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<mods:title id="79EB947E9D9FD35B8FCDD0939C1F9454">Taxonomic revision of the genus Glochidion (Phyllanthaceae) in Taiwan, China</mods:title>
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<mods:namePart id="9A68B760843DBE535C57F63B13A3668E">Yao, Gang</mods:namePart>
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<mods:namePart id="B07735624E062523CB6215EA44224492">Song, Zhu-Qiu</mods:namePart>
<mods:affiliation id="CE82C8D200D8796A42242512E208FA3B">Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 516650, China</mods:affiliation>
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<mods:namePart id="C269ED5C44DFC8C35E5CC121C3CB06A8">Xue, Bin-E</mods:namePart>
<mods:affiliation id="A673ACB01AA052477DFE9568C73C273B">College of Horticulture and Landscape Architecture, Zhongkai University of Agriculture and Engineering, Guangzhou 510225, Guangdong, China</mods:affiliation>
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<mods:namePart id="88DCD5A8AFC8466589A4B0C76A5E5DEB">Shi, Shi</mods:namePart>
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<mods:namePart id="79DAC3A8BEB5D18856EE956876C8CB86">Li, Yu-Ling</mods:namePart>
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<mods:namePart id="1DF85FD7C4B73E2FEE7C4D776D2538F3">Luo, Shi-Xiao</mods:namePart>
<mods:affiliation id="CB4E5A4EDF175068AF106431F5E51AB9">Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 516650, China</mods:affiliation>
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<treatment id="DB527E019340582F9FF1EE1ABA0F769A" ID-GBIF-Taxon="167514866" LSID="urn:lsid:plazi:treatment:DB527E019340582F9FF1EE1ABA0F769A" httpUri="http://treatment.plazi.org/id/DB527E019340582F9FF1EE1ABA0F769A" lastPageNumber="137" pageId="0" pageNumber="137" scope_family="Phyllanthaceae">
<subSubSection id="372242459D777CAAB6DDE48CE071489E" pageId="0" pageNumber="137" type="nomenclature">
<paragraph id="6F1E8C9AE9062DDB7C4D88A27A81E7F6" pageId="0" pageNumber="137">
3. 
<taxonomicName id="D3D2E9451B30A92C0C220DB0D9E5324F" ID-CoL="6KK59" LSID="DB527E01-9340-582F-9FF1-EE1ABA0F769A" authority="Hayata in J. Coll. Sci. Univ. Tokyo 20: 16. 1904" authorityName="Hayata in J. Coll. Sci. Univ. Tokyo 20: 16." authorityYear="1904" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Glochidion lanceolatum" order="Malpighiales" pageId="0" pageNumber="137" phylum="Tracheophyta" rank="species" species="lanceolatum">Glochidion lanceolatum Hayata in J. Coll. Sci. Univ. Tokyo 20: 16. 1904</taxonomicName>
<figureCitation id="C5F04C309710C195A477EEBF9D4E76CC" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. A holotype of Glochidion assamicum var. magnicapsulum Croiatz &amp; Hara (A. Henry 117, A) B holotype of G. kusukusense Hayata (B. Hayata &amp; S. Sasaki s. n., TI) C lectotype of G. lanceolatum Hayata (T. Makino s. n., TI) D holotype of G. chademenosocarpum Hayata (B. Hayata s. n., TI) E isotype of G. rubrum Blume (C. L. von Blume s. n., NY) F isolectotype of G. suishaense Hayata (B. Hayata s. n., TI)." figureDoi="10.3897/phytokeys.159.54839.figure1" httpUri="https://binary.pensoft.net/fig/449558" pageId="0" pageNumber="137">Figures 1B, C</figureCitation>
<figureCitation id="04B6697D00D0591E5F6BBC7D5DFB1951" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. General morphology of Glochidion A-C G. acuminatum var. acuminatum Muell. Arg D, E, H G. ellipticum Wight F, G G. lanceolatum Hayata I-K G. philippicum (Cav.) C. B. Rob L, P G. puberum (L.) Hutch M-O G. rubrum Blume Q-S G. zeylanicum var. zeylanicum (Gaertn.) A. Juss T-V G. zeylanicum var. tomentosum Trimen A, D, F, I, M, R, T female flowers B, E, J, S, U male flowers C, G, H, K, N-Q, V fruits L female flowers and male flowers N male flowers and fruits. Photographs: A-C, F, G, I-K, M-O, Q-U by A. Kawakita (Kyoto University, Japan) D-E, H, L, P by G. Yao V Z. Q. Song." figureDoi="10.3897/phytokeys.159.54839.figure2" httpUri="https://binary.pensoft.net/fig/449559" pageId="0" pageNumber="137">, 2F, G</figureCitation>
<figureCitation id="EB61CD4388B22E48D32FDBE584470B60" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphological comparison between Glochidion lanyuense Gang Yao &amp; S. X. Luo and G. lanceolatum Hayata A, C, E, G, I, L G. lanyuense B, D, F, H, J, K, M G. lanceolatum A, B female flower C, D ovary and style E, F pedicle of female flower G, H general view of style I-K female flowers (shown by arrowheads) L, M fruit. Scar bars: 1 mm (A-H); 3 mm (I-K); 5 mm (L)." figureDoi="10.3897/phytokeys.159.54839.figure5" httpUri="https://binary.pensoft.net/fig/449562" pageId="0" pageNumber="137">, 5A, C, E, G, I, L</figureCitation>
</paragraph>
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<subSubSection id="779466E804E43A9C150FA7E48EECC51D" pageId="0" pageNumber="137" type="reference_group">
<paragraph id="0DADAEA3455C7798B976526E7C2CB128" pageId="0" pageNumber="137">
<taxonomicName id="31F52D10C1634E7D8B21FFBDBD14C325" authorityName="Hayata" authorityYear="1920" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Glochidion kotoense" order="Malpighiales" pageId="0" pageNumber="137" phylum="Tracheophyta" rank="species" species="kotoense">Glochidion kotoense</taxonomicName>
Hayata in Icon. Pl. Form. 9: 96. 1920. Type: China. Formosa (now Taiwan), 
<normalizedToken id="4FA08C7D416D2373E24946FA11D92E05" originalValue="Kôtôshô">Kotosho</normalizedToken>
, 
<emphasis id="F0505242ED168C5AC3FD17FE0B6653CB" italics="true" pageId="0" pageNumber="137">Anonymous s.n.</emphasis>
(holotype: TI-01807, photo!).
