♂, length 2.2 mm; 7) T. sanfelipe, ♀, length 2.5 mm; 8) T. finegan, ♀, length 4.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5334786" httpUri="https://zenodo.org/record/5334786/files/figure.png" pageId="18">8</figureCitation>
, 
<figureCitation id="CD9777B73464E962A9DD6556588FFD9B" box="[399,425,603,624]" captionStart="Figs" captionStartId="9.[197,237,1306,1325]" captionTargetBox="[173,1193,194,1283]" captionTargetId="figure-180@9.[172,1194,193,1283]" captionTargetPageId="9" captionText="Figs. 11–12. Typhloelmis species, adult wet specimens. 11) T. sanfelipe, lateral view; 12) T. finegan, a) Dorsal view, b) Ventral view." figureDoi="http://doi.org/10.5281/zenodo.5334790" httpUri="https://zenodo.org/record/5334790/files/figure.png" pageId="18">12</figureCitation>
, 
<figureCitation id="CD9777B73464E962A9E6655658E8FD9B" box="[436,462,603,624]" captionStart="Figs" captionStartId="10.[175,215,1239,1258]" captionTargetBox="[151,1171,194,1216]" captionTargetId="figure-212@10.[151,1173,193,1216]" captionTargetPageId="10" captionText="Figs. 13–15. Typhloelmis species, adult wet specimens, anterior dorsal view. 13) T. caroline; 14) T. sanfelipe;" figureDoi="http://doi.org/10.5281/zenodo.5334794" httpUri="https://zenodo.org/record/5334794/files/figure.png" pageId="18">15</figureCitation>
, 
<figureCitation id="CD9777B73464E962A988655658D4FD9B" box="[474,498,603,624]" captionStart="Figs" captionStartId="11.[197,237,1493,1512]" captionTargetBox="[188,1178,193,1470]" captionTargetId="figure-73@11.[187,1178,193,1470]" captionTargetPageId="11" captionText="Figs. 16–19. Typhloelmis species, adult wet specimens, ventral view. 16) T. caroline, anterior; 17) T. sanfelipe, anterior; 18) T. finegan, anterior; 19) T. sanfelipe, fused elytra." figureDoi="http://doi.org/10.5281/zenodo.5334796" httpUri="https://zenodo.org/record/5334796/files/figure.png" pageId="18">18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1DB638B93464E962A8FD658B5B58FCF4" pageId="18" type="materials_examined">
<paragraph id="55136B323464E962A8FD658B5B58FCF4" blockId="18.[151,638,645,1729]" pageId="18">
<emphasis id="67D8B7203464E962A8FD658B5855FD70" bold="true" box="[175,371,646,667]" pageId="18">Material Examined.</emphasis>
<materialsCitation id="E5C4616F3464E962A92D658B58D7FCED" collectionCode="EMEC, R" collectorName="P. H. Diaz" country="United States of America" county="Val Verde Co." elevation="405" location="Devils River" municipality="Devils River State Natural" pageId="18" specimenCount="1" specimenCount-female="1" stateProvince="Texas" typeStatus="holotype">
<emphasis id="67D8B7203464E962A92D658B5B25FD70" bold="true" box="[383,515,646,667]" pageId="18">
<typeStatus id="8A17D5903464E962A92D658B5B25FD70" box="[383,515,646,667]" pageId="18" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
<specimenCount id="43AAA0BB3464E962AA5A65885B3CFD70" box="[520,538,645,667]" pageId="18" type="female">♀</specimenCount>
in 
<collectionCode id="33BDF3F73464E962AA65658B5B5CFD70" box="[567,634,646,667]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="18" type="University or college">EMEC</collectionCode>
, labeled “ 
<collectingCountry id="2DBB2BA23464E962A8B965AD583BFD5E" box="[235,285,672,693]" name="United States of America" pageId="18">USA</collectingCountry>
: 
<collectingRegion id="9768A5D03464E962A97A65AD5846FD5E" box="[296,352,672,693]" country="United States of America" name="Texas" pageId="18">Texas</collectingRegion>
: 
<collectingCounty id="BC7213BE3464E962A93965AD58D7FD5E" box="[363,497,672,693]" pageId="18">Val Verde Co.</collectingCounty>
/ 
<collectingMunicipality id="B577F1483464E962AA5565AD5839FD3B" pageId="18">
<location id="50733DE93464E962AA5565AD5B58FD5E" LSID="urn:lsid:plazi:treatment:DD05DA243464E963A895654D5885F9D7:50733DE93464E962AA5565AD5B58FD5E" box="[519,638,672,693]" country="United States of America" county="Val Verde Co." municipality="Devils River State Natural" name="Devils River" pageId="18" stateProvince="Texas">
Devils 
<location id="50733DE93464E962AA1965AD5B58FD5E" LSID="urn:lsid:plazi:treatment:DD05DA243464E963A895654D5885F9D7:50733DE93464E962AA1965AD5B58FD5E" box="[587,638,672,693]" country="United States of America" county="Val Verde Co." municipality="Devils River State Natural" name="River State Natural" pageId="18" stateProvince="Texas">River</location>
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State Natural
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/ 
<location id="50733DE93464E962A96865B6584BFD3B" LSID="urn:lsid:plazi:treatment:DD05DA243464E963A895654D5885F9D7:50733DE93464E962A96865B6584BFD3B" box="[314,365,699,720]" country="United States of America" county="Val Verde Co." municipality="Devils River State Natural" name="Area" pageId="18" stateProvince="Texas">Area</location>
, 
<location id="50733DE93464E962A92E65B65B00FD3B" LSID="urn:lsid:plazi:treatment:DD05DA243464E963A895654D5885F9D7:50733DE93464E962A92E65B65B00FD3B" box="[380,550,699,720]" country="United States of America" county="Val Verde Co." municipality="Devils River State Natural" name="Finegan Springs" pageId="18" stateProvince="Texas">Finegan Springs</location>
, 
<quantity id="9254C6D73464E962AA6665B65B4BFD3B" box="[564,621,699,720]" metricMagnitude="2" metricUnit="m" metricValue="4.05" pageId="18" unit="m" value="405.0">
<elevation id="DE818C013464E962AA6665B65B4BFD3B" box="[564,621,699,720]" metricMagnitude="2" metricUnit="m" metricValue="4.05" pageId="18" unit="m" value="405.0">405m</elevation>
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/ 29.9011°,-100.9987°, 25-II- / 2010, J. 
<collectionCode id="33BDF3F73464E962AA5165D85B33FD01" box="[515,533,725,746]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="18">R</collectionCode>
. Gibson &amp; 
<collectorName id="F8590EE43464E962A8C565FD5825FCEE" box="[151,259,752,773]" pageId="18">P. H. Diaz</collectorName>
“ // [genitalia vial] // “
</materialsCitation>
<typeStatus id="8A17D5903464E962A9AE65FD5B58FCEE" box="[508,638,752,773]" pageId="18" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="92AC10B13464E962A8C5640658A1FCF4" authority="Barr" authorityName="Barr &amp; Gibson &amp; Diaz" authorityYear="2015" box="[151,391,778,800]" class="Insecta" family="Elmidae" genus="Typhloelmis" kingdom="Animalia" order="Coleoptera" pageId="18" phylum="Arthropoda" rank="species" species="finegan">
<emphasis id="67D8B7203464E962A8C564065871FCCB" box="[151,343,779,800]" italics="true" pageId="18">Typhloelmis finegan</emphasis>
Barr
</taxonomicName>
” [red label, handwritten].
