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<document id="683935C2C5514E160C8CC5403CBFBCE0" ID-DOI="10.1080/002229301300323910" ID-ISSN="1464-5262" ID-Zenodo-Dep="5277129" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630022732653" checkinUser="felipe" docAuthor="Gussmann, S. M. V. &amp; Scholtz, C. H." docDate="2001" docId="DA651E6CFF8BFB2AFE723506FF38FBD4" docLanguage="en" docName="JNatHist.35.7.1013-1084.pdf" docOrigin="Journal of Natural History 35 (7)" docSource="http://www.tandfonline.com/doi/abs/10.1080/002229301300323910" docStyle="DocumentStyle:B6EF65DE5B4D2965D4D236BA736D172F.2:JNatHist.2000-2002.journal_article.2cover" docStyleId="B6EF65DE5B4D2965D4D236BA736D172F" docStyleName="JNatHist.2000-2002.journal_article.2cover" docStyleVersion="2" docTitle="Bolbaoeer dudleyi Gussmann &amp; Scholtz 2001, sp. nov." docType="treatment" docVersion="3" lastPageNumber="1051" masterDocId="265C6614FFADFB02FFDA3151FFDBFFCD" masterDocTitle="Systematic revision of the Afrotropical genus BolbaOEer Vulcano, Martinez and Pereira (Coleoptera: Scarabaeoidea: Bolboceratidae: Bolboceratinae), with descriptions of eight new species" masterLastPageNumber="1084" masterPageNumber="1013" pageNumber="1049" updateTime="1699137266390" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="40237C39A6768E2CAE99FD308A00EDF9">Systematic revision of the Afrotropical genus BolbaOEer Vulcano, Martinez and Pereira (Coleoptera: Scarabaeoidea: Bolboceratidae: Bolboceratinae), with descriptions of eight new species</mods:title>
</mods:titleInfo>
<mods:name id="76E1AE7295C4736F47FEB044CF78A89D" type="personal">
<mods:role id="44948C4762B6C82B82C639DA059575AE">
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</mods:role>
<mods:namePart id="806C8757356EC697E47284CA2793999B">Gussmann, S. M. V.</mods:namePart>
</mods:name>
<mods:name id="5CB6E0B571832AF87662A637FC7372C7" type="personal">
<mods:role id="45A9F643FEA72C002DBC05CEE744A8A2">
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<mods:namePart id="DA133610468BA72F80A128D9F1203615">Scholtz, C. H.</mods:namePart>
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<mods:typeOfResource id="FFBC96C68D965733E35B0CF6A0CA6A35">text</mods:typeOfResource>
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<mods:title id="8C493CA5F3A85F0289E34AD22E2A4ED4">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="1D9C8266545DAFE4A73CD69B1455369D">
<mods:date id="8FEC10E2C0851EDBB8C46FAFD9AD9F4F">2001</mods:date>
<mods:detail id="AD00C59D8E019BF5AF59DEC60B2330E3" type="pubDate">
<mods:number id="029F5964C9857BC684DF2A077BB49263">2001-07-31</mods:number>
</mods:detail>
<mods:detail id="08E55164ECEA3F64AEF81C1B8CA49904" type="volume">
<mods:number id="AB01C1F972B6FED1768EAC986D1D7966">35</mods:number>
</mods:detail>
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<mods:number id="6811EBF67ADD87E8EDD16D0956FDFF97">7</mods:number>
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<mods:start id="ABF8D3EF693636EF99B7F3120B79BC91">1013</mods:start>
<mods:end id="FF30CBE968278769543A2B3A26BAF333">1084</mods:end>
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</mods:location>
<mods:classification id="B80E9EB5CC7C574B846A0BD512632A0A">journal article</mods:classification>
<mods:identifier id="5008D9E21EC9A0B635612139B8E42A32" type="DOI">10.1080/002229301300323910</mods:identifier>
<mods:identifier id="7683A25ABE7DD312F56CE43F1043F9FF" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="01B4CBF41CAA98906633FFAAD1F2AF83" type="Zenodo-Dep">5277129</mods:identifier>
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<treatment id="DA651E6CFF8BFB2AFE723506FF38FBD4" LSID="urn:lsid:plazi:treatment:DA651E6CFF8BFB2AFE723506FF38FBD4" httpUri="http://treatment.plazi.org/id/DA651E6CFF8BFB2AFE723506FF38FBD4" lastPageId="40" lastPageNumber="1051" pageId="38" pageNumber="1049">
<subSubSection id="1AD6FCF1FF8BFB24FE723506FCDDFB42" pageId="38" pageNumber="1049" type="nomenclature">
<paragraph id="5273AF7AFF8BFB24FE723506FD11FBA2" blockId="38.[424,714,1111,1135]" box="[424,714,1111,1135]" pageId="38" pageNumber="1049">
<heading id="093B1816FF8BFB24FE723506FD11FBA2" box="[424,714,1111,1135]" centered="true" fontSize="10" level="2" pageId="38" pageNumber="1049" reason="2">
<taxonomicName id="95CCD4F9FF8BFB24FE723506FDBDFBA2" authority="Gussmann &amp; Scholtz, 2001" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[424,614,1111,1135]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="1049" phylum="Arthropoda" rank="species" species="dudleyi" status="sp. nov.">
<emphasis id="60B87368FF8BFB24FE723506FDBDFBA2" bold="true" box="[424,614,1111,1135]" italics="true" pageId="38" pageNumber="1049">BolbaOEer dudleyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7B8BCE13FF8BFB24FDB53509FD11FBA2" box="[623,714,1112,1135]" pageId="38" pageNumber="1049" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="5273AF7AFF8BFB24FEB63526FCDDFB42" blockId="38.[364,774,1143,1168]" box="[364,774,1143,1168]" pageId="38" pageNumber="1049">
(®gures 1a, 3k±n, 12h±m,
<quantity id="9534029FFF8BFB24FD4E3526FD67FB42" box="[660,700,1143,1167]" metricMagnitude="-2" metricUnit="kg" metricValue="1.3" pageId="38" pageNumber="1049" unit="g" value="13.0">13g</quantity>
±i, 14)
</paragraph>
</subSubSection>
<subSubSection id="1AD6FCF1FF8BFB2AFF863590FF38FBD4" lastPageId="40" lastPageNumber="1051" pageId="38" pageNumber="1049" type="description">
<paragraph id="5273AF7AFF8BFB24FF863590FEC3FB15" blockId="38.[92,1048,1216,1686]" box="[92,280,1216,1240]" pageId="38" pageNumber="1049">
<emphasis id="60B87368FF8BFB24FF863590FEC3FB15" box="[92,280,1216,1240]" italics="true" pageId="38" pageNumber="1049">Description male</emphasis>
</paragraph>
<paragraph id="5273AF7AFF8BFB25FF5E35B1FCA6FD97" blockId="38.[92,1048,1216,1686]" lastBlockId="39.[92,1047,132,602]" lastPageId="39" lastPageNumber="1050" pageId="38" pageNumber="1049">
<emphasis id="60B87368FF8BFB24FF5E35B1FED7FB35" box="[132,268,1248,1272]" italics="true" pageId="38" pageNumber="1049">Body length</emphasis>
18.6±20.0 mm (
<specimenCount id="44CA64F3FF8BFB24FE1735B1FD82FB35" box="[461,601,1248,1272]" pageId="38" pageNumber="1049" type="generic">6 specimens</specimenCount>
).
