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<document id="65F4EA3C4C60AD7DAFA17A5B7CE9E8DF" ID-DOI="10.1080/002229301300323910" ID-ISSN="1464-5262" ID-Zenodo-Dep="5277129" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630022732653" checkinUser="felipe" docAuthor="Gussmann, S. M. V. &amp; Scholtz, C. H." docDate="2001" docId="DA651E6CFFEFFB46FE4A32E6FF04FA53" docLanguage="en" docName="JNatHist.35.7.1013-1084.pdf" docOrigin="Journal of Natural History 35 (7)" docSource="http://www.tandfonline.com/doi/abs/10.1080/002229301300323910" docStyle="DocumentStyle:B6EF65DE5B4D2965D4D236BA736D172F.2:JNatHist.2000-2002.journal_article.2cover" docStyleId="B6EF65DE5B4D2965D4D236BA736D172F" docStyleName="JNatHist.2000-2002.journal_article.2cover" docStyleVersion="2" docTitle="Bolbaoeer namibiensis Gussmann &amp; Scholtz 2001, sp. nov." docType="treatment" docVersion="4" lastPageNumber="1079" masterDocId="265C6614FFADFB02FFDA3151FFDBFFCD" masterDocTitle="Systematic revision of the Afrotropical genus BolbaOEer Vulcano, Martinez and Pereira (Coleoptera: Scarabaeoidea: Bolboceratidae: Bolboceratinae), with descriptions of eight new species" masterLastPageNumber="1084" masterPageNumber="1013" pageNumber="1077" updateTime="1699137266390" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="FE3C1D4D363F3A4C092F5CD3AB849E16">
<mods:title id="0CD241A27FF833AC27F603D90109EF4D">Systematic revision of the Afrotropical genus BolbaOEer Vulcano, Martinez and Pereira (Coleoptera: Scarabaeoidea: Bolboceratidae: Bolboceratinae), with descriptions of eight new species</mods:title>
</mods:titleInfo>
<mods:name id="1D7958206C815B6262FC37A54F7A2E5A" type="personal">
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<mods:namePart id="1383D5B6F0E6BC9045B9C8729D170194">Gussmann, S. M. V.</mods:namePart>
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<mods:namePart id="AE423F290F77D2D37C64122401762794">Scholtz, C. H.</mods:namePart>
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<mods:title id="6190F5B40B4F25AD15CDFDA27516AC7C">Journal of Natural History</mods:title>
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<mods:part id="5C4D4B41FA3A552CDFCF5618B0DFD2DE">
<mods:date id="EC75017CF9536E0A4C7334033BADA9D2">2001</mods:date>
<mods:detail id="106413FACFD2F1908B4788E68D8606ED" type="pubDate">
<mods:number id="4A0241F0A6F72FD80E86AEE872861F0E">2001-07-31</mods:number>
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<mods:detail id="26465407A9514AB6D60D895434B62636" type="volume">
<mods:number id="0176D498D6BA4FFCD5E589A10540CCF4">35</mods:number>
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<mods:number id="7048019013F74AA26087D799EBB343D6">7</mods:number>
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<mods:start id="F1FE7DA968E393069D66AE6F8791AF31">1013</mods:start>
<mods:end id="2F4FBAB0B383F4C01E95D5C87444B469">1084</mods:end>
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<mods:classification id="A5EB6CC069765C44EB7942388A84790E">journal article</mods:classification>
<mods:identifier id="650572844F3850366092E2786894C1E0" type="DOI">10.1080/002229301300323910</mods:identifier>
<mods:identifier id="48D853FB8EB8F9947649AF654720F742" type="ISSN">1464-5262</mods:identifier>
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<treatment id="DA651E6CFFEFFB46FE4A32E6FF04FA53" ID-DOI="http://doi.org/10.5281/zenodo.5279298" ID-Zenodo-Dep="5279298" LSID="urn:lsid:plazi:treatment:DA651E6CFFEFFB46FE4A32E6FF04FA53" httpUri="http://treatment.plazi.org/id/DA651E6CFFEFFB46FE4A32E6FF04FA53" lastPageId="68" lastPageNumber="1079" pageId="66" pageNumber="1077">
<subSubSection id="1AD6FCF1FFEFFB40FE4A32E6FD64FC22" pageId="66" pageNumber="1077" type="nomenclature">
<paragraph id="5273AF7AFFEFFB40FE4A32E6FD39FC02" blockId="66.[400,738,951,1008]" box="[400,738,951,975]" pageId="66" pageNumber="1077">
<heading id="093B1816FFEFFB40FE4A32E6FD39FC02" box="[400,738,951,975]" centered="true" fontSize="10" level="2" pageId="66" pageNumber="1077" reason="3">
<taxonomicName id="95CCD4F9FFEFFB40FE4A32E6FDA5FC02" authority="Gussmann &amp; Scholtz, 2001" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[400,638,951,975]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1077" phylum="Arthropoda" rank="species" species="namibiensis" status="sp. nov.">
<emphasis id="60B87368FFEFFB40FE4A32E6FDA5FC02" bold="true" box="[400,638,951,975]" italics="true" pageId="66" pageNumber="1077">BolbaOEer namibiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7B8BCE13FFEFFB40FD5232E9FD39FC02" box="[648,738,952,975]" pageId="66" pageNumber="1077" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="5273AF7AFFEFFB40FE693286FD64FC22" blockId="66.[400,738,951,1008]" box="[435,703,983,1008]" pageId="66" pageNumber="1077">(®gures 4r±t, 23i±p, 25)</paragraph>
</subSubSection>
<subSubSection id="1AD6FCF1FFEFFB46FF863573FF04FA53" lastPageId="68" lastPageNumber="1079" pageId="66" pageNumber="1077" type="description">
<paragraph id="5273AF7AFFEFFB40FF863573FEC3FBF4" blockId="66.[92,1047,1057,1686]" box="[92,280,1057,1081]" pageId="66" pageNumber="1077">
<emphasis id="60B87368FFEFFB40FF863573FEC3FBF4" box="[92,280,1057,1081]" italics="true" pageId="66" pageNumber="1077">Description male</emphasis>
</paragraph>
<paragraph id="5273AF7AFFEFFB41FF5E3510FEA4FE76" blockId="66.[92,1047,1057,1686]" lastBlockId="67.[92,1047,132,443]" lastPageId="67" lastPageNumber="1078" pageId="66" pageNumber="1077">
<emphasis id="60B87368FFEFFB40FF5E3510FED7FB94" box="[132,268,1089,1113]" italics="true" pageId="66" pageNumber="1077">Body length</emphasis>
<quantity id="9534029FFFEFFB40FEC23510FE6EFB94" box="[280,437,1089,1113]" metricMagnitude="-2" metricUnit="m" metricValue="2.23" metricValueMax="2.38" metricValueMin="2.08" pageId="66" pageNumber="1077" unit="mm" value="22.3" valueMax="23.8" valueMin="20.8">20.8±23.8 mm</quantity>
(
<specimenCount id="44CA64F3FFEFFB40FE173510FD82FB94" box="[461,601,1089,1113]" pageId="66" pageNumber="1077" type="generic">5 specimens</specimenCount>
).