</paragraph>
</subSubSection>
<subSubSection id="1DB638B93464E962A8FD64285ABAFA83" pageId="18" type="description">
<paragraph id="55136B323464E962A8FD64285D0BFAA6" blockId="18.[151,638,645,1729]" lastBlockId="18.[685,1173,194,1730]" pageId="18">
<emphasis id="67D8B7203464E962A8FD6428580DFCD1" bold="true" box="[175,299,805,826]" pageId="18">Description.</emphasis>
<typeStatus id="8A17D5903464E962A96B642858B7FCD1" box="[313,401,805,826]" pageId="18" type="holotype">Holotype</typeStatus>
female. Length 4.1 mm (elytra + pronotum), width 1.2 mm. Cuticle redbrown, barely translucent, with extensive whitish dorsal and ventral plastron; surface smooth, lacking dorsal carinae, striae, and punctures (
<figureCitation id="CD9777B73464E962AA6064825B5FFC4F" box="[562,633,911,932]" captionStart="Figs" captionStartId="7.[197,237,1661,1680]" captionTargetBox="[173,1193,194,1638]" captionTargetId="figure-1@7.[172,1194,193,1638]" captionTargetPageId="7" captionText="Figs. 6–8. Typhloelmis species, adult dry specimens, dorsal view (length = pronotum + elytra). 6) T. caroline, ♂, length 2.2 mm; 7) T. sanfelipe, ♀, length 2.5 mm; 8) T. finegan, ♀, length 4.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5334786" httpUri="https://zenodo.org/record/5334786/files/figure.png" pageId="18">Figs. 8</figureCitation>
, 
<figureCitation id="CD9777B73464E962A8C564A75997FC54" box="[151,177,938,959]" captionStart="Figs" captionStartId="9.[197,237,1306,1325]" captionTargetBox="[173,1193,194,1283]" captionTargetId="figure-180@9.[172,1194,193,1283]" captionTargetPageId="9" captionText="Figs. 11–12. Typhloelmis species, adult wet specimens. 11) T. sanfelipe, lateral view; 12) T. finegan, a) Dorsal view, b) Ventral view." figureDoi="http://doi.org/10.5281/zenodo.5334790" httpUri="https://zenodo.org/record/5334790/files/figure.png" pageId="18">12</figureCitation>
). 
<emphasis id="67D8B7203464E962A89664A75805FC54" bold="true" box="[196,291,938,959]" pageId="18">Antenna:</emphasis>
Color medium red-brown, antennomeres 1–10 narrowly cone-shaped, wider distally than basally; antennomere 11 narrowly fusiform and acutely pointed (
<figureCitation id="CD9777B73464E962A93964F75894FBE4" box="[363,434,1018,1039]" captionStart="Figs" captionStartId="7.[197,237,1661,1680]" captionTargetBox="[173,1193,194,1638]" captionTargetId="figure-1@7.[172,1194,193,1638]" captionTargetPageId="7" captionText="Figs. 6–8. Typhloelmis species, adult dry specimens, dorsal view (length = pronotum + elytra). 6) T. caroline, ♂, length 2.2 mm; 7) T. sanfelipe, ♀, length 2.5 mm; 8) T. finegan, ♀, length 4.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5334786" httpUri="https://zenodo.org/record/5334786/files/figure.png" pageId="18">Figs. 8</figureCitation>
, 
<figureCitation id="CD9777B73464E962A99364F758FDFBE4" box="[449,475,1018,1039]" captionStart="Figs" captionStartId="9.[197,237,1306,1325]" captionTargetBox="[173,1193,194,1283]" captionTargetId="figure-180@9.[172,1194,193,1283]" captionTargetPageId="9" captionText="Figs. 11–12. Typhloelmis species, adult wet specimens. 11) T. sanfelipe, lateral view; 12) T. finegan, a) Dorsal view, b) Ventral view." figureDoi="http://doi.org/10.5281/zenodo.5334790" httpUri="https://zenodo.org/record/5334790/files/figure.png" pageId="18">12</figureCitation>
, 
<figureCitation id="CD9777B73464E962A9BA64F75B24FBE4" box="[488,514,1018,1039]" captionStart="Figs" captionStartId="10.[175,215,1239,1258]" captionTargetBox="[151,1171,194,1216]" captionTargetId="figure-212@10.[151,1173,193,1216]" captionTargetPageId="10" captionText="Figs. 13–15. Typhloelmis species, adult wet specimens, anterior dorsal view. 13) T. caroline; 14) T. sanfelipe;" figureDoi="http://doi.org/10.5281/zenodo.5334794" httpUri="https://zenodo.org/record/5334794/files/figure.png" pageId="18">15</figureCitation>
, 
<figureCitation id="CD9777B73464E962AA4264F75B0DFBE4" box="[528,555,1018,1039]" captionStart="Figs" captionStartId="11.[197,237,1493,1512]" captionTargetBox="[188,1178,193,1470]" captionTargetId="figure-73@11.[187,1178,193,1470]" captionTargetPageId="11" captionText="Figs. 16–19. Typhloelmis species, adult wet specimens, ventral view. 16) T. caroline, anterior; 17) T. sanfelipe, anterior; 18) T. finegan, anterior; 19) T. sanfelipe, fused elytra." figureDoi="http://doi.org/10.5281/zenodo.5334796" httpUri="https://zenodo.org/record/5334796/files/figure.png" pageId="18">18</figureCitation>
). 
<emphasis id="67D8B7203464E962AA6D64F75B5BFBE4" bold="true" box="[575,637,1018,1039]" pageId="18">Head:</emphasis>
Color medium red-brown, darker at clypeal margin; surface bearing evenly spaced, small granules and short setae. Eyes absent (
<figureCitation id="CD9777B73464E962A9D5634458F1FBB5" box="[391,471,1097,1118]" captionStart="Figs" captionStartId="9.[197,237,1306,1325]" captionTargetBox="[173,1193,194,1283]" captionTargetId="figure-180@9.[172,1194,193,1283]" captionTargetPageId="9" captionText="Figs. 11–12. Typhloelmis species, adult wet specimens. 11) T. sanfelipe, lateral view; 12) T. finegan, a) Dorsal view, b) Ventral view." figureDoi="http://doi.org/10.5281/zenodo.5334790" httpUri="https://zenodo.org/record/5334790/files/figure.png" pageId="18">Figs. 12</figureCitation>
, 
<figureCitation id="CD9777B73464E962A9B0634458DDFBB5" box="[482,507,1097,1118]" captionStart="Figs" captionStartId="10.[175,215,1239,1258]" captionTargetBox="[151,1171,194,1216]" captionTargetId="figure-212@10.[151,1173,193,1216]" captionTargetPageId="10" captionText="Figs. 13–15. Typhloelmis species, adult wet specimens, anterior dorsal view. 13) T. caroline; 14) T. sanfelipe;" figureDoi="http://doi.org/10.5281/zenodo.5334794" httpUri="https://zenodo.org/record/5334794/files/figure.png" pageId="18">15</figureCitation>
, 
<figureCitation id="CD9777B73464E962AA5563445B07FBB5" box="[519,545,1097,1118]" captionStart="Figs" captionStartId="11.[197,237,1493,1512]" captionTargetBox="[188,1178,193,1470]" captionTargetId="figure-73@11.[187,1178,193,1470]" captionTargetPageId="11" captionText="Figs. 16–19. Typhloelmis species, adult wet specimens, ventral view. 16) T. caroline, anterior; 17) T. sanfelipe, anterior; 18) T. finegan, anterior; 19) T. sanfelipe, fused elytra." figureDoi="http://doi.org/10.5281/zenodo.5334796" httpUri="https://zenodo.org/record/5334796/files/figure.png" pageId="18">18</figureCitation>
). Labrum with long, pale setae on distal 1/2. Mandible dark brown with 2 short teeth; upper tooth longest, narrow with blunt tip; lower tooth much shorter and broader. 
