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<document id="F186F6DB3AF9288FECC52943D298EE62" ID-DOI="10.1080/002229301300323910" ID-ISSN="1464-5262" ID-Zenodo-Dep="5277129" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630022732653" checkinUser="felipe" docAuthor="Gussmann, S. M. V. &amp; Scholtz, C. H." docDate="2001" docId="DA651E6CFF8FFB21FE7A3427FDC3FAA2" docLanguage="en" docName="JNatHist.35.7.1013-1084.pdf" docOrigin="Journal of Natural History 35 (7)" docSource="http://www.tandfonline.com/doi/abs/10.1080/002229301300323910" docStyle="DocumentStyle:B6EF65DE5B4D2965D4D236BA736D172F.2:JNatHist.2000-2002.journal_article.2cover" docStyleId="B6EF65DE5B4D2965D4D236BA736D172F" docStyleName="JNatHist.2000-2002.journal_article.2cover" docStyleVersion="2" docTitle="Bolbaoeer barbatus Gussmann &amp; Scholtz 2001, sp. nov." docType="treatment" docVersion="4" lastPageNumber="1046" masterDocId="265C6614FFADFB02FFDA3151FFDBFFCD" masterDocTitle="Systematic revision of the Afrotropical genus BolbaOEer Vulcano, Martinez and Pereira (Coleoptera: Scarabaeoidea: Bolboceratidae: Bolboceratinae), with descriptions of eight new species" masterLastPageNumber="1084" masterPageNumber="1013" pageNumber="1045" updateTime="1699137266390" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="146653D8E9B11304131C68D833CE7D40">Systematic revision of the Afrotropical genus BolbaOEer Vulcano, Martinez and Pereira (Coleoptera: Scarabaeoidea: Bolboceratidae: Bolboceratinae), with descriptions of eight new species</mods:title>
</mods:titleInfo>
<mods:name id="EECBC6AEBF8A7CEF7396C9ACCC284E34" type="personal">
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<mods:namePart id="B0BF26EA1025A3F29ADDF62AB455944B">Gussmann, S. M. V.</mods:namePart>
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<mods:namePart id="EF29FA931D9C621AD00DA18857E15684">Scholtz, C. H.</mods:namePart>
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<mods:title id="162E59450BC338431E7BE86A00ECC639">Journal of Natural History</mods:title>
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<mods:part id="D980A12CB27A999DCAB08F1127A74A51">
<mods:date id="0082C3CFF17FAC6A47FA431C98C92897">2001</mods:date>
<mods:detail id="4551991822D0D2BBF2333CEAF4329CDE" type="pubDate">
<mods:number id="02C9D38A6CF10F879A7B4C1154FDE8A8">2001-07-31</mods:number>
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<mods:number id="F0E1D0C76BA3887B66DAA8585F22558D">35</mods:number>
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<mods:number id="8DB09C89DB7B14A330CDF42F87A4CE6F">7</mods:number>
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<mods:start id="A24251246A1BBEF1F5BD51B6A8664896">1013</mods:start>
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<mods:classification id="924DEA1AE6FA0426B44F85299BB46665">journal article</mods:classification>
<mods:identifier id="2B5E0205BF2E7EAB005A318B9C011961" type="DOI">10.1080/002229301300323910</mods:identifier>
<mods:identifier id="A6991E62812A4476E40D0F8381EAF902" type="ISSN">1464-5262</mods:identifier>
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<treatment id="DA651E6CFF8FFB21FE7A3427FDC3FAA2" ID-DOI="http://doi.org/10.5281/zenodo.5279272" ID-Zenodo-Dep="5279272" LSID="urn:lsid:plazi:treatment:DA651E6CFF8FFB21FE7A3427FDC3FAA2" httpUri="http://treatment.plazi.org/id/DA651E6CFF8FFB21FE7A3427FDC3FAA2" lastPageId="35" lastPageNumber="1046" pageId="34" pageNumber="1045">
<subSubSection id="1AD6FCF1FF8FFB20FE7A3427FD6EFA63" pageId="34" pageNumber="1045" type="nomenclature">
<paragraph id="5273AF7AFF8FFB20FE7A3427FD09FA43" blockId="34.[416,722,1398,1455]" box="[416,722,1398,1422]" pageId="34" pageNumber="1045">
<heading id="093B1816FF8FFB20FE7A3427FD09FA43" box="[416,722,1398,1422]" centered="true" fontSize="10" level="2" pageId="34" pageNumber="1045" reason="3">