<emphasis id="60B87368FFEFFB40FDAF3510FD1BFB94" box="[629,704,1089,1113]" italics="true" pageId="66" pageNumber="1077">Colour</emphasis>
light brown to dark brown, head and pronotum usually slightly darker.
<emphasis id="60B87368FFEFFB40FDA23531FC5CFBB5" box="[632,903,1120,1144]" italics="true" pageId="66" pageNumber="1077">Outer margin mandible</emphasis>
very feebly sinuate.
<emphasis id="60B87368FFEFFB40FF6435D1FECDFB55" box="[190,278,1152,1176]" italics="true" pageId="66" pageNumber="1077">Clypeus</emphasis>
with bituberculate, straight to feebly arcuate transverse posterior carina; tubercles in line with clypeo-frontal transition; posterior carina more or less as long as anterior carina, sometimes obsolete between tubercles.
<emphasis id="60B87368FFEFFB40FC883590FC4BFB15" box="[850,912,1217,1240]" italics="true" pageId="66" pageNumber="1077">Frons</emphasis>
very feebly depressed posterior to posterior clypeal carina.
<emphasis id="60B87368FFEFFB40FDA435B1FCCCFB35" box="[638,791,1248,1272]" italics="true" pageId="66" pageNumber="1077">Antennal club</emphasis>
with glabrous area of basal segment about one-third of exposed surface.
<emphasis id="60B87368FFEFFB40FD773450FCC1FAD5" box="[685,794,1281,1304]" italics="true" pageId="66" pageNumber="1077">Pronotum</emphasis>
with anterior margin medially raised and bituberculate
<emphasis id="60B87368FFEFFB40FE053471FE33FAF3" bold="true" box="[479,488,1312,1342]" pageId="66" pageNumber="1077">/</emphasis>
bihorned (®gure 4r, s), in major males in dorsal view slightly projecting over frons; with pair of long, slightly inward curved, outer horns along posterior margin in major males (®gure 4r) or with pair of short, more approximated horns in minor males (®gure 4s) and with transverse swollen noncarinate areas in between (as
<taxonomicName id="95CCD4F9FFEFFB40FE6034F0FDF2FA75" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[442,553,1440,1464]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1077" phylum="Arthropoda" rank="species" species="sasakii">
<emphasis id="60B87368FFEFFB40FE6034F0FDF2FA75" box="[442,553,1440,1464]" italics="true" pageId="66" pageNumber="1077">B. sasakii</emphasis>
</taxonomicName>
, see ®gure 6a); disc with ®ne, well-spaced punctures, these increasingly interspersed by large, often conūent punctures towards sides and along anterior margin of pronotum.
<taxonomicName id="95CCD4F9FFEFFB40FD45348EFCD0FA3A" box="[671,779,1503,1527]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="66" pageNumber="1077" phylum="Arthropoda" rank="genus">
<emphasis id="60B87368FFEFFB40FD45348EFCD0FA3A" box="[671,779,1503,1527]" italics="true" pageId="66" pageNumber="1077">Scutellum</emphasis>
</taxonomicName>
with surface coarsely punctate.
<emphasis id="60B87368FFEFFB40FF1734AEFEFBF9DA" box="[205,288,1535,1559]" italics="true" pageId="66" pageNumber="1077">Elytron</emphasis>
with strial punctures separated by four to six puncture diameters; intervals with approximately ®ve to six punctures between two striae.
<emphasis id="60B87368FFEFFB40FC59374EFC05F9FA" box="[899,990,1567,1591]" italics="true" pageId="66" pageNumber="1077">Protibia</emphasis>
®veto six-dentate.
<emphasis id="60B87368FFEFFB40FED7376EFE70F99A" box="[269,427,1599,1623]" italics="true" pageId="66" pageNumber="1077">Protibial spur</emphasis>
more or less as long as ®fth tarsomere, acuminate.
<emphasis id="60B87368FFEFFB40FF86370EFED3F9BA" box="[92,264,1631,1655]" italics="true" pageId="66" pageNumber="1077">Metatrochanter</emphasis>
of unmodi®ed shape; entire surface with dense, moderately long setation.
<emphasis id="60B87368FFEFFB40FF1C372FFE9AF95B" box="[198,321,1662,1686]" italics="true" pageId="66" pageNumber="1077">Metafemur</emphasis>
in ventral view in posterior third with line of densely spaced, moderately long and long setae; with dense, moderately long and long setae in basal area and basal third along posterior margin; with fairly dense, long setae in anterior third and apical area; remaining areas with sparse, long setation.
<emphasis id="60B87368FFEEFB41FCE73195FC71FF11" box="[829,938,196,220]" italics="true" pageId="67" pageNumber="1078">Metatibia</emphasis>
in lateral view with subapical carina bilobed; with unmodi®ed spurs.
<emphasis id="60B87368FFEEFB41FCDD31B5FCAEFF31" box="[775,885,228,252]" italics="true" pageId="67" pageNumber="1078">Underside</emphasis>
with sternites 4, 5 and 6 strongly modi®ed (®gure 23h±k); processes on sternite 4 and 5 gradually narrowing from broad base to rounded apex (®gure 23k); apex of pygidium sharply incised (®gure 23h); median incision dividing apical ®fth of pygidium; the two halves of pygidium in basal four ®fth fused to a clearly visible suture (stippled lines in ®gure 23h).
<emphasis id="60B87368FFEEFB41FF3C30D2FE90FE56" box="[230,331,387,411]" italics="true" pageId="67" pageNumber="1078">Genitalia</emphasis>
with aedeagus as in ®gure 23l±n; genital capsule apically with moderately long setation.