<emphasis id="67D8B7203464E962A94D63B958B6FB22" bold="true" box="[287,400,1204,1225]" pageId="18">Pronotum:</emphasis>
Color dark red-brown, covered with whitish plastron except at anterior margin and antero-medially on disc where abraded (
<figureCitation id="CD9777B73464E962A8CC620E59FCFAF3" box="[158,218,1283,1304]" captionStart="Figs" captionStartId="7.[197,237,1661,1680]" captionTargetBox="[173,1193,194,1638]" captionTargetId="figure-1@7.[172,1194,193,1638]" captionTargetPageId="7" captionText="Figs. 6–8. Typhloelmis species, adult dry specimens, dorsal view (length = pronotum + elytra). 6) T. caroline, ♂, length 2.2 mm; 7) T. sanfelipe, ♀, length 2.5 mm; 8) T. finegan, ♀, length 4.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5334786" httpUri="https://zenodo.org/record/5334786/files/figure.png" pageId="18">Fig. 8</figureCitation>
). Longer than wide; length 1.4 mm, width 1.1 mm at widest point just anterior to base (
<figureCitation id="CD9777B73464E962A8CC623559C6FAA6" box="[158,224,1336,1357]" captionStart="Figs" captionStartId="7.[197,237,1661,1680]" captionTargetBox="[173,1193,194,1638]" captionTargetId="figure-1@7.[172,1194,193,1638]" captionTargetPageId="7" captionText="Figs. 6–8. Typhloelmis species, adult dry specimens, dorsal view (length = pronotum + elytra). 6) T. caroline, ♂, length 2.2 mm; 7) T. sanfelipe, ♀, length 2.5 mm; 8) T. finegan, ♀, length 4.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5334786" httpUri="https://zenodo.org/record/5334786/files/figure.png" pageId="18">Figs. 8</figureCitation>
, 
<figureCitation id="CD9777B73464E962A8B862355825FAA6" box="[234,259,1336,1357]" captionStart="Figs" captionStartId="9.[197,237,1306,1325]" captionTargetBox="[173,1193,194,1283]" captionTargetId="figure-180@9.[172,1194,193,1283]" captionTargetPageId="9" captionText="Figs. 11–12. Typhloelmis species, adult wet specimens. 11) T. sanfelipe, lateral view; 12) T. finegan, a) Dorsal view, b) Ventral view." figureDoi="http://doi.org/10.5281/zenodo.5334790" httpUri="https://zenodo.org/record/5334790/files/figure.png" pageId="18">12</figureCitation>
, 
<figureCitation id="CD9777B73464E962A95C6235580EFAA6" box="[270,296,1336,1357]" captionStart="Figs" captionStartId="10.[175,215,1239,1258]" captionTargetBox="[151,1171,194,1216]" captionTargetId="figure-212@10.[151,1173,193,1216]" captionTargetPageId="10" captionText="Figs. 13–15. Typhloelmis species, adult wet specimens, anterior dorsal view. 13) T. caroline; 14) T. sanfelipe;" figureDoi="http://doi.org/10.5281/zenodo.5334794" httpUri="https://zenodo.org/record/5334794/files/figure.png" pageId="18">15</figureCitation>
). Lateral margins distinctly reflexed; crenulate with small, rounded, evenly spaced teeth; basolateral angles blunt and rounded. Disc with short, sparse setae; small, protuberant granules emergent from plastron adjacent to apicolateral and lateral margins. 
<emphasis id="67D8B7203464E962A92662B058E8FA39" bold="true" box="[372,462,1469,1490]" pageId="18">Elytron:</emphasis>
Color dark redbrown, covered with whitish plastron except for at basal margin and suture, and where scratched and abraded medially (
<figureCitation id="CD9777B73464E962A91C610058ACF9C9" box="[334,394,1549,1570]" captionStart="Figs" captionStartId="7.[197,237,1661,1680]" captionTargetBox="[173,1193,194,1638]" captionTargetId="figure-1@7.[172,1194,193,1638]" captionTargetPageId="7" captionText="Figs. 6–8. Typhloelmis species, adult dry specimens, dorsal view (length = pronotum + elytra). 6) T. caroline, ♂, length 2.2 mm; 7) T. sanfelipe, ♀, length 2.5 mm; 8) T. finegan, ♀, length 4.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5334786" httpUri="https://zenodo.org/record/5334786/files/figure.png" pageId="18">Fig. 8</figureCitation>
). Length 2.7 mm, width 0.6 mm at widest point. Disc with short, sparse setae; widely spaced, small, round granules emergent from plastron, more protuberant laterally; loosely arranged into longitudinal rows more closely spaced laterally. Outer 1/3 of disc in posterior 1/2 wrinkled, with irregular grooves perpendicular to lateral margin. Lateral margin serrate with evenly spaced teeth (
<figureCitation id="CD9777B73464E962AC4367D05D7AFF19" box="[1041,1116,221,242]" captionStart="Figs" captionStartId="7.[197,237,1661,1680]" captionTargetBox="[173,1193,194,1638]" captionTargetId="figure-1@7.[172,1194,193,1638]" captionTargetPageId="7" captionText="Figs. 6–8. Typhloelmis species, adult dry specimens, dorsal view (length = pronotum + elytra). 6) T. caroline, ♂, length 2.2 mm; 7) T. sanfelipe, ♀, length 2.5 mm; 8) T. finegan, ♀, length 4.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5334786" httpUri="https://zenodo.org/record/5334786/files/figure.png" pageId="18">Figs. 8</figureCitation>
, 
<figureCitation id="CD9777B73464E962AC3E67D05DAEFF19" box="[1132,1160,221,242]" captionStart="Figs" captionStartId="9.[197,237,1306,1325]" captionTargetBox="[173,1193,194,1283]" captionTargetId="figure-180@9.[172,1194,193,1283]" captionTargetPageId="9" captionText="Figs. 11–12. Typhloelmis species, adult wet specimens. 11) T. sanfelipe, lateral view; 12) T. finegan, a) Dorsal view, b) Ventral view." figureDoi="http://doi.org/10.5281/zenodo.5334790" httpUri="https://zenodo.org/record/5334790/files/figure.png" pageId="18">12</figureCitation>
). 
<emphasis id="67D8B7203464E962AAFF67FA5BFBFEE7" bold="true" box="[685,733,247,268]" pageId="18">Leg:</emphasis>
Color of coxa, trochanter, femur, and tibia dark red-brown; tarsus medium red-brown. Pro- and mesothoracic legs without plastron, shiny; metathoracic leg with plastron present on coxa, trochanter, and femur, absent on tibia and tarsus. Femur with low granules and fine setae dorsally, flattened and granulate ventrally; pro- and mesofemora with numerous, spinose granules at margins of ventral flattened area; metafemur much less granulate and spinose. Tibia with large granules bearing prominent spines on all surfaces, spines largest, most numerous, at apex (
<figureCitation id="CD9777B73464E962AB2065165A9AFDDB" box="[882,956,539,560]" captionStart="Figs" captionStartId="9.[197,237,1306,1325]" captionTargetBox="[173,1193,194,1283]" captionTargetId="figure-180@9.[172,1194,193,1283]" captionTargetPageId="9" captionText="Figs. 11–12. Typhloelmis species, adult wet specimens. 11) T. sanfelipe, lateral view; 12) T. finegan, a) Dorsal view, b) Ventral view." figureDoi="http://doi.org/10.5281/zenodo.5334790" httpUri="https://zenodo.org/record/5334790/files/figure.png" pageId="18">Fig. 12</figureCitation>
