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<document id="B60DF1A6DDEBF51BCDD845A4C332EA55" ID-DOI="10.5281/zenodo.5341440" ID-ISSN="0374-1036" ID-Zenodo-Dep="5341440" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630190347714" checkinUser="felipe" docAuthor="Ribes, Jordi, Pagola-Carte, Santiago &amp; Heiss, Ernst" docDate="2008" docId="DC6D87A0FFE4286375FDFE90FC98F9FE" docLanguage="en" docName="ActaEntMusNatPra.2008.48.2.423-431.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 48 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Systellonotus stysi Ribes &amp; Pagola-Carte &amp; Heiss 2008, sp. nov." docType="treatment" docVersion="3" lastPageNumber="427" masterDocId="2054FFD8FFE52867743AFF83FFB0FFA0" masterDocTitle="Two new Phylinae (Hemiptera: Heteroptera: Miridae) from the Canary Islands" masterLastPageNumber="431" masterPageNumber="423" pageNumber="424" updateTime="1699181410245" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="84B0734C067A8931B2F6FF7CBDFEEAF9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="ECE76E65371F51F5F019E2F4FCB0EC5E">
<mods:title id="A8FB976E00E56DB5024C6E3C8AD6631E">Two new Phylinae (Hemiptera: Heteroptera: Miridae) from the Canary Islands</mods:title>
</mods:titleInfo>
<mods:name id="C0C4659E8EED78B665BACD64D95A5D34" type="personal">
<mods:role id="612F3ECAB8AA4CF63FFC0CB02895F020">
<mods:roleTerm id="5315F9D850B266AEF206CB952BE783E4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B247DE177ABA266908524C0B8FE3BDCB">Ribes, Jordi</mods:namePart>
<mods:affiliation id="EB27E252DA7CBFA4791ED9ED47958816">València 123 - 125, ent., 3 a, E- 08011 Barcelona, Catalonia, Spain; e-mail: 4354 jrr @ comb. es</mods:affiliation>
</mods:name>
<mods:name id="D230D26E1C99C14281CF6E8CE87275F9" type="personal">
<mods:role id="DE362EDCB1BDCB54BDE924BDE7F28D7A">
<mods:roleTerm id="2B4F0D4A938F33EB431889EF40EA5322">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F1A1FE3B92391313A748192FE221CD9E">Pagola-Carte, Santiago</mods:namePart>
<mods:affiliation id="8897FF8EF8906C14D6BC35134ABF19B6">Azpeitia 3, 7. D, E- 20010 Donostia (Gipuzkoa), Spain; e-mail: pagolaxpc @ telefonica. net</mods:affiliation>
</mods:name>
<mods:name id="E427D00D4C25F09A89A8C5F3E4851129" type="personal">
<mods:role id="B7358C8A8DCF2873F9CB39F0FEFC7972">
<mods:roleTerm id="0B3E5B32A6729EEE31E704AFC9F12974">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2AD00E079B0E151CF245E4C5510A978B">Heiss, Ernst</mods:namePart>
<mods:affiliation id="318011BBBA5E9E72AE832539453B4B90">Entomologische Forschungsgruppe, Tiroler Landesmuseum, Josef-Schraffl-Strasse 2 a, A- 6020 Innsbruck, Austria; e-mail: aradus @ aon. at</mods:affiliation>
</mods:name>
<mods:typeOfResource id="715FB257359CFB8B9A5BC9995461C2C1">text</mods:typeOfResource>
<mods:relatedItem id="2325BB5E7D247D1B142F138FEA546DA7" type="host">
<mods:titleInfo id="838DBD69490B3741FD8318EF7A339258">
<mods:title id="9597CF656CF253C9BAAA319AD9C207C6">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="3EB675E17598E016A45EDCCDCC09F0EF">
<mods:date id="5714772D960789C0AEF55DA993A0D90E">2008</mods:date>
<mods:detail id="9578C72F99485B20EC7AB6FE4853D443" type="pubDate">
<mods:number id="F3706222650CAF4D57DADEBEB703923D">2008-12-08</mods:number>
</mods:detail>
<mods:detail id="43D7E1B618B5434D08E0F999C5F0A2D3" type="volume">
<mods:number id="3AD8D440FB9C808489C52F98B2DD86AC">48</mods:number>
</mods:detail>
<mods:detail id="13F461EA3C95DA91887832D103A68861" type="issue">
<mods:number id="57BE981AFC03612E86F5E17DAD4EC4A2">2</mods:number>
</mods:detail>
<mods:extent id="0057C77A38F574C67F448504C88E3A63" unit="page">
<mods:start id="60C8BD610131E2D0A8FE0FD2266D3613">423</mods:start>
<mods:end id="C0D3457AB31E8D26AF6B640D3A77ED51">431</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="93DC4CF437E125DFB09E1D6E1B83AAC1">journal article</mods:classification>
<mods:identifier id="8EF63494DB3C77714F1B90E1CF3B7BD5" type="DOI">10.5281/zenodo.5341440</mods:identifier>
<mods:identifier id="A73F5B71CDF02713F8EDB4B5AFBAECC1" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="2F13B03B6EB1DDAF483E6C6A7E0700E5" type="Zenodo-Dep">5341440</mods:identifier>
</mods:mods>
<treatment id="DC6D87A0FFE4286375FDFE90FC98F9FE" LSID="urn:lsid:plazi:treatment:DC6D87A0FFE4286375FDFE90FC98F9FE" httpUri="http://treatment.plazi.org/id/DC6D87A0FFE4286375FDFE90FC98F9FE" lastPageId="4" lastPageNumber="427" pageId="1" pageNumber="424">
<subSubSection id="1CDE653DFFE4286675FDFE90FD5BFE8C" box="[455,747,275,300]" pageId="1" pageNumber="424" type="nomenclature">
<paragraph id="547B36B6FFE4286675FDFE90FD5BFE8C" blockId="1.[455,747,275,300]" box="[455,747,275,300]" pageId="1" pageNumber="424">
<heading id="0F3381DAFFE4286675FDFE90FD5BFE8C" bold="true" box="[455,747,275,300]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="424" reason="3">
<emphasis id="66B0EAA4FFE4286675FDFE90FD5BFE8C" bold="true" box="[455,747,275,300]" pageId="1" pageNumber="424">
<taxonomicName id="93C44D35FFE4286675FDFE90FD3EFE8B" ID-CoL="544NX" authority="Ribes &amp; Pagola-Carte &amp; Heiss, 2008" authorityName="Ribes &amp; Pagola-Carte &amp; Heiss" authorityYear="2008" box="[455,654,275,300]" class="Insecta" family="Miridae" genus="Systellonotus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="424" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="66B0EAA4FFE4286675FDFE90FD3EFE8B" bold="true" box="[455,654,275,300]" italics="true" pageId="1" pageNumber="424">Systellonotus stysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7D8357DFFFE4286676AFFE97FD5BFE8C" box="[661,747,276,300]" pageId="1" pageNumber="424" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1CDE653DFFE428667616FEB1FD36FEE5" box="[556,646,306,325]" pageId="1" pageNumber="424" type="description">
<paragraph id="547B36B6FFE428667616FEB1FD36FEE5" blockId="1.[113,1090,306,588]" box="[556,646,306,325]" pageId="1" pageNumber="424">
(
<figureCitation id="CCFF2A33FFE428667608FEB1FDCFFEE5" box="[562,639,306,325]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="2.[151,182,754,773]" captionStartId-1="3.[113,144,869,889]" captionStartId-2="4.[751,782,748,768]" captionTargetBox-0="[255,1031,217,730]" captionTargetBox-1="[131,1048,226,847]" captionTargetBox-2="[181,720,233,847]" captionTargetId-0="figure-288@2.[245,1031,217,731]" captionTargetId-1="figure-307@3.[113,1088,216,857]" captionTargetId-2="figure-352@4.[151,743,219,884]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Fig. 1. Systellonotus stysi sp. nov., paratype ♀ from Tenerife. a – habitus, b – head and thorax." captionText-1="Fig. 2. Systellonotus stysi sp. nov. a – head and anterior portion of thorax in lateral view (vestiture omitted); b – head in frontal view (vestiture and antennae omitted); c – anterior half of body in lateral view; d – tarsus of middle leg." captionText-2="Fig. 3. Female genitalia of Systellonotus stysi sp. nov.: dorsal wall of gynatrial complex. Scale bar = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5341442" figureDoi-1="http://doi.org/10.5281/zenodo.5341444" figureDoi-2="http://doi.org/10.5281/zenodo.5341446" httpUri-0="https://zenodo.org/record/5341442/files/figure.png" httpUri-1="https://zenodo.org/record/5341444/files/figure.png" httpUri-2="https://zenodo.org/record/5341446/files/figure.png" pageId="1" pageNumber="424">Figs. 1-3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1CDE653DFFE42866744BFED7FC06FDEC" pageId="1" pageNumber="424" type="materials_examined">
<paragraph id="547B36B6FFE42866744BFED7FDFDFEC7" blockId="1.[113,1090,306,588]" box="[113,589,339,360]" pageId="1" pageNumber="424">
<emphasis id="66B0EAA4FFE42866744BFED7FF5AFEC7" bold="true" box="[113,234,339,359]" pageId="1" pageNumber="424">Type locality.</emphasis>
<collectingRegion id="9600F854FFE4286674D5FED7FEC1FEC7" box="[239,369,340,359]" country="Spain" name="Canarias" pageId="1" pageNumber="424">Canary Islands</collectingRegion>
, Tenerife, Los Cristianos.