</paragraph>
<paragraph id="5273AF7AFFEEFB41FF8630B5FEF6FE36" blockId="67.[92,1047,483,1686]" box="[92,301,483,507]" pageId="67" pageNumber="1078">
<emphasis id="60B87368FFEEFB41FF8630B5FEF6FE36" box="[92,301,483,507]" italics="true" pageId="67" pageNumber="1078">Description female</emphasis>
</paragraph>
<paragraph id="5273AF7AFFEEFB41FF5E3352FEECFB15" blockId="67.[92,1047,483,1686]" pageId="67" pageNumber="1078">
<emphasis id="60B87368FFEEFB41FF5E3352FED7FDD6" box="[132,268,515,539]" italics="true" pageId="67" pageNumber="1078">Body length</emphasis>
<quantity id="9534029FFFEEFB41FEC23352FE6EFDD6" box="[280,437,515,539]" metricMagnitude="-2" metricUnit="m" metricValue="2.195" metricValueMax="2.31" metricValueMin="2.08" pageId="67" pageNumber="1078" unit="mm" value="21.95" valueMax="23.1" valueMin="20.8">20.8±23.1 mm</quantity>
(
<specimenCount id="44CA64F3FFEEFB41FE173352FD82FDD6" box="[461,601,515,539]" pageId="67" pageNumber="1078" type="generic">6 specimens</specimenCount>
).
<emphasis id="60B87368FFEEFB41FDAE3352FD64FDD6" box="[628,703,515,539]" italics="true" pageId="67" pageNumber="1078">Colour</emphasis>
as in male.
<emphasis id="60B87368FFEEFB41FC813352FF65FDF6" italics="true" pageId="67" pageNumber="1078">Outer margin of mandible</emphasis>
rounded.
<emphasis id="60B87368FFEEFB41FEED3375FEAEFDF6" box="[311,373,548,571]" italics="true" pageId="67" pageNumber="1078">Frons</emphasis>
with strongly raised bituberculate, transverse carina; position of carina variable, from between eye-canthi to slightly anterior to posterolateral genal angle; carina straight and slightly shorter than anterior clypeal carina.
<emphasis id="60B87368FFEEFB41FF8633D3FF2EFD57" box="[92,245,642,666]" italics="true" pageId="67" pageNumber="1078">Antennal club</emphasis>
as in male.
<emphasis id="60B87368FFEEFB41FE5833D2FE34FD57" box="[386,495,643,666]" italics="true" pageId="67" pageNumber="1078">Pronotum</emphasis>
with anterior margin medially raised, in frontal view slightly bisinuate (as in ®gure
<quantity id="9534029FFFEEFB41FE3333F3FDDAFD77" box="[489,513,674,698]" metricMagnitude="3" metricUnit="kg" metricValue="4.0" pageId="67" pageNumber="1078" unit="t" value="4.0">4t</quantity>
), in dorsal view feebly arcuate; with transverse distinctly carinate swelling at median portion of disc; carina in frontal view as in ®gure
<quantity id="9534029FFFEEFB41FF7933B3FF63FD37" box="[163,184,738,762]" metricMagnitude="3" metricUnit="kg" metricValue="4.0" pageId="67" pageNumber="1078" unit="t" value="4.0">4t</quantity>
, in dorsal view feebly bisinuate; area anterior to carina (shaded area in ®gure
<quantity id="9534029FFFEEFB41FF793253FF63FCD7" box="[163,184,770,794]" metricMagnitude="3" metricUnit="kg" metricValue="4.0" pageId="67" pageNumber="1078" unit="t" value="4.0">4t</quantity>
) with very densely spaced small, deep punctures; on feebly swollen areas lateral to densely punctate area and also along posterior margin with densely spaced large punctures; otherwise with very densely spaced, partially conūent, large punctures.
<emphasis id="60B87368FFEEFB41FF723233FE73FCB7" box="[168,424,866,890]" italics="true" pageId="67" pageNumber="1078">
<taxonomicName id="95CCD4F9FFEEFB41FF723233FECEFCB7" box="[168,277,866,890]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="67" pageNumber="1078" phylum="Arthropoda" rank="genus">Scutellum</taxonomicName>
and elytron
</emphasis>
as in male.
<emphasis id="60B87368FFEEFB41FD903233FC59FCB7" box="[586,898,866,890]" italics="true" pageId="67" pageNumber="1078">Protibia and protibial spur</emphasis>
as in male.
<emphasis id="60B87368FFEEFB41FF8632D3FED3FC57" box="[92,264,898,922]" italics="true" pageId="67" pageNumber="1078">Metatrochanter</emphasis>
of unmodi®ed shape; with dense, moderately long setation in basal third and along posterior margin; remaining surface asetose.
<emphasis id="60B87368FFEEFB41FCF832F3FC46FC77" box="[802,925,930,954]" italics="true" pageId="67" pageNumber="1078">Metafemur</emphasis>
in ventral view in posterior third with line of fairly dense, moderately long and long setae; with fairly dense and long setae in anterior third and apical area; with fairly dense, moderately long setae along posterior margin; remaining areas with few isolated setae.
<emphasis id="60B87368FFEEFB41FF7C3570FEC8FBF4" box="[166,275,1057,1081]" italics="true" pageId="67" pageNumber="1078">Metatibia</emphasis>
as in male.
<emphasis id="60B87368FFEEFB41FE713570FDC2FBF4" box="[427,537,1057,1081]" italics="true" pageId="67" pageNumber="1078">Underside</emphasis>
with sternite 5 postero-medially feebly but discernibly swollen (not illustrated); apex of sternite 6 feebly arcuately emarginate (®gure 23p) or narrowly truncate; apex of pygidium sharply triangularly incised (®gure 23o); median incision dividing approximatel y apical seventh of pygidium; the two halves of pygidium in basal six seventh fused to a clearly visible suture (stippled lines in ®gure 23o).
</paragraph>
<paragraph id="5273AF7AFFEEFB41FF5E35B1FDA0FAD5" blockId="67.[92,1047,483,1686]" pageId="67" pageNumber="1078">
<emphasis id="60B87368FFEEFB41FF5E35B1FED3FB35" box="[132,264,1248,1272]" italics="true" pageId="67" pageNumber="1078">Distribution</emphasis>
(®gure 25). Northeastern parts of
<collectingCountry id="2ADBEFEAFFEEFB41FD1235B1FCF0FB35" box="[712,811,1248,1272]" name="Namibia" pageId="67" pageNumber="1078">Namibia</collectingCountry>
, northern parts of
<collectingCountry id="2ADBEFEAFFEEFB41FF863450FF17FAD5" box="[92,204,1281,1304]" name="Botswana" pageId="67" pageNumber="1078">Botswana</collectingCountry>
(`Ngamiland) and southern
<collectingCountry id="2ADBEFEAFFEEFB41FDC63451FDAFFAD5" box="[540,628,1280,1304]" name="Zambia" pageId="67" pageNumber="1078">Zambia</collectingCountry>
.