); protibia with spines most well-developed (
<figureCitation id="CD9777B73464E962ABDF653B5AC7FDA0" box="[909,993,566,587]" captionStart="Figs" captionStartId="9.[197,237,1306,1325]" captionTargetBox="[173,1193,194,1283]" captionTargetId="figure-180@9.[172,1194,193,1283]" captionTargetPageId="9" captionText="Figs. 11–12. Typhloelmis species, adult wet specimens. 11) T. sanfelipe, lateral view; 12) T. finegan, a) Dorsal view, b) Ventral view." figureDoi="http://doi.org/10.5281/zenodo.5334790" httpUri="https://zenodo.org/record/5334790/files/figure.png" pageId="18">Figs. 12</figureCitation>
, 
<figureCitation id="CD9777B73464E962ABBC653B5D2EFDA0" box="[1006,1032,566,587]" captionStart="Figs" captionStartId="10.[175,215,1239,1258]" captionTargetBox="[151,1171,194,1216]" captionTargetId="figure-212@10.[151,1173,193,1216]" captionTargetPageId="10" captionText="Figs. 13–15. Typhloelmis species, adult wet specimens, anterior dorsal view. 13) T. caroline; 14) T. sanfelipe;" figureDoi="http://doi.org/10.5281/zenodo.5334794" httpUri="https://zenodo.org/record/5334794/files/figure.png" pageId="18">15</figureCitation>
, 
<figureCitation id="CD9777B73464E962AC47653B5D16FDA0" box="[1045,1072,566,587]" captionStart="Figs" captionStartId="11.[197,237,1493,1512]" captionTargetBox="[188,1178,193,1470]" captionTargetId="figure-73@11.[187,1178,193,1470]" captionTargetPageId="11" captionText="Figs. 16–19. Typhloelmis species, adult wet specimens, ventral view. 16) T. caroline, anterior; 17) T. sanfelipe, anterior; 18) T. finegan, anterior; 19) T. sanfelipe, fused elytra." figureDoi="http://doi.org/10.5281/zenodo.5334796" httpUri="https://zenodo.org/record/5334796/files/figure.png" pageId="18">18</figureCitation>
). Tibia of all legs with fringe of long, pale setae (tomentum) on medial surface (
<figureCitation id="CD9777B73464E962AB2E65665AEFFD6B" box="[892,969,619,640]" captionStart="Figs" captionStartId="7.[197,237,1661,1680]" captionTargetBox="[173,1193,194,1638]" captionTargetId="figure-1@7.[172,1194,193,1638]" captionTargetPageId="7" captionText="Figs. 6–8. Typhloelmis species, adult dry specimens, dorsal view (length = pronotum + elytra). 6) T. caroline, ♂, length 2.2 mm; 7) T. sanfelipe, ♀, length 2.5 mm; 8) T. finegan, ♀, length 4.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5334786" httpUri="https://zenodo.org/record/5334786/files/figure.png" pageId="18">Figs. 8</figureCitation>
, 
<figureCitation id="CD9777B73464E962AB8A65665AD2FD6B" box="[984,1012,619,640]" captionStart="Figs" captionStartId="9.[197,237,1306,1325]" captionTargetBox="[173,1193,194,1283]" captionTargetId="figure-180@9.[172,1194,193,1283]" captionTargetPageId="9" captionText="Figs. 11–12. Typhloelmis species, adult wet specimens. 11) T. sanfelipe, lateral view; 12) T. finegan, a) Dorsal view, b) Ventral view." figureDoi="http://doi.org/10.5281/zenodo.5334790" httpUri="https://zenodo.org/record/5334790/files/figure.png" pageId="18">12</figureCitation>
); protibia with tomentum at middle 1/3, meso- and metatibiae with tomentum extending most of length except for at base and apex. Tarsomeres 1–4 with short, stout setae at ventral apex; tarsomere 5 with few setae along length of ventral surface. Claws moderately slender and barely curved. 
<emphasis id="67D8B7203464E962AC5B64075D72FCF4" bold="true" box="[1033,1108,778,799]" pageId="18">Venter:</emphasis>
Color light red-brown, covered with plastron; setae short and sparse. Prosternum (
<figureCitation id="CD9777B73464E962ABC7644D5AFCFCBE" box="[917,986,832,853]" captionStart="Figs" captionStartId="11.[197,237,1493,1512]" captionTargetBox="[188,1178,193,1470]" captionTargetId="figure-73@11.[187,1178,193,1470]" captionTargetPageId="11" captionText="Figs. 16–19. Typhloelmis species, adult wet specimens, ventral view. 16) T. caroline, anterior; 17) T. sanfelipe, anterior; 18) T. finegan, anterior; 19) T. sanfelipe, fused elytra." figureDoi="http://doi.org/10.5281/zenodo.5334796" httpUri="https://zenodo.org/record/5334796/files/figure.png" pageId="18">Fig. 18</figureCitation>
) covered with variably spaced granules; prosternal process narrowly triangular, apex broadly rounded, nearly truncate. Metaventrite (
<figureCitation id="CD9777B73464E962AB6C64825AABFC4F" box="[830,909,911,932]" captionStart="Figs" captionStartId="9.[197,237,1306,1325]" captionTargetBox="[173,1193,194,1283]" captionTargetId="figure-180@9.[172,1194,193,1283]" captionTargetPageId="9" captionText="Figs. 11–12. Typhloelmis species, adult wet specimens. 11) T. sanfelipe, lateral view; 12) T. finegan, a) Dorsal view, b) Ventral view." figureDoi="http://doi.org/10.5281/zenodo.5334790" httpUri="https://zenodo.org/record/5334790/files/figure.png" pageId="18">Fig.12b</figureCitation>
) covered with numerous, closely spaced granules; anterior margin truncate between mesocoxae; metakatepisternal suture slightly arcuate, discrimen obvious just anterior to intersection only. Abdominal ventrites (
<figureCitation id="CD9777B73464E962AC6A64F75DABFBE4" box="[1080,1165,1018,1039]" captionStart="Figs" captionStartId="9.[197,237,1306,1325]" captionTargetBox="[173,1193,194,1283]" captionTargetId="figure-180@9.[172,1194,193,1283]" captionTargetPageId="9" captionText="Figs. 11–12. Typhloelmis species, adult wet specimens. 11) T. sanfelipe, lateral view; 12) T. finegan, a) Dorsal view, b) Ventral view." figureDoi="http://doi.org/10.5281/zenodo.5334790" httpUri="https://zenodo.org/record/5334790/files/figure.png" pageId="18">Fig. 12b</figureCitation>
) with widely spaced, low granules and setae; ventrite 1 with granules present on entire surface; ventrites 2–4 with granules present mainly laterad; ventrite 5 with apical 1/2 granulate and setose, with shallow median depression near apex, apex bearing fringe of fine, moderately long setae; intercoxal process of ventrite 1 triangularly shaped, apex acute. 
<emphasis id="67D8B7203464E962AC1863B95A36FB08" bold="true" pageId="18">Female genitalia:</emphasis>
Each longitudinal bacula subequal in length to gonocoxite; stylus short, narrowly ovoid. Median sclerite present between bases of gonocoxites, both dorsally and ventrally. Vagina extruding between the gonocoxites well beyond tips of styli.