</paragraph>
<paragraph id="547B36B6FFE42866744BFEFDFC41FE5C" blockId="1.[113,1090,306,588]" pageId="1" pageNumber="424">
<emphasis id="66B0EAA4FFE42866744BFEFDFF43FE31" bold="true" box="[113,243,381,401]" pageId="1" pageNumber="424">Type material.</emphasis>
<materialsCitation id="E4AC3CEBFFE4286674CCFEFDFC01FE0C" collectionCode="HOLO-, TYPE" collectorName="Los Cristianos &amp; J. de Ferrer' &amp; J. Ribes &amp; Pagola-Carte &amp; Heiss" pageId="1" pageNumber="424" specimenCode="II-57" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="8B7F8814FFE4286674CCFEFDFEF8FE30" box="[246,328,382,401]" pageId="1" pageNumber="424" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCount id="42C2FD3FFFE428667568FEFBFED2FE31" box="[338,354,376,401]" pageId="1" pageNumber="424" type="female">♀</specimenCount>
, ‘TENERIFE /
<collectorName id="F9315360FFE4286675D9FEFDFDEBFE31" box="[483,603,382,401]" pageId="1" pageNumber="424">Los Cristianos</collectorName>
/ 7-
<specimenCode id="04629ECDFFE428667643FEFDFD15FE31" box="[633,677,382,401]" collectionCode="II-" pageId="1" pageNumber="424">II-57</specimenCode>
<collectorName id="F9315360FFE428667692FEFDFCBBFE31" box="[680,779,382,401]" pageId="1" pageNumber="424">J. de Ferrer’</collectorName>
[white, handwritten label] // ‘
<collectionCode id="32D5AE73FFE4286677C1FEFDFF16FE0C" pageId="1" pageNumber="424">HOLO- TYPE</collectionCode>
/
<taxonomicName id="93C44D35FFE42866748DFE19FEE3FE0C" authority="Ribes &amp; Pagola-Carte &amp; Heiss, 2008" authorityName="Ribes &amp; Pagola-Carte &amp; Heiss" authorityYear="2008" box="[183,339,410,428]" class="Insecta" family="Miridae" genus="Systellonotus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="424" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="66B0EAA4FFE42866748DFE19FEE3FE0C" box="[183,339,410,428]" italics="true" pageId="1" pageNumber="424">Systellonotus stysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7D8357DFFFE428667562FE1AFE37FE0C" box="[344,391,409,428]" pageId="1" pageNumber="424" rank="species">n. sp.</taxonomicNameLabel>
/
<collectorName id="F9315360FFE4286675ADFE1AFE6DFE0C" box="[407,477,409,428]" pageId="1" pageNumber="424">J. Ribes</collectorName>
,
<collectorName id="F9315360FFE4286675DFFE1AFDE4FE0C" box="[485,596,409,428]" pageId="1" pageNumber="424">Pagola-Carte</collectorName>
/ &amp;
<collectorName id="F9315360FFE428667641FE1AFD1CFE0C" box="[635,684,409,428]" pageId="1" pageNumber="424">Heiss</collectorName>
, 2008’ [red, typewritten label]
</materialsCitation>
.
<materialsCitation id="E4AC3CEBFFE428667783FE1AFEBAFE41" collectingDate="1992-04-09" collectorName="Zerche'" county="Euphorbia" elevation="400" location="Isl. Can." municipality="Adeje" pageId="1" pageNumber="424" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="8B7F8814FFE428667783FE1AFBA1FE0A" box="[953,1041,409,428]" pageId="1" pageNumber="424" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="42C2FD3FFFE428667026FE1AFB8CFE0C" box="[1052,1084,403,428]" pageId="1" pageNumber="424" type="female">1 ♀</specimenCount>
, ‘
<location id="511B606DFFE42866744FFE30FF72FE66" LSID="urn:lsid:plazi:treatment:DC6D87A0FFE4286375FDFE90FC98F9FE:511B606DFFE42866744FFE30FF72FE66" box="[117,194,435,454]" county="Euphorbia" municipality="Adeje" name="Isl. Can." pageId="1" pageNumber="424">Isl. Can.</location>
,
<location id="511B606DFFE4286674F3FE30FEA0FE66" LSID="urn:lsid:plazi:treatment:DC6D87A0FFE4286375FDFE90FC98F9FE:511B606DFFE4286674F3FE30FEA0FE66" box="[201,272,435,454]" county="Euphorbia" municipality="Adeje" name="Tenerife" pageId="1" pageNumber="424">Tenerife</location>
/
<collectingMunicipality id="B41FACCCFFE428667524FE30FEE3FE66" box="[286,339,435,454]" pageId="1" pageNumber="424">Adeje</collectingMunicipality>
, Bco. del In- / fierno,
<quantity id="933C9B53FFE428667633FE30FD8BFE66" box="[521,571,435,454]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="424" unit="m" value="400.0">
<elevation id="DFE9D185FFE428667633FE30FD8BFE66" box="[521,571,435,454]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="424" unit="m" value="400.0">400m</elevation>
</quantity>
, /
<collectingCounty id="BD1A4E3AFFE428667674FE30FD17FE66" box="[590,679,435,454]" pageId="1" pageNumber="424">Euphorbia</collectingCounty>
carariensis [sic!] /
<date id="207A1076FFE42866777DFE30FC2AFE66" box="[839,922,435,454]" pageId="1" pageNumber="424" value="1992-04-09">
<collectingDate id="303EE99EFFE42866777DFE30FC2AFE66" box="[839,922,435,454]" pageId="1" pageNumber="424" value="1992-04-09">9.IV.1992</collectingDate>
</date>
,leg.
<collectorName id="F9315360FFE428667781FE30FBB0FE66" box="[955,1024,435,454]" pageId="1" pageNumber="424">Zerche’</collectorName>
[white, typewritten label]
</materialsCitation>
;
<specimenCount id="42C2FD3FFFE428667529FE4DFE82FE41" box="[275,306,456,481]" pageId="1" pageNumber="424" type="female">1 ♀</specimenCount>
, ‘La Palma / Volcán Martín / VIII-86 / P. Oromí’ [white, handwritten label]. Both
<typeStatus id="8B7F8814FFE4286677D3FE4DFB8EFE41" box="[1001,1086,462,481]" pageId="1" pageNumber="424" type="paratype">paratypes</typeStatus>
: ‘
<typeStatus id="8B7F8814FFE42866744EFE6AFF52FE5C" box="[116,226,489,508]" pageId="1" pageNumber="424" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="93C44D35FFE4286674C9FE69FE3FFE5C" authority="Ribes &amp; Pagola-Carte &amp; Heiss, 2008" authorityName="Ribes &amp; Pagola-Carte &amp; Heiss" authorityYear="2008" box="[243,399,490,508]" class="Insecta" family="Miridae" genus="Systellonotus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="424" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="66B0EAA4FFE4286674C9FE69FE3FFE5C" box="[243,399,490,508]" italics="true" pageId="1" pageNumber="424">Systellonotus stysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7D8357DFFFE4286675AEFE6AFE73FE5C" box="[404,451,489,508]" pageId="1" pageNumber="424" rank="species">n. sp.</taxonomicNameLabel>
/ J. Ribes, Pagola-Carte / &amp; Heiss, 2008’ [red, typewritten label].
</paragraph>
<paragraph id="547B36B6FFE4286674AAFD80FC06FDEC" blockId="1.[113,1090,306,588]" pageId="1" pageNumber="424">
<typeStatus id="8B7F8814FFE4286674AAFD80FF6FFDB6" box="[144,223,515,534]" pageId="1" pageNumber="424" type="holotype">Holotype</typeStatus>
and
<typeStatus id="8B7F8814FFE428667531FD87FEEDFDB7" box="[267,349,516,535]" pageId="1" pageNumber="424" type="paratype">paratypes</typeStatus>
mounted on a card; the female
<typeStatus id="8B7F8814FFE42866764BFD87FD0AFDB7" box="[625,698,516,535]" pageId="1" pageNumber="424" type="paratype">paratype</typeStatus>
from La Palma with the genitalia glued on a second card below.