</paragraph>
<paragraph id="5273AF7AFFEEFB41FF5E3470FEF5F9BA" blockId="67.[92,1047,483,1686]" pageId="67" pageNumber="1078">
<emphasis id="60B87368FFEEFB41FF5E3470FE45FAF5" box="[132,414,1312,1336]" italics="true" pageId="67" pageNumber="1078">Type material examined</emphasis>
.
<typeStatus id="8D7711D8FFEEFB41FE6F3471FDF3FAF5" box="[437,552,1312,1336]" pageId="67" pageNumber="1078" type="holotype">HOLOTYPE</typeStatus>
l[diss.]:`[
<collectingCountry id="2ADBEFEAFFEEFB41FD1B3471FCFFFAF5" box="[705,804,1312,1336]" name="Namibia" pageId="67" pageNumber="1078">Namibia</collectingCountry>
], Abachaus [
<geoCoordinate id="37F8C9BDFFEEFB41FC163471FBCDFAF5" box="[972,1046,1312,1336]" degrees="19.47" direction="south" orientation="latitude" pageId="67" pageNumber="1078" precision="555" value="-19.47">19.47S</geoCoordinate>
<geoCoordinate id="37F8C9BDFFEEFB41FF863411FF71FA9A" box="[92,170,1344,1368]" degrees="16.44" direction="east" orientation="longitude" pageId="67" pageNumber="1078" precision="555" value="16.44">16.44E</geoCoordinate>
], iii.53, G. Hobohm
<emphasis id="60B87368FFEEFB41FE773411FE67FA95" bold="true" box="[429,444,1344,1374]" pageId="67" pageNumber="1078">/`</emphasis>
<taxonomicName id="95CCD4F9FFEEFB41FE673411FD7DFA95" box="[445,678,1344,1368]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="1078" phylum="Arthropoda" rank="species" species="princeps">Bolboceras princeps</taxonomicName>
, det. G. Frey 1960 (TMSA);
<typeStatus id="8D7711D8FFEEFB41FF863431FF1CFAB5" box="[92,199,1376,1400]" pageId="67" pageNumber="1078" type="allotype">ALLOTYPE</typeStatus>
m: same data (TMSA); 5
<typeStatus id="8D7711D8FFEEFB41FDD83431FDAEFAB5" box="[514,629,1376,1400]" pageId="67" pageNumber="1078" type="paratype">PARATYPES</typeStatus>
: 1 l[diss.],`S.W. Protect., Narugas [misprint, Nuragas
<geoCoordinate id="37F8C9BDFFEEFB41FE9A34D1FE51FA55" box="[320,394,1408,1432]" degrees="19.16" direction="south" orientation="latitude" pageId="67" pageNumber="1078" precision="555" value="-19.16">19.16S</geoCoordinate>
<geoCoordinate id="37F8C9BDFFEEFB41FE4C34D1FE3FFA5A" box="[406,484,1408,1432]" degrees="18.52" direction="east" orientation="longitude" pageId="67" pageNumber="1078" precision="555" value="18.52">18.52E</geoCoordinate>
],
<date id="267289BAFFEEFB41FDDB34D1FDB4FA55" box="[513,623,1408,1432]" pageId="67" pageNumber="1078" value="1919-01">Jan. 1919</date>
, R. Lightfoot
<emphasis id="60B87368FFEEFB41FCC734D1FCF6FA55" bold="true" box="[797,813,1408,1438]" pageId="67" pageNumber="1078">/`</emphasis>
<emphasis id="60B87368FFEEFB41FC9C34D1FC8EFA55" bold="true" box="[838,853,1408,1438]" pageId="67" pageNumber="1078">/`</emphasis>
<taxonomicName id="95CCD4F9FFEEFB41FC8F34D1FBDEFA55" authority="Kolbe" authorityName="Kolbe" authorityYear="1894" box="[853,1029,1408,1432]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="1078" phylum="Arthropoda" rank="species" species="princeps">princeps Kolbe</taxonomicName>
? [handwritten by Andreae, ex curator at SAMC] (SAMC); 1, same data but:`Nuragas,`m (SAMC); 1,`
<collectingCountry id="2ADBEFEAFFEEFB41FE603491FDC6FA15" box="[442,541,1472,1496]" name="Namibia" pageId="67" pageNumber="1078">Namibia</collectingCountry>
,
<collectingRegion id="90086198FFEEFB41FDEB3491FD5DFA15" box="[561,646,1472,1496]" country="Namibia" name="Zambezi" pageId="67" pageNumber="1078">Caprivi</collectingRegion>
Nakatwa, Kwando River,
<geoCoordinate id="37F8C9BDFFEEFB41FC163491FBCDFA15" box="[972,1046,1472,1496]" degrees="18.10" direction="south" orientation="latitude" pageId="67" pageNumber="1078" precision="555" value="-18.1">18.10S</geoCoordinate>
<geoCoordinate id="37F8C9BDFFEEFB41FF86348EFF73FA3A" box="[92,168,1503,1527]" degrees="23.25" direction="east" orientation="longitude" pageId="67" pageNumber="1078" precision="555" value="23.25">23.25E</geoCoordinate>
,
<quantity id="9534029FFFEEFB41FF1F348EFED9FA3A" box="[197,258,1503,1527]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="67" pageNumber="1078" unit="m" value="950.0">950m</quantity>
,
<date id="267289BAFFEEFB41FEC5348EFE4AFA3A" box="[287,401,1503,1527]" pageId="67" pageNumber="1078" value="1992-03-09">09.iii.1992</date>
, R. Oberprieler (SANC); 1 l[diss.
<date id="267289BAFFEEFB41FCA6348EFC3CFA3A" box="[892,999,1503,1527]" pageId="67" pageNumber="1078" value="1994-05-18">18.v.1994</date>
by Riaan Stals but no aedeagus found],`S. Africa:
<collectingCountry id="2ADBEFEAFFEEFB41FD0C34AEFCAEF9DA" box="[726,885,1535,1559]" name="Botswana" pageId="67" pageNumber="1078">Bechuanaland</collectingCountry>
. Ngamiland.