</paragraph>
<paragraph id="55136B323464E962AA97625E5ABAFA83" blockId="18.[685,1173,194,1730]" box="[709,924,1363,1384]" pageId="18">
<emphasis id="67D8B7203464E962AA97625E5A0CFA83" bold="true" box="[709,810,1363,1384]" pageId="18">Variation.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="1DB638B93464E963AA976263583FFE47" lastPageId="19" pageId="18" type="diagnosis">
<paragraph id="55136B323464E963AA976263583FFE47" blockId="18.[685,1173,194,1730]" lastBlockId="19.[173,660,194,1596]" lastPageId="19" pageId="18">
<emphasis id="67D8B7203464E962AA9762635A14FA68" bold="true" box="[709,818,1390,1411]" pageId="18">Diagnosis.</emphasis>
<taxonomicName id="92AC10B13464E962AB1262635D3DFA68" authorityName="Barr &amp; Gibson &amp; Diaz" authorityYear="2015" box="[832,1051,1390,1411]" class="Insecta" family="Elmidae" genus="Typhloelmis" kingdom="Animalia" order="Coleoptera" pageId="18" phylum="Arthropoda" rank="species" species="finegan">
<emphasis id="67D8B7203464E962AB1262635D3DFA68" box="[832,1051,1390,1411]" italics="true" pageId="18">Typhloelmis finegan</emphasis>
</taxonomicName>
(
<figureCitation id="CD9777B73464E962AC7F62635D56FA68" box="[1069,1136,1390,1411]" captionStart="Figs" captionStartId="7.[197,237,1661,1680]" captionTargetBox="[173,1193,194,1638]" captionTargetId="figure-1@7.[172,1194,193,1638]" captionTargetPageId="7" captionText="Figs. 6–8. Typhloelmis species, adult dry specimens, dorsal view (length = pronotum + elytra). 6) T. caroline, ♂, length 2.2 mm; 7) T. sanfelipe, ♀, length 2.5 mm; 8) T. finegan, ♀, length 4.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5334786" httpUri="https://zenodo.org/record/5334786/files/figure.png" pageId="18">Fig. 8</figureCitation>
) is unlikely to be confused with either 
<taxonomicName id="92AC10B13464E962AC7662855DB2FA76" authorityName="Barr &amp; Gibson &amp; Diaz" authorityYear="2015" box="[1060,1172,1416,1437]" class="Insecta" family="Elmidae" genus="Typhloelmis" kingdom="Animalia" order="Coleoptera" pageId="18" phylum="Arthropoda" rank="species" species="caroline">
<emphasis id="67D8B7203464E962AC7662855DB2FA76" box="[1060,1172,1416,1437]" italics="true" pageId="18">T. caroline</emphasis>
</taxonomicName>
(
<figureCitation id="CD9777B73464E962AAE662AE5BC8FA53" box="[692,750,1443,1464]" captionStart="Figs" captionStartId="7.[197,237,1661,1680]" captionTargetBox="[173,1193,194,1638]" captionTargetId="figure-1@7.[172,1194,193,1638]" captionTargetPageId="7" captionText="Figs. 6–8. Typhloelmis species, adult dry specimens, dorsal view (length = pronotum + elytra). 6) T. caroline, ♂, length 2.2 mm; 7) T. sanfelipe, ♀, length 2.5 mm; 8) T. finegan, ♀, length 4.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5334786" httpUri="https://zenodo.org/record/5334786/files/figure.png" pageId="18">Fig. 6</figureCitation>
) or 
<taxonomicName id="92AC10B13464E962AB4862AE5AAEFA53" authorityName="Barr &amp; Gibson &amp; Diaz" authorityYear="2015" box="[794,904,1443,1464]" class="Insecta" family="Elmidae" genus="Typhloelmis" kingdom="Animalia" order="Coleoptera" pageId="18" phylum="Arthropoda" rank="species" species="sanfelipe">
<emphasis id="67D8B7203464E962AB4862AE5AAEFA53" box="[794,904,1443,1464]" italics="true" pageId="18">T. sanfelipe</emphasis>
</taxonomicName>
(
<figureCitation id="CD9777B73464E962ABC562AE5AF7FA53" box="[919,977,1443,1464]" captionStart="Figs" captionStartId="7.[197,237,1661,1680]" captionTargetBox="[173,1193,194,1638]" captionTargetId="figure-1@7.[172,1194,193,1638]" captionTargetPageId="7" captionText="Figs. 6–8. Typhloelmis species, adult dry specimens, dorsal view (length = pronotum + elytra). 6) T. caroline, ♂, length 2.2 mm; 7) T. sanfelipe, ♀, length 2.5 mm; 8) T. finegan, ♀, length 4.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5334786" httpUri="https://zenodo.org/record/5334786/files/figure.png" pageId="18">Fig. 7</figureCitation>
) because it is much larger (length&gt; 
<quantity id="9254C6D73464E962AB0762B05AB5FA39" box="[853,915,1469,1490]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="18" unit="mm" value="4.0">4 mm</quantity>
vs. &lt;
<quantity id="9254C6D73464E962AB8662B05D34FA39" box="[980,1042,1469,1490]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="18" unit="mm" value="3.0">3 mm</quantity>
) and darker colored. In addition, in 
<taxonomicName id="92AC10B13464E962ABF662D55D2AFA06" authorityName="Barr &amp; Gibson &amp; Diaz" authorityYear="2015" box="[932,1036,1496,1517]" class="Insecta" family="Elmidae" genus="Typhloelmis" kingdom="Animalia" order="Coleoptera" pageId="18" phylum="Arthropoda" rank="species" species="finegan">
<emphasis id="67D8B7203464E962ABF662D55D2AFA06" box="[932,1036,1496,1517]" italics="true" pageId="18">T. finegan</emphasis>
</taxonomicName>
(
<figureCitation id="CD9777B73464E962AC4F62D55D41FA06" box="[1053,1127,1496,1517]" captionStart="Figs" captionStartId="7.[197,237,1661,1680]" captionTargetBox="[173,1193,194,1638]" captionTargetId="figure-1@7.[172,1194,193,1638]" captionTargetPageId="7" captionText="Figs. 6–8. Typhloelmis species, adult dry specimens, dorsal view (length = pronotum + elytra). 6) T. caroline, ♂, length 2.2 mm; 7) T. sanfelipe, ♀, length 2.5 mm; 8) T. finegan, ♀, length 4.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5334786" httpUri="https://zenodo.org/record/5334786/files/figure.png" pageId="18">Figs. 8</figureCitation>
, 
<figureCitation id="CD9777B73464E962AC2762D55DB6FA06" box="[1141,1168,1496,1517]" captionStart="Figs" captionStartId="9.[197,237,1306,1325]" captionTargetBox="[173,1193,194,1283]" captionTargetId="figure-180@9.[172,1194,193,1283]" captionTargetPageId="9" captionText="Figs. 11–12. Typhloelmis species, adult wet specimens. 11) T. sanfelipe, lateral view; 12) T. finegan, a) Dorsal view, b) Ventral view." figureDoi="http://doi.org/10.5281/zenodo.5334790" httpUri="https://zenodo.org/record/5334790/files/figure.png" pageId="18">12</figureCitation>
, 
<figureCitation id="CD9777B73464E962AAFF62FF5BE1F9EC" box="[685,711,1522,1543]" captionStart="Figs" captionStartId="10.[175,215,1239,1258]" captionTargetBox="[151,1171,194,1216]" captionTargetId="figure-212@10.[151,1173,193,1216]" captionTargetPageId="10" captionText="Figs. 13–15. Typhloelmis species, adult wet specimens, anterior dorsal view. 13) T. caroline; 14) T. sanfelipe;" figureDoi="http://doi.org/10.5281/zenodo.5334794" httpUri="https://zenodo.org/record/5334794/files/figure.png" pageId="18">15</figureCitation>
, 
<figureCitation id="CD9777B73464E962AA8662FF5BCBF9EC" box="[724,749,1522,1543]" captionStart="Figs" captionStartId="11.[197,237,1493,1512]" captionTargetBox="[188,1178,193,1470]" captionTargetId="figure-73@11.[187,1178,193,1470]" captionTargetPageId="11" captionText="Figs. 16–19. Typhloelmis species, adult wet specimens, ventral view. 16) T. caroline, anterior; 17) T. sanfelipe, anterior; 18) T. finegan, anterior; 19) T. sanfelipe, fused elytra." figureDoi="http://doi.org/10.5281/zenodo.5334796" httpUri="https://zenodo.org/record/5334796/files/figure.png" pageId="18">18</figureCitation>