<emphasis id="60B87368FF8BFB24FDAF35B1FD1BFB35" box="[629,704,1248,1272]" italics="true" pageId="38" pageNumber="1049">Colour</emphasis>
light brown to dark brown.
<emphasis id="60B87368FF8BFB24FF863451FEAFFAD5" box="[92,372,1280,1304]" italics="true" pageId="38" pageNumber="1049">Outer margin of mandibl e</emphasis>
distinctly sinuate.
<emphasis id="60B87368FF8BFB24FD9E3451FD47FAD5" box="[580,668,1280,1304]" italics="true" pageId="38" pageNumber="1049">Clypeus</emphasis>
with bituberculate, feebly arcuate or feebly bisinuate transverse posterior carina; position of tubercles variable, from in line with antennal insertions to in line with clypeo-frontal transition; posterior carina as long as anterior carina.
<emphasis id="60B87368FF8BFB24FE243430FDE7FAB5" box="[510,572,1377,1400]" italics="true" pageId="38" pageNumber="1049">Frons</emphasis>
feebly depressed posterior to posterior clypeal carina.
<emphasis id="60B87368FF8BFB24FED234D1FE7AFA55" box="[264,417,1408,1432]" italics="true" pageId="38" pageNumber="1049">Antennal club</emphasis>
with glabrous area of basal segment about one-half of exposed surface.
<emphasis id="60B87368FF8BFB24FEFF34F0FE49FA75" box="[293,402,1441,1464]" italics="true" pageId="38" pageNumber="1049">Pronotum</emphasis>
with anterior margin medially feebly raised, in dorsal view arcuate and not projecting over head; with four protrusions (®gure 3k±m): in major males outer horns distinctly longer and more strongly inward curved than inner horns, the latter ones sometimes reduced to feeble swellings (®gure 3k); in minor males all four horns much reduced in size, little curved, more or less equal in length and more approximated (®gure 3l, m); surface with large, densely spaced and often conūent punctures; punctures decreasing in density and size and also interspersed with some ®ne punctures often in between and always on pronotal protrusions.
<taxonomicName id="95CCD4F9FF8AFB25FF3431D5FE81FF51" box="[238,346,132,156]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="39" pageNumber="1050" phylum="Arthropoda" rank="genus">
<emphasis id="60B87368FF8AFB25FF3431D5FE81FF51" box="[238,346,132,156]" italics="true" pageId="39" pageNumber="1050">Scutellum</emphasis>
</taxonomicName>
with surface coarsely punctate.
<emphasis id="60B87368FF8AFB25FD1531D5FCF9FF51" box="[719,802,132,156]" italics="true" pageId="39" pageNumber="1050">Elytron</emphasis>
with strial punctures separated by four to ®ve and a half puncture diameters; intervals with approximatel y ®ve to six punctures between two striae.
<emphasis id="60B87368FF8AFB25FDED3195FD49FF11" box="[567,658,196,220]" italics="true" pageId="39" pageNumber="1050">Protibia</emphasis>
®ve-dentate.
<emphasis id="60B87368FF8AFB25FCEB3195FC10FF11" box="[817,971,196,220]" italics="true" pageId="39" pageNumber="1050">Protibial spur</emphasis>
about as long as ®fth tarsomere, apex shaped as in ®gure 12h, sometimes as in ®gure
<specimenCount id="44CA64F3FF8AFB25FC0531B5FBCCFF36" box="[991,1047,228,252]" pageId="39" pageNumber="1050" type="generic">
<quantity id="9534029FFF8AFB25FC0531B5FBD4FF31" box="[991,1039,228,252]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="39" pageNumber="1050" unit="m" value="12.0">12m</quantity>
.
</specimenCount>
<emphasis id="60B87368FF8AFB25FF863055FED3FED1" box="[92,264,260,284]" italics="true" pageId="39" pageNumber="1050">Metatrochanter</emphasis>
postero-medially with distinct angle and setation as in ®gure 12i.
<emphasis id="60B87368FF8AFB25FF863075FF0CFEF1" box="[92,215,292,316]" italics="true" pageId="39" pageNumber="1050">Metafemur</emphasis>
in ventral in posterior third with line of densely spaced short and moderately long setae; with fairly dense, moderately long and long setation in anterior third and apical area; with very dense short setation in basal area and along basal half of posterior margin; remaining areas with few isolated setae.
<emphasis id="60B87368FF8AFB25FC5630D2FC22FE56" box="[908,1017,387,411]" italics="true" pageId="39" pageNumber="1050">Metatibia</emphasis>
in lateral view with subapical carina feebly bilobed or with one distinct, lateral lobe only; with unmodi®ed spurs.