<date id="267289BAFFEEFB41FF86374EFF1CF9FA" box="[92,199,1567,1591]" pageId="67" pageNumber="1078" value="1930-11">Nov.1930</date>
±
<date id="267289BAFFEEFB41FF02374EFEE1F9FA" box="[216,314,1567,1591]" pageId="67" pageNumber="1078" value="1931-01">Jan.1931</date>
, G.D. Hale Carpenter, B.M. 1931-160. (BMNH); 1 l[diss.],`S.W. Protectorate
<emphasis id="60B87368FFEEFB41FEE5376EFE94F99A" bold="true" box="[319,335,1599,1629]" pageId="67" pageNumber="1078">/`</emphasis>
l
<emphasis id="60B87368FFEEFB41FEB2376EFEACF99A" bold="true" box="[360,375,1599,1629]" pageId="67" pageNumber="1078">/`</emphasis>
<taxonomicName id="95CCD4F9FFEEFB41FEA2376EFDF5F99A" authority="Kolbe" authorityName="Kolbe" authorityYear="1894" box="[376,558,1599,1623]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="1078" phylum="Arthropoda" rank="species" species="princeps">princeps Kolbe</taxonomicName>
? [handwritten by Andreae, ex curator SAMC] (SAMC).
</paragraph>
<paragraph id="5273AF7AFFEEFB46FF5E372FFD39FE4F" blockId="67.[92,1047,483,1686]" lastBlockId="68.[92,1048,132,1438]" lastPageId="68" lastPageNumber="1079" pageId="67" pageNumber="1078">
<emphasis id="60B87368FFEEFB41FF5E372FFE0AF95B" box="[132,465,1662,1686]" italics="true" pageId="67" pageNumber="1078">Additional material examined</emphasis>
(
<specimenCount id="44CA64F3FFEEFB41FE3D372FFDA9F95B" box="[487,626,1662,1686]" pageId="67" pageNumber="1078" type="generic">4 specimens</specimenCount>
).
<materialsCitation id="E2A4A527FFEEFB46FD56372FFE56FEAC" collectionCode="ZMHB, SAMC" collectorName="D. S. W. Afrika &amp; Dr. Liesegang S. G. &amp; Deutsch S. W. Afrika &amp; Dabegabis &amp; Borchmann S. &amp; Rietfontei &amp; Abend &amp; Licht &amp; K. Borchmann' &amp; Kolbe &amp; Type' &amp; Grootfontein &amp; S. W. Protect. &amp; Andreae" country="Namibia" elevation="1" lastPageId="68" lastPageNumber="1079" latitude="-19.35" location="Rietfontein" longLatPrecision="763" longitude="18.07" pageId="67" pageNumber="1078" specimenCount="1">
<emphasis id="60B87368FFEEFB41FD56372FFD2AF95B" bold="true" box="[652,753,1662,1686]" pageId="67" pageNumber="1078">
<collectingCountry id="2ADBEFEAFFEEFB41FD56372FFD36F95B" box="[652,749,1662,1686]" name="Namibia" pageId="67" pageNumber="1078">Namibia</collectingCountry>
:
</emphasis>
1,`
<collectorName id="FF39CAACFFEEFB41FCEF372FFC06F95B" box="[821,989,1662,1686]" pageId="67" pageNumber="1078">D.S.W. Afrika</collectorName>
,
<collectorName id="FF39CAACFFEEFB46FC35372EFEDEFF51" lastPageId="68" lastPageNumber="1079" pageId="67" pageNumber="1078">Dr. Liesegang S.G.</collectorName>
<emphasis id="60B87368FFE9FB46FED731D5FEC0FF51" bold="true" box="[269,283,132,162]" pageId="68" pageNumber="1079">/`</emphasis>
BolbaOEer` (
<collectionCode id="34DD37BFFFE9FB46FE4431D5FE2FFF51" box="[414,500,132,156]" pageId="68" pageNumber="1079">ZMHB</collectionCode>
);
<quantity id="9534029FFFE9FB46FDD031D5FDFEFF51" box="[522,549,132,156]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="68" pageNumber="1079" unit="m" value="1.0">
<elevation id="D9E14849FFE9FB46FDD031D5FDFEFF51" box="[522,549,132,156]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="68" pageNumber="1079" unit="m" value="1.0">1m</elevation>
</quantity>
,`
<collectorName id="FF39CAACFFE9FB46FDE331D5FCC6FF51" box="[569,797,132,156]" pageId="68" pageNumber="1079">Deutsch S.W.Afrika</collectorName>
,
<location id="5713F9A1FFE9FB46FCF631D5FC77FF51" LSID="urn:lsid:plazi:treatment:DA651E6CFFEFFB46FE4A32E6FF04FA53:5713F9A1FFE9FB46FCF631D5FC77FF51" box="[812,940,132,156]" country="Namibia" latitude="-19.35" longLatPrecision="763" longitude="18.07" name="Rietfontein" pageId="68" pageNumber="1079">Rietfontein</location>
[ambiguous but probably`Fort Rietfontein at
<geoCoordinate id="37F8C9BDFFE9FB46FDC731F4FDB3FF70" box="[541,616,165,189]" degrees="21.52" direction="south" orientation="latitude" pageId="68" pageNumber="1079" precision="555" value="-21.52">21.52S</geoCoordinate>
<geoCoordinate id="37F8C9BDFFE9FB46FDAB31F4FD64FF70" box="[625,703,165,189]" degrees="20.53" direction="east" orientation="longitude" pageId="68" pageNumber="1079" precision="555" value="20.53">20.53E</geoCoordinate>
and not
<collectorName id="FF39CAACFFE9FB46FCF631F4FC7FFF70" box="[812,932,165,189]" pageId="68" pageNumber="1079">Dabegabis</collectorName>
at
<geoCoordinate id="37F8C9BDFFE9FB46FC1631F4FBCDFF70" box="[972,1046,165,189]" degrees="28.15" direction="south" orientation="latitude" pageId="68" pageNumber="1079" precision="555" value="-28.15">28.15S</geoCoordinate>
<geoCoordinate id="37F8C9BDFFE9FB46FF863197FF71FF10" box="[92,170,198,222]" degrees="18.34" direction="east" orientation="longitude" pageId="68" pageNumber="1079" precision="555" value="18.34">18.34E</geoCoordinate>
], iii.97,
<collectorName id="FF39CAACFFE9FB46FECE3197FE67FF13" box="[276,444,198,222]" pageId="68" pageNumber="1079">Borchmann S.</collectorName>
<emphasis id="60B87368FFE9FB46FE1E3197FE17FF29" bold="true" box="[452,460,198,228]" pageId="68" pageNumber="1079">/</emphasis>
`
<collectorName id="FF39CAACFFE9FB46FE093197FD99FF13" box="[467,578,198,222]" pageId="68" pageNumber="1079">Rietfontei</collectorName>
n d. 5.3.97 (am
<collectorName id="FF39CAACFFE9FB46FCD53197FC80FF13" box="[783,859,198,222]" pageId="68" pageNumber="1079">Abend</collectorName>
bei
<collectorName id="FF39CAACFFE9FB46FC4C3197FC09FF13" box="[918,978,198,222]" pageId="68" pageNumber="1079">Licht</collectorName>
) [ill.]
