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<document ID-DOI="http://dx.doi.org/10.3897/jhr.70.34206" ID-GBIF-Dataset="2be0b95e-7f6b-4817-b733-c6e2a4ebd224" ID-Pensoft-Pub="1314-2607-70-41" ID-Pensoft-UUID="043FFFE5D662FFB1FF8BFF88FF94FF84" ID-ZBK="B8F3FB3E723749E79AA71AAC9EB4C787" ID-Zenodo-Dep="3269963" ID-ZooBank="B8F3FB3E723749E79AA71AAC9EB4C787" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1314-2607-70-41" ModsDocOrigin="Journal of Hymenoptera Research 70" ModsDocTitle="Revision of the south Asian amisegine genus Cladobethylus Kieffer, 1922 (Hymenoptera, Chrysididae, Amiseginae)" checkinTime="1561791832741" checkinUser="pensoft" docAuthor="Kimsey, Lynn S." docDate="2019" docId="DB0087DF602FDFE044ACF3168EEF1A82" docLanguage="en" docName="JourHymenoptRes 70: 41-64" docOrigin="Journal of Hymenoptera Research 70" docPubDate="2019-06-28" docSource="http://dx.doi.org/10.3897/jhr.70.34206" docTitle="Cladobethylus gilbus Kimsey 1986" docType="treatment" docVersion="4" id="043FFFE5D662FFB1FF8BFF88FF94FF84" lastPageNumber="41" masterDocId="043FFFE5D662FFB1FF8BFF88FF94FF84" masterDocTitle="Revision of the south Asian amisegine genus Cladobethylus Kieffer, 1922 (Hymenoptera, Chrysididae, Amiseginae)" masterLastPageNumber="64" masterPageNumber="41" pageNumber="41" updateTime="1678751192564" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the south Asian amisegine genus Cladobethylus Kieffer, 1922 (Hymenoptera, Chrysididae, Amiseginae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kimsey, Lynn S.</mods:namePart>
<mods:affiliation>Bohart Museum of Entomology, Department of Entomology and Nematology, University of California, Davis, California, 95616, USA</mods:affiliation>
<mods:nameIdentifier type="email">lskimsey@ucdavis.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Journal of Hymenoptera Research</mods:title>
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<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2019-06-28</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>70</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>41</mods:start>
<mods:end>64</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/jhr.70.34206</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.70.34206</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2607-70-41</mods:identifier>
<mods:identifier type="ZooBank">B8F3FB3E723749E79AA71AAC9EB4C787</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3269963</mods:identifier>
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<treatment ID-GBIF-Taxon="158471889" LSID="urn:lsid:plazi:treatment:DB0087DF602FDFE044ACF3168EEF1A82" httpUri="http://treatment.plazi.org/id/DB0087DF602FDFE044ACF3168EEF1A82" lastPageNumber="41" pageId="0" pageNumber="41">
<subSubSection pageId="0" pageNumber="41" type="nomenclature">
<paragraph pageId="0" pageNumber="41">
<taxonomicName ID-CoL="VNLF" LSID="0D0AAE8B-98ED-5450-B7FF-C15EEEB159B3" authority="Kimsey" authorityName="Kimsey" authorityYear="1986" class="Insecta" family="Chrysididae" genus="Cladobethylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cladobethylus gilbus" order="Hymenoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="gilbus">Cladobethylus gilbus Kimsey</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="description">
<paragraph pageId="0" pageNumber="41">
<figureCitation captionStart="Figures 2025" captionStartId="F4" captionText="Figures 20 - 25. Cladobethylus gilbus 20 graph of seasonal abundance of males and females in Malaysia 21, 22 males 23 - 25 females 21, 23 lateral view 22, 24 dorsal view 25 ventral view of head." figureDoi="10.3897/jhr.70.34206.figures20-25" httpUri="https://binary.pensoft.net/fig/313611" pageId="0" pageNumber="41">Figs 20-25</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="reference_group">
<paragraph pageId="0" pageNumber="41">
<taxonomicName authorityName="Kimsey" authorityYear="1986" class="Insecta" family="Chrysididae" genus="Cladobethylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cladobethylus gilbus" order="Hymenoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="gilbus">Cladobethylus gilbus</taxonomicName>
<bibRefCitation DOI="https://doi.org/10.1155/1986/31631" author="Kimsey, LS" journalOrPublisher="Psyche" pageId="0" pageNumber="41" pagination="153 - 165" refId="B2" refString="Kimsey, LS, 1986. New species and genera of Asian Amiseginae. Psyche 93: 153 - 165, DOI: https://doi.org/10.1155/1986/31631" title="New species and genera of Asian Amiseginae." url="https://doi.org/10.1155/1986/31631" volume="93" year="1986">Kimsey 1986</bibRefCitation>
:157. Holotype male; Malaysia: Pasoh Forest Res., Negri Sembilan (AEI).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="diagnosis">
<paragraph pageId="0" pageNumber="41">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="41">
<taxonomicName authorityName="Kimsey" authorityYear="1986" class="Insecta" family="Chrysididae" genus="Cladobethylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cladobethylus gilbus" order="Hymenoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="gilbus">
<emphasis italics="true" pageId="0" pageNumber="41">Cladobethylus gilbus</emphasis>
</taxonomicName>
shares the nearly impunctate second metasomal tergum with