<emphasis id="60B87368FF8AFB25FE773092FDC0FE16" box="[429,539,451,475]" italics="true" pageId="39" pageNumber="1050">Underside</emphasis>
with posterior margin of sternite 5 laterally with concave curvature, medially triangularly emarginate (®gure 12l); posterior margin of sternite 6 broadly triangularly emarginate (®gure 12l); pygidium unmodi- ®ed (®gure 12k).
<emphasis id="60B87368FF8AFB25FEC63372FE5AFDF6" box="[284,385,547,571]" italics="true" pageId="39" pageNumber="1050">Genitalia</emphasis>
with aedeagus as in ®gure
<quantity id="9534029FFF8AFB25FD6A3372FD03FDF6" box="[688,728,547,571]" metricMagnitude="-2" metricUnit="kg" metricValue="1.3" pageId="39" pageNumber="1050" unit="g" value="13.0">13g</quantity>
±i; aedeagus strongly curved in lateral view; genital capsule apically with moderately long setation.
</paragraph>
<paragraph id="5273AF7AFF8AFB25FF8633D2FEF6FD57" blockId="39.[92,1048,642,1686]" box="[92,301,642,666]" pageId="39" pageNumber="1050">
<emphasis id="60B87368FF8AFB25FF8633D2FEF6FD57" box="[92,301,642,666]" italics="true" pageId="39" pageNumber="1050">Description female</emphasis>
</paragraph>
<paragraph id="5273AF7AFF8AFB25FF5E33F3FC8AFB75" blockId="39.[92,1048,642,1686]" pageId="39" pageNumber="1050">
<emphasis id="60B87368FF8AFB25FF5E33F3FED7FD77" box="[132,268,674,698]" italics="true" pageId="39" pageNumber="1050">Body length</emphasis>
<quantity id="9534029FFF8AFB25FEC233F3FE6EFD77" box="[280,437,674,698]" metricMagnitude="-2" metricUnit="m" metricValue="1.97" metricValueMax="2.05" metricValueMin="1.89" pageId="39" pageNumber="1050" unit="mm" value="19.7" valueMax="20.5" valueMin="18.9">18.9±20.5 mm</quantity>
(
<specimenCount id="44CA64F3FF8AFB25FE1733F3FD82FD77" box="[461,601,674,698]" pageId="39" pageNumber="1050" type="generic">6 specimens</specimenCount>
).
<emphasis id="60B87368FF8AFB25FDAE33F3FD64FD77" box="[628,703,674,698]" italics="true" pageId="39" pageNumber="1050">Colour</emphasis>
as in male.
<emphasis id="60B87368FF8AFB25FC8133F3FF65FD17" italics="true" pageId="39" pageNumber="1050">Outer margin of mandible</emphasis>
feebly sinuate.
<emphasis id="60B87368FF8AFB25FEAF3392FE68FD17" box="[373,435,707,730]" italics="true" pageId="39" pageNumber="1050">Frons</emphasis>
with strongly raised bituberculate, transverse carina; position of carina variable, from slightly anterior to to in line with postero-lateral genal angle; carina straight and slightly shorter than anterior clypeal carina.
<emphasis id="60B87368FF8AFB25FC693253FF52FCF7" italics="true" pageId="39" pageNumber="1050">Antennal club</emphasis>
as in male.
<emphasis id="60B87368FF8AFB25FEC13272FE53FCF7" box="[283,392,803,826]" italics="true" pageId="39" pageNumber="1050">Pronotum</emphasis>
with anterior margin as in male; with transverse, feebly carinate swelling at median portion of disc; carina in frontal view medially straight, rarely very feebly depressed, and laterally arcuate (®gure 3n); swelling with very few ®ne punctures, interspersed with large punctures; with large, mostly conūent punctures anterior, lateral and posterior to carinate swelling.
<emphasis id="60B87368FF8AFB25FD3E32F3FC02FC77" box="[740,985,930,954]" italics="true" pageId="39" pageNumber="1050">
<taxonomicName id="95CCD4F9FF8AFB25FD3E32F3FC8AFC77" box="[740,849,930,954]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="39" pageNumber="1050" phylum="Arthropoda" rank="genus">Scutellum</taxonomicName>
and elytron
</emphasis>
as in male.
<emphasis id="60B87368FF8AFB25FF7E3290FF24FC14" box="[164,255,961,985]" italics="true" pageId="39" pageNumber="1050">Protibia</emphasis>
as in male.
<emphasis id="60B87368FF8AFB25FE4F3290FDEAFC14" box="[405,561,961,985]" italics="true" pageId="39" pageNumber="1050">Protibial spur</emphasis>
slightly longer than ®fth tarsomere, acuminate.
<emphasis id="60B87368FF8AFB25FF6632B0FEB3FC34" box="[188,360,993,1017]" italics="true" pageId="39" pageNumber="1050">Metatrochanter</emphasis>
postero-medially with discernible angle and setation as in ®gure 12j.
<emphasis id="60B87368FF8AFB25FF0F3550FE8BFBD4" box="[213,336,1025,1049]" italics="true" pageId="39" pageNumber="1050">Metafemur</emphasis>
in ventral view in posterior third with line of densely spaced moderately long and long setae; in basal area and anterior third with fairly dense, long setation; in posterior third with fairly dense but moderately long setation; remaining median area with sparse setation.
<emphasis id="60B87368FF8AFB25FDB33531FD0DFBB5" box="[617,726,1120,1144]" italics="true" pageId="39" pageNumber="1050">Metatibia</emphasis>
as in male.
<emphasis id="60B87368FF8AFB25FCB63531FC01FBB5" box="[876,986,1120,1144]" italics="true" pageId="39" pageNumber="1050">Underside</emphasis>
with posterior margin of sternite 4 with distinct bisinuate curvature; sternite 5 unmodi®ed; apex of sternite 6 unmodi®ed, i.e. rounded; pygidium unmodi®ed.
</paragraph>
<paragraph id="5273AF7AFF8AFB25FF5E3591FE19FB35" blockId="39.[92,1048,642,1686]" pageId="39" pageNumber="1050">
<emphasis id="60B87368FF8AFB25FF5E3591FED3FB15" box="[132,264,1216,1240]" italics="true" pageId="39" pageNumber="1050">Distribution</emphasis>
(®gure 14). Known only from localities in
<collectingCountry id="2ADBEFEAFF8AFB25FC923591FC45FB15" box="[840,926,1216,1240]" name="Malawi" pageId="39" pageNumber="1050">Malawi</collectingCountry>
and one untraced locality in the D.R.C.