<collectorName id="FF39CAACFFE9FB46FF8631B6FED8FF32" box="[92,259,231,255]" pageId="68" pageNumber="1079">K.Borchmann</collectorName>
[handwritten]
<emphasis id="60B87368FFE9FB46FE7031B6FE68FEC8" bold="true" box="[426,435,231,261]" pageId="68" pageNumber="1079">/</emphasis>
`copridoide s
<collectorName id="FF39CAACFFE9FB46FD9231B6FD4BFF32" box="[584,656,231,255]" pageId="68" pageNumber="1079">Kolbe</collectorName>
* [handwritten on green label by unknown curator]
<emphasis id="60B87368FFE9FB46FEF33056FEE3FED2" bold="true" box="[297,312,263,293]" pageId="68" pageNumber="1079">/`</emphasis>
<collectorName id="FF39CAACFFE9FB46FEE23056FEAEFED2" box="[312,373,263,287]" pageId="68" pageNumber="1079">Type</collectorName>
[printed on red label]
<emphasis id="60B87368FFE9FB46FDA93056FDA7FEE8" bold="true" box="[627,636,263,293]" pageId="68" pageNumber="1079">/</emphasis>
`70732 (
<collectionCode id="34DD37BFFFE9FB46FD053056FCEEFED2" box="[735,821,263,287]" pageId="68" pageNumber="1079">ZMHB</collectionCode>
); 1,`
<collectorName id="FF39CAACFFE9FB46FC5A3056FBCCFED2" box="[896,1047,263,287]" pageId="68" pageNumber="1079">Grootfontein</collectorName>
[
<geoCoordinate id="37F8C9BDFFE9FB46FFBF3079FF74FE8D" box="[101,175,296,320]" degrees="19.35" direction="south" orientation="latitude" pageId="68" pageNumber="1079" precision="555" value="-19.35">19.35S</geoCoordinate>
<geoCoordinate id="37F8C9BDFFE9FB46FF603079FED3FE8D" box="[186,264,296,320]" degrees="18.07" direction="east" orientation="longitude" pageId="68" pageNumber="1079" precision="555" value="18.07">18.07E</geoCoordinate>
],
<collectorName id="FF39CAACFFE9FB46FEFF3079FE1BFE8D" box="[293,448,296,320]" pageId="68" pageNumber="1079">S.W. Protect.</collectorName>
<emphasis id="60B87368FFE9FB46FE123079FE2EFE8B" bold="true" box="[456,501,296,326]" pageId="68" pageNumber="1079">/`m/</emphasis>
`princep s
<collectorName id="FF39CAACFFE9FB46FDBC3079FD77FE8D" box="[614,684,296,320]" pageId="68" pageNumber="1079">Kolbe</collectorName>
? [handwritten by
<collectorName id="FF39CAACFFE9FB46FC563079FC30FE8D" box="[908,1003,296,320]" pageId="68" pageNumber="1079">Andreae</collectorName>
, ex curator
<collectionCode id="34DD37BFFFE9FB46FF663018FED7FEAC" box="[188,268,329,353]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="68" pageNumber="1079" type="Museum">SAMC</collectionCode>
] (
<collectionCode id="34DD37BFFFE9FB46FEF63018FEA4FEAC" box="[300,383,329,353]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="68" pageNumber="1079" type="Museum">SAMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E2A4A527FFE9FB46FE7A3018FD01FE4F" collectingDate="1997-04-07" collectionCode="TMSA" collectorName="WD Haacke, E" country="Zambia" elevation="1" latitude="-17.28" location="Zambia" longLatPrecision="767" longitude="24.56" pageId="68" pageNumber="1079" specimenCount="1">
<emphasis id="60B87368FFE9FB46FE7A3018FE26FEAC" bold="true" box="[416,509,329,353]" pageId="68" pageNumber="1079">
<collectingCountry id="2ADBEFEAFFE9FB46FE7A3018FE23FEAC" box="[416,504,329,353]" name="Zambia" pageId="68" pageNumber="1079">Zambia</collectingCountry>
:
</emphasis>
<quantity id="9534029FFFE9FB46FDD03018FDFDFEAC" box="[522,550,329,353]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="68" pageNumber="1079" unit="m" value="1.0">
<elevation id="D9E14849FFE9FB46FDD03018FDFDFEAC" box="[522,550,329,353]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="68" pageNumber="1079" unit="m" value="1.0">1m</elevation>
</quantity>
,`
<location id="5713F9A1FFE9FB46FD9B3018FD42FEAC" LSID="urn:lsid:plazi:treatment:DA651E6CFFEFFB46FE4A32E6FF04FA53:5713F9A1FFE9FB46FD9B3018FD42FEAC" box="[577,665,329,353]" country="Zambia" latitude="-17.28" longLatPrecision="767" longitude="24.56" name="Zambia" pageId="68" pageNumber="1079">Zambia</location>
,
<quantity id="9534029FFFE9FB46FD743018FD23FEAC" box="[686,760,329,353]" metricMagnitude="3" metricUnit="m" metricValue="1.023" pageId="68" pageNumber="1079" unit="m" value="1023.0">
<elevation id="D9E14849FFE9FB46FD743018FD23FEAC" box="[686,760,329,353]" metricMagnitude="3" metricUnit="m" metricValue="1.023" pageId="68" pageNumber="1079" unit="m" value="1023.0">1023m</elevation>
</quantity>
,
<locationDeviation id="43C1B658FFE9FB46FCD73018FBD6FEAC" box="[781,1037,329,353]" location="Kasangula" pageId="68" pageNumber="1079">
<quantity id="9534029FFFE9FB46FCD73018FC97FEAC" box="[781,844,329,353]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="68" pageNumber="1079" unit="km" value="50.0">50km</quantity>
NW Kasangula
</locationDeviation>
,
<geoCoordinate id="37F8C9BDFFE9FB46FF86303BFF7EFE4F" box="[92,165,362,386]" degrees="17.28" direction="south" orientation="latitude" pageId="68" pageNumber="1079" precision="555" value="-17.28">17.28S</geoCoordinate>
±
<geoCoordinate id="37F8C9BDFFE9FB46FF6C303BFED9FE4F" box="[182,258,362,386]" degrees="24.56" direction="east" orientation="longitude" pageId="68" pageNumber="1079" precision="555" value="24.56">24.56E</geoCoordinate>
,
<date id="267289BAFFE9FB46FECE303BFEABFE4F" box="[276,368,362,386]" pageId="68" pageNumber="1079" value="1997-04-07">
<collectingDate id="36367052FFE9FB46FECE303BFEABFE4F" box="[276,368,362,386]" pageId="68" pageNumber="1079" value="1997-04-07">7.4.1997</collectingDate>
</date>
,
<collectorName id="FF39CAACFFE9FB46FE58303AFDE8FE4F" box="[386,563,362,386]" pageId="68" pageNumber="1079">WD Haacke, E</collectorName>
.Erb (
<collectionCode id="34DD37BFFFE9FB46FD5B303BFD08FE4F" box="[641,723,362,386]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="68" pageNumber="1079">TMSA</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="5273AF7AFFE9FB46FF5E30DAFF1DFE0E" blockId="68.[92,1048,132,1438]" pageId="68" pageNumber="1079">
<emphasis id="60B87368FFE9FB46FF5E30DAFF20FE6E" box="[132,251,395,419]" italics="true" pageId="68" pageNumber="1079">Etymology</emphasis>
. Named after the country where most of the
<typeStatus id="8D7711D8FFE9FB46FC9B30DAFCA9FE6E" box="[833,882,395,419]" pageId="68" pageNumber="1079">type</typeStatus>
material was collected.