</paragraph>
<paragraph id="F1F92325354DFA1ED9F6530FEC2FF706" pageId="0" pageNumber="137">
<taxonomicName id="A0B173C3A6E1E4A817F16EF59699CC5E" authorityName="Hayata" authorityYear="1920" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Glochidion sphaerostigmum" order="Malpighiales" pageId="0" pageNumber="137" phylum="Tracheophyta" rank="species" species="sphaerostigmum">Glochidion sphaerostigmum</taxonomicName>
Hayata in Icon. Pl. Form. 9: 96. 1920. Type: China. Formosa (now Taiwan), Suisha, 
<emphasis id="AFD11F28F7B1A9B6F22E244B92A97C30" italics="true" pageId="0" pageNumber="137">Anonymous s.n.</emphasis>
(holotype: TI-01817, photo!).
</paragraph>
<paragraph id="DCA57CF46E96C997D9B59FF9EE32AEFE" pageId="0" pageNumber="137">
<taxonomicName id="F20F877C62B6A3302123B865029B8D64" authorityName="Hayata" authorityYear="1920" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Glochidion kusukusense" order="Malpighiales" pageId="0" pageNumber="137" phylum="Tracheophyta" rank="species" species="kusukusense">Glochidion kusukusense</taxonomicName>
Hayata in Icon. Pl. Formos. 9: 96. 1920. syn. nov. Type: China. Formosa (now Taiwan), Kusukusu, July 1912, 
<emphasis id="355D50BB6A0EB7ACC9868488DFF41876" italics="true" pageId="0" pageNumber="137">B. Hayata</emphasis>
&amp; 
<emphasis id="44643478332A885F72A25B66389F2205" italics="true" pageId="0" pageNumber="137">S. Sasaki s.n.</emphasis>
(holotype: TI-01808, photo!, Figure 
<figureCitation id="305A5A7DE74C04CCDDBE6D8C68DFFF84" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. A holotype of Glochidion assamicum var. magnicapsulum Croiatz &amp; Hara (A. Henry 117, A) B holotype of G. kusukusense Hayata (B. Hayata &amp; S. Sasaki s. n., TI) C lectotype of G. lanceolatum Hayata (T. Makino s. n., TI) D holotype of G. chademenosocarpum Hayata (B. Hayata s. n., TI) E isotype of G. rubrum Blume (C. L. von Blume s. n., NY) F isolectotype of G. suishaense Hayata (B. Hayata s. n., TI)." figureDoi="10.3897/phytokeys.159.54839.figure1" httpUri="https://binary.pensoft.net/fig/449558" pageId="0" pageNumber="137">1B</figureCitation>
).
</paragraph>
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<subSubSection id="C6756510A09360D173D2AC5F3767B543" pageId="0" pageNumber="137" type="materials_examined">
<paragraph id="E128D5B37D1982C981401CBD5DDA4B0B" pageId="0" pageNumber="137">Type.</paragraph>
<paragraph id="3CF3CE026FDD604098D07CC2B7FA8AEC" pageId="0" pageNumber="137">
China. Formosa (now Taiwan), Kelung, 31 October 1896, 
<emphasis id="1391D5E763D4A06364538BA7E8A00AB3" italics="true" pageId="0" pageNumber="137">T. Makino s.n.</emphasis>
(lectotype: TI-01810, photo!, Figure 
<figureCitation id="5F17DE6FE9DBC8570B9C337C19678DE7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. A holotype of Glochidion assamicum var. magnicapsulum Croiatz &amp; Hara (A. Henry 117, A) B holotype of G. kusukusense Hayata (B. Hayata &amp; S. Sasaki s. n., TI) C lectotype of G. lanceolatum Hayata (T. Makino s. n., TI) D holotype of G. chademenosocarpum Hayata (B. Hayata s. n., TI) E isotype of G. rubrum Blume (C. L. von Blume s. n., NY) F isolectotype of G. suishaense Hayata (B. Hayata s. n., TI)." figureDoi="10.3897/phytokeys.159.54839.figure1" httpUri="https://binary.pensoft.net/fig/449558" pageId="0" pageNumber="137">1C</figureCitation>
; here designated); Remaining syntype: Taiwan, Kelung, 1 November 1896, 
<emphasis id="86EAB39324D5C2FEFECD43CF585CEF62" italics="true" pageId="0" pageNumber="137">C. Owatari s.n.</emphasis>
(TI-01811, photo!).
</paragraph>
</subSubSection>
<subSubSection id="E08332A8292E8ADCDD0E83817D36CAE2" pageId="0" pageNumber="137" type="distribution">
<paragraph id="DE67E6618F8D1A1765479F596DC71025" pageId="0" pageNumber="137">Distribution and habitat.</paragraph>
<paragraph id="FFDD1C868C7689D0DA69E2240FBF827E" pageId="0" pageNumber="137">
<taxonomicName id="2CAD2BB05B6A5A9D2A6F88A74DA15BCC" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Glochidion lanceolatum" order="Malpighiales" pageId="0" pageNumber="137" phylum="Tracheophyta" rank="species" species="lanceolatum">
<emphasis id="943524D6790F774A0951B25933F0E8DE" italics="true" pageId="0" pageNumber="137">Glochidion lanceolatum</emphasis>
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is distributed in China (only in Taiwan) and south Japan, and also recorded from the Philippines (
<bibRefCitation id="0452E56786A58D4C1762EDA4278B5EF4" author="Govaerts, R" journalOrPublisher="Botany" pageId="0" pageNumber="137" publicationUrl="http://wcsp.science.kew.org" refId="B14" refString="Govaerts, R, Dransfield, J, Zona, S, Hodel, DR, Henderson, A, 2019. World Checklist of Phyllanthaceae. Facilitated by the Royal Botanic Gardens, Kew. http://wcsp.science.kew.org" title="World Checklist of Phyllanthaceae. Facilitated by the Royal Botanic Gardens, Kew." url="http://wcsp.science.kew.org" year="2019">Govaerts et al. 2019</bibRefCitation>
). It usually occurs in open forests, roadsides, and at low altitudes. The species is widely distributed from northern to southern Taiwan.