</paragraph>
<paragraph id="5273AF7AFF8AFB25FF5E3450FC02F99A" blockId="39.[92,1048,642,1686]" pageId="39" pageNumber="1050">
<emphasis id="60B87368FF8AFB25FF5E3450FE57FAD5" box="[132,396,1280,1304]" italics="true" pageId="39" pageNumber="1050">Type material examined</emphasis>
.
<materialsCitation id="E2A4A527FF8AFB25FE433450FF0CFAB5" collectingDate="1973-02-11" collectionCode="MMCM" collectorName="C. O. Dudley &amp; C. O. Dudley'" country="Malawi" latitude="-15.49" location="Malawi" longLatPrecision="771" longitude="35.03" pageId="39" pageNumber="1050" specimenCount="1" typeStatus="holotype">
<typeStatus id="8D7711D8FF8AFB25FE433450FDD7FAD5" box="[409,524,1280,1304]" pageId="39" pageNumber="1050" type="holotype">HOLOTYPE</typeStatus>
:`M2680 [¯ew to light, in area of cultivation, degraded woodland and blue gum plantationsÐpersonal comment by
<collectorName id="FF39CAACFF8AFB25FCAC3471FBD3FAF5" box="[886,1032,1312,1336]" pageId="39" pageNumber="1050">C.O. Dudley</collectorName>
], Limbe [
<geoCoordinate id="37F8C9BDFF8AFB25FF623411FED9FA95" box="[184,258,1344,1368]" degrees="15.49" direction="south" orientation="latitude" pageId="39" pageNumber="1050" precision="555" value="-15.49">15.49S</geoCoordinate>
<geoCoordinate id="37F8C9BDFF8AFB25FED73411FE87FA9A" box="[269,348,1344,1368]" degrees="35.03" direction="east" orientation="longitude" pageId="39" pageNumber="1050" precision="555" value="35.03">35.03E</geoCoordinate>
],
<collectingCountry id="2ADBEFEAFF8AFB25FEA23411FE15FA95" box="[376,462,1344,1368]" name="Malawi" pageId="39" pageNumber="1050">Malawi</collectingCountry>
,
<date id="267289BAFF8AFB25FE053411FD91FA95" box="[479,586,1344,1368]" pageId="39" pageNumber="1050" value="1973-02-11">
<collectingDate id="36367052FF8AFB25FE053411FD91FA95" box="[479,586,1344,1368]" pageId="39" pageNumber="1050" value="1973-02-11">11.ii.1973</collectingDate>
</date>
,
<collectorName id="FF39CAACFF8AFB25FD873411FD30FA95" box="[605,747,1344,1368]" pageId="39" pageNumber="1050">C.O.Dudley</collectorName>
[handwritten by Dudley] (
<collectionCode id="34DD37BFFF8AFB25FFBC3431FF13FAB5" box="[102,200,1376,1400]" country="Malawi" httpUri="http://grbio.org/cool/g4ts-fgzt" name="Museum of Malawi" pageId="39" pageNumber="1050">MMCM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E2A4A527FF8AFB25FF313431FE52FA55" collectingDate="1973-02-11" collectionCode="MMCM" collectorName="C. O. Dudley &amp; C. O. Dudley'" country="Malawi" latitude="-15.49" location="Malawi" longLatPrecision="771" longitude="35.03" pageId="39" pageNumber="1050" specimenCount="2" typeStatus="allotype">
<typeStatus id="8D7711D8FF8AFB25FF313431FE8DFAB5" box="[235,342,1376,1400]" pageId="39" pageNumber="1050" type="allotype">ALLOTYPE</typeStatus>
m: same data as holotype but`M2685 [same data as for
<typeStatus id="8D7711D8FF8AFB25FF8634D1FF1FFA55" box="[92,196,1408,1432]" pageId="39" pageNumber="1050" type="holotype">holotype</typeStatus>
]... (
<collectionCode id="34DD37BFFF8AFB25FF2034D1FE87FA55" box="[250,348,1408,1432]" country="Malawi" httpUri="http://grbio.org/cool/g4ts-fgzt" name="Museum of Malawi" pageId="39" pageNumber="1050">MMCM</collectionCode>
); 6
</materialsCitation>
<materialsCitation id="E2A4A527FF8AFB25FE4E34D1FC81FA3A" collectingDate="1971-12" collectionCode="TMSA" collectorName="C. O. Dudley &amp; Dudley" country="Malawi" elevation="1" latitude="-15.49" location="So. Region" longLatPrecision="771" longitude="35.03" pageId="39" pageNumber="1050" specimenCount="1" stateProvince="Blantyre" typeStatus="paratype">
<typeStatus id="8D7711D8FF8AFB25FE4E34D1FDDCFA55" box="[404,519,1408,1432]" pageId="39" pageNumber="1050" type="paratype">PARATYPES</typeStatus>
: 1,`M3965 [notes stolen but probably collected in the Shire Highlands near
<collectingRegion id="90086198FF8AFB25FDDF34F1FDBDFA75" box="[517,614,1440,1464]" country="Malawi" name="Blantyre" pageId="39" pageNumber="1050">Blantyre</collectingRegion>
<emphasis id="60B87368FF8AFB25FDBC34F1FDB4FA73" bold="true" box="[614,623,1440,1470]" pageId="39" pageNumber="1050">/</emphasis>
LimbeÐpersonal comment by
<collectorName id="FF39CAACFF8AFB25FC3B34F1FF69FA15" pageId="39" pageNumber="1050">C.O. Dudley</collectorName>
],
<location id="5713F9A1FF8AFB25FF173491FE8BFA15" LSID="urn:lsid:plazi:treatment:DA651E6CFF8BFB2AFE723506FF38FBD4:5713F9A1FF8AFB25FF173491FE8BFA15" box="[205,336,1472,1496]" country="Malawi" name="So. Region" pageId="39" pageNumber="1050" stateProvince="Blantyre">So. Region</location>