</paragraph>
<paragraph id="5273AF7AFFE9FB46FF5E309CFD71FCA3" blockId="68.[92,1048,132,1438]" pageId="68" pageNumber="1079">
<emphasis id="60B87368FFE9FB46FF5E309CFF23FE28" box="[132,248,461,485]" italics="true" pageId="68" pageNumber="1079">Comments</emphasis>
. Two of the male specimens of
<taxonomicName id="95CCD4F9FFE9FB46FDAC309FFCC2FE28" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[630,793,461,485]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="1079" phylum="Arthropoda" rank="species" species="namibiensis">
<emphasis id="60B87368FFE9FB46FDAC309FFCC2FE28" box="[630,793,461,485]" italics="true" pageId="68" pageNumber="1079">B. namibiensis</emphasis>
</taxonomicName>
have previously been confused with
<taxonomicName id="95CCD4F9FFE9FB46FEDE30BFFEA4FDC8" authorityName="Kolbe" authorityYear="1894" box="[260,383,494,517]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="1079" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="60B87368FFE9FB46FEDE30BFFEA4FDC8" box="[260,383,494,517]" italics="true" pageId="68" pageNumber="1079">B. princeps</emphasis>
</taxonomicName>
although the strong modi®cations on sternites 4 to 6 and on the pygidium in male
<taxonomicName id="95CCD4F9FFE9FB46FE48335EFDEDFDEB" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[402,566,526,550]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="1079" phylum="Arthropoda" rank="species" species="namibiensis">
<emphasis id="60B87368FFE9FB46FE48335EFDEDFDEB" box="[402,566,526,550]" italics="true" pageId="68" pageNumber="1079">B. namibiensis</emphasis>
</taxonomicName>
bear no similarity to the comparatively feeble modi®cations in male
<taxonomicName id="95CCD4F9FFE9FB46FE7A3361FDC1FD8A" authorityName="Kolbe" authorityYear="1894" box="[416,538,560,583]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="1079" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="60B87368FFE9FB46FE7A3361FDC1FD8A" box="[416,538,560,583]" italics="true" pageId="68" pageNumber="1079">B. princeps</emphasis>
</taxonomicName>
. At most the dorsal aspects of males of both species show some super®cial resemblance although the pronotal punctation diOEers signi®cantly between the two species (compare descriptions). The modi®cations on sternites 4, 5 and 6 place
<taxonomicName id="95CCD4F9FFE9FB46FE5E33C3FDFEFD64" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[388,549,657,681]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="1079" phylum="Arthropoda" rank="species" species="namibiensis">
<emphasis id="60B87368FFE9FB46FE5E33C3FDFEFD64" box="[388,549,657,681]" italics="true" pageId="68" pageNumber="1079">B. namibiensis</emphasis>
</taxonomicName>
in the
<taxonomicName id="95CCD4F9FFE9FB46FDA033C3FCCCFD64" box="[634,791,657,681]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="1079" phylum="Arthropoda" rank="species" species="abyssinicus">
<emphasis id="60B87368FFE9FB46FDA033C3FCCCFD64" box="[634,791,657,681]" italics="true" pageId="68" pageNumber="1079">B. abyssinicus</emphasis>
</taxonomicName>
species-group. This is supported by the females of
<taxonomicName id="95CCD4F9FFE9FB46FE7933E2FD9FFD07" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[419,580,690,714]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="1079" phylum="Arthropoda" rank="species" species="namibiensis">
<emphasis id="60B87368FFE9FB46FE7933E2FD9FFD07" box="[419,580,690,714]" italics="true" pageId="68" pageNumber="1079">B. namibiensis</emphasis>
</taxonomicName>
which have the typical pronotal punctation that is found in most females of the
<taxonomicName id="95CCD4F9FFE9FB46FD923385FD3CFD26" box="[584,743,723,747]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="1079" phylum="Arthropoda" rank="species" species="abyssinicus">
<emphasis id="60B87368FFE9FB46FD923385FD3CFD26" box="[584,743,723,747]" italics="true" pageId="68" pageNumber="1079">B. abyssinicus</emphasis>
</taxonomicName>
species-group. The above male
<emphasis id="60B87368FFE9FB46FF4B33A5FF41FCDF" bold="true" box="[145,154,756,786]" pageId="68" pageNumber="1079">/</emphasis>
female association is certain as two combinations, consisting of a male and female each, were collected at two diOEerent localities. We did, however, decide to include only the females from these combinations into the
<typeStatus id="8D7711D8FFE9FB46FCD13267FCE7FC83" box="[779,828,822,846]" pageId="68" pageNumber="1079">type</typeStatus>
series while listing the remaining single females as additional material.