<taxonomicName id="95CCD4F9FF8FFB20FE7A3427FDB6FA43" authority="Gussmann &amp; Scholtz, 2001" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[416,621,1398,1422]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="1045" phylum="Arthropoda" rank="species" species="barbatus" status="sp. nov.">
<emphasis id="60B87368FF8FFB20FE7A3427FDB6FA43" bold="true" box="[416,621,1398,1422]" italics="true" pageId="34" pageNumber="1045">BolbaOEer barbatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7B8BCE13FF8FFB20FDA23426FD09FA43" box="[632,722,1399,1422]" pageId="34" pageNumber="1045" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="5273AF7AFF8FFB20FE6434C7FD6EFA63" blockId="34.[416,722,1398,1455]" box="[446,693,1430,1455]" pageId="34" pageNumber="1045">(®gures 3j, 10a±e, 11)</paragraph>
</subSubSection>
<subSubSection id="1AD6FCF1FF8FFB21FF8634B1FDC3FAA2" lastPageId="35" lastPageNumber="1046" pageId="34" pageNumber="1045" type="description">
<paragraph id="5273AF7AFF8FFB20FF8634B1FEA4FA3A" blockId="34.[92,1047,1503,1686]" box="[92,383,1503,1527]" pageId="34" pageNumber="1045">
<emphasis id="60B87368FF8FFB20FF8634B1FEA4FA3A" box="[92,383,1503,1527]" italics="true" pageId="34" pageNumber="1045">
Description male
<typeStatus id="8D7711D8FF8FFB20FEFB348EFEA4FA3A" box="[289,383,1503,1527]" pageId="34" pageNumber="1045" type="holotype">holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="5273AF7AFF8FFB21FF5E34AEFF67FD1A" blockId="34.[92,1047,1503,1686]" lastBlockId="35.[92,1047,132,1391]" lastPageId="35" lastPageNumber="1046" pageId="34" pageNumber="1045">
<emphasis id="60B87368FF8FFB20FF5E34AEFED7F9DA" box="[132,268,1535,1559]" italics="true" pageId="34" pageNumber="1045">Body length</emphasis>
<quantity id="9534029FFF8FFB20FEC234AEFEA1F9DA" box="[280,378,1535,1559]" metricMagnitude="-2" metricUnit="m" metricValue="1.86" pageId="34" pageNumber="1045" unit="mm" value="18.6">18.6 mm</quantity>
.
<emphasis id="60B87368FF8FFB20FE5034AEFE0EF9DA" box="[394,469,1535,1559]" italics="true" pageId="34" pageNumber="1045">Colour</emphasis>
mid-brown, head and pronotum slightly darker.
<emphasis id="60B87368FF8FFB20FF86374EFEA1F9FA" box="[92,378,1567,1591]" italics="true" pageId="34" pageNumber="1045">Outer margin of mandible</emphasis>
rounded.
<emphasis id="60B87368FF8FFB20FE35374EFD9CF9FA" box="[495,583,1567,1591]" italics="true" pageId="34" pageNumber="1045">Clypeus</emphasis>
with strongly arcuate, transverse posterior carina; carina medially raised to single horn; lateral endpoints of carina anteriad to antennal insertions but not reaching lateral clypeal margins; horn medially connected to anterior clypeal margin with short carina.
<emphasis id="60B87368FF8FFB20FD44372EFD07F95B" box="[670,732,1663,1686]" italics="true" pageId="34" pageNumber="1045">Frons</emphasis>
distinctly concave posterior to posterior clypeal carina.
<emphasis id="60B87368FF8EFB21FE4E31D5FDF7FF51" box="[404,556,132,156]" italics="true" pageId="35" pageNumber="1046">Antennal club</emphasis>
with glabrous area of basal segment about one-third of exposed surface.
<emphasis id="60B87368FF8EFB21FE7531F4FDC7FF71" box="[431,540,165,188]" italics="true" pageId="35" pageNumber="1046">Pronotum</emphasis>
with anterior margin not raised, very feebly curved in dorsal view; with one pair of outer horns along posterior margin and one pair of horns along anterior margin (®gure 3j); disc with large fairly densely spaced punctures, these denser and often conūent towards sides, along posterior and anterior margin of pronotum.
<taxonomicName id="95CCD4F9FF8EFB21FE673072FDF2FEF6" box="[445,553,291,315]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="35" pageNumber="1046" phylum="Arthropoda" rank="genus">
<emphasis id="60B87368FF8EFB21FE673072FDF2FEF6" box="[445,553,291,315]" italics="true" pageId="35" pageNumber="1046">Scutellum</emphasis>
</taxonomicName>
with surface moderately punctate.