<materialsCitation id="E4AC3CEBFFE428667520FD9DFC02FDEC" collectorName="J. Ribes &amp; E. Heiss" country="Spain" county="Donostia" location="Innsbruck" municipality="Pagola-Zabalegui" pageId="1" pageNumber="424" specimenCount="2" typeStatus="holotype">
<typeStatus id="8B7F8814FFE428667520FD9DFED9FD91" box="[282,361,542,561]" pageId="1" pageNumber="424" type="holotype">Holotype</typeStatus>
deposited in the collection of
<collectorName id="F9315360FFE428667657FD9DFD01FD91" box="[621,689,542,561]" pageId="1" pageNumber="424">J. Ribes</collectorName>
(Barcelona,
<collectingCountry id="2CD37626FFE428667725FD9DFCE3FD91" box="[799,851,542,561]" name="Spain" pageId="1" pageNumber="424">Spain</collectingCountry>
);
<typeStatus id="8B7F8814FFE428667758FD9DFC04FD91" box="[866,948,542,561]" pageId="1" pageNumber="424" type="paratype">paratypes</typeStatus>
deposited in the collections of
<collectorName id="F9315360FFE4286674D0FDBAFE81FDEC" box="[234,305,569,588]" pageId="1" pageNumber="424">E. Heiss</collectorName>
(
<location id="511B606DFFE428667506FDBAFE25FDEC" LSID="urn:lsid:plazi:treatment:DC6D87A0FFE4286375FDFE90FC98F9FE:511B606DFFE428667506FDBAFE25FDEC" box="[316,405,569,588]" country="Spain" county="Donostia" municipality="Pagola-Zabalegui" name="Innsbruck" pageId="1" pageNumber="424">Innsbruck</location>
,
<collectingCountry id="2CD37626FFE4286675A7FDBAFE6DFDEC" box="[413,477,569,588]" name="Austria" pageId="1" pageNumber="424">Austria</collectingCountry>
) and
<collectingMunicipality id="B41FACCCFFE428667636FDBAFD12FDEC" box="[524,674,569,588]" pageId="1" pageNumber="424">Pagola-Zabalegui</collectingMunicipality>
(
<collectingCounty id="BD1A4E3AFFE428667694FDBAFD4EFDEC" box="[686,766,569,588]" pageId="1" pageNumber="424">Donostia</collectingCounty>
,
<collectingCountry id="2CD37626FFE42866773DFDBAFC8BFDEC" box="[775,827,569,588]" name="Spain" pageId="1" pageNumber="424">Spain</collectingCountry>
), respectively
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="1CDE653DFFE42865744BFDE7FEEEFABC" lastPageId="2" lastPageNumber="425" pageId="1" pageNumber="424" type="description">
<paragraph id="547B36B6FFE42866744BFDE7FE4AFDDC" blockId="1.[113,1089,612,1564]" box="[113,506,612,636]" pageId="1" pageNumber="424">
<emphasis id="66B0EAA4FFE42866744BFDE7FF4CFDDC" bold="true" box="[113,252,612,636]" pageId="1" pageNumber="424">Description.</emphasis>
Length:
<quantity id="933C9B53FFE428667566FDE7FE46FDDC" box="[348,502,612,636]" metricMagnitude="-3" metricUnit="m" metricValue="3.675" metricValueMax="3.85" metricValueMin="3.5" pageId="1" pageNumber="424" unit="mm" value="3.675" valueMax="3.85" valueMin="3.5">3.50-3.85 mm</quantity>
.
</paragraph>
<paragraph id="547B36B6FFE4286674AAFD07FDB9FCBC" blockId="1.[113,1089,612,1564]" pageId="1" pageNumber="424">
Body surface shiny. Dorsal colouration reddish brown or golden brown, with abdomen darker brown (
<figureCitation id="CCFF2A33FFE428667528FD27FED0FD1C" box="[274,352,676,700]" captionStart="Fig" captionStartId="2.[151,182,754,773]" captionTargetBox="[255,1031,217,730]" captionTargetId="figure-288@2.[245,1031,217,731]" captionTargetPageId="2" captionText="Fig. 1. Systellonotus stysi sp. nov., paratype ♀ from Tenerife. a – habitus, b – head and thorax." figureDoi="http://doi.org/10.5281/zenodo.5341442" httpUri="https://zenodo.org/record/5341442/files/figure.png" pageId="1" pageNumber="424">Fig. 1a</figureCitation>
). Dorsal vestiture of head, pronotum, scutellum and hemelytra consisting of erect, pale setae, shorter than or as long as the diameter of anterior tibiae. Dorsal vestiture of abdomen consisting of semierect, stout, pale uniformly scattered hairs, twice as long as the diameter of anterior tibiae.
</paragraph>
<paragraph id="547B36B6FFE4286674AAFCA7FD9FFC5C" blockId="1.[113,1089,612,1564]" pageId="1" pageNumber="424">
Head in dorsal view (
<figureCitation id="CCFF2A33FFE428667543FCA7FE76FC9C" box="[377,454,804,828]" captionStart="Fig" captionStartId="2.[151,182,754,773]" captionTargetBox="[255,1031,217,730]" captionTargetId="figure-288@2.[245,1031,217,731]" captionTargetPageId="2" captionText="Fig. 1. Systellonotus stysi sp. nov., paratype ♀ from Tenerife. a – habitus, b – head and thorax." figureDoi="http://doi.org/10.5281/zenodo.5341442" httpUri="https://zenodo.org/record/5341442/files/figure.png" pageId="1" pageNumber="424">Fig. 1b</figureCitation>
) almost spherical, 1.2 times as long and 1.6 times as wide as pronotum. Eyes relatively small and flattened, postocular region rounded and strongly constricted towards pronotal collar. Head in frontal view (
<figureCitation id="CCFF2A33FFE4286676E7FCE7FC9BFCDC" box="[733,811,868,892]" captionStart="Fig" captionStartId="3.[113,144,869,889]" captionTargetBox="[131,1048,226,847]" captionTargetId="figure-307@3.[113,1088,216,857]" captionTargetPageId="3" captionText="Fig. 2. Systellonotus stysi sp. nov. a – head and anterior portion of thorax in lateral view (vestiture omitted); b – head in frontal view (vestiture and antennae omitted); c – anterior half of body in lateral view; d – tarsus of middle leg." figureDoi="http://doi.org/10.5281/zenodo.5341444" httpUri="https://zenodo.org/record/5341444/files/figure.png" pageId="1" pageNumber="424">Fig. 2b</figureCitation>
) subtriangular and approximately as high as wide; antennal fossae equally distant from the inferior margin of eye and the base of clypeus. Head in lateral view (
<figureCitation id="CCFF2A33FFE428667603FC27FD35FC1C" box="[569,645,932,956]" captionStart="Fig" captionStartId="3.[113,144,869,889]" captionTargetBox="[131,1048,226,847]" captionTargetId="figure-307@3.[113,1088,216,857]" captionTargetPageId="3" captionText="Fig. 2. Systellonotus stysi sp. nov. a – head and anterior portion of thorax in lateral view (vestiture omitted); b – head in frontal view (vestiture and antennae omitted); c – anterior half of body in lateral view; d – tarsus of middle leg." figureDoi="http://doi.org/10.5281/zenodo.5341444" httpUri="https://zenodo.org/record/5341444/files/figure.png" pageId="1" pageNumber="424">Fig. 2a</figureCitation>
) ovoid, gula slightly curved and covered by erect hairs like the juga; clypeus weakly protruding and separated from frons by a slight transversal sulcus; maxillary plates small.
</paragraph>
<paragraph id="547B36B6FFE4286674AAFB86FF55FBDC" blockId="1.[113,1089,612,1564]" pageId="1" pageNumber="424">
Rostrum reaching or slightly surpassing metacoxae, reddish brown in colour. Segment I thick and nearly as wide as segment I of antennae, its basal half concealed by small bucculae, the latter antero-laterally projected and visible in frontal view (
<figureCitation id="CCFF2A33FFE428667734FBC7FCEDFBFC" box="[782,861,1092,1116]" captionStart="Fig" captionStartId="3.[113,144,869,889]" captionTargetBox="[131,1048,226,847]" captionTargetId="figure-307@3.[113,1088,216,857]" captionTargetPageId="3" captionText="Fig. 2. Systellonotus stysi sp. nov. a – head and anterior portion of thorax in lateral view (vestiture omitted); b – head in frontal view (vestiture and antennae omitted); c – anterior half of body in lateral view; d – tarsus of middle leg." figureDoi="http://doi.org/10.5281/zenodo.5341444" httpUri="https://zenodo.org/record/5341444/files/figure.png" pageId="1" pageNumber="424">Fig. 2b</figureCitation>
); segments II, III and IV thinner.