</paragraph>
</subSubSection>
<subSubSection id="EF35C70173721949C82B25C4A4124943" pageId="0" pageNumber="137" type="taxonomic discussion">
<paragraph id="CC841C60921CB0544557B19BC2B80E3E" pageId="0" pageNumber="137">Taxonomic discussion.</paragraph>
<paragraph id="26D8157443AFFEF185E2F163A46FBAD3" pageId="0" pageNumber="137">
The morphological description of 
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<emphasis id="185CBE4982E8CE033639A3F204237B8C" italics="true" pageId="0" pageNumber="137">G. kusukusense</emphasis>
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provided by 
<bibRefCitation id="80492E4457FFAF8771DFF5C7E5325E56" author="Hayata, B" journalOrPublisher="Taipei" pageId="0" pageNumber="137" pagination="94 - 97" refId="B17" refString="Hayata, B, 1920. Icones Plantarum Formosanarum Bureau Productive Industris. Taipei 9: 94 - 97" title="Icones Plantarum Formosanarum Bureau Productive Industris." volume="9" year="1920">Hayata (1920)</bibRefCitation>
is brief and short, and it is in accordance with its type that has only a short branch and several leaves (Figure 
<figureCitation id="221FB9BCA6042013A3F0C3032598B359" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. A holotype of Glochidion assamicum var. magnicapsulum Croiatz &amp; Hara (A. Henry 117, A) B holotype of G. kusukusense Hayata (B. Hayata &amp; S. Sasaki s. n., TI) C lectotype of G. lanceolatum Hayata (T. Makino s. n., TI) D holotype of G. chademenosocarpum Hayata (B. Hayata s. n., TI) E isotype of G. rubrum Blume (C. L. von Blume s. n., NY) F isolectotype of G. suishaense Hayata (B. Hayata s. n., TI)." figureDoi="10.3897/phytokeys.159.54839.figure1" httpUri="https://binary.pensoft.net/fig/449558" pageId="0" pageNumber="137">1B</figureCitation>
). In the protologue, 
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<emphasis id="D6ADB69956A8BCA8F83FEA3C98EBDB12" italics="true" pageId="0" pageNumber="137">G. kusukusense</emphasis>
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was compared morphologically with 
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<emphasis id="1F43C5FD265C715AE4223412E0AC4273" italics="true" pageId="0" pageNumber="137">G. wrightii</emphasis>
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Benth, a species widely distributed in southern and southwestern China. After observing the type of 
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<emphasis id="BD6726AB53429990B51D73A08BDEB54E" italics="true" pageId="0" pageNumber="137">G. kusukusense</emphasis>
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deposited in herbarium TI (Figure 
<figureCitation id="5AFEBD81C1DCA600111AFF2AC7B51828" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. A holotype of Glochidion assamicum var. magnicapsulum Croiatz &amp; Hara (A. Henry 117, A) B holotype of G. kusukusense Hayata (B. Hayata &amp; S. Sasaki s. n., TI) C lectotype of G. lanceolatum Hayata (T. Makino s. n., TI) D holotype of G. chademenosocarpum Hayata (B. Hayata s. n., TI) E isotype of G. rubrum Blume (C. L. von Blume s. n., NY) F isolectotype of G. suishaense Hayata (B. Hayata s. n., TI)." figureDoi="10.3897/phytokeys.159.54839.figure1" httpUri="https://binary.pensoft.net/fig/449558" pageId="0" pageNumber="137">1B</figureCitation>
), we found that its glabrous habit and lanceolate leaves were identical to that of the species 
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<emphasis id="73EC04BCF6D2E45847CDA9CAF9507DB8" italics="true" pageId="0" pageNumber="137">G. lanceolatum</emphasis>
</taxonomicName>
(Figure 
<figureCitation id="DCA7DEF030B6FD69D3DD108281DC4A63" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. A holotype of Glochidion assamicum var. magnicapsulum Croiatz &amp; Hara (A. Henry 117, A) B holotype of G. kusukusense Hayata (B. Hayata &amp; S. Sasaki s. n., TI) C lectotype of G. lanceolatum Hayata (T. Makino s. n., TI) D holotype of G. chademenosocarpum Hayata (B. Hayata s. n., TI) E isotype of G. rubrum Blume (C. L. von Blume s. n., NY) F isolectotype of G. suishaense Hayata (B. Hayata s. n., TI)." figureDoi="10.3897/phytokeys.159.54839.figure1" httpUri="https://binary.pensoft.net/fig/449558" pageId="0" pageNumber="137">1C</figureCitation>
), which is widely distributed in Taiwan. Thus, we considered that 
<taxonomicName id="69F212EB671295D96BDB8CA3A4030899" lsidName="G. kusukusense" pageId="0" pageNumber="137" rank="species" species="kusukusense">
<emphasis id="BB350EE6C5981086BA58D888B0F833BB" italics="true" pageId="0" pageNumber="137">G. kusukusense</emphasis>
</taxonomicName>
is conspecific with 
<taxonomicName id="8946807111FE005C3029379392AB0AD6" lsidName="G. lanceolatum" pageId="0" pageNumber="137" rank="species" species="lanceolatum">
<emphasis id="3D99C22C27A4E9A1A2B8D75776820B8C" italics="true" pageId="0" pageNumber="137">G. lanceolatum</emphasis>
</taxonomicName>
(Figure 
<figureCitation id="3005871EA30937DFDDC84AD733DDBCF4" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. A holotype of Glochidion assamicum var. magnicapsulum Croiatz &amp; Hara (A. Henry 117, A) B holotype of G. kusukusense Hayata (B. Hayata &amp; S. Sasaki s. n., TI) C lectotype of G. lanceolatum Hayata (T. Makino s. n., TI) D holotype of G. chademenosocarpum Hayata (B. Hayata s. n., TI) E isotype of G. rubrum Blume (C. L. von Blume s. n., NY) F isolectotype of G. suishaense Hayata (B. Hayata s. n., TI)." figureDoi="10.3897/phytokeys.159.54839.figure1" httpUri="https://binary.pensoft.net/fig/449558" pageId="0" pageNumber="137">1C</figureCitation>
) and reduced it to be a new synonym of the latter.