<emphasis id="60B87368FF8EFB21FC193072FBCDFEF6" box="[963,1046,291,315]" italics="true" pageId="35" pageNumber="1046">Elytron</emphasis>
with strial punctures separated by four to ®ve puncture diameters; intervals with approximately four to six punctures between two striae.
<emphasis id="60B87368FF8EFB21FCF33033FC5FFEB7" box="[809,900,354,378]" italics="true" pageId="35" pageNumber="1046">Protibia</emphasis>
six-dentate.
<emphasis id="60B87368FF8EFB21FF8630D3FED2FE57" box="[92,265,386,410]" italics="true" pageId="35" pageNumber="1046">Protibial spur</emphasis>
possibly longer than missing ®fth tarsomere, acuminate.
<emphasis id="60B87368FF8EFB21FF8630F3FED3FE77" box="[92,264,418,442]" italics="true" pageId="35" pageNumber="1046">Metatrochanter</emphasis>
of unmodi®ed shape, with very dense and very long setal tuft along posterior margin (®gure 10a).
<emphasis id="60B87368FF8EFB21FE693093FDF5FE17" box="[435,558,450,474]" italics="true" pageId="35" pageNumber="1046">Metafemur</emphasis>
in ventral view in posterior third with line of densely spaced long setae; with dense, long setation in anterior third, in basal area and along posterior margin; remaining areas asetose.
<emphasis id="60B87368FF8EFB21FD243350FCB0FDD4" box="[766,875,513,537]" italics="true" pageId="35" pageNumber="1046">Metatibia</emphasis>
in lateral view with subapical carina feebly bilobed; with unmodi®ed spurs.
<emphasis id="60B87368FF8EFB21FCC93370FC5AFDF4" box="[787,897,545,569]" italics="true" pageId="35" pageNumber="1046">Underside</emphasis>
with sternite 4 unmodi®ed; sternite 5 broad and postero-medially strongly declivous and membranous (see area enclosed by stippled line in ®gure 10b); posterior margin of sternite 6 feebly roundedly incised (®gure 10b); pygidium unmodi®ed.
<emphasis id="60B87368FF8EFB21FCEB33D1FC4DFD55" box="[817,918,640,664]" italics="true" pageId="35" pageNumber="1046">Genitalia</emphasis>
with aedeagus as in ®gure 10c±e, large relative to body size; genital capsule apically with short setation.
</paragraph>
<paragraph id="5273AF7AFF8EFB21FF5E338EFE96FD3A" blockId="35.[92,1047,132,1391]" box="[132,333,735,759]" pageId="35" pageNumber="1046">
<emphasis id="60B87368FF8EFB21FF5E338EFF09FD3A" box="[132,210,735,759]" italics="true" pageId="35" pageNumber="1046">Female</emphasis>
unknown.
</paragraph>
<paragraph id="5273AF7AFF8EFB21FF5E33AFFF6BFCFB" blockId="35.[92,1047,132,1391]" pageId="35" pageNumber="1046">
<emphasis id="60B87368FF8EFB21FF5E33AFFED3FCDB" box="[132,264,766,790]" italics="true" pageId="35" pageNumber="1046">Distribution</emphasis>
(®gure 11). The species is only known from the
<typeStatus id="8D7711D8FF8EFB21FCBF33AEFC4DFCDA" box="[869,918,767,791]" pageId="35" pageNumber="1046">type</typeStatus>
locality in
<collectingCountry id="2ADBEFEAFF8EFB21FF86324FFF77FCFB" box="[92,172,798,822]" name="Ghana" pageId="35" pageNumber="1046">Ghana</collectingCountry>
.
</paragraph>
<paragraph id="5273AF7AFF8EFB21FF5E326EFCC4FCB8" blockId="35.[92,1047,132,1391]" pageId="35" pageNumber="1046">
<emphasis id="60B87368FF8EFB21FF5E326EFE41FC9B" box="[132,410,830,854]" italics="true" pageId="35" pageNumber="1046">Type material examined</emphasis>
.