</paragraph>
<paragraph id="547B36B6FFE4286674AAFB07FED2FABC" blockId="1.[113,1089,612,1564]" pageId="1" pageNumber="424">
Antennae stout; segment I thick, cylindrical, pale brown; segment II brownish with darker apex, slightly to distinctly arched in its basal half and somewhat enlarged apically (
<figureCitation id="CCFF2A33FFE42866702CFB27FF3DFB7C" captionStart="Fig" captionStartId="3.[113,144,869,889]" captionTargetBox="[131,1048,226,847]" captionTargetId="figure-307@3.[113,1088,216,857]" captionTargetPageId="3" captionText="Fig. 2. Systellonotus stysi sp. nov. a – head and anterior portion of thorax in lateral view (vestiture omitted); b – head in frontal view (vestiture and antennae omitted); c – anterior half of body in lateral view; d – tarsus of middle leg." figureDoi="http://doi.org/10.5281/zenodo.5341444" httpUri="https://zenodo.org/record/5341444/files/figure.png" pageId="1" pageNumber="424">Fig. 2a</figureCitation>
); segment III cylindrical, its basal half whitish and apex blackish; segment IV fusiform, black (
<figureCitation id="CCFF2A33FFE428667483FB67FEB6FB5C" box="[185,262,1252,1276]" captionStart="Fig" captionStartId="2.[151,182,754,773]" captionTargetBox="[255,1031,217,730]" captionTargetId="figure-288@2.[245,1031,217,731]" captionTargetPageId="2" captionText="Fig. 1. Systellonotus stysi sp. nov., paratype ♀ from Tenerife. a – habitus, b – head and thorax." figureDoi="http://doi.org/10.5281/zenodo.5341442" httpUri="https://zenodo.org/record/5341442/files/figure.png" pageId="1" pageNumber="424">Fig. 1a</figureCitation>
). Length of antennal segments I-II-III-IV = 0.30-1.05-0.65-
<quantity id="933C9B53FFE428667747FB67FC51FB5C" box="[893,993,1252,1276]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="424" unit="mm" value="0.5">0.50 mm</quantity>
. Ratio of segments III/II = 0.62.
</paragraph>
<paragraph id="547B36B6FFE4286674AAFAA7FE14FA9C" blockId="1.[113,1089,612,1564]" box="[144,420,1316,1340]" pageId="1" pageNumber="424">Ocular index = 4.32-4.65.</paragraph>
<paragraph id="547B36B6FFE4286674AAFAC6FD04FA1C" blockId="1.[113,1089,612,1564]" pageId="1" pageNumber="424">
Pronotum 1.18 times as long as wide and 1.4-1.5 times as long as scutellum, dorsally globose with rounded lateral margins (
<figureCitation id="CCFF2A33FFE4286675D4FAE7FD8CFADC" box="[494,572,1380,1404]" captionStart="Fig" captionStartId="2.[151,182,754,773]" captionTargetBox="[255,1031,217,730]" captionTargetId="figure-288@2.[245,1031,217,731]" captionTargetPageId="2" captionText="Fig. 1. Systellonotus stysi sp. nov., paratype ♀ from Tenerife. a – habitus, b – head and thorax." figureDoi="http://doi.org/10.5281/zenodo.5341442" httpUri="https://zenodo.org/record/5341442/files/figure.png" pageId="1" pageNumber="424">Fig. 1b</figureCitation>
); collar (
<figureCitation id="CCFF2A33FFE4286676A5FAE7FD5CFADC" box="[671,748,1380,1404]" captionStart="Fig" captionStartId="3.[113,144,869,889]" captionTargetBox="[131,1048,226,847]" captionTargetId="figure-307@3.[113,1088,216,857]" captionTargetPageId="3" captionText="Fig. 2. Systellonotus stysi sp. nov. a – head and anterior portion of thorax in lateral view (vestiture omitted); b – head in frontal view (vestiture and antennae omitted); c – anterior half of body in lateral view; d – tarsus of middle leg." figureDoi="http://doi.org/10.5281/zenodo.5341444" httpUri="https://zenodo.org/record/5341444/files/figure.png" pageId="1" pageNumber="424">Fig. 2a</figureCitation>
) depressed, flattened and finely granulate; posterior margin emarginate and depressed, finely granulate. Supracoxal lobes of fore legs visible from above on the sides of pronotum.
</paragraph>
<paragraph id="547B36B6FFE4286674AAFA47FE3DF9BC" blockId="1.[113,1089,612,1564]" pageId="1" pageNumber="424">
Scutellum triangular, nearly as wide as long, lateral margins slightly convex. Disc with a distinctive conical hump on posterior half, its apex blunt, as high as the upper level of head and pronotum (
<figureCitation id="CCFF2A33FFE42866752FF987FE30F9BC" box="[277,384,1540,1564]" captionStart="Fig" captionStartId="3.[113,144,869,889]" captionTargetBox="[131,1048,226,847]" captionTargetId="figure-307@3.[113,1088,216,857]" captionTargetPageId="3" captionText="Fig. 2. Systellonotus stysi sp. nov. a – head and anterior portion of thorax in lateral view (vestiture omitted); b – head in frontal view (vestiture and antennae omitted); c – anterior half of body in lateral view; d – tarsus of middle leg." figureDoi="http://doi.org/10.5281/zenodo.5341444" httpUri="https://zenodo.org/record/5341444/files/figure.png" pageId="1" pageNumber="424">Figs. 2a,c</figureCitation>
).
</paragraph>
<caption id="00BB663EFFE7286574ADFD71FC0BFCA5" ID-DOI="http://doi.org/10.5281/zenodo.5341442" ID-Zenodo-Dep="5341442" box="[151,955,754,774]" httpUri="https://zenodo.org/record/5341442/files/figure.png" pageId="2" pageNumber="425" startId="2.[151,182,754,773]" targetBox="[255,1031,217,730]" targetPageId="2">
<paragraph id="547B36B6FFE7286574ADFD71FC0BFCA5" blockId="2.[151,955,754,774]" box="[151,955,754,774]" pageId="2" pageNumber="425">
Fig. 1.
<taxonomicName id="93C44D35FFE7286574EEFD70FEC0FCA5" authority="Ribes &amp; Pagola-Carte &amp; Heiss, 2008" authorityName="Ribes &amp; Pagola-Carte &amp; Heiss" authorityYear="2008" box="[212,368,755,773]" class="Insecta" family="Miridae" genus="Systellonotus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="425" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="66B0EAA4FFE7286574EEFD70FEC0FCA5" box="[212,368,755,773]" italics="true" pageId="2" pageNumber="425">Systellonotus stysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7D8357DFFFE72865754FFD70FE0AFCA6" box="[373,442,755,774]" pageId="2" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
, paratype ♀ from Tenerife. a – habitus, b – head and thorax.
</paragraph>
</caption>
<paragraph id="547B36B6FFE72865748FFCC7FE03FC7C" blockId="2.[151,1127,836,1564]" pageId="2" pageNumber="425">
Hemelytra without membrane, shiny, distinctly surpassing the apex of scutellum (
<figureCitation id="CCFF2A33FFE728657036FCC7FBE8FCFC" box="[1036,1112,836,860]" captionStart="Fig" captionStartId="2.[151,182,754,773]" captionTargetBox="[255,1031,217,730]" captionTargetId="figure-288@2.[245,1031,217,731]" captionTargetPageId="2" captionText="Fig. 1. Systellonotus stysi sp. nov., paratype ♀ from Tenerife. a – habitus, b – head and thorax." figureDoi="http://doi.org/10.5281/zenodo.5341442" httpUri="https://zenodo.org/record/5341442/files/figure.png" pageId="2" pageNumber="425">Fig. 1b</figureCitation>
), subparallel at base and then convex, posterior part of triangular shape and strongly turned upward (
<figureCitation id="CCFF2A33FFE7286574CCFC07FED2FC3C" box="[246,354,900,924]" captionStart="Fig" captionStartId="3.[113,144,869,889]" captionTargetBox="[131,1048,226,847]" captionTargetId="figure-307@3.[113,1088,216,857]" captionTargetPageId="3" captionText="Fig. 2. Systellonotus stysi sp. nov. a – head and anterior portion of thorax in lateral view (vestiture omitted); b – head in frontal view (vestiture and antennae omitted); c – anterior half of body in lateral view; d – tarsus of middle leg." figureDoi="http://doi.org/10.5281/zenodo.5341444" httpUri="https://zenodo.org/record/5341444/files/figure.png" pageId="2" pageNumber="425">Figs. 2a,c</figureCitation>
). Clavus indistinct, claval suture short (0.35 times as long as scutellum) but distinct. Exocorium with a longitudinal whitish spot, not reaching anterior and posterior apices of corium (
<figureCitation id="CCFF2A33FFE728657562FC47FE16FC7C" box="[344,422,964,988]" captionStart="Fig" captionStartId="2.[151,182,754,773]" captionTargetBox="[255,1031,217,730]" captionTargetId="figure-288@2.[245,1031,217,731]" captionTargetPageId="2" captionText="Fig. 1. Systellonotus stysi sp. nov., paratype ♀ from Tenerife. a – habitus, b – head and thorax." figureDoi="http://doi.org/10.5281/zenodo.5341442" httpUri="https://zenodo.org/record/5341442/files/figure.png" pageId="2" pageNumber="425">Fig. 1b</figureCitation>
).
</paragraph>
<paragraph id="547B36B6FFE72865748FFC67FCBEFBBC" blockId="2.[151,1127,836,1564]" pageId="2" pageNumber="425">
Abdomen transversally arched or swollen in semicircle; colouration brown with darker terminal segments; connexivum reflexed dorsally (
<figureCitation id="CCFF2A33FFE72865768EFB87FCB0FBBC" box="[692,768,1028,1052]" captionStart="Fig" captionStartId="2.[151,182,754,773]" captionTargetBox="[255,1031,217,730]" captionTargetId="figure-288@2.[245,1031,217,731]" captionTargetPageId="2" captionText="Fig. 1. Systellonotus stysi sp. nov., paratype ♀ from Tenerife. a – habitus, b – head and thorax." figureDoi="http://doi.org/10.5281/zenodo.5341442" httpUri="https://zenodo.org/record/5341442/files/figure.png" pageId="2" pageNumber="425">Fig. 1a</figureCitation>
).