</paragraph>
<paragraph id="98B5049D33964CCEF29EBA81C1D6EEE7" pageId="0" pageNumber="137">
In 
<bibRefCitation id="E6EAE50235C3C4D066E666BEE4F1B701" author="Hsu, YS" journalOrPublisher="Quarterly Journal of Forest Research" pageId="0" pageNumber="137" pagination="49 - 68" refId="B20" refString="Hsu, YS, Lu, FY, Deng, SL, Liaw, JC, 2006. A taxonomic study of the Glochidion (Euphorbiaceae) of Taiwan. Quarterly Journal of Forest Research 28: 49 - 68" title="A taxonomic study of the Glochidion (Euphorbiaceae) of Taiwan." volume="28" year="2006">Hsu et al. (2006)</bibRefCitation>
study, some specimens collected from Nantou and Taoyuan of Taiwan (out of the type locality of 
<taxonomicName id="77F6DA853E5DB0418D384C0862E62699" lsidName="G. kusukusense" pageId="0" pageNumber="137" rank="species" species="kusukusense">
<emphasis id="9DFB893AB963EF2EF751DDBAA136C96F" italics="true" pageId="0" pageNumber="137">G. kusukusense</emphasis>
</taxonomicName>
) cannot be identified as 
<taxonomicName id="386975E7C97FD56BA85122C8718E7C14" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Glochidion" order="Malpighiales" pageId="0" pageNumber="137" phylum="Tracheophyta" rank="genus">
<emphasis id="93BE1251A72C9F617032A791240BBCCB" italics="true" pageId="0" pageNumber="137">Glochidion</emphasis>
</taxonomicName>
species usually known to Taiwan, while their lanceolate leaves and glabrous pedicel of male flowers observed seemed to be consistent with the diagnostic traits of 
<taxonomicName id="9F5B35AA686238E00F389DB1C416E31D" lsidName="G. kusukusense" pageId="0" pageNumber="137" rank="species" species="kusukusense">
<emphasis id="B6E549A406ABA8C1B74FAC1DD6D68FC6" italics="true" pageId="0" pageNumber="137">G. kusukusense</emphasis>
</taxonomicName>
, when compared with another dubious species 
<taxonomicName id="E80241230438935553813FA5BB1330D9" lsidName="G. chademenosocarpum" pageId="0" pageNumber="137" rank="species" species="chademenosocarpum">
<emphasis id="BCE0DEFA550B3545886652EBB295C52A" italics="true" pageId="0" pageNumber="137">G. chademenosocarpum</emphasis>
</taxonomicName>
also described in Taiwan, as suggested by 
<bibRefCitation id="B1AC67AF0AE5EB55332521C6A58BCF17" author="Li, PT" journalOrPublisher="Taiwania" pageId="0" pageNumber="137" refId="B28" refString="Li, PT, 1994. Glochidion. Flora Republica Popularis Sinicae. Tomus 44. AngiospermaeDicotyledoneaeEuphorbiaceae. Science Press, Beijing, 133-162." title="Glochidion. Flora Republica Popularis Sinicae. Tomus 44. AngiospermaeDicotyledoneaeEuphorbiaceae. Science Press, Beijing, 133 - 162." year="1994">Li (1994)</bibRefCitation>
based on the protologues of the two species. Thus the distribution of 
<taxonomicName id="ECCDB136689F49CA11339DF0EFA56521" lsidName="G. kusukusense" pageId="0" pageNumber="137" rank="species" species="kusukusense">
<emphasis id="67BABF6F0BDA7253B33572265D5E7AA9" italics="true" pageId="0" pageNumber="137">G. kusukusense</emphasis>
</taxonomicName>
in Taiwan was accepted by 
<bibRefCitation id="293A7688299402D3A473BA29393A1082" author="Hsu, YS" journalOrPublisher="Quarterly Journal of Forest Research" pageId="0" pageNumber="137" pagination="49 - 68" refId="B20" refString="Hsu, YS, Lu, FY, Deng, SL, Liaw, JC, 2006. A taxonomic study of the Glochidion (Euphorbiaceae) of Taiwan. Quarterly Journal of Forest Research 28: 49 - 68" title="A taxonomic study of the Glochidion (Euphorbiaceae) of Taiwan." volume="28" year="2006">Hsu et al. (2006)</bibRefCitation>
, although the type of the species was unavailable in their study. However, the detailed morphological description and line drawing of 
<taxonomicName id="A5B91631E855F324575489AD5301D1DE" lsidName="G. kusukusense" pageId="0" pageNumber="137" rank="species" species="kusukusense">
<emphasis id="9BC900C8379F589DE53C3ED926B3A53B" italics="true" pageId="0" pageNumber="137">G. kusukusense</emphasis>
</taxonomicName>
provided in 
<bibRefCitation id="1B49F3465B573F5F3CEECDA6BB2347B9" author="Hsu, YS" journalOrPublisher="Quarterly Journal of Forest Research" pageId="0" pageNumber="137" pagination="49 - 68" refId="B20" refString="Hsu, YS, Lu, FY, Deng, SL, Liaw, JC, 2006. A taxonomic study of the Glochidion (Euphorbiaceae) of Taiwan. Quarterly Journal of Forest Research 28: 49 - 68" title="A taxonomic study of the Glochidion (Euphorbiaceae) of Taiwan." volume="28" year="2006">Hsu et al. (2006)</bibRefCitation>
are very different from the type of the species but well identical with 
<taxonomicName id="0C84EABF4F4A712095731F806A66AEC9" lsidName="G. ellipticum" pageId="0" pageNumber="137" rank="species" species="ellipticum">
<emphasis id="42BBA72638E16BB3F9C75FD5A680B2D3" italics="true" pageId="0" pageNumber="137">G. ellipticum</emphasis>
</taxonomicName>
, a species distributed in Taiwan but omitted in most literature of Taiwanese 
<taxonomicName id="696C34D113ECCE802F6E87C1CF2CF602" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Glochidion" order="Malpighiales" pageId="0" pageNumber="137" phylum="Tracheophyta" rank="genus">