<materialsCitation id="E2A4A527FF8EFB21FE75326EFCC4FCB8" collectionCode="ZMHB" collectorName="W. Afrika &amp; Graf Zech S. Gr." country="Ghana" latitude="7.48" location="Kete Kratje" longLatPrecision="781" longitude="-0.01" pageId="35" pageNumber="1046" specimenCount="1" typeStatus="holotype">
<typeStatus id="8D7711D8FF8EFB21FE75326EFDFAFC9B" box="[431,545,830,854]" pageId="35" pageNumber="1046" type="holotype">HOLOTYPE</typeStatus>
:`
<collectorName id="FF39CAACFF8EFB21FD87326FFD12FC9B" box="[605,713,830,854]" pageId="35" pageNumber="1046">W.Afrika</collectorName>
,
<location id="5713F9A1FF8EFB21FD07326EFCB1FC9B" LSID="urn:lsid:plazi:treatment:DA651E6CFF8FFB21FE7A3427FDC3FAA2:5713F9A1FF8EFB21FD07326EFCB1FC9B" box="[733,874,830,854]" country="Ghana" latitude="7.48" longLatPrecision="781" longitude="-0.01" name="Kete Kratje" pageId="35" pageNumber="1046">Kete Kratje</location>
[
<collectingCountry id="2ADBEFEAFF8EFB21FCA5326FFC17FC9B" box="[895,972,830,854]" name="Ghana" pageId="35" pageNumber="1046">Ghana</collectingCountry>
, Kete Krachi,
<geoCoordinate id="37F8C9BDFF8EFB21FF61320CFED6FCB8" box="[187,269,861,885]" degrees="07.48" direction="north" orientation="latitude" pageId="35" pageNumber="1046" precision="555" value="7.48">07.48N</geoCoordinate>
<geoCoordinate id="37F8C9BDFF8EFB21FECC320CFEB7FCB8" box="[278,364,861,885]" degrees="00.01" direction="west" orientation="longitude" pageId="35" pageNumber="1046" precision="555" value="-0.01">00.01W</geoCoordinate>
],
<collectorName id="FF39CAACFF8EFB21FE52320CFD9EFCB8" box="[392,581,861,885]" pageId="35" pageNumber="1046">Graf Zech S.Gr.</collectorName>
<emphasis id="60B87368FF8EFB21FD97320CFD87FCB8" bold="true" box="[589,604,861,891]" pageId="35" pageNumber="1046">/`</emphasis>
69727 (
<collectionCode id="34DD37BFFF8EFB21FD60320CFCCBFCB8" box="[698,784,861,885]" pageId="35" pageNumber="1046">ZMHB</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="5273AF7AFF8EFB21FF5E322CFC08FC79" blockId="35.[92,1047,132,1391]" pageId="35" pageNumber="1046">
<emphasis id="60B87368FF8EFB21FF5E322CFF20FC58" box="[132,251,893,917]" italics="true" pageId="35" pageNumber="1046">Etymology</emphasis>
. The term`barbatus, Latin for`bearded, refers to the very dense, long hairtuft on the metatrochanter, a unique character state for this species.
</paragraph>
<paragraph id="5273AF7AFF8EFB21FF5E32EDFDC3FAA2" blockId="35.[92,1047,132,1391]" pageId="35" pageNumber="1046">
<emphasis id="60B87368FF8EFB21FF5E32EDFF23FC19" box="[132,248,956,980]" italics="true" pageId="35" pageNumber="1046">Comments</emphasis>
. The clypeal armature of the
<typeStatus id="8D7711D8FF8EFB21FD8332EDFD66FC19" box="[601,701,956,980]" pageId="35" pageNumber="1046" type="holotype">holotype</typeStatus>
of
<taxonomicName id="95CCD4F9FF8EFB21FD3232ECFCB3FC19" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[744,872,956,980]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="1046" phylum="Arthropoda" rank="species" species="barbatus">
<emphasis id="60B87368FF8EFB21FD3232ECFCB3FC19" box="[744,872,956,980]" italics="true" pageId="35" pageNumber="1046">B. barbatus</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="95CCD4F9FF8EFB21FFA7328CFF02FC39" box="[125,217,989,1012]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="1046" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="60B87368FF8EFB21FFA7328CFF02FC39" box="[125,217,989,1012]" italics="true" pageId="35" pageNumber="1046">B. gigas</emphasis>
</taxonomicName>
: in males of both species the posterior carina is produced into a single horn, a character state not found in any other male
<taxonomicName id="95CCD4F9FF8EFB21FD6132AAFCFAFBDE" box="[699,801,1019,1043]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="1046" phylum="Arthropoda" rank="genus">
<emphasis id="60B87368FF8EFB21FD6132AAFCFAFBDE" box="[699,801,1019,1043]" italics="true" pageId="35" pageNumber="1046">BolbaOEer</emphasis>
</taxonomicName>
. There are, however, signi®cant diOEerences between males of the two species. As in most male