</paragraph>
<paragraph id="547B36B6FFE72865748FFBA7FC48FBFB" blockId="2.[151,1127,836,1564]" pageId="2" pageNumber="425">
Evaporatory area of metathoracic scent glands very prominent, transverse, of ivory-whitish colour (
<figureCitation id="CCFF2A33FFE7286574D0FBC7FE88FBFC" box="[234,312,1092,1116]" captionStart="Fig" captionStartId="3.[113,144,869,889]" captionTargetBox="[131,1048,226,847]" captionTargetId="figure-307@3.[113,1088,216,857]" captionTargetPageId="3" captionText="Fig. 2. Systellonotus stysi sp. nov. a – head and anterior portion of thorax in lateral view (vestiture omitted); b – head in frontal view (vestiture and antennae omitted); c – anterior half of body in lateral view; d – tarsus of middle leg." figureDoi="http://doi.org/10.5281/zenodo.5341444" httpUri="https://zenodo.org/record/5341444/files/figure.png" pageId="2" pageNumber="425">Fig. 2c</figureCitation>
), with a robust peritreme and a large orifice bordering metacoxae.
</paragraph>
<paragraph id="547B36B6FFE72865748FFBE7FC00FB1C" blockId="2.[151,1127,836,1564]" pageId="2" pageNumber="425">
Legs long and slender. Tibiae with short, reclining, brown hairs and short, isolated spines arranged along the inner sides. Length of metatarsus =
<quantity id="933C9B53FFE7286576DFFB07FCF7FB3B" box="[741,839,1156,1180]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="2" pageNumber="425" unit="mm" value="0.43">0.43 mm</quantity>
; claw of metatarsus =
<quantity id="933C9B53FFE728657002FB07FF74FB1B" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="425" unit="mm" value="0.08">0.08 mm</quantity>
; length of metatarsal segments I-II-III = 0.17-0.23-
<quantity id="933C9B53FFE7286576DEFB27FCF4FB1B" box="[740,836,1188,1212]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="425" unit="mm" value="0.17">0.17 mm</quantity>
(
<figureCitation id="CCFF2A33FFE728657769FB27FC13FB1C" box="[851,931,1188,1212]" captionStart="Fig" captionStartId="3.[113,144,869,889]" captionTargetBox="[131,1048,226,847]" captionTargetId="figure-307@3.[113,1088,216,857]" captionTargetPageId="3" captionText="Fig. 2. Systellonotus stysi sp. nov. a – head and anterior portion of thorax in lateral view (vestiture omitted); b – head in frontal view (vestiture and antennae omitted); c – anterior half of body in lateral view; d – tarsus of middle leg." figureDoi="http://doi.org/10.5281/zenodo.5341444" httpUri="https://zenodo.org/record/5341444/files/figure.png" pageId="2" pageNumber="425">Fig. 2d</figureCitation>
).
</paragraph>
<paragraph id="547B36B6FFE72865748FFB47FF5FFB5C" blockId="2.[151,1127,836,1564]" pageId="2" pageNumber="425">
Female genitalia. Dorsal wall of gynatrial complex as in
<figureCitation id="CCFF2A33FFE728657721FB47FCEDFB7C" box="[795,861,1220,1244]" captionStart="Fig" captionStartId="4.[751,782,748,768]" captionTargetBox="[181,720,233,847]" captionTargetId="figure-352@4.[151,743,219,884]" captionTargetPageId="4" captionText="Fig. 3. Female genitalia of Systellonotus stysi sp. nov.: dorsal wall of gynatrial complex. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5341446" httpUri="https://zenodo.org/record/5341446/files/figure.png" pageId="2" pageNumber="425">Fig. 3</figureCitation>
. See remarks in the Discussion.
</paragraph>
<paragraph id="547B36B6FFE72865748FFA87FEEEFABC" blockId="2.[151,1127,836,1564]" box="[181,350,1284,1308]" pageId="2" pageNumber="425">Male unknown.</paragraph>
</subSubSection>
<subSubSection id="1CDE653DFFE7286574ADFAA7FED1FADB" pageId="2" pageNumber="425" type="etymology">
<paragraph id="547B36B6FFE7286574ADFAA7FED1FADB" blockId="2.[151,1127,836,1564]" pageId="2" pageNumber="425">
<emphasis id="66B0EAA4FFE7286574ADFAA7FEA8FA9C" bold="true" box="[151,280,1316,1340]" pageId="2" pageNumber="425">Etymology.</emphasis>
We are pleased to dedicate this species to our friend Pavel Štys on the occasion of his 75
<superScript id="A3B19BFEFFE7286574CDFAC1FEB3FAF1" attach="left" box="[247,259,1346,1361]" fontSize="6" pageId="2" pageNumber="425">th</superScript>
birthday as a recognition of his many and great contributions to our knowledge of the
<taxonomicName id="93C44D35FFE7286574E1FAE7FEECFADB" box="[219,348,1380,1403]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heteroptera" pageId="2" pageNumber="425" phylum="Arthropoda" rank="order">Heteroptera</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="1CDE653DFFE7286574ADFA00FBE0FA3C" box="[151,1104,1411,1436]" pageId="2" pageNumber="425" type="distribution">
<paragraph id="547B36B6FFE7286574ADFA00FBE0FA3C" blockId="2.[151,1127,836,1564]" box="[151,1104,1411,1436]" pageId="2" pageNumber="425">
<emphasis id="66B0EAA4FFE7286574ADFA00FE9AFA3B" bold="true" box="[151,298,1411,1435]" pageId="2" pageNumber="425">Distribution.</emphasis>
So far only known from two of the
<collectingRegion id="9600F854FFE728657690FA07FCFDFA3C" box="[682,845,1412,1436]" country="Spain" name="Canarias" pageId="2" pageNumber="425">Canary Islands</collectingRegion>
: Tenerife and La Palma.
</paragraph>
</subSubSection>
<subSubSection id="1CDE653DFFE7286374ADFA20FC98F9FE" lastPageId="4" lastPageNumber="427" pageId="2" pageNumber="425" type="discussion">
<paragraph id="547B36B6FFE7286474ADFA20FD69FB21" blockId="2.[151,1127,836,1564]" lastBlockId="3.[112,1095,969,1633]" lastPageId="3" lastPageNumber="426" pageId="2" pageNumber="425">
<emphasis id="66B0EAA4FFE7286574ADFA20FEA6FA1B" bold="true" box="[151,278,1443,1467]" pageId="2" pageNumber="425">Discussion.</emphasis>
<taxonomicName id="93C44D35FFE728657527FA26FE6FFA1C" authority="Ribes &amp; Pagola-Carte &amp; Heiss, 2008" authorityName="Ribes &amp; Pagola-Carte &amp; Heiss" authorityYear="2008" box="[285,479,1445,1468]" class="Insecta" family="Miridae" genus="Systellonotus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="425" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="66B0EAA4FFE728657527FA26FE6FFA1C" box="[285,479,1445,1468]" italics="true" pageId="2" pageNumber="425">Systellonotus stysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7D8357DFFFE7286575DCFA27FD89FA1B" box="[486,569,1444,1467]" pageId="2" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
shares the set of diagnostic characters of the mainly Mediterranean genus
<taxonomicName id="93C44D35FFE7286575B9FA46FDA0FA7C" authorityName="Fieber" authorityYear="1858" box="[387,528,1477,1500]" class="Insecta" family="Miridae" genus="Systellonotus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="425" phylum="Arthropoda" rank="genus">
<emphasis id="66B0EAA4FFE7286575B9FA46FDA0FA7C" box="[387,528,1477,1500]" italics="true" pageId="2" pageNumber="425">Systellonotus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="30554B47FFE728657676FA47FD57FA7C" author="WAGNER E." box="[588,743,1476,1500]" pageId="2" pageNumber="425" pagination="1 - 421" refId="ref3873" refString="WAGNER E. 1974: Die Miridae Hahn, 1831, des Mittelmeerraumes und der Makaronesischen Inseln (Hemiptera, Heteroptera). Teil 2. Entomologische Abhandlungen Staatlichen Museum fur Tierkunde Dresden 39 (Suppl.) (1973): 1 - 421." type="journal article" year="1974">WAGNER 1974</bibRefCitation>
), except the ratio of segment III to segment II of antennae, which is only
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contrast to the minimum value of 0.75 given by
<bibRefCitation id="30554B47FFE7286574ADF987FEF1F9BC" author="WAGNER E." box="[151,321,1540,1564]" pageId="2" pageNumber="425" pagination="1 - 421" refId="ref3873" refString="WAGNER E. 1974: Die Miridae Hahn, 1831, des Mittelmeerraumes und der Makaronesischen Inseln (Hemiptera, Heteroptera). Teil 2. Entomologische Abhandlungen Staatlichen Museum fur Tierkunde Dresden 39 (Suppl.) (1973): 1 - 421." type="journal article" year="1974">WAGNER (1974)</bibRefCitation>
. In our view, this is a variable character and we suggest extending the range of the ratio down to 0.60. In his key,
<bibRefCitation id="30554B47FFE6286475D5FC49FD25FC42" author="WAGNER E." box="[495,661,970,994]" pageId="3" pageNumber="426" pagination="1 - 421" refId="ref3873" refString="WAGNER E. 1974: Die Miridae Hahn, 1831, des Mittelmeerraumes und der Makaronesischen Inseln (Hemiptera, Heteroptera). Teil 2. Entomologische Abhandlungen Staatlichen Museum fur Tierkunde Dresden 39 (Suppl.) (1973): 1 - 421." type="journal article" year="1974">WAGNER (1974)</bibRefCitation>
stated: ‘Selten ist das 3. Glied kürzer als das 2., dann sind die Augen vom Vorderrande des Pronotum entfernt’ [= Rarely is segment III shorter than II, but then the eyes are distant from the anterior pronotal margin]. However, the ratio is smaller than
<quantity id="933C9B53FFE62864754FFBA9FE2FFBE1" box="[373,415,1065,1090]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="426" unit="in" value="1.0">1 in</quantity>
most species of
<taxonomicName id="93C44D35FFE628647669FBA8FD50FBE2" authorityName="Fieber" authorityYear="1858" box="[595,736,1067,1090]" class="Insecta" family="Miridae" genus="Systellonotus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="426" phylum="Arthropoda" rank="genus">
<emphasis id="66B0EAA4FFE628647669FBA8FD50FBE2" box="[595,736,1067,1090]" italics="true" pageId="3" pageNumber="426">Systellonotus</emphasis>
</taxonomicName>
(mostly in the range of 0.8-0.9).