<taxonomicName lsidName="Cl. darlingi" pageId="0" pageNumber="41" rank="species" species="darlingi">
<emphasis italics="true" pageId="0" pageNumber="41">Cl. darlingi</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="Cl. acehensis" pageId="0" pageNumber="41" rank="species" species="acehensis">
<emphasis italics="true" pageId="0" pageNumber="41">Cl. acehensis</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="Cl. densepunctatus" pageId="0" pageNumber="41" rank="species" species="densepunctatus">
<emphasis italics="true" pageId="0" pageNumber="41">Cl. densepunctatus</emphasis>
</taxonomicName>
. In males it can be distinguished from those species by the narrow malar space (less than 2.8 midocellar diameters), flagellomere I less than 4
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad, flagellomere II more than 2.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad, flagellomere XI 7
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad and postocular distance less than 1 MOD. Females have the basal three flagellar segments paler than the rest of the flagellum, very short flagellomeres I and II and malar space 4.5 midocellar diameters wide (shared with
<taxonomicName lsidName="Cl. aquilus" pageId="0" pageNumber="41" rank="species" species="aquilus">
<emphasis italics="true" pageId="0" pageNumber="41">Cl. aquilus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="description">
<paragraph pageId="0" pageNumber="41">Male description.</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis italics="true" pageId="0" pageNumber="41">Body</emphasis>
(Figs
<figureCitation captionStart="Figures 2025" captionStartId="F4" captionText="Figures 20 - 25. Cladobethylus gilbus 20 graph of seasonal abundance of males and females in Malaysia 21, 22 males 23 - 25 females 21, 23 lateral view 22, 24 dorsal view 25 ventral view of head." figureDoi="10.3897/jhr.70.34206.figures20-25" httpUri="https://binary.pensoft.net/fig/313611" pageId="0" pageNumber="41">21</figureCitation>
,
<figureCitation captionStart="Figures 2025" captionStartId="F4" captionText="Figures 20 - 25. Cladobethylus gilbus 20 graph of seasonal abundance of males and females in Malaysia 21, 22 males 23 - 25 females 21, 23 lateral view 22, 24 dorsal view 25 ventral view of head." figureDoi="10.3897/jhr.70.34206.figures20-25" httpUri="https://binary.pensoft.net/fig/313611" pageId="0" pageNumber="41">22</figureCitation>
). Length 3.5-4.0 mm.
<emphasis italics="true" pageId="0" pageNumber="41">Head</emphasis>
. Scapal basin with narrow band of cross-ridges separated by smooth medial band; head venter without ovoid foveae along midline of gular area, gular area punctate; malar space 2.6 MOD long; head as wide as long; interocular distance 1.5
<normalizedToken originalValue="×">x</normalizedToken>
eye width in front view; midocellus 1.4 MOD from ocular margin; ocelli arranged in a nearly equilateral triangle; hindocelli separated from ocular margin by 1 diameter; subantennal distance 0.6 MOD; clypeal apex truncate to slightly concave; flagellomere I 3.6
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad, setae length 0.4
<normalizedToken originalValue="×">x</normalizedToken>
flagellomere breadth; flagellomere II 2.6
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad; flagellomere IX 7
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad; flagellomeres VIII-X slightly lobate submedially.
<emphasis italics="true" pageId="0" pageNumber="41">Mesosoma</emphasis>
. Pronotum 0.9
<normalizedToken originalValue="×">x</normalizedToken>
as long as scutum; mesopleural punctures 0.2-0.5 puncture diameters apart; metapleuron with zone of cross ridging below hindwing base; propodeum laterally cross-ridged, posteromedially with subrectangular enclosure.
<emphasis italics="true" pageId="0" pageNumber="41">Metasoma</emphasis>
: terga I and II impunctate; terga III-V with basal band of dense, minute, contiguous punctures.
<emphasis italics="true" pageId="0" pageNumber="41">Color</emphasis>
: body black, except pronotum, scutum, scutellum and metanotal triangle with metallic blue highlights; antenna and legs including coxae yellow; wing membrane light brown tinted, with brown veins.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="description">
<paragraph pageId="0" pageNumber="41">Female description.</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis italics="true" pageId="0" pageNumber="41">Body</emphasis>
(Figs
<figureCitation captionStart="Figures 2025" captionStartId="F4" captionText="Figures 20 - 25. Cladobethylus gilbus 20 graph of seasonal abundance of males and females in Malaysia 21, 22 males 23 - 25 females 21, 23 lateral view 22, 24 dorsal view 25 ventral view of head." figureDoi="10.3897/jhr.70.34206.figures20-25" httpUri="https://binary.pensoft.net/fig/313611" pageId="0" pageNumber="41">23</figureCitation>
,
<figureCitation captionStart="Figures 2025" captionStartId="F4" captionText="Figures 20 - 25. Cladobethylus gilbus 20 graph of seasonal abundance of males and females in Malaysia 21, 22 males 23 - 25 females 21, 23 lateral view 22, 24 dorsal view 25 ventral view of head." figureDoi="10.3897/jhr.70.34206.figures20-25" httpUri="https://binary.pensoft.net/fig/313611" pageId="0" pageNumber="41">24</figureCitation>
). Length. 4.0-5.0 mm.