) the prosternal process is broader, triangular, and not medially constricted; the pronotum is margined with rounded, evenly spaced teeth; antennomeres 1–10 are similarly cone-shaped; the metaventrite is very granulate; the femora are ventrally flattened; the tibia of all legs possesses fringes of tomentum; and protarsomere 5 is not bulbous. In both 
<taxonomicName id="92AC10B13464E962AB0F61A05AECF929" authorityName="Barr &amp; Gibson &amp; Diaz" authorityYear="2015" box="[861,970,1709,1730]" class="Insecta" family="Elmidae" genus="Typhloelmis" kingdom="Animalia" order="Coleoptera" pageId="18" phylum="Arthropoda" rank="species" species="caroline">
<emphasis id="67D8B7203464E962AB0F61A05AECF929" box="[861,970,1709,1730]" italics="true" pageId="18">T. caroline</emphasis>
</taxonomicName>
(
<figureCitation id="CD9777B73464E962AB8861A15D04F92A" box="[986,1058,1708,1729]" captionStart="Figs" captionStartId="7.[197,237,1661,1680]" captionTargetBox="[173,1193,194,1638]" captionTargetId="figure-1@7.[172,1194,193,1638]" captionTargetPageId="7" captionText="Figs. 6–8. Typhloelmis species, adult dry specimens, dorsal view (length = pronotum + elytra). 6) T. caroline, ♂, length 2.2 mm; 7) T. sanfelipe, ♀, length 2.5 mm; 8) T. finegan, ♀, length 4.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5334786" httpUri="https://zenodo.org/record/5334786/files/figure.png" pageId="18">Figs. 6</figureCitation>
, 
<figureCitation id="CD9777B73464E962AC6261A15D18F92A" box="[1072,1086,1708,1729]" captionStart="Figs" captionStartId="8.[175,215,1686,1705]" captionTargetBox="[188,1135,194,1663]" captionTargetId="figure-1@8.[187,1135,193,1663]" captionTargetPageId="8" captionText="Figs. 9–10. Typhloelmis species, adult wet specimens. 9) T. caroline, a) Dorsal view, b) Ventral view; 10) T. sanfelipe, a) Dorsal view, b) Ventral view." figureDoi="http://doi.org/10.5281/zenodo.5334788" httpUri="https://zenodo.org/record/5334788/files/figure.png" pageId="18">9</figureCitation>
, 
<figureCitation id="CD9777B73464E962AC1961A15D43F92A" box="[1099,1125,1708,1729]" captionStart="Figs" captionStartId="10.[175,215,1239,1258]" captionTargetBox="[151,1171,194,1216]" captionTargetId="figure-212@10.[151,1173,193,1216]" captionTargetPageId="10" captionText="Figs. 13–15. Typhloelmis species, adult wet specimens, anterior dorsal view. 13) T. caroline; 14) T. sanfelipe;" figureDoi="http://doi.org/10.5281/zenodo.5334794" httpUri="https://zenodo.org/record/5334794/files/figure.png" pageId="18">13</figureCitation>
, 
<figureCitation id="CD9777B73464E962AC2161A15DAAF92A" box="[1139,1164,1708,1729]" captionStart="Figs" captionStartId="11.[197,237,1493,1512]" captionTargetBox="[188,1178,193,1470]" captionTargetId="figure-73@11.[187,1178,193,1470]" captionTargetPageId="11" captionText="Figs. 16–19. Typhloelmis species, adult wet specimens, ventral view. 16) T. caroline, anterior; 17) T. sanfelipe, anterior; 18) T. finegan, anterior; 19) T. sanfelipe, fused elytra." figureDoi="http://doi.org/10.5281/zenodo.5334796" httpUri="https://zenodo.org/record/5334796/files/figure.png" pageId="18">16</figureCitation>
) and 
<taxonomicName id="92AC10B13465E963A88767CF5865FF3C" authorityName="Barr &amp; Gibson &amp; Diaz" authorityYear="2015" box="[213,323,194,215]" class="Insecta" family="Elmidae" genus="Typhloelmis" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="species" species="sanfelipe">
<emphasis id="67D8B7203465E963A88767CF5865FF3C" box="[213,323,194,215]" italics="true" pageId="19">T. sanfelipe</emphasis>
</taxonomicName>
(
<figureCitation id="CD9777B73465E963A90367CF58B5FF3C" box="[337,403,194,215]" captionStart="Figs" captionStartId="7.[197,237,1661,1680]" captionTargetBox="[173,1193,194,1638]" captionTargetId="figure-1@7.[172,1194,193,1638]" captionTargetPageId="7" captionText="Figs. 6–8. Typhloelmis species, adult dry specimens, dorsal view (length = pronotum + elytra). 6) T. caroline, ♂, length 2.2 mm; 7) T. sanfelipe, ♀, length 2.5 mm; 8) T. finegan, ♀, length 4.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5334786" httpUri="https://zenodo.org/record/5334786/files/figure.png" pageId="19">Figs. 7</figureCitation>
, 
<figureCitation id="CD9777B73465E963A9CD67CF589EFF3C" box="[415,440,194,215]" captionStart="Figs" captionStartId="8.[175,215,1686,1705]" captionTargetBox="[188,1135,194,1663]" captionTargetId="figure-1@8.[187,1135,193,1663]" captionTargetPageId="8" captionText="Figs. 9–10. Typhloelmis species, adult wet specimens. 9) T. caroline, a) Dorsal view, b) Ventral view; 10) T. sanfelipe, a) Dorsal view, b) Ventral view." figureDoi="http://doi.org/10.5281/zenodo.5334788" httpUri="https://zenodo.org/record/5334788/files/figure.png" pageId="19">10</figureCitation>
, 
<figureCitation id="CD9777B73465E963A99167CF58FAFF3C" box="[451,476,194,215]" captionStart="Figs" captionStartId="10.[175,215,1239,1258]" captionTargetBox="[151,1171,194,1216]" captionTargetId="figure-212@10.[151,1173,193,1216]" captionTargetPageId="10" captionText="Figs. 13–15. Typhloelmis species, adult wet specimens, anterior dorsal view. 13) T. caroline; 14) T. sanfelipe;" figureDoi="http://doi.org/10.5281/zenodo.5334794" httpUri="https://zenodo.org/record/5334794/files/figure.png" pageId="19">14</figureCitation>
, 
<figureCitation id="CD9777B73465E963A9B567CF5B26FF3C" box="[487,512,194,215]" captionStart="Figs" captionStartId="11.[197,237,1493,1512]" captionTargetBox="[188,1178,193,1470]" captionTargetId="figure-73@11.[187,1178,193,1470]" captionTargetPageId="11" captionText="Figs. 16–19. Typhloelmis species, adult wet specimens, ventral view. 16) T. caroline, anterior; 17) T. sanfelipe, anterior; 18) T. finegan, anterior; 19) T. sanfelipe, fused elytra." figureDoi="http://doi.org/10.5281/zenodo.5334796" httpUri="https://zenodo.org/record/5334796/files/figure.png" pageId="19">17</figureCitation>
), the prosternal process is narrow and elongate; the pronotal margins have irregular granules or teeth; antennomeres 1–10 are not similarly cone-shaped, and antennomere 2 is larger than all but antennomeres 1 and 11; the metaventrite is sparsely granulate; the femora are not ventrally flattened; a fringe of tomentum is present on the protibiae only; and protarsomere 5 is bulbous.
</paragraph>
</subSubSection>
<subSubSection id="1DB638B93465E963A89766BC5885F9D7" pageId="19" type="etymology">
<paragraph id="55136B323465E963A89766BC58D7FE0A" blockId="19.[173,660,194,1596]" pageId="19">
<emphasis id="67D8B7203465E963A89766BC5811FE2D" bold="true" box="[197,311,433,454]" pageId="19">Etymology.</emphasis>
<emphasis id="67D8B7203465E963A91666BC58B7FE2D" box="[324,401,433,454]" italics="true" pageId="19">Finegan</emphasis>
, a noun in apposition, from the 
<typeStatus id="8A17D5903465E963A88366C159DFFE0A" box="[209,249,460,481]" pageId="19">type</typeStatus>
locality, Finegan Springs.