<emphasis id="2A599B1C7F4E05D2FF72A39F25D62C01" italics="true" pageId="0" pageNumber="137">Glochidion</emphasis>
</taxonomicName>
, including 
<bibRefCitation id="F1DB5D0B4632CC3968CA78338F3A8A08" author="Hsu, YS" journalOrPublisher="Quarterly Journal of Forest Research" pageId="0" pageNumber="137" pagination="49 - 68" refId="B20" refString="Hsu, YS, Lu, FY, Deng, SL, Liaw, JC, 2006. A taxonomic study of the Glochidion (Euphorbiaceae) of Taiwan. Quarterly Journal of Forest Research 28: 49 - 68" title="A taxonomic study of the Glochidion (Euphorbiaceae) of Taiwan." volume="28" year="2006">Hsu et al. (2006)</bibRefCitation>
. The result from rechecking the specimens cited as 
<taxonomicName id="9ED5C6EE02F7FA1332CACEC2DBEBE2C7" lsidName="G. kusukusense" pageId="0" pageNumber="137" rank="species" species="kusukusense">
<emphasis id="D4EB06A5FEF186A7283E5267C8FD5087" italics="true" pageId="0" pageNumber="137">G. kusukusense</emphasis>
</taxonomicName>
in 
<bibRefCitation id="2409355511B4BF40E4B0E4527DE0D4A7" author="Hsu, YS" journalOrPublisher="Quarterly Journal of Forest Research" pageId="0" pageNumber="137" pagination="49 - 68" refId="B20" refString="Hsu, YS, Lu, FY, Deng, SL, Liaw, JC, 2006. A taxonomic study of the Glochidion (Euphorbiaceae) of Taiwan. Quarterly Journal of Forest Research 28: 49 - 68" title="A taxonomic study of the Glochidion (Euphorbiaceae) of Taiwan." volume="28" year="2006">Hsu et al. (2006)</bibRefCitation>
further confirmed our conclusion. More specimens of 
<taxonomicName id="B0F36530652D8F17FF6B9AEF701FA8AF" lsidName="G. ellipticum" pageId="0" pageNumber="137" rank="species" species="ellipticum">
<emphasis id="ED70BBA5DF4B528710D89BAE42FC8787" italics="true" pageId="0" pageNumber="137">G. ellipticum</emphasis>
</taxonomicName>
collected from Chiayi, Kaosiung, Keelung, Pingtung, Nantou, Taichung and Tainan of Taiwan were also found and studied in the current study (see 'Representative specimens 
<normalizedToken id="F39B59B8982ED9AF716401EB8FB1FF84" originalValue="examined’">examined'</normalizedToken>
under the species 
<taxonomicName id="AD40E91F0D2AB4D1EC2B3399395B9186" lsidName="G. ellipticum" pageId="0" pageNumber="137" rank="species" species="ellipticum">
<emphasis id="C2FE4A71E312CD748DBC18AA5601025F" italics="true" pageId="0" pageNumber="137">G. ellipticum</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="6ED9D5E283A26A94EC1D79A0E5E4A9EE" pageId="0" pageNumber="137">
Morphologically, the species 
<taxonomicName id="172F46943D77950B1942E5C14E113DDE" lsidName="G. lanceolatum" pageId="0" pageNumber="137" rank="species" species="lanceolatum">
<emphasis id="C4238FB2EBD613C42601941098A79113" italics="true" pageId="0" pageNumber="137">G. lanceolatum</emphasis>
</taxonomicName>
is similar to the typical variety of 
<taxonomicName id="D53B710718C36A3CE871964EE090E407" lsidName="G. zeylanicum" pageId="0" pageNumber="137" rank="species" species="zeylanicum">
<emphasis id="2F4057DF4D3C2130DCDEA88088B5795B" italics="true" pageId="0" pageNumber="137">G. zeylanicum</emphasis>
</taxonomicName>
, but differs by its smaller leaves (6-13 
<normalizedToken id="D49165AC5FEBC296A9DF15F9E14F68F4" originalValue="×">x</normalizedToken>
2.5-4 cm), ovaries 4-6-locular, and capsules 6-7 mm in diameter (Figure 
<figureCitation id="A24E6697A3B446009E3299A8AC6E3C9B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. General morphology of Glochidion A-C G. acuminatum var. acuminatum Muell. Arg D, E, H G. ellipticum Wight F, G G. lanceolatum Hayata I-K G. philippicum (Cav.) C. B. Rob L, P G. puberum (L.) Hutch M-O G. rubrum Blume Q-S G. zeylanicum var. zeylanicum (Gaertn.) A. Juss T-V G. zeylanicum var. tomentosum Trimen A, D, F, I, M, R, T female flowers B, E, J, S, U male flowers C, G, H, K, N-Q, V fruits L female flowers and male flowers N male flowers and fruits. Photographs: A-C, F, G, I-K, M-O, Q-U by A. Kawakita (Kyoto University, Japan) D-E, H, L, P by G. Yao V Z. Q. Song." figureDoi="10.3897/phytokeys.159.54839.figure2" httpUri="https://binary.pensoft.net/fig/449559" pageId="0" pageNumber="137">2G</figureCitation>
). In contrast, 
<taxonomicName id="19B63BAB63A30FD8E9FF2C7B75A3E34E" authority="var. zeylanicum" lsidName="G. zeylanicum subsp. var. var. zeylanicum" pageId="0" pageNumber="137" rank="variety" species="zeylanicum" subSpecies="var." variety="zeylanicum">G. zeylanicum var. zeylanicum</taxonomicName>
has larger leaves (8.5-23.5 
<normalizedToken id="EB417F0DD8FC12B21FF4EEA89949E542" originalValue="×">x</normalizedToken>
5-9 cm), ovaries 6-8-locular, and capsules 8-12 mm in diameter (Figure 
<figureCitation id="3A83661E5F612DCD3F12917FE67DDBDE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. General morphology of Glochidion A-C G. acuminatum var. acuminatum Muell. Arg D, E, H G. ellipticum Wight F, G G. lanceolatum Hayata I-K G. philippicum (Cav.) C. B. Rob L, P G. puberum (L.) Hutch M-O G. rubrum Blume Q-S G. zeylanicum var. zeylanicum (Gaertn.) A. Juss T-V G. zeylanicum var. tomentosum Trimen A, D, F, I, M, R, T female flowers B, E, J, S, U male flowers C, G, H, K, N-Q, V fruits L female flowers and male flowers N male flowers and fruits. Photographs: A-C, F, G, I-K, M-O, Q-U by A. Kawakita (Kyoto University, Japan) D-E, H, L, P by G. Yao V Z. Q. Song." figureDoi="10.3897/phytokeys.159.54839.figure2" httpUri="https://binary.pensoft.net/fig/449559" pageId="0" pageNumber="137">2Q</figureCitation>