<emphasis italics="true" pageId="0" pageNumber="41">Head</emphasis>
. Frons punctures contiguous to 0.5 puncture diameters apart; scapal basin primarily smooth with short strip of cross-ridges on either side of broad medial stripe; head venter with ovoid fovea on either side of gular midline (Fig.
<figureCitation captionStart="Figures 2025" captionStartId="F4" captionText="Figures 20 - 25. Cladobethylus gilbus 20 graph of seasonal abundance of males and females in Malaysia 21, 22 males 23 - 25 females 21, 23 lateral view 22, 24 dorsal view 25 ventral view of head." figureDoi="10.3897/jhr.70.34206.figures20-25" httpUri="https://binary.pensoft.net/fig/313611" pageId="0" pageNumber="41">25</figureCitation>
), gular area punctate; clypeus subtruncate apically; subantennal distance 0.8 midocellar diameter; malar space 4.5 midocellar diameters long; head about as long as wide; midocellus 2
<normalizedToken originalValue="×">x</normalizedToken>
midocellar diameters from ocular margin; ocelli arranged in nearly equilateral triangle; hindocelli separated from ocular margin by one diameter; flagellomere I as long as broad; flagellomere II 0.6
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad; flagellomere X twice as long as broad.
<emphasis italics="true" pageId="0" pageNumber="41">Mesosoma</emphasis>
. Pronotum about as long as scutum; mesopleuron with irregular scrobal sulcus, punctures contiguous to 0.5 puncture diameter apart; metapleuron smooth and impunctate; propodeum laterally polished, impunctate, posteromedially with subrectangular enclosure smooth but somewhat irregular.
<emphasis italics="true" pageId="0" pageNumber="41">Metasoma</emphasis>
. Tergum I impunctate; tergum II punctures 10 puncture diameters apart becoming slightly denser laterally; terga II-IV punctures 1-2 puncture diameters apart with impunctate rim.
<emphasis italics="true" pageId="0" pageNumber="41">Color</emphasis>
. Body black, except head and pronotum with faint metallic blue highlights; legs including coxae yellow; scape yellowish brown becoming darker ventrally; pedicel and flagellomeres I-III yellow, remaining flagellomeres brown dorsally becoming yellow ventrally; wing membrane brown-tinted, veins dark brown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="distribution">
<paragraph pageId="0" pageNumber="41">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="41">
MALAYSIA:
<emphasis bold="true" pageId="0" pageNumber="41">Negeri Sembilan</emphasis>
, Pasoh Forest Res.; 106 males and 50 females were examined (AEI, BME). Males and females were found in nearly every month over a two-year period of Malaise trap collecting at this site (Fig.
<figureCitation captionStart="Figures 2025" captionStartId="F4" captionText="Figures 20 - 25. Cladobethylus gilbus 20 graph of seasonal abundance of males and females in Malaysia 21, 22 males 23 - 25 females 21, 23 lateral view 22, 24 dorsal view 25 ventral view of head." figureDoi="10.3897/jhr.70.34206.figures20-25" httpUri="https://binary.pensoft.net/fig/313611" pageId="0" pageNumber="41">20</figureCitation>
).
</paragraph>
<caption ID-Zenodo-Dep="3270347" doi="10.3897/jhr.70.34206.figures20-25" httpUri="https://binary.pensoft.net/fig/313611" pageId="0" pageNumber="41" start="Figures 2025" startId="F4">
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Figures 20-25.</emphasis>
<taxonomicName authorityName="Kimsey" authorityYear="1986" class="Insecta" family="Chrysididae" genus="Cladobethylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cladobethylus gilbus" order="Hymenoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="gilbus">
<emphasis italics="true" pageId="0" pageNumber="41">Cladobethylus gilbus</emphasis>
</taxonomicName>
20 graph of seasonal abundance of males and females in Malaysia
<emphasis bold="true" pageId="0" pageNumber="41">21, 22</emphasis>
males
<emphasis bold="true" pageId="0" pageNumber="41">23-25</emphasis>
females
<emphasis bold="true" pageId="0" pageNumber="41">21, 23</emphasis>
lateral view
<emphasis bold="true" pageId="0" pageNumber="41">22, 24</emphasis>
dorsal view
<emphasis bold="true" pageId="0" pageNumber="41">25</emphasis>
ventral view of head.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>