,
<collectingCountry id="2ADBEFEAFF8AFB25FEBE3491FE61FA15" box="[356,442,1472,1496]" name="Malawi" pageId="39" pageNumber="1050">Malawi</collectingCountry>
,
<date id="267289BAFF8AFB25FE063491FDE8FA15" box="[476,563,1472,1496]" pageId="39" pageNumber="1050" value="1971-12">
<collectingDate id="36367052FF8AFB25FE063491FDE8FA15" box="[476,563,1472,1496]" pageId="39" pageNumber="1050" value="1971-12">xii.1971</collectingDate>
</date>
[handwritten by
<collectorName id="FF39CAACFF8AFB25FCCB3491FCBFFA15" box="[785,868,1472,1496]" pageId="39" pageNumber="1050">Dudley</collectorName>
] (
<collectionCode id="34DD37BFFF8AFB25FC5F3491FC0CFA15" box="[901,983,1472,1496]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="39" pageNumber="1050">TMSA</collectionCode>
);
<quantity id="9534029FFF8AFB25FC2E3491FBD4FA15" box="[1012,1039,1472,1496]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="39" pageNumber="1050" unit="m" value="1.0">
<elevation id="D9E14849FF8AFB25FC2E3491FBD4FA15" box="[1012,1039,1472,1496]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="39" pageNumber="1050" unit="m" value="1.0">1m</elevation>
</quantity>
, same data but:`M3955 [same comment as for M3965]... (
<collectionCode id="34DD37BFFF8AFB25FCDB348EFC88FA3A" box="[769,851,1503,1527]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="39" pageNumber="1050">TMSA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E2A4A527FF8AFB25FCB6348EFC0FF9DA" collectionCode="BMNH" collectorName="Elsa &amp; Distant Coll." country="Malawi" latitude="-15.47" location="Blantyre" longLatPrecision="771" longitude="35.0" pageId="39" pageNumber="1050" specimenCount="1" stateProvince="Blantyre" typeStatus="paratype">
1 l[diss.],
<quantity id="9534029FFF8AFB25FC3F348EFBCBFA3A" box="[997,1040,1503,1527]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="39" pageNumber="1050" unit="mm" value="2.0">2mm</quantity>
,`
<collectingRegion id="90086198FF8AFB25FFB834AEFF18F9DA" box="[98,195,1535,1559]" country="Malawi" name="Blantyre" pageId="39" pageNumber="1050">Blantyre</collectingRegion>
[
<geoCoordinate id="37F8C9BDFF8AFB25FF3E34AEFEF5F9DA" box="[228,302,1535,1559]" degrees="15.47" direction="south" orientation="latitude" pageId="39" pageNumber="1050" precision="555" value="-15.47">15.47S</geoCoordinate>
<geoCoordinate id="37F8C9BDFF8AFB25FE9F34AEFE48F9DA" box="[325,403,1535,1559]" degrees="35.00" direction="east" orientation="longitude" pageId="39" pageNumber="1050" precision="555" value="35.0">35.00E</geoCoordinate>
] (
<collectorName id="FF39CAACFF8AFB25FE6534AEFE2AF9DA" box="[447,497,1535,1559]" pageId="39" pageNumber="1050">Elsa</collectorName>
),
<collectorName id="FF39CAACFF8AFB25FDC234AEFD60F9DA" box="[536,699,1535,1559]" pageId="39" pageNumber="1050">Distant Coll.</collectorName>
, 1911-383. (
<collectionCode id="34DD37BFFF8AFB25FCB43751FC1DF9DA" box="[878,966,1536,1559]" pageId="39" pageNumber="1050">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E2A4A527FF8AFB25FC2B34AEFE30F99A" collectionCode="HAHC" collectorName="E. S. Ross &amp; R. E. Leech" country="Malawi" latitude="-16.04" location="7 mi. S of Cholo" longLatPrecision="769" longitude="35.08" pageId="39" pageNumber="1050" specimenCount="1" stateProvince="Thyolo" typeStatus="paratype">
1,`
<collectingCountry id="2ADBEFEAFF8AFB25FFB8374EFF00F9FA" box="[98,219,1567,1591]" name="Malawi" pageId="39" pageNumber="1050">Nyasaland</collectingCountry>
,
<location id="5713F9A1FF8AFB25FF363771FE76F9FA" LSID="urn:lsid:plazi:treatment:DA651E6CFF8BFB2AFE723506FF38FBD4:5713F9A1FF8AFB25FF363771FE76F9FA" box="[236,429,1567,1591]" country="Malawi" latitude="-16.04" longLatPrecision="769" longitude="35.08" name="7 mi. S of Cholo" pageId="39" pageNumber="1050" stateProvince="Thyolo">
<quantity id="9534029FFF8AFB25FF363771FEFCF9FA" box="[236,295,1567,1591]" metricMagnitude="4" metricUnit="m" metricValue="1.1265408" pageId="39" pageNumber="1050" unit="mi" value="7.0">7 mi.</quantity>
S of Cholo
</location>
[
<collectingRegion id="90086198FF8AFB25FE1B374EFDC8F9FA" box="[449,531,1567,1591]" country="Malawi" name="Thyolo" pageId="39" pageNumber="1050">Thyolo</collectingRegion>
,
<geoCoordinate id="37F8C9BDFF8AFB25FDF9374EFDB6F9FA" box="[547,621,1567,1591]" degrees="16.04" direction="south" orientation="latitude" pageId="39" pageNumber="1050" precision="555" value="-16.04">16.04S</geoCoordinate>