). Additionally, as revealed in previous studies, pollen morphology of the two taxa also showed differences in terms of pollen size and aperture system (
<bibRefCitation id="1D24169374F9EB2A984F89CB3424B99A" author="Deng, MZ" journalOrPublisher="MSc thesis, Taiwan" pageId="0" pageNumber="137" refId="B9" refString="Deng, MZ, 1992. A taxonomic study of Phyllantheae (Euphorbiaceae) of Taiwan. MSc thesis, Taiwan" title="A taxonomic study of Phyllantheae (Euphorbiaceae) of Taiwan." year="1992">Deng 1992</bibRefCitation>
; 
<bibRefCitation id="914568DBA8A59DBF9C029F3E041DE1A8" DOI="https://doi.org/10.1111/njb.00912" author="Yao, G" journalOrPublisher="Nordic Journal of Botany" pageId="0" pageNumber="137" pagination="102 - 110" refId="B45" refString="Yao, G, Zhang, DX, 2016. Pollen morphology of Chinese Glochidion (Phyllanthaceae) and its taxonomic implications. Nordic Journal of Botany 34 (1): 102 - 110, DOI: https://doi.org/10.1111/njb.00912" title="Pollen morphology of Chinese Glochidion (Phyllanthaceae) and its taxonomic implications." url="https://doi.org/10.1111/njb.00912" volume="34" year="2016">Yao and Zhang 2016</bibRefCitation>
). Pollen grains of 
<taxonomicName id="8399D0DCC8593BD51BAB47DD8A2699F1" lsidName="G. lanceolatum" pageId="0" pageNumber="137" rank="species" species="lanceolatum">
<emphasis id="905B0D99EDF3AE147FA2E8005FF8C6C5" italics="true" pageId="0" pageNumber="137">G. lanceolatum</emphasis>
</taxonomicName>
are smaller in size [polar axis (P) = 17.25 
<normalizedToken id="47B58754D7A6A302B79A17FABD1B8F5F" originalValue="μm">μm</normalizedToken>
, equatorial axis (E) = 15.47 
<normalizedToken id="0D8D33D0898174B59AC40C65D3D24192" originalValue="μm">μm</normalizedToken>
] and showed a 3-4-colporate aperture pattern (
<bibRefCitation id="E53104D9A6BF84A5E9688BC139B46DA5" DOI="https://doi.org/10.1111/njb.00912" author="Yao, G" journalOrPublisher="Nordic Journal of Botany" pageId="0" pageNumber="137" pagination="102 - 110" refId="B45" refString="Yao, G, Zhang, DX, 2016. Pollen morphology of Chinese Glochidion (Phyllanthaceae) and its taxonomic implications. Nordic Journal of Botany 34 (1): 102 - 110, DOI: https://doi.org/10.1111/njb.00912" title="Pollen morphology of Chinese Glochidion (Phyllanthaceae) and its taxonomic implications." url="https://doi.org/10.1111/njb.00912" volume="34" year="2016">Yao and Zhang 2016</bibRefCitation>
). In contrast, pollen grains of 
<taxonomicName id="8B1E27018F0B8604BD2F412E013414AA" authority="var. zeylanicum" lsidName="G. zeylanicum subsp. var. var. zeylanicum" pageId="0" pageNumber="137" rank="variety" species="zeylanicum" subSpecies="var." variety="zeylanicum">G. zeylanicum var. zeylanicum</taxonomicName>
are larger in size (P = 22.28 
<normalizedToken id="1CE2117D1ECC2D2215E647D41E2C6700" originalValue="μm">μm</normalizedToken>
, E = 19.49μm) and showed a 4-colporate aperture pattern (
<bibRefCitation id="6C373359D6002CE6477DC959B55B2972" DOI="https://doi.org/10.1111/njb.00912" author="Yao, G" journalOrPublisher="Nordic Journal of Botany" pageId="0" pageNumber="137" pagination="102 - 110" refId="B45" refString="Yao, G, Zhang, DX, 2016. Pollen morphology of Chinese Glochidion (Phyllanthaceae) and its taxonomic implications. Nordic Journal of Botany 34 (1): 102 - 110, DOI: https://doi.org/10.1111/njb.00912" title="Pollen morphology of Chinese Glochidion (Phyllanthaceae) and its taxonomic implications." url="https://doi.org/10.1111/njb.00912" volume="34" year="2016">Yao and Zhang 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F7DBBAAB7F4158CC8E79AFCD4D40659F" pageId="0" pageNumber="137" type="materials_examined">
<paragraph id="DDB4E008D5D120536C978A2E412214D5" pageId="0" pageNumber="137">Representative specimens examined.</paragraph>
<paragraph id="959F03730E6FF6447CF28768F186224D" pageId="0" pageNumber="137">
China. Taiwan. Hsinchu Hsien, Peipu, 
<geoCoordinate id="AB018E9E1FA79A5EB4259CDD69BA5D42" degrees="24" direction="north" minutes="42" orientation="latitude" precision="15" seconds="0" value="24.7">24°42'0&quot;N</geoCoordinate>
, 
<geoCoordinate id="729DE3F392540F0EA89A80208F70419A" degrees="121" direction="east" minutes="3" orientation="longitude" precision="15" seconds="5" value="121.05139">121°3'5&quot;E</geoCoordinate>
, 12 January 1908, 
<emphasis id="0307C1F063EC47662C9A56476C7F3E8D" italics="true" pageId="0" pageNumber="137">U. Mori s.n.</emphasis>
(TAI); Hualien Hsien, Tungmen, Wunlan, at an elevation of 180-250 m, 23 November 1982, 
<emphasis id="7C773114B84FB57A98B59EC928217073" italics="true" pageId="0" pageNumber="137">Y. Tateishi 16250</emphasis>
(HAST); Keelung, 