</paragraph>
<paragraph id="55136B323465E963A89766EB5808FC54" blockId="19.[173,660,194,1596]" pageId="19">
<emphasis id="67D8B7203465E963A89766EB583FFE10" bold="true" box="[197,281,486,507]" pageId="19">Habitat.</emphasis>
Finegan Springs (
<figureCitation id="CD9777B73465E963A98666EB5B3FFE10" box="[468,537,486,507]" captionStart="Figs" captionStartId="3.[197,237,1662,1681]" captionTargetBox="[250,1154,192,1639]" captionTargetId="figure-1@3.[187,1178,191,1640]" captionTargetPageId="3" captionText="Figs. 1–2. 1) Map of Terrell and Val Verde counties, Texas, showing the type localities of Typhoelmis caroline at Caroline Springs, Typhoelmis sanfelipe at San Felipe Springs, and Typhoelmis finegan at Finegan Springs; 2) Map of the Edwards and Trinity Aquifer regions of Texas. GWD = groundwater divide; black dots = type locality springs." figureDoi="http://doi.org/10.5281/zenodo.5334776" httpUri="https://zenodo.org/record/5334776/files/figure.png" pageId="19">Figs. 1</figureCitation>
, 
<figureCitation id="CD9777B73465E963AA7566EB5B41FE10" box="[551,615,486,507]" captionStart="Figs" captionStartId="20.[175,215,1656,1675]" captionTargetBox="[151,1171,193,1632]" captionTargetId="figure-1@20.[151,1173,192,1632]" captionTargetPageId="20" captionText="Figs. 27–28. Finegan Springs, type locality of Typhloelmis finegan. 27) Spring run with outflow of small springs entering on left; 28) Spring head with three drift nets." figureDoi="http://doi.org/10.5281/zenodo.5334812" httpUri="https://zenodo.org/record/5334812/files/figure.png" pageId="19">27, 28</figureCitation>
) are located in the Devils River State Natural Area (
<collectingRegion id="9768A5D03465E963A8E1651659CFFDDB" box="[179,233,539,560]" country="United States of America" name="Texas" pageId="19">Texas</collectingRegion>
Parks and Wildlife Department) west of Loma Alta, 
<collectingRegion id="9768A5D03465E963A8BA653B5803FDA0" box="[232,293,566,587]" country="United States of America" name="Texas" pageId="19">Texas</collectingRegion>
, in the Edwards Plateau Ecoregion (
<bibRefCitation id="313D16C33465E963A8E1655C5B32FD8D" author="Bureau of Economic Geology &amp; Ecoregions of Texas. Map. Jackson School of Geosciences, University of Texas &amp; Austin" box="[179,532,593,614]" pageId="19" refId="ref15783" refString="Bureau of Economic Geology. 2010. Ecoregions of Texas. Map. Jackson School of Geosciences, University of Texas at Austin, TX." type="book" year="2010">Bureau of Economic Geology 2010</bibRefCitation>
). At the site, a series of small springs (
<figureCitation id="CD9777B73465E963A9CA656658FDFD6B" box="[408,475,619,640]" captionStart="Figs" captionStartId="20.[175,215,1656,1675]" captionTargetBox="[151,1171,193,1632]" captionTargetId="figure-1@20.[151,1173,192,1632]" captionTargetPageId="20" captionText="Figs. 27–28. Finegan Springs, type locality of Typhloelmis finegan. 27) Spring run with outflow of small springs entering on left; 28) Spring head with three drift nets." figureDoi="http://doi.org/10.5281/zenodo.5334812" httpUri="https://zenodo.org/record/5334812/files/figure.png" pageId="19">Fig. 28</figureCitation>
) emerge from early Cretaceous Edwards Limestone (
<bibRefCitation id="313D16C33465E963A9A5658B5859FD5E" author="Bureau of Economic Geology &amp; Geologic" pageId="19" refId="ref15763" refString="Bureau of Economic Geology. 1992. Geologic map of Texas. University of Texas at Austin, TX." type="book" year="1992">Bureau of Economic Geology 1992</bibRefCitation>
) at the base of a limestone bluff adjacent to the east bank of the upper Devils River, joining to form a stream that flows over impermeable chert bedrock (B. F. Schwartz, 
<emphasis id="67D8B7203465E963AA0165FD5BB5FCEE" box="[595,659,752,773]" italics="true" pageId="19">in litt.)</emphasis>
before emptying into the river (
<figureCitation id="CD9777B73465E963A9BA64075B14FCF4" box="[488,562,778,799]" captionStart="Figs" captionStartId="20.[175,215,1656,1675]" captionTargetBox="[151,1171,193,1632]" captionTargetId="figure-1@20.[151,1173,192,1632]" captionTargetPageId="20" captionText="Figs. 27–28. Finegan Springs, type locality of Typhloelmis finegan. 27) Spring run with outflow of small springs entering on left; 28) Spring head with three drift nets." figureDoi="http://doi.org/10.5281/zenodo.5334812" httpUri="https://zenodo.org/record/5334812/files/figure.png" pageId="19">Fig. 27</figureCitation>
). Historic discharges from the springs ranged between 99– 760 L/s (
<bibRefCitation id="313D16C33465E963A95E644D58AEFCBE" author="Brune" box="[268,392,832,853]" pageId="19" refId="ref15739" refString="Brune, G. 1981. Springs of Texas, Volume I. Branch- Smith, Inc., Fort Worth, TX." type="book" year="1981">Brune 1981</bibRefCitation>
) from five measurements taken in the years 1928–1971. The single specimen of 
<taxonomicName id="92AC10B13465E963A8AE64785847FC61" authorityName="Barr &amp; Gibson &amp; Diaz" authorityYear="2015" box="[252,353,885,906]" class="Insecta" family="Elmidae" genus="Typhloelmis" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="species" species="finegan">
<emphasis id="67D8B7203465E963A8AE64785847FC61" box="[252,353,885,906]" italics="true" pageId="19">T. finegan</emphasis>
</taxonomicName>
was collected from the spring outlet termed “Finegan Spring 1” in 
<bibRefCitation id="313D16C33465E963AA5E6482580CFC54" author="Kulkoyluoglu" pageId="19" pagination="47 - 58" refId="ref16484" refString="Kulkoyluoglu, O., R. Gibson, P. H. Diaz, and J. - P. Colin. 2011. Bicornucandona gen. nov., sp. nov. (Crustacea, Ostracoda) from Finegan Spring (Texas, U. S. A.). Zootaxa 3059: 47 - 58." type="journal article" year="2011">
Külköyluoğlü 
<emphasis id="67D8B7203465E963A8FF64A759C6FC54" box="[173,224,938,959]" italics="true" pageId="19">et al.</emphasis>
(2011)
</bibRefCitation>
.
</paragraph>
<paragraph id="55136B323465E963A89764C95885F9D7" blockId="19.[173,660,194,1596]" pageId="19">
<emphasis id="67D8B7203465E963A89764C958A4FC32" bold="true" box="[197,386,964,985]" pageId="19">Associated Fauna.</emphasis>
Finegan Springs are the 
<typeStatus id="8A17D5903465E963A8FF64D259FEFC1F" box="[173,216,991,1012]" pageId="19">type</typeStatus>
locality for the spring-endemic ostracod 
<taxonomicName id="92AC10B13465E963A8FF64F758CAFBC2" authority="Kulkoyluoglu, Gibson, Diaz, and Colin (2011)" authorityName="Kulkoyluoglu, Gibson, Diaz, and Colin" authorityYear="2011" class="Ostracoda" genus="Bicornucandona" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" phylum="Arthropoda" rank="species" species="fineganensis">
<emphasis id="67D8B7203465E963A8FF64F758CBFBE4" box="[173,493,1018,1039]" italics="true" pageId="19">Bicornucandona fineganensis</emphasis>
Külköyluoğlü, Gibson, Diaz, and Colin (2011)
</taxonomicName>
. Other subterranean aquatic fauna present in the outflow of the springs includes the isopods (Crustacea) 
<emphasis id="67D8B7203465E963AA7B63475BB2FBB4" box="[553,660,1098,1119]" italics="true" pageId="19">
<taxonomicName id="92AC10B13465E963AA7B63475BB6FBB4" authorityName="Benedict" authorityYear="1896" box="[553,656,1098,1119]" class="Malacostraca" family="Cirolanidae" genus="Cirolanides" kingdom="Animalia" order="Isopoda" pageId="19" phylum="Arthropoda" rank="species" species="texensis">C. texensis</taxonomicName>
,
</emphasis>
<taxonomicName id="92AC10B13465E963A8FF6369581FFB92" box="[173,313,1124,1145]" class="Malacostraca" family="Asellidae" genus="Lirceolus" kingdom="Animalia" order="Isopoda" pageId="19" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="67D8B7203465E963A8FF63695829FB92" box="[173,271,1124,1145]" italics="true" pageId="19">Lirceolus</emphasis>
sp.