<taxonomicName id="93C44D35FFE62864744AFBC8FE82FBC2" authority="Ribes &amp; Pagola-Carte &amp; Heiss, 2008" authorityName="Ribes &amp; Pagola-Carte &amp; Heiss" authorityYear="2008" box="[112,306,1099,1122]" class="Insecta" family="Miridae" genus="Systellonotus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="426" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="66B0EAA4FFE62864744AFBC8FE82FBC2" box="[112,306,1099,1122]" italics="true" pageId="3" pageNumber="426">Systellonotus stysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7D8357DFFFE628647503FBC9FE3CFBC1" box="[313,396,1098,1121]" pageId="3" pageNumber="426" rank="species">sp. nov.</taxonomicNameLabel>
shares both characters as the antennal segment III is shorter than segment II and the eyes quite distant from pronotal collar.
</paragraph>
<caption id="00BB663EFFE62864744BFCE6FB8CFC34" ID-DOI="http://doi.org/10.5281/zenodo.5341444" ID-Zenodo-Dep="5341444" httpUri="https://zenodo.org/record/5341444/files/figure.png" pageId="3" pageNumber="426" startId="3.[113,144,869,889]" targetBox="[131,1048,226,847]" targetPageId="3">
<paragraph id="547B36B6FFE62864744BFCE6FB8CFC34" blockId="3.[113,1088,869,916]" pageId="3" pageNumber="426">
<emphasis id="66B0EAA4FFE62864744BFCE6FF18FCD9" bold="true" box="[113,168,869,889]" pageId="3" pageNumber="426">Fig. 2.</emphasis>
<taxonomicName id="93C44D35FFE628647497FCE4FEF8FCD9" authority="Ribes &amp; Pagola-Carte &amp; Heiss, 2008" authorityName="Ribes &amp; Pagola-Carte &amp; Heiss" authorityYear="2008" box="[173,328,871,889]" class="Insecta" family="Miridae" genus="Systellonotus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="426" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="66B0EAA4FFE628647497FCE4FEF8FCD9" box="[173,328,871,889]" italics="true" pageId="3" pageNumber="426">Systellonotus stysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7D8357DFFFE628647576FCE4FE3EFCDA" box="[332,398,871,890]" pageId="3" pageNumber="426" rank="species">sp. nov.</taxonomicNameLabel>
a – head and anterior portion of thorax in lateral view (vestiture omitted); b – head in frontal view (vestiture and antennae omitted); c – anterior half of body in lateral view; d – tarsus of middle leg.
</paragraph>
</caption>
<paragraph id="547B36B6FFE6286474B4FB0AFD94FA81" blockId="3.[112,1095,969,1633]" pageId="3" pageNumber="426">
Within the
<taxonomicName id="93C44D35FFE628647536FB08FE29FB02" authorityName="Fieber" authorityYear="1858" box="[268,409,1163,1186]" class="Insecta" family="Miridae" genus="Systellonotus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="426" phylum="Arthropoda" rank="genus">
<emphasis id="66B0EAA4FFE628647536FB08FE29FB02" box="[268,409,1163,1186]" italics="true" pageId="3" pageNumber="426">Systellonotus</emphasis>
</taxonomicName>
group of genera sensu
<bibRefCitation id="30554B47FFE62864769BFB09FC86FB02" author="SCHUH R. T." box="[673,822,1162,1186]" pageId="3" pageNumber="426" pagination="1 - 332" refId="ref3764" refString="SCHUH R. T. 1974: The Orthotylinae and Phylinae (Hemiptera: Miridae) of South Africa with a phylogenetic analysis of the ant-mimetic tribes of the two subfamilies for the world. Entomologica Americana 47: 1 - 332." type="journal article" year="1974">SCHUH (1974)</bibRefCitation>
and
<bibRefCitation id="30554B47FFE62864774AFB09FB8BFB01" author="LINNAVUORI R. E." box="[880,1083,1161,1185]" pageId="3" pageNumber="426" pagination="1 - 84" refId="ref3683" refString="LINNAVUORI R. E. 1996: Miridae of West and Central Africa (Hemiptera, Heteroptera). Acta Zoologica Fennica 202: 1 - 84." type="journal article" year="1996">LINNAVUORI (1996)</bibRefCitation>
,
<taxonomicName id="93C44D35FFE62864744AFB28FF4DFB62" authorityName="Fieber" authorityYear="1858" box="[112,253,1195,1218]" class="Insecta" family="Miridae" genus="Systellonotus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="426" phylum="Arthropoda" rank="genus">
<emphasis id="66B0EAA4FFE62864744AFB28FF4DFB62" box="[112,253,1195,1218]" italics="true" pageId="3" pageNumber="426">Systellonotus</emphasis>
</taxonomicName>
is included in the first subgroup of genera (
<bibRefCitation id="30554B47FFE6286476E9FB29FC3CFB61" author="LINNAVUORI R. E." box="[723,908,1193,1218]" pageId="3" pageNumber="426" pagination="1 - 84" refId="ref3683" refString="LINNAVUORI R. E. 1996: Miridae of West and Central Africa (Hemiptera, Heteroptera). Acta Zoologica Fennica 202: 1 - 84." type="journal article" year="1996">LINNAVUORI 1996</bibRefCitation>
), which displays a striking tendency towards an ant-like appearance. The myrmecomorphic features include the development of a scutellar hump, observed in a few genera, which resembles a hump on the basal abdominal tergite of some ants.