<geoCoordinate id="37F8C9BDFF8AFB25FDAC374EFD1EF9FA" box="[630,709,1567,1591]" degrees="35.08" direction="east" orientation="longitude" pageId="39" pageNumber="1050" precision="555" value="35.08">35.08E</geoCoordinate>
], II-25-58, 940 m
<emphasis id="60B87368FF8AFB25FC42374EFC7BF9F0" bold="true" box="[920,928,1567,1597]" pageId="39" pageNumber="1050">/</emphasis>
`
<collectorName id="FF39CAACFF8AFB25FC72374EFBCDF9FA" box="[936,1046,1567,1591]" pageId="39" pageNumber="1050">E.S. Ross</collectorName>
&amp;
<collectorName id="FF39CAACFF8AFB25FFA33711FF22F99A" box="[121,249,1599,1623]" pageId="39" pageNumber="1050">R.E. Leech</collectorName>
collectors (
<collectionCode id="34DD37BFFF8AFB25FE50376EFE06F99A" box="[394,477,1599,1623]" pageId="39" pageNumber="1050">HAHC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E2A4A527FF8AFB25FE20376EFC0FF99A" box="[506,980,1599,1623]" collectionCode="RMCI" country="Democratic Republic of the Congo" location="Maru" pageId="39" pageNumber="1050" specimenCount="1" typeStatus="paratype">
1,`
<collectingCountry id="2ADBEFEAFF8AFB25FDEA376EFDA6F99A" box="[560,637,1599,1623]" name="Democratic Republic of the Congo" pageId="39" pageNumber="1050">Congo</collectingCountry>
,
<location id="5713F9A1FF8AFB25FD573711FD16F99A" LSID="urn:lsid:plazi:treatment:DA651E6CFF8BFB2AFE723506FF38FBD4:5713F9A1FF8AFB25FD573711FD16F99A" box="[653,717,1600,1623]" country="Democratic Republic of the Congo" name="Maru" pageId="39" pageNumber="1050">Maru</location>
[not traced] (
<collectionCode id="34DD37BFFF8AFB25FCA3376EFC1EF99A" box="[889,965,1599,1623]" pageId="39" pageNumber="1050">RMCI</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="5273AF7AFF8AFB2AFF5E370EFE8AFF11" blockId="39.[92,1048,642,1686]" lastBlockId="40.[92,1048,132,1049]" lastPageId="40" lastPageNumber="1051" pageId="39" pageNumber="1050">
<emphasis id="60B87368FF8AFB25FF5E370EFE03F9BA" box="[132,472,1631,1655]" italics="true" pageId="39" pageNumber="1050">Additional material examined</emphasis>
(
<specimenCount id="44CA64F3FF8AFB25FE28370EFD5BF9BA" box="[498,640,1631,1655]" pageId="39" pageNumber="1050" type="generic">3 specimens</specimenCount>
).
<emphasis id="60B87368FF8AFB25FD44370EFD21F9BA" bold="true" box="[670,762,1631,1655]" pageId="39" pageNumber="1050">
<collectingCountry id="2ADBEFEAFF8AFB25FD44370EFD2EF9BA" box="[670,757,1631,1655]" name="Malawi" pageId="39" pageNumber="1050">Malawi</collectingCountry>
:
</emphasis>
<materialsCitation id="E2A4A527FF8AFB2AFCD2370EFF0CFF51" collectingDate="1966-12" collectionCode="MMCM" collectorName="Coll. R. F. Lee &amp; C. I. E. Coll." country="Malawi" elevation="1" lastPageId="40" lastPageNumber="1051" latitude="-15.22" location="Region" longLatPrecision="772" longitude="35.22" pageId="39" pageNumber="1050" specimenCount="1" stateProvince="Zomba">
<quantity id="9534029FFF8AFB25FCD2370EFCFFF9BA" box="[776,804,1631,1655]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="39" pageNumber="1050" unit="m" value="1.0">
<elevation id="D9E14849FF8AFB25FCD2370EFCFFF9BA" box="[776,804,1631,1655]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="39" pageNumber="1050" unit="m" value="1.0">1m</elevation>
</quantity>
, S.
<location id="5713F9A1FF8AFB25FC87370EFC75F9BA" LSID="urn:lsid:plazi:treatment:DA651E6CFF8BFB2AFE723506FF38FBD4:5713F9A1FF8AFB25FC87370EFC75F9BA" box="[861,942,1631,1655]" country="Malawi" latitude="-15.22" longLatPrecision="772" longitude="35.22" name="Region" pageId="39" pageNumber="1050" stateProvince="Zomba">Region</location>
,
<collectingRegion id="90086198FF8AFB25FC1E370EFBCDF9BA" box="[964,1046,1631,1655]" country="Malawi" name="Zomba" pageId="39" pageNumber="1050">Zomba</collectingRegion>
[
<geoCoordinate id="37F8C9BDFF8AFB25FFBF372FFF74F95B" box="[101,175,1662,1686]" degrees="15.22" direction="south" orientation="latitude" pageId="39" pageNumber="1050" precision="555" value="-15.22">15.22S</geoCoordinate>
<geoCoordinate id="37F8C9BDFF8AFB25FF64372FFED7F95B" box="[190,268,1662,1686]" degrees="35.22" direction="east" orientation="longitude" pageId="39" pageNumber="1050" precision="555" value="35.22">35.22E</geoCoordinate>
],
<date id="267289BAFF8AFB25FEF7372FFE39F95B" box="[301,482,1662,1686]" pageId="39" pageNumber="1050" value="1966-12">
<collectingDate id="36367052FF8AFB25FEF7372FFE39F95B" box="[301,482,1662,1686]" pageId="39" pageNumber="1050" value="1966-12">December 1966</collectingDate>
</date>
, at light,