</taxonomicName>
, and 
<taxonomicName id="92AC10B13465E963A92A636958C0FB92" authorityName="Bowman" authorityYear="1992" box="[376,486,1124,1145]" class="Malacostraca" family="Cirolanidae" genus="Speocirolana" kingdom="Animalia" order="Isopoda" pageId="19" phylum="Arthropoda" rank="species" species="hardeni">
<emphasis id="67D8B7203465E963A92A636958C0FB92" box="[376,486,1124,1145]" italics="true" pageId="19">S. hardeni</emphasis>
</taxonomicName>
and the amphipods (Crustacea) 
<taxonomicName id="92AC10B13465E963A90D637258D3FB7F" authorityName="Holsinger" authorityYear="1980" box="[351,501,1151,1172]" class="Malacostraca" family="Artesiidae" genus="Artesia" kingdom="Animalia" order="Amphipoda" pageId="19" phylum="Arthropoda" rank="species" species="subterranea">
<emphasis id="67D8B7203465E963A90D637258D3FB7F" box="[351,501,1151,1172]" italics="true" pageId="19">A. subterranea</emphasis>
</taxonomicName>
, 
<emphasis id="67D8B7203465E963AA57637258E1FB45" italics="true" pageId="19">
<taxonomicName id="92AC10B13465E963AA576372586DFB45" authority=", Paraholsingerius" authorityName="Paraholsingerius" class="Malacostraca" family="Bogidiellidae" genus="Parabogidiella" kingdom="Animalia" order="Amphipoda" pageId="19" phylum="Arthropoda" rank="species" species="americana">P. americana, Paraholsingerius</taxonomicName>
smaragdinus
</emphasis>
(Holsinger), 
<taxonomicName id="92AC10B13465E963AA1363945BABFB45" box="[577,653,1177,1198]" class="Insecta" family="Elmidae" genus="Phanocerus" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="species" species="ruffoi">
<emphasis id="67D8B7203465E963AA1363945BABFB45" box="[577,653,1177,1198]" italics="true" pageId="19">P. ruffoi</emphasis>
</taxonomicName>
, 
<taxonomicName id="92AC10B13465E963A8FF63B95817FB22" box="[173,305,1204,1225]" class="Malacostraca" family="Seborgiidae" genus="Seborgia" kingdom="Animalia" order="Amphipoda" pageId="19" phylum="Arthropoda" rank="species" species="hershleri">
<emphasis id="67D8B7203465E963A8FF63B95817FB22" box="[173,305,1204,1225]" italics="true" pageId="19">S. hershleri</emphasis>
</taxonomicName>
, and 
<taxonomicName id="92AC10B13465E963A92963B95807FB08" authority="(Holsinger)" baseAuthorityName="Holsinger" baseAuthorityYear="1966" class="Malacostraca" family="Crangonyctidae" genus="Stygobromus" kingdom="Animalia" order="Amphipoda" pageId="19" phylum="Arthropoda" rank="species" species="hadenoecus">
<emphasis id="67D8B7203465E963A92963B95BB5FB22" box="[379,659,1204,1225]" italics="true" pageId="19">Stygobromus hadenoecus</emphasis>
(Holsinger)
</taxonomicName>
. Epigean riffle beetles present in the springs include 
<taxonomicName id="92AC10B13465E963A96563E458AAFB15" baseAuthorityName="Horn" baseAuthorityYear="1870" box="[311,396,1257,1278]" class="Insecta" family="Elmidae" genus="Heterelmis" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="67D8B7203465E963A96563E458AAFB15" box="[311,396,1257,1278]" italics="true" pageId="19">H. glabra</emphasis>
</taxonomicName>
, unidentified 
<taxonomicName id="92AC10B13465E963AA5763E45BABFB15" authorityName="Hinton" authorityYear="1935" box="[517,653,1257,1278]" class="Insecta" family="Elmidae" genus="Microcylloepus" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="genus">
<emphasis id="67D8B7203465E963AA5763E45BABFB15" box="[517,653,1257,1278]" italics="true" pageId="19">Microcylloepus</emphasis>
</taxonomicName>
, 
<taxonomicName id="92AC10B13465E963A8FF620E58B0FAF3" authorityName="Sharp" authorityYear="1882" box="[173,406,1283,1304]" class="Insecta" family="Elmidae" genus="Phanocerus" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="species" species="clavicornis">
<emphasis id="67D8B7203465E963A8FF620E5817FAF3" box="[173,305,1283,1304]" italics="true" pageId="19">P. clavicornis</emphasis>
(Elmidae)
</taxonomicName>
and 
<taxonomicName id="92AC10B13465E963A996620E5BB5FAF3" authority="Haldeman" authorityName="Haldeman" authorityYear="1853" box="[452,659,1283,1304]" class="Insecta" family="Psephenidae" genus="Psephenus" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="genus">
<emphasis id="67D8B7203465E963A996620E5B0CFAF3" box="[452,554,1283,1304]" italics="true" pageId="19">Psephenus</emphasis>
Haldeman
</taxonomicName>
(almost certainly 
<taxonomicName id="92AC10B13465E963A90A62135898FAD8" box="[344,446,1310,1331]" class="Insecta" family="Psephenidae" genus="Psephenus" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="species" species="texanus">
<emphasis id="67D8B7203465E963A90A62135898FAD8" box="[344,446,1310,1331]" italics="true" pageId="19">P. texanus</emphasis>
</taxonomicName>
although adults have not been collected) (
<taxonomicName id="92AC10B13465E963A925623558D5FAA6" box="[375,499,1336,1357]" class="Insecta" family="Psephenidae" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="family">Psephenidae</taxonomicName>
); the population of 
<taxonomicName id="92AC10B13465E963A895625E5805FA83" baseAuthorityName="Horn" baseAuthorityYear="1870" box="[199,291,1363,1384]" class="Insecta" family="Elmidae" genus="Heterelmis" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="67D8B7203465E963A895625E5805FA83" box="[199,291,1363,1384]" italics="true" pageId="19">H. glabra</emphasis>
</taxonomicName>
at the springs was included in a population genetics study of 
<taxonomicName id="92AC10B13465E963A9F662635B2BFA68" authorityName="Sharp" authorityYear="1882" box="[420,525,1390,1411]" class="Insecta" family="Elmidae" genus="Heterelmis" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="genus">
<emphasis id="67D8B7203465E963A9F662635B2BFA68" box="[420,525,1390,1411]" italics="true" pageId="19">Heterelmis</emphasis>
</taxonomicName>
by 
<bibRefCitation id="313D16C33465E963AA6A626359C8FA76" author="Gonzales" pageId="19" refId="ref15997" refString="Gonzales, T. K. 2008. Conservation genetics of the Comal Springs riffle beetle (Heterelmis comalensis) population in central Texas, with examination of molecular and morphological variation in Heterelmis sp. throughout Texas. MSc thesis. Texas State University, San Marcos, TX." type="book" year="2008">Gonzales (2008)</bibRefCitation>
. An unidentified species of marsh beetle in the genus 
<taxonomicName id="92AC10B13465E963A94662AE5BB5FA53" authority="Redtenbacher" authorityName="Redtenbacher" authorityYear="1858" box="[276,659,1443,1464]" class="Insecta" family="Scirtidae" genus="Prionocyphon" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="genus">
<emphasis id="67D8B7203465E963A94662AE58BDFA53" box="[276,411,1443,1464]" italics="true" pageId="19">Prionocyphon</emphasis>
Redtenbacher (Scirtidae)
</taxonomicName>
has also been identified from the springs. The spring-associated roundnose minnows 
<taxonomicName id="92AC10B13465E963AA7C62D55BB5FA06" authorityName="Hubbs and Brown (USFWS" authorityYear="2007" box="[558,659,1496,1517]" class="Actinopterygii" family="Cyprinidae" genus="Dionda" kingdom="Animalia" order="Cypriniformes" pageId="19" phylum="Chordata" rank="species" species="diaboli">
<emphasis id="67D8B7203465E963AA7C62D55BB5FA06" box="[558,659,1496,1517]" italics="true" pageId="19">D. diaboli</emphasis>
</taxonomicName>
and 
<taxonomicName id="92AC10B13465E963A88562FE5873F9E3" authorityName="Girard" authorityYear="1856" box="[215,341,1523,1544]" class="Actinopterygii" family="Cyprinidae" genus="Dionda" kingdom="Animalia" order="Cypriniformes" pageId="19" phylum="Chordata" rank="species" species="argentosa">
<emphasis id="67D8B7203465E963A88562FE5873F9E3" box="[215,341,1523,1544]" italics="true" pageId="19">D. argentosa</emphasis>
</taxonomicName>
regularly occur in the stretch of the Devils River from Finegan Springs to Dolan Falls 1.5 km downstream.
</paragraph>
</subSubSection>
</treatment>
</document>