</paragraph>
<paragraph id="547B36B6FFE6286474B4FAA9FCFDF9C1" blockId="3.[112,1095,969,1633]" pageId="3" pageNumber="426">
Due to the possession of such a hump and some other morphological adaptations (small eyes, general habitus, etc.),
<taxonomicName id="93C44D35FFE6286475ACFAC8FE50FAC2" authority="Ribes &amp; Pagola-Carte &amp; Heiss, 2008" authorityName="Ribes &amp; Pagola-Carte &amp; Heiss" authorityYear="2008" box="[406,480,1355,1378]" class="Insecta" family="Miridae" genus="Systellonotus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="426" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="66B0EAA4FFE6286475ACFAC8FE50FAC2" box="[406,480,1355,1378]" italics="true" pageId="3" pageNumber="426">S. stysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7D8357DFFFE6286475DCFAC9FD89FAC1" box="[486,569,1354,1377]" pageId="3" pageNumber="426" rank="species">sp. nov.</taxonomicNameLabel>
could be misplaced, at first glance, in the mainly African, ant-mimetic genus
<taxonomicName id="93C44D35FFE62864759DFAE8FD6EFA22" authority="Reuter, 1901" authorityName="Reuter" authorityYear="1901" box="[423,734,1386,1410]" class="Insecta" family="Miridae" genus="Aspidacanthus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="426" phylum="Arthropoda" rank="genus">
<emphasis id="66B0EAA4FFE62864759DFAE8FDF5FA22" box="[423,581,1387,1410]" italics="true" pageId="3" pageNumber="426">Aspidacanthus</emphasis>
Reuter, 1901
</taxonomicName>
. Nevertheless, it clearly lacks a number of generic diagnostic features of the latter genus (according to
<bibRefCitation id="30554B47FFE62864774BFA09FB8CFA01" author="LINNAVUORI R. E." box="[881,1084,1417,1441]" pageId="3" pageNumber="426" pagination="1 - 84" refId="ref3683" refString="LINNAVUORI R. E. 1996: Miridae of West and Central Africa (Hemiptera, Heteroptera). Acta Zoologica Fennica 202: 1 - 84." type="journal article" year="1996">LINNAVUORI (1996)</bibRefCitation>
: antennal fossae closer to eyes, neck narrower, rostrum shorter, posterior part of pronotum strongly widened, claval suture not developed, tarsal segment III longer than II). In fact, various myrmecomorphic characters have evolved separately in different lineages. For instance, the genus
<taxonomicName id="93C44D35FFE6286474E0F988FE59F982" authority="Reuter, 1879" authorityName="Reuter" authorityYear="1879" box="[218,489,1546,1570]" class="Insecta" family="Miridae" genus="Laemocoris" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="426" phylum="Arthropoda" rank="genus">
<emphasis id="66B0EAA4FFE6286474E0F988FEE8F982" box="[218,344,1547,1570]" italics="true" pageId="3" pageNumber="426">Laemocoris</emphasis>
Reuter, 1879
</taxonomicName>
, which also includes both African and European species, belongs to the
<taxonomicName id="93C44D35FFE628647539F9A8FECDF9E2" authorityName="Fieber" authorityYear="1858" box="[259,381,1579,1602]" class="Insecta" family="Miridae" genus="Hallodapus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="426" phylum="Arthropoda" rank="genus">
<emphasis id="66B0EAA4FFE628647539F9A8FECDF9E2" box="[259,381,1579,1602]" italics="true" pageId="3" pageNumber="426">Hallodapus</emphasis>
</taxonomicName>
group of genera (
<bibRefCitation id="30554B47FFE628647615F9A9FD50F9E1" author="LINNAVUORI R. E." box="[559,736,1577,1602]" pageId="3" pageNumber="426" pagination="1 - 84" refId="ref3683" refString="LINNAVUORI R. E. 1996: Miridae of West and Central Africa (Hemiptera, Heteroptera). Acta Zoologica Fennica 202: 1 - 84." type="journal article" year="1996">LINNAVUORI 1996</bibRefCitation>
) but shows a strongly ant-mimetic appearance as well, and most of its species possess a scutellar hump.
</paragraph>
<caption id="00BB663EFFE1286376D5FD6FFC2DFC95" ID-DOI="http://doi.org/10.5281/zenodo.5341446" ID-Zenodo-Dep="5341446" httpUri="https://zenodo.org/record/5341446/files/figure.png" pageId="4" pageNumber="427" startId="4.[751,782,748,768]" targetBox="[181,720,233,847]" targetPageId="4">
<paragraph id="547B36B6FFE1286376D5FD6FFC2DFC95" blockId="4.[751,1128,748,821]" pageId="4" pageNumber="427">
<emphasis id="66B0EAA4FFE1286376D5FD6FFC95FD5F" bold="true" box="[751,805,748,768]" pageId="4" pageNumber="427">Fig. 3.</emphasis>
Female genitalia of
<taxonomicName id="93C44D35FFE1286377F4FD6DFBD7FCA0" authority="Ribes &amp; Pagola-Carte &amp; Heiss, 2008" authorityName="Ribes &amp; Pagola-Carte &amp; Heiss" authorityYear="2008" box="[974,1127,750,768]" class="Insecta" family="Miridae" genus="Systellonotus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="427" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="66B0EAA4FFE1286377F4FD6DFBD7FCA0" box="[974,1127,750,768]" italics="true" pageId="4" pageNumber="427">Systellonotus stysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7D8357DFFFE1286376D5FC8BFC88FCBB" box="[751,824,776,795]" pageId="4" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
: dorsal wall of gynatrial complex. Scale bar = 0.2 mm.
</paragraph>
</caption>
<paragraph id="547B36B6FFE12863748FFC04FC55FB5E" blockId="4.[151,1128,902,1631]" pageId="4" pageNumber="427">
<bibRefCitation id="30554B47FFE12863748FFC04FED5FC3F" author="MAGNIEN P." box="[181,357,903,927]" pageId="4" pageNumber="427" pagination="51 - 67" refId="ref3711" refString="MAGNIEN P. 2000: Revision du genre Cremnocephalus Fieber, 1860; description de deux nouvelles especes du sud de la France et de Calabre; note sur les genitalia femelle (Heteroptera, Miridae). Nouvelle Revue d'Entomologie (N. S.) 17: 51 - 67." type="journal article" year="2000">MAGNIEN (2000)</bibRefCitation>
, in his revision of the genus
<taxonomicName id="93C44D35FFE1286376ADFC0BFC6BFC3E" authority="Fieber, 1860" authorityName="Fieber" authorityYear="1860" box="[663,987,902,927]" class="Insecta" family="Miridae" genus="Cremnocephalus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="427" phylum="Arthropoda" rank="genus">
<emphasis id="66B0EAA4FFE1286376ADFC0BFCFBFC3F" box="[663,843,904,927]" italics="true" pageId="4" pageNumber="427">Cremnocephalus</emphasis>
Fieber, 1860
</taxonomicName>
, was the first to deal with the female genitalia in the tribe
<taxonomicName id="93C44D35FFE12863765AFC25FD51FC1E" authorityName="Van Duzee" authorityYear="1916" box="[608,737,934,958]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="427" phylum="Arthropoda" rank="tribe" tribe="Hallodapini">Hallodapini</taxonomicName>
. More recently,
<bibRefCitation id="30554B47FFE1286377B2FC24FB83FC1E" author="WYNIGER D." box="[904,1075,934,958]" pageId="4" pageNumber="427" pagination="1 - 1184" refId="ref3999" refString="WYNIGER D. 2006: The Central European Hallodapini: studies of the female genitalia (Heteroptera, Phylinae, Miridae). Pp. 711 - 720. In: RABITSCH W. (ed.): Hug the bug - For love of true bugs. Festschrift zum 70. Geburtstag von Ernst Heiss. Denisia 19: 1 - 1184." type="journal article" year="2006">WYNIGER (2006)</bibRefCitation>
examined and illustrated the dorsal and posterior walls, the ventral sac and the vestibulum of 10 species belonging to five genera in a comprehensive study of Central European
<taxonomicName id="93C44D35FFE1286377D8FC65FBD3FC5E" authorityName="Van Duzee" authorityYear="1916" box="[994,1123,998,1022]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="427" phylum="Arthropoda" rank="tribe" tribe="Hallodapini">Hallodapini</taxonomicName>
. Concerning the dorsal view of the gynatrial complex, she has shown that the three species of
<taxonomicName id="93C44D35FFE128637489FBABFEF0FB9F" authorityName="Fieber" authorityYear="1858" box="[179,320,1064,1087]" class="Insecta" family="Miridae" genus="Systellonotus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="427" phylum="Arthropoda" rank="genus">
<emphasis id="66B0EAA4FFE128637489FBABFEF0FB9F" box="[179,320,1064,1087]" italics="true" pageId="4" pageNumber="427">Systellonotus</emphasis>
</taxonomicName>
studied (
<taxonomicName id="93C44D35FFE128637598FBABFD53FB9F" authority="Frey-Gessner, 1871" authorityName="Frey-Gessner" authorityYear="1871" box="[418,739,1063,1087]" class="Insecta" family="Miridae" genus="Systellonotus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="427" phylum="Arthropoda" rank="species" species="alpinus">
<emphasis id="66B0EAA4FFE128637598FBABFDBBFB9F" box="[418,523,1064,1087]" italics="true" pageId="4" pageNumber="427">S. alpinus</emphasis>
Frey-Gessner, 1871
</taxonomicName>
,
<taxonomicName id="93C44D35FFE1286376D7FBABFBACFB9F" authority="Horvath, 1894" authorityName="Horvath" authorityYear="1894" box="[749,1052,1062,1087]" class="Insecta" family="Miridae" genus="Systellonotus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="427" phylum="Arthropoda" rank="species" species="discoidalis">
<emphasis id="66B0EAA4FFE1286376D7FBABFCCBFB9F" box="[749,891,1064,1087]" italics="true" pageId="4" pageNumber="427">S. discoidalis</emphasis>
Horváth, 1894
</taxonomicName>
, and
<taxonomicName id="93C44D35FFE128637068FBABFE7BFBFE" authority="(Linnaeus, 1767))" authorityName="" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" class="Insecta" family="Miridae" genus="Systellonotus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="427" phylum="Arthropoda" rank="species" species="triguttatus">
<emphasis id="66B0EAA4FFE128637068FBABFEB6FBFF" italics="true" pageId="4" pageNumber="427">S. triguttatus</emphasis>
(Linnaeus, 1767))
</taxonomicName>
are diverse in the shape of their sclerotized rings (more or less circular or triangular, pointed or rounded apically, and with or without an auricle-like apex) but always have voluminous lateral oviducts. In
<taxonomicName id="93C44D35FFE1286376ADFB0BFD50FB3F" authority="Ribes &amp; Pagola-Carte &amp; Heiss, 2008" authorityName="Ribes &amp; Pagola-Carte &amp; Heiss" authorityYear="2008" box="[663,736,1160,1183]" class="Insecta" family="Miridae" genus="Systellonotus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="427" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="66B0EAA4FFE1286376ADFB0BFD50FB3F" box="[663,736,1160,1183]" italics="true" pageId="4" pageNumber="427">S. stysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7D8357DFFFE1286376DDFB04FC8BFB3E" box="[743,827,1159,1182]" pageId="4" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
, those structures (
<figureCitation id="CCFF2A33FFE1286377C3FB05FB87FB3F" box="[1017,1079,1158,1183]" captionStart="Fig" captionStartId="4.[751,782,748,768]" captionTargetBox="[181,720,233,847]" captionTargetId="figure-352@4.[151,743,219,884]" captionTargetPageId="4" captionText="Fig. 3. Female genitalia of Systellonotus stysi sp. nov.: dorsal wall of gynatrial complex. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5341446" httpUri="https://zenodo.org/record/5341446/files/figure.png" pageId="4" pageNumber="427">Fig. 3</figureCitation>
) are more similar to WYNIGER’ s (2006) drawing for
<taxonomicName id="93C44D35FFE1286376B6FB2BFF66FB7E" authority="(Kirschbaum, 1856)" baseAuthorityName="Kirschbaum" baseAuthorityYear="1856" class="Insecta" family="Miridae" genus="Omphalonotus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="427" phylum="Arthropoda" rank="species" species="quadriguttatus">
<emphasis id="66B0EAA4FFE1286376B6FB2BFC7DFB1F" box="[652,973,1192,1215]" italics="true" pageId="4" pageNumber="427">Omphalonotus quadriguttatus</emphasis>
(Kirschbaum, 1856)
</taxonomicName>
due to the more roundish sclerotized rings, less voluminous lateral oviducts, general shape of the gynatrial complex and membranous, U-shaped antero-medial fold.