</paragraph>
<paragraph id="5273AF7AFFE9FB46FF5E3226FDFFFAD7" blockId="68.[92,1048,132,1438]" pageId="68" pageNumber="1079">
Illustrations of the pronotal armature (®gure 4r, s), the sternal and pygidial modi®cations (®gure 23h±k) and illustrations of the aedeagus (®gure 23l±n) for males of
<taxonomicName id="95CCD4F9FFE9FB46FF5A32EBFEFEFC1C" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[128,293,953,977]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="1079" phylum="Arthropoda" rank="species" species="namibiensis">
<emphasis id="60B87368FFE9FB46FF5A32EBFEFEFC1C" box="[128,293,953,977]" italics="true" pageId="68" pageNumber="1079">B. namibiensis</emphasis>
</taxonomicName>
do not need any further explanation as they clearly show the diOEerences to males of other members of the
<taxonomicName id="95CCD4F9FFE9FB46FDBE328AFCDBFC3F" box="[612,768,986,1010]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="1079" phylum="Arthropoda" rank="species" species="abyssinicus">
<emphasis id="60B87368FFE9FB46FDBE328AFCDBFC3F" box="[612,768,986,1010]" italics="true" pageId="68" pageNumber="1079">B. abyssinicus</emphasis>
</taxonomicName>
species-group. At most, males of
<taxonomicName id="95CCD4F9FFE9FB46FF1032ADFEB5FBDE" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[202,366,1019,1043]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="1079" phylum="Arthropoda" rank="species" species="namibiensis">
<emphasis id="60B87368FFE9FB46FF1032ADFEB5FBDE" box="[202,366,1019,1043]" italics="true" pageId="68" pageNumber="1079">B. namibiensis</emphasis>
</taxonomicName>
could be confused with males of
<taxonomicName id="95CCD4F9FFE9FB46FCDF32ADFCAEFBDE" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[773,885,1019,1043]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="1079" phylum="Arthropoda" rank="species" species="sasakii">
<emphasis id="60B87368FFE9FB46FCDF32ADFCAEFBDE" box="[773,885,1019,1043]" italics="true" pageId="68" pageNumber="1079">B. sasakii</emphasis>
</taxonomicName>
. Their dorsal aspects, in particular with respect to the pronotal armature, are virtually identical. The sternal and pygidial modi®cations and the genitalia, however, diOEer between males of the two species (compare ®gure 23h±n with 22i±k). The
<typeStatus id="8D7711D8FFE9FB46FCB1350CFC14FBB8" box="[875,975,1117,1141]" pageId="68" pageNumber="1079" type="holotype">holotype</typeStatus>
of
<taxonomicName id="95CCD4F9FFE9FB46FC24350FFF01FB5B" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="1079" phylum="Arthropoda" rank="species" species="namibiensis">
<emphasis id="60B87368FFE9FB46FC24350FFF01FB5B" italics="true" pageId="68" pageNumber="1079">B. namibiensis</emphasis>
</taxonomicName>
represents a major male. Females of
<taxonomicName id="95CCD4F9FFE9FB46FD40352EFCE6FB5B" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[666,829,1150,1174]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="1079" phylum="Arthropoda" rank="species" species="namibiensis">
<emphasis id="60B87368FFE9FB46FD40352EFCE6FB5B" box="[666,829,1150,1174]" italics="true" pageId="68" pageNumber="1079">B. namibiensis</emphasis>
</taxonomicName>
are best identi®ed by the following modi®cations: apex of sternite 6 feebly arcuately emarginated or narrowly truncate and apex of pygidium with sharp triangular incision. DiOEerences between females of
<taxonomicName id="95CCD4F9FFE9FB46FEE035B3FE02FB34" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[314,473,1249,1273]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="1079" phylum="Arthropoda" rank="species" species="namibiensis">
<emphasis id="60B87368FFE9FB46FEE035B3FE02FB34" box="[314,473,1249,1273]" italics="true" pageId="68" pageNumber="1079">B. namibiensis</emphasis>
</taxonomicName>
and the very similar females of
<taxonomicName id="95CCD4F9FFE9FB46FC9E35B3FBCDFB34" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[836,1046,1249,1273]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="1079" phylum="Arthropoda" rank="species" species="mozambiquensis">
<emphasis id="60B87368FFE9FB46FC9E35B3FBCDFB34" box="[836,1046,1249,1273]" italics="true" pageId="68" pageNumber="1079">B. mozambiquensis</emphasis>
</taxonomicName>
are already explained under the former.
</paragraph>
<paragraph id="5273AF7AFFE9FB46FF5E3472FF04FA53" blockId="68.[92,1048,132,1438]" pageId="68" pageNumber="1079">
The distribution of
<taxonomicName id="95CCD4F9FFE9FB46FE843475FE27FAF6" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[350,508,1315,1339]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="1079" phylum="Arthropoda" rank="species" species="namibiensis">
<emphasis id="60B87368FFE9FB46FE843475FE27FAF6" box="[350,508,1315,1339]" italics="true" pageId="68" pageNumber="1079">B. namibiensis</emphasis>
</taxonomicName>
, based on a small number of specimens, extends from northern
<collectingCountry id="2ADBEFEAFFE9FB46FED13415FEB4FA91" box="[267,367,1348,1372]" name="Namibia" pageId="68" pageNumber="1079">Namibia</collectingCountry>
westwards into northern
<collectingCountry id="2ADBEFEAFFE9FB46FD463414FCD7FA91" box="[668,780,1349,1372]" name="Botswana" pageId="68" pageNumber="1079">Botswana</collectingCountry>
and southern
<collectingCountry id="2ADBEFEAFFE9FB46FC623415FBCBFA91" box="[952,1040,1348,1372]" name="Zambia" pageId="68" pageNumber="1079">Zambia</collectingCountry>
. Thus
<taxonomicName id="95CCD4F9FFE9FB46FF7B3437FE9FFAB0" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[161,324,1381,1405]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="1079" phylum="Arthropoda" rank="species" species="namibiensis">
<emphasis id="60B87368FFE9FB46FF7B3437FE9FFAB0" box="[161,324,1381,1405]" italics="true" pageId="68" pageNumber="1079">B. namibiensis</emphasis>
</taxonomicName>
is sympatric only with populations of the widely distributed
<taxonomicName id="95CCD4F9FFE9FB46FF8634D6FF0CFA53" authorityName="Kolbe" authorityYear="1894" box="[92,215,1415,1438]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="1079" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="60B87368FFE9FB46FF8634D6FF0CFA53" box="[92,215,1415,1438]" italics="true" pageId="68" pageNumber="1079">B. princeps</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>