<collectorName id="FF39CAACFF8AFB25FDBC372FFCC3F95B" box="[614,792,1662,1686]" pageId="39" pageNumber="1050">Coll. R.F. Lee</collectorName>
,
<collectorName id="FF39CAACFF8AFB25FCF7372FFC6CF95B" box="[813,951,1662,1686]" pageId="39" pageNumber="1050">C.I.E. Coll.</collectorName>
A.1815 (
<collectionCode id="34DD37BFFF85FB2AFFBC31D5FF13FF51" box="[102,200,132,156]" country="Malawi" httpUri="http://grbio.org/cool/g4ts-fgzt" name="Museum of Malawi" pageId="40" pageNumber="1051">MMCM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E2A4A527FF85FB2AFF3231D5FDFEFF71" collectingDate="1990-02" collectionCode="MMCM" collectorName="Mughese &amp; M. Chivura" country="Malawi" elevation="1" latitude="-9.41" location="Chitipa" longLatPrecision="780" longitude="33.19" pageId="40" pageNumber="1051" specimenCount="1" stateProvince="Chitipa">
<quantity id="9534029FFF85FB2AFF3231D5FEDFFF51" box="[232,260,132,156]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="40" pageNumber="1051" unit="m" value="1.0">
<elevation id="D9E14849FF85FB2AFF3231D5FEDFFF51" box="[232,260,132,156]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="40" pageNumber="1051" unit="m" value="1.0">1m</elevation>
</quantity>
,
<collectingRegion id="90086198FF85FB2AFECD31D5FEB7FF51" box="[279,364,132,156]" country="Malawi" name="Chitipa" pageId="40" pageNumber="1051">Chitipa</collectingRegion>
[
<geoCoordinate id="37F8C9BDFF85FB2AFE5B31D5FE10FF51" box="[385,459,132,156]" degrees="09.41" direction="south" orientation="latitude" pageId="40" pageNumber="1051" precision="555" value="-9.41">09.41S</geoCoordinate>
<geoCoordinate id="37F8C9BDFF85FB2AFE0C31D5FDFEFF51" box="[470,549,132,156]" degrees="33.19" direction="east" orientation="longitude" pageId="40" pageNumber="1051" precision="555" value="33.19">33.19E</geoCoordinate>
],
<collectorName id="FF39CAACFF85FB2AFD9931D5FD72FF51" box="[579,681,132,156]" pageId="40" pageNumber="1051">Mughese</collectorName>
[?, illegible, not traced] forest,
<date id="267289BAFF85FB2AFF8631F5FED8FF71" box="[92,259,164,188]" pageId="40" pageNumber="1051" value="1990-02">
<collectingDate id="36367052FF85FB2AFF8631F5FED8FF71" box="[92,259,164,188]" pageId="40" pageNumber="1051" value="1990-02">February 1990</collectingDate>
</date>
,
<collectorName id="FF39CAACFF85FB2AFECC31F4FE45FF71" box="[278,414,164,188]" pageId="40" pageNumber="1051">M. Chivura</collectorName>
(
<collectionCode id="34DD37BFFF85FB2AFE6E31F5FDCDFF71" box="[436,534,164,188]" country="Malawi" httpUri="http://grbio.org/cool/g4ts-fgzt" name="Museum of Malawi" pageId="40" pageNumber="1051">MMCM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E2A4A527FF85FB2AFDEE31F5FE93FF11" collectionCode="DMSA" collectorName="Museum Collection'" country="Malawi" location="Untraced" pageId="40" pageNumber="1051" specimenCount="1">
<emphasis id="60B87368FF85FB2AFDEE31F5FD7BFF71" bold="true" box="[564,672,164,188]" pageId="40" pageNumber="1051">
<location id="5713F9A1FF85FB2AFDEE31F5FD40FF71" LSID="urn:lsid:plazi:treatment:DA651E6CFF8BFB2AFE723506FF38FBD4:5713F9A1FF85FB2AFDEE31F5FD40FF71" box="[564,667,164,188]" country="Malawi" name="Untraced" pageId="40" pageNumber="1051">Untraced</location>
:
</emphasis>
1,`unlabelled in old
<collectorName id="FF39CAACFF85FB2AFC6E31F4FF0DFF11" pageId="40" pageNumber="1051">Museum Collection</collectorName>
(
<collectionCode id="34DD37BFFF85FB2AFF363195FE9AFF11" box="[236,321,196,220]" country="South Africa" httpUri="http://grbio.org/cool/i1wq-4y1w" name="Durban Museum" pageId="40" pageNumber="1051">DMSA</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="5273AF7AFF85FB2AFF5E31B5FCB9FEF6" blockId="40.[92,1048,132,1049]" pageId="40" pageNumber="1051">
<emphasis id="60B87368FF85FB2AFF5E31B5FF20FF31" box="[132,251,228,252]" italics="true" pageId="40" pageNumber="1051">Etymology</emphasis>
. The species is named after C.O. Dudley, Research Associate and Honorary Curator of Entomology at the Museums of
<collectingCountry id="2ADBEFEAFF85FB2AFD0B3055FCFCFED1" box="[721,807,260,284]" name="Malawi" pageId="40" pageNumber="1051">Malawi</collectingCountry>
, who collected some of the above specimens and kindly provided the material for study.
</paragraph>
<paragraph id="5273AF7AFF85FB2AFF5E3015FEAFFD17" blockId="40.[92,1048,132,1049]" pageId="40" pageNumber="1051">
<emphasis id="60B87368FF85FB2AFF5E3015FF23FE91" box="[132,248,324,348]" italics="true" pageId="40" pageNumber="1051">Comments</emphasis>
.