<geoCoordinate id="1B2C837F2BA9F8B2D629FE3E41AD71FF" degrees="25" direction="north" minutes="7" orientation="latitude" precision="15" seconds="43" value="25.12861">25°7'43&quot;N</geoCoordinate>
, 
<geoCoordinate id="2107EBB655652F09074B8900E8BF508B" degrees="121" direction="east" minutes="44" orientation="longitude" precision="15" seconds="9" value="121.73583">121°44'9&quot;E</geoCoordinate>
, 8 September 1928, 
<emphasis id="A25920E84B47B0CE1B61D104CDB07A67" italics="true" pageId="0" pageNumber="137">S. Sasaki s.n.</emphasis>
(TAI); Kaohsiung, Shoushan, al. 400 m, 16 October 1985, 
<emphasis id="48A462C99C1A5CE4E211842F516B3AAD" italics="true" pageId="0" pageNumber="137">S.Y. Lu 17379</emphasis>
(HAST); Keelung, Ensorei, 
<geoCoordinate id="82588A3FD692726510E46033F9978602" degrees="25" direction="north" minutes="7" orientation="latitude" precision="15" seconds="32" value="25.125555">25°7'32&quot;N</geoCoordinate>
, 
<geoCoordinate id="4A6E932FF9E8047AF6B8E2C537AE2A48" degrees="121" direction="east" minutes="45" orientation="longitude" precision="15" seconds="56" value="121.76556">121°45'56&quot;E</geoCoordinate>
, 14 April 1929, 
<emphasis id="D5C523336B30B7031A8965946A12AB61" italics="true" pageId="0" pageNumber="137">S. Suzuki s.n.</emphasis>
(TAI); Keelung, Hopingtao, 
<geoCoordinate id="9B24CD11FF098A19C5D8C56FF6FEFBDC" degrees="25" direction="north" minutes="9" orientation="latitude" precision="15" seconds="33" value="25.159166">25°9'33&quot;N</geoCoordinate>
, 
<geoCoordinate id="A1235323ACADEE444CE49D0B34522C5C" degrees="121" direction="east" minutes="45" orientation="longitude" precision="15" seconds="5" value="121.75139">121°45'5&quot;E</geoCoordinate>
, 8 December 1963, 
<emphasis id="7A6584F5C972F0C8A62E204AE1EB899E" italics="true" pageId="0" pageNumber="137">C.C. Chuang &amp; M.T. Kao 5541</emphasis>
(PH, TAI); Ilan Hsien, Lotung, 
<geoCoordinate id="E81C5268BC16D465266821CA20713181" degrees="24" direction="north" minutes="40" orientation="latitude" precision="15" seconds="58" value="24.682777">24°40'58&quot;N</geoCoordinate>
, 
<geoCoordinate id="5442F18FC08264951DC42FE608E8DB25" degrees="121" direction="east" minutes="47" orientation="longitude" precision="15" seconds="13" value="121.78694">121°47'13&quot;E</geoCoordinate>
, 13 November 1932, 
<emphasis id="C8EA545B444A60EBFA220A002442E039" italics="true" pageId="0" pageNumber="137">S. Suziki 12348</emphasis>
(TAI); Ilan Hsien, Suao, 
<geoCoordinate id="9EA4E6C31F6DE22DD33DD1905F6FA825" degrees="24" direction="north" minutes="35" orientation="latitude" precision="15" seconds="34" value="24.59278">24°35'34&quot;N</geoCoordinate>
, 
<geoCoordinate id="9223B9D4A8CC8A2AA84B8D28FF32F547" degrees="121" direction="east" minutes="50" orientation="longitude" precision="15" seconds="38" value="121.843895">121°50'38&quot;E</geoCoordinate>
, 
<emphasis id="4E702DEB4A1D2D8F5C3FC74D336FC51E" italics="true" pageId="0" pageNumber="137">Y.M. Hsu 113</emphasis>
(TAI); Ilan Hsien, Lungtanhu, along the paved road surrounding the lake, at an elevation of 100 m, 23 January 1997, 
<emphasis id="E771043525865EDAB3BDC4456A1C87E9" italics="true" pageId="0" pageNumber="137">S.M. Liu 556</emphasis>
(HAST); Ilan Hsien, Toucheng Town, TaHSI, Taoyuanku trail, at an elevation of 50 m, 16 November 2000, 
<emphasis id="80B0157BFDED42C43945790B75DBA422" italics="true" pageId="0" pageNumber="137">J.J. Chen 539</emphasis>
(HAST); Miaoli Hsien, Zhuolan, at an elevation of 0-300 m, 10 July 2001, 
<emphasis id="75B4E09A00FDA4DE4591B5E3B0FC2A16" italics="true" pageId="0" pageNumber="137">C.M. Wang 5060</emphasis>
(IBSC); Pingtung Hsien, Manchou Hsiang, Chunhsing Bridge-Chiatulu, 
<geoCoordinate id="6F30547160A0AC096933CA08B5792A76" degrees="22" direction="north" minutes="01" orientation="latitude" precision="15" seconds="17" value="22.021389">22°01'17&quot;N</geoCoordinate>
, 
<geoCoordinate id="CC904D491589CD23DD795D68817A7BEA" degrees="120" direction="east" minutes="48" orientation="longitude" precision="15" seconds="29" value="120.80806">120°48'29&quot;E</geoCoordinate>
, at an elevation of 60-100, on broadleaf forest, 11 April 1998, 
<emphasis id="2E1BB7354658C5B7C90B70684124F432" italics="true" pageId="0" pageNumber="137">C.M. Wang et al. 03069</emphasis>
(PE); Pingtung Hsien, Peiyeh-Shanpaiwan, 
<geoCoordinate id="03BC4E477231EBC464CD3830F5482E25" degrees="22" direction="north" minutes="42" orientation="latitude" precision="15" seconds="6" value="22.701668">22°42'6&quot;N</geoCoordinate>
, 
<geoCoordinate id="B7BF115451B9A4690A6EBDB2A0368CCC" degrees="120" direction="east" minutes="38" orientation="longitude" precision="15" seconds="31" value="120.641945">120°38'31&quot;E</geoCoordinate>
, 25 December 1930, 
<emphasis id="641B8CC65FA27707744C7D17B3534F36" italics="true" pageId="0" pageNumber="137">S. Suzuki 6798</emphasis>
(TAI); Pingtung Hsien, Shihtzu Hsiang, Shouchia-Mutan, 
<geoCoordinate id="4D87B245418810F0052B82A9711FEDC6" degrees="22" direction="north" minutes="14" orientation="latitude" precision="15" seconds="46" value="22.24611">22°14'46&quot;N</geoCoordinate>