</paragraph>
<paragraph id="547B36B6FFE12863748FFA85FC98F9FE" blockId="4.[151,1128,902,1631]" pageId="4" pageNumber="427">
Despite these similarities in the female genitalia,
<taxonomicName id="93C44D35FFE1286376F4FA8BFCA9FABF" authority="Ribes &amp; Pagola-Carte &amp; Heiss, 2008" authorityName="Ribes &amp; Pagola-Carte &amp; Heiss" authorityYear="2008" box="[718,793,1288,1311]" class="Insecta" family="Miridae" genus="Systellonotus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="427" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="66B0EAA4FFE1286376F4FA8BFCA9FABF" box="[718,793,1288,1311]" italics="true" pageId="4" pageNumber="427">S. stysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7D8357DFFFE12863771BFA84FCC6FABE" box="[801,886,1287,1310]" pageId="4" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
cannot be included in the genus
<taxonomicName id="93C44D35FFE12863753FFAABFD85FA9E" authority="Reuter, 1876" authorityName="Reuter" authorityYear="1876" box="[261,565,1318,1343]" class="Insecta" family="Miridae" genus="Omphalonotus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="427" phylum="Arthropoda" rank="genus">
<emphasis id="66B0EAA4FFE12863753FFAABFE12FA9F" box="[261,418,1320,1343]" italics="true" pageId="4" pageNumber="427">Omphalonotus</emphasis>
Reuter, 1876
</taxonomicName>
, because of the quite different external morphology. In
<taxonomicName id="93C44D35FFE12863748EFACBFEE1FAFF" authorityName="Reuter" authorityYear="1876" box="[180,337,1352,1375]" class="Insecta" family="Miridae" genus="Omphalonotus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="427" phylum="Arthropoda" rank="genus">
<emphasis id="66B0EAA4FFE12863748EFACBFEE1FAFF" box="[180,337,1352,1375]" italics="true" pageId="4" pageNumber="427">Omphalonotus</emphasis>
</taxonomicName>
, the eyes are in contact with the anterior margin of the pronotum, the ratio of segments III and II of the antennae is close to 1, and the body shape is rather different. However, one morphological feature of the new species deserves attention, since it is shared with
<taxonomicName id="93C44D35FFE1286374F6FA2BFED9FA1F" authorityName="Reuter" authorityYear="1876" box="[204,361,1448,1471]" class="Insecta" family="Miridae" genus="Omphalonotus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="427" phylum="Arthropoda" rank="genus">
<emphasis id="66B0EAA4FFE1286374F6FA2BFED9FA1F" box="[204,361,1448,1471]" italics="true" pageId="4" pageNumber="427">Omphalonotus</emphasis>
</taxonomicName>
: the prominent bucculae, which are similar to those of
<taxonomicName id="93C44D35FFE128637790FA2BFBD6FA1F" baseAuthorityName="Kirschbaum" baseAuthorityYear="1856" box="[938,1126,1448,1471]" class="Insecta" family="Miridae" genus="Omphalonotus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="427" phylum="Arthropoda" rank="species" species="quadriguttatus">
<emphasis id="66B0EAA4FFE128637790FA2BFBD6FA1F" box="[938,1126,1448,1471]" italics="true" pageId="4" pageNumber="427">O. quadriguttatus</emphasis>
</taxonomicName>
as illustrated by
<bibRefCitation id="30554B47FFE12863757CFA44FE58FA7F" author="WAGNER E." box="[326,488,1479,1503]" pageId="4" pageNumber="427" pagination="1 - 421" refId="ref3873" refString="WAGNER E. 1974: Die Miridae Hahn, 1831, des Mittelmeerraumes und der Makaronesischen Inseln (Hemiptera, Heteroptera). Teil 2. Entomologische Abhandlungen Staatlichen Museum fur Tierkunde Dresden 39 (Suppl.) (1973): 1 - 421." type="journal article" year="1974">WAGNER (1974</bibRefCitation>
: p. 325: Figs. 571c-d), but different to those previously outlined by
<bibRefCitation id="30554B47FFE1286374CEFA64FDB1FA5E" author="WAGNER E. &amp; WEBER H. H." box="[244,513,1510,1535]" pageId="4" pageNumber="427" refId="ref3965" refString="WAGNER E. &amp; WEBER H. H. 1964: Heteropteres Miridae. Faune de France Vol. 67. Federation Francaise des Societes de Sciences Naturelles, Paris, 591 pp." type="book" year="1964">WAGNER &amp; WEBER (1964</bibRefCitation>
: p. 369: Figs. 205c-d). Although the prominent structures are referred to as the maxillary plates (= ‘brides’ in French; ‘Zügel’ in German) in both books (
<bibRefCitation id="30554B47FFE1286374A5F9A4FE2FF99E" author="WAGNER E. &amp; WEBER H. H." box="[159,415,1574,1599]" pageId="4" pageNumber="427" refId="ref3965" refString="WAGNER E. &amp; WEBER H. H. 1964: Heteropteres Miridae. Faune de France Vol. 67. Federation Francaise des Societes de Sciences Naturelles, Paris, 591 pp." type="book" year="1964">WAGNER &amp; WEBER 1964</bibRefCitation>
,
<bibRefCitation id="30554B47FFE128637592F9A4FDF0F99F" author="WAGNER E." box="[424,576,1575,1599]" pageId="4" pageNumber="427" pagination="1 - 421" refId="ref3873" refString="WAGNER E. 1974: Die Miridae Hahn, 1831, des Mittelmeerraumes und der Makaronesischen Inseln (Hemiptera, Heteroptera). Teil 2. Entomologische Abhandlungen Staatlichen Museum fur Tierkunde Dresden 39 (Suppl.) (1973): 1 - 421." type="journal article" year="1974">WAGNER 1974</bibRefCitation>
), they actually represent bucculae in the more recent book as well as in our specimens of
<taxonomicName id="93C44D35FFE128637620F9CBFDD3F9FF" authority="Ribes &amp; Pagola-Carte &amp; Heiss, 2008" authorityName="Ribes &amp; Pagola-Carte &amp; Heiss" authorityYear="2008" box="[538,611,1608,1631]" class="Insecta" family="Miridae" genus="Systellonotus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="427" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="66B0EAA4FFE128637620F9CBFDD3F9FF" box="[538,611,1608,1631]" italics="true" pageId="4" pageNumber="427">S. stysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7D8357DFFFE128637651F9C4FD0DF9FE" box="[619,701,1607,1630]" pageId="4" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="CCFF2A33FFE1286376F1F9C5FCABF9FE" box="[715,795,1606,1630]" captionStart="Fig" captionStartId="3.[113,144,869,889]" captionTargetBox="[131,1048,226,847]" captionTargetId="figure-307@3.[113,1088,216,857]" captionTargetPageId="3" captionText="Fig. 2. Systellonotus stysi sp. nov. a – head and anterior portion of thorax in lateral view (vestiture omitted); b – head in frontal view (vestiture and antennae omitted); c – anterior half of body in lateral view; d – tarsus of middle leg." figureDoi="http://doi.org/10.5281/zenodo.5341444" httpUri="https://zenodo.org/record/5341444/files/figure.png" pageId="4" pageNumber="427">Fig. 2b</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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