<taxonomicName id="95CCD4F9FF8EFB21FC73354AFBD4FBFE" box="[937,1039,1051,1075]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="1046" phylum="Arthropoda" rank="genus">
<emphasis id="60B87368FF8EFB21FC73354AFBD4FBFE" box="[937,1039,1051,1075]" italics="true" pageId="35" pageNumber="1046">BolbaOEer</emphasis>
</taxonomicName>
, the pronotal protrusions in
<taxonomicName id="95CCD4F9FF8EFB21FE6B356DFDCAFB9E" box="[433,529,1084,1107]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="1046" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="60B87368FF8EFB21FE6B356DFDCAFB9E" box="[433,529,1084,1107]" italics="true" pageId="35" pageNumber="1046">B. gigas</emphasis>
</taxonomicName>
are positioned along the posterior margin whereas
<taxonomicName id="95CCD4F9FF8EFB21FF18350DFE9FFBBE" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[194,324,1115,1139]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="1046" phylum="Arthropoda" rank="species" species="barbatus">
<emphasis id="60B87368FF8EFB21FF18350DFE9FFBBE" box="[194,324,1115,1139]" italics="true" pageId="35" pageNumber="1046">B. barbatus</emphasis>
</taxonomicName>
has an additional pair of horns positioned near the anterior margin, a unique character for the latter species. Apart from that, males of both species diOEer strongly in the shape of the anterior pronotal margin (unmodi®ed in
<taxonomicName id="95CCD4F9FF8EFB21FF8635EBFF06FB1C" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[92,221,1209,1233]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="1046" phylum="Arthropoda" rank="species" species="barbatus">
<emphasis id="60B87368FF8EFB21FF8635EBFF06FB1C" box="[92,221,1209,1233]" italics="true" pageId="35" pageNumber="1046">B. barbatus</emphasis>
</taxonomicName>
but bisinuate and projecting over head in
<taxonomicName id="95CCD4F9FF8EFB21FD0235EBFCEEFB1C" box="[728,821,1210,1233]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="1046" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="60B87368FF8EFB21FD0235EBFCEEFB1C" box="[728,821,1210,1233]" italics="true" pageId="35" pageNumber="1046">B. gigas</emphasis>
</taxonomicName>
), the modi®cations on the abdominal sternites (compare ®gure 10b with
<quantity id="9534029FFF8EFB21FD193588FD31FB3C" box="[707,746,1241,1265]" metricMagnitude="-2" metricUnit="kg" metricValue="1.9" pageId="35" pageNumber="1046" unit="g" value="19.0">19g</quantity>
) and shape and structure of the aedeagi (compare ®gure 10c±e with 19h±j). Lastly the extremely long setal tuft on the metatrochanter of
<taxonomicName id="95CCD4F9FF8EFB21FE1E3448FD9DFAFD" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[452,582,1304,1328]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="1046" phylum="Arthropoda" rank="species" species="barbatus">
<emphasis id="60B87368FF8EFB21FE1E3448FD9DFAFD" box="[452,582,1304,1328]" italics="true" pageId="35" pageNumber="1046">B. barbatus</emphasis>
</taxonomicName>
is not shared by male
<taxonomicName id="95CCD4F9FF8EFB21FCBA3448FC66FAFD" box="[864,957,1305,1328]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="1046" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="60B87368FF8EFB21FCBA3448FC66FAFD" box="[864,957,1305,1328]" italics="true" pageId="35" pageNumber="1046">B. gigas</emphasis>
</taxonomicName>
or any other male
<taxonomicName id="95CCD4F9FF8EFB21FF3E3466FE91FA82" box="[228,330,1335,1359]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="1046" phylum="Arthropoda" rank="genus">
<emphasis id="60B87368FF8EFB21FF3E3466FE91FA82" box="[228,330,1335,1359]" italics="true" pageId="35" pageNumber="1046">BolbaOEer</emphasis>
</taxonomicName>
. Hence we have no doubt that
<taxonomicName id="95CCD4F9FF8EFB21FD1D3469FC92FA82" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[711,841,1335,1359]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="1046" phylum="Arthropoda" rank="species" species="barbatus">
<emphasis id="60B87368FF8EFB21FD1D3469FC92FA82" box="[711,841,1335,1359]" italics="true" pageId="35" pageNumber="1046">B. barbatus</emphasis>
</taxonomicName>
is a new species. The female of this species is unknown.
</paragraph>
</subSubSection>
</treatment>
</document>