, 
<geoCoordinate id="3D167FE5451EB6AA6E8312F18D562B30" degrees="120" direction="east" minutes="49" orientation="longitude" precision="15" seconds="49" value="120.83028">120°49'49&quot;E</geoCoordinate>
, at an elevation of 420 m, roadside, 26 March 1999, 
<emphasis id="FDAB2108FC832A10F5FDAEA9DDE1281D" italics="true" pageId="0" pageNumber="137">C.M. Wang 03936</emphasis>
(PE); Taipei, Muchihshan, 
<geoCoordinate id="FB15093822CDBD8A1782E3DB2F4EF805" degrees="25" direction="north" minutes="1" orientation="latitude" precision="15" seconds="15" value="25.020834">25°1'15&quot;N</geoCoordinate>
, 
<geoCoordinate id="DAA5DF73BA74F7EBC663C80CBE2D5495" degrees="121" direction="east" minutes="35" orientation="longitude" precision="15" seconds="3" value="121.58417">121°35'3&quot;E</geoCoordinate>
, 12 April 1985, 
<emphasis id="9543DD77F5459AF4EEA036F9054C650B" italics="true" pageId="0" pageNumber="137">S.F. Huang 2780</emphasis>
(TAI); Taipei, Peitou, 
<geoCoordinate id="AF73684729ACF7B1D418114B75D03126" degrees="25" direction="north" minutes="7" orientation="latitude" precision="15" seconds="42" value="25.128334">25°7'42&quot;N</geoCoordinate>
, 
<geoCoordinate id="28722766CAFB8AA9152E0BB4A507FE4C" degrees="121" direction="east" minutes="29" orientation="longitude" precision="15" seconds="42" value="121.494995">121°29'42&quot;E</geoCoordinate>
, 13 December 1931, 
<emphasis id="589F3B814244E684AEF3B75D8364364E" italics="true" pageId="0" pageNumber="137">T. Suzuki 5904</emphasis>
(TAI); Taipei, Tatungshan, 
<geoCoordinate id="D3BF197C6FF37D7A35CAA524BE0A83E8" degrees="25" direction="north" minutes="10" orientation="latitude" precision="15" seconds="22" value="25.172777">25°10'22&quot;N</geoCoordinate>
, 
<geoCoordinate id="7C8515F4906B7192349AE2E9335ED8D7" degrees="121" direction="east" minutes="31" orientation="longitude" precision="15" seconds="33" value="121.52583">121°31'33&quot;E</geoCoordinate>
, 30 December 1929, 
<emphasis id="5D43673AE8C0F3A2844C60B7D61EE029" italics="true" pageId="0" pageNumber="137">Y. Simada 1743C</emphasis>
(TAI); Taipei, Wantan, 
<geoCoordinate id="1BF7B03D8B1AE7CC750C3EF761BCBEAF" degrees="24" direction="north" minutes="56" orientation="latitude" precision="15" seconds="39" value="24.944166">24°56'39&quot;N</geoCoordinate>
, 
<geoCoordinate id="E6B0BC787E545ED1F51A253695B2F627" degrees="121" direction="east" minutes="31" orientation="longitude" precision="15" seconds="49" value="121.53028">121°31'49&quot;E</geoCoordinate>
, 21 March 1949, 
<emphasis id="63CA2373FE113D3B1BFA783A470BDA80" italics="true" pageId="0" pageNumber="137">H. Keng 1008</emphasis>
(TAI); Taipei, Chungho Shih: Yuan-Tung-Ssu, at an elevation of 50 m, 6 October 1989, 
<emphasis id="A7B043723839CB2020ACC8665E7B4D46" italics="true" pageId="0" pageNumber="137">C.H. Lin 258</emphasis>
(HAST); Taipei, Linkou Hsien, Hou-hu, roadside, at an elevation of 100-200 m, 23 September 2000, 
<emphasis id="42F03453223E49D1C6B2D5E29C6D6349" italics="true" pageId="0" pageNumber="137">C.L. Huang &amp; H.M. Chang 134</emphasis>
(HAST); Taitung Hsien, Hungtou river, Lanyu, 
<geoCoordinate id="18056EB6B1B1895A8F7990AD92055A06" degrees="22" direction="north" minutes="1" orientation="latitude" precision="15" seconds="49" value="22.030277">22°1'49&quot;N</geoCoordinate>
, 
<geoCoordinate id="FD28073503D52DEDD70D3797DA0CAFA6" degrees="121" direction="east" minutes="33" orientation="longitude" precision="15" seconds="13" value="121.55361">121°33'13&quot;E</geoCoordinate>
, 
<emphasis id="7367D55C12FD5D9BA4F731A184FCE0E1" italics="true" pageId="0" pageNumber="137">T. Hosokawa 8048</emphasis>
(TAI); Taitung Hsien, Lanyu Hsiang, Bridge Chungaichiao, roadside, 4 December 1996, 
<emphasis id="75A9ED4D3D7E254B77A601F97A655656" italics="true" pageId="0" pageNumber="137">T.Y.A. Yang et al. 07749</emphasis>
(KUN); Taitung Hsien, Lanyu Hsiang, Langtao, Pond Hsiaotienchih, at an elevation of 180 m, roadside, 18 December 1997, 
<emphasis id="756EFFE73814343C058B543CAF45B123" italics="true" pageId="0" pageNumber="137">T.Y.A. Yang et al. 09881</emphasis>
(IBSC); Taitung Hsien, Lanyu Hsiang, Langtao, Pond Hsiaotienchih, at an elevation of 150-180 m, 9 July 1997, 
<emphasis id="DF8DE25740DB3790ED3DD51C02B91B79" italics="true" pageId="0" pageNumber="137">T.Y.A. Yang et al. 08598</emphasis>
(IBSC); Taitung, Lanyu, Orchid Is., 
<geoCoordinate id="6A1A213D51AB36AA04DCCCAB69982579" degrees="22" direction="north" minutes="3" orientation="latitude" precision="15" seconds="23" value="22.056389">22°3'23&quot;N</geoCoordinate>
, 
<geoCoordinate id="32DA059556BFD540A5EB9B1E72196918" degrees="121" direction="east" minutes="30" orientation="longitude" precision="15" seconds="52" value="121.51444">121°30'52&quot;E</geoCoordinate>
, 
<emphasis id="39255A3D0E28B044E3F235E8B5696786" italics="true" pageId="0" pageNumber="137">T.C Huang et al. 10552</emphasis>
(TAI).
</paragraph>
</subSubSection>
</treatment>
</document>