<taxonomicName id="95CCD4F9FF85FB2AFED53015FE09FE91" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[271,466,324,348]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1051" phylum="Arthropoda" rank="species" species="dudleyi">
<emphasis id="60B87368FF85FB2AFED53015FE09FE91" box="[271,466,324,348]" italics="true" pageId="40" pageNumber="1051">BolbaOEer dudleyi</emphasis>
</taxonomicName>
is placed within the
<taxonomicName id="95CCD4F9FF85FB2AFD3B3014FCB5FE91" box="[737,878,325,348]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1051" phylum="Arthropoda" rank="species" species="coriaceus">
<emphasis id="60B87368FF85FB2AFD3B3014FCB5FE91" box="[737,878,325,348]" italics="true" pageId="40" pageNumber="1051">B. coriaceus</emphasis>
</taxonomicName>
species-group mainly on the basis of sternal and pygidial modi®cations in the males. The laterally concave and medially triangularly emarginate posterior margin of sternite ®ve and the broadly triangularl y emarginate posterior margin of sternite six easily distinguish the males of
<taxonomicName id="95CCD4F9FF85FB2AFF343095FE86FE16" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[238,349,451,475]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1051" phylum="Arthropoda" rank="species" species="dudleyi">
<emphasis id="60B87368FF85FB2AFF343095FE86FE16" box="[238,349,451,475]" italics="true" pageId="40" pageNumber="1051">B. dudleyi</emphasis>
</taxonomicName>
from the males of
<taxonomicName id="95CCD4F9FF85FB2AFDE23095FD64FE16" box="[568,703,452,475]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1051" phylum="Arthropoda" rank="species" species="coriaceus">
<emphasis id="60B87368FF85FB2AFDE23095FD64FE16" box="[568,703,452,475]" italics="true" pageId="40" pageNumber="1051">B. coriaceus</emphasis>
</taxonomicName>
(compare ®gure 12l with
<quantity id="9534029FFF85FB2AFC3E3092FBD7FE16" box="[996,1036,451,475]" metricMagnitude="-2" metricUnit="kg" metricValue="1.2" pageId="40" pageNumber="1051" unit="g" value="12.0">12g</quantity>
) but also from the males of
<taxonomicName id="95CCD4F9FF85FB2AFE7830B5FDCFFE36" box="[418,532,483,507]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1051" phylum="Arthropoda" rank="species" species="abditus">
<emphasis id="60B87368FF85FB2AFE7830B5FDCFFE36" box="[418,532,483,507]" italics="true" pageId="40" pageNumber="1051">B. abditus</emphasis>
</taxonomicName>
(compare ®gure 12l with 12q). Males of
<taxonomicName id="95CCD4F9FF85FB2AFC2430B5FF71FDD6" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1051" phylum="Arthropoda" rank="species" species="dudleyi">
<emphasis id="60B87368FF85FB2AFC2430B5FF71FDD6" italics="true" pageId="40" pageNumber="1051">B. dudleyi</emphasis>
</taxonomicName>
and
<taxonomicName id="95CCD4F9FF85FB2AFF363355FE86FDD6" box="[236,349,515,539]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1051" phylum="Arthropoda" rank="species" species="abditus">
<emphasis id="60B87368FF85FB2AFF363355FE86FDD6" box="[236,349,515,539]" italics="true" pageId="40" pageNumber="1051">B. abditus</emphasis>
</taxonomicName>
furthermore diOEer by the shape of the pygidium (compare ®gure 12k with 12p) and by the shape of the aedeagal parameres (compare ®gure
<quantity id="9534029FFF85FB2AFF793313FF11FD97" box="[163,202,578,602]" metricMagnitude="-2" metricUnit="kg" metricValue="1.3" pageId="40" pageNumber="1051" unit="g" value="13.0">13g</quantity>
±i with 13j±l). Of importance, on the other hand, is the strong similarity of
<taxonomicName id="95CCD4F9FF85FB2AFFA63332FF30FDB7" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[124,235,610,634]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1051" phylum="Arthropoda" rank="species" species="dudleyi">
<emphasis id="60B87368FF85FB2AFFA63332FF30FDB7" box="[124,235,610,634]" italics="true" pageId="40" pageNumber="1051">B. dudleyi</emphasis>
</taxonomicName>
and southern populations of
<taxonomicName id="95CCD4F9FF85FB2AFD9E3332FD17FDB7" box="[580,716,611,634]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1051" phylum="Arthropoda" rank="species" species="coriaceus">
<emphasis id="60B87368FF85FB2AFD9E3332FD17FDB7" box="[580,716,611,634]" italics="true" pageId="40" pageNumber="1051">B. coriaceus</emphasis>
</taxonomicName>
in the shape of the aedeagal parameres (compare ®gure
<quantity id="9534029FFF85FB2AFE4633D3FE1FFD57" box="[412,452,642,666]" metricMagnitude="-2" metricUnit="kg" metricValue="1.3" pageId="40" pageNumber="1051" unit="g" value="13.0">13g</quantity>
±i with 13a±c). This and the lack of any distinct diOEerences between females, as already discussed, might indicate close relationship between the two species.
</paragraph>
<paragraph id="5273AF7AFF85FB2AFF5E33B3FF38FBD4" blockId="40.[92,1048,132,1049]" pageId="40" pageNumber="1051">
<taxonomicName id="95CCD4F9FF85FB2AFF5E33B3FE9FFD37" authorityName="Gussmann &amp; Scholtz" authorityYear="2001" box="[132,324,738,762]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1051" phylum="Arthropoda" rank="species" species="dudleyi">
<emphasis id="60B87368FF85FB2AFF5E33B3FE9FFD37" box="[132,324,738,762]" italics="true" pageId="40" pageNumber="1051">BolbaOEer dudleyi</emphasis>
</taxonomicName>
is known mainly from localities in
<collectingCountry id="2ADBEFEAFF85FB2AFD2833B3FC93FD37" box="[754,840,738,762]" name="Malawi" pageId="40" pageNumber="1051">Malawi</collectingCountry>
and most female specimens were collected together with males, thus leaving no doubt about the correct male
<emphasis id="60B87368FF85FB2AFF323273FF2AFC8D" bold="true" box="[232,241,802,832]" pageId="40" pageNumber="1051">/</emphasis>
female association for these combinations. Two females, however, were caught alone, and, as in the case of unassociate d
<taxonomicName id="95CCD4F9FF85FB2AFD593212FCD2FC97" box="[643,777,835,858]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1051" phylum="Arthropoda" rank="species" species="coriaceus">
<emphasis id="60B87368FF85FB2AFD593212FCD2FC97" box="[643,777,835,858]" italics="true" pageId="40" pageNumber="1051">B. coriaceus</emphasis>
</taxonomicName>
females, the determination labels bear a question mark. The
<specimenCount id="44CA64F3FF85FB2AFDD73233FD48FCB7" box="[525,659,866,890]" pageId="40" pageNumber="1051" type="female">two females</specimenCount>
were excluded from
<typeStatus id="8D7711D8FF85FB2AFC5B3233FC69FCB7" box="[897,946,866,890]" pageId="40" pageNumber="1051">type</typeStatus>
designation. One rather badly preserved male specimen was also excluded from
<typeStatus id="8D7711D8FF85FB2AFC3C32D3FBCCFC57" box="[998,1047,898,922]" pageId="40" pageNumber="1051">type</typeStatus>
designation (see above) because the posterior margins of sternites 5 and 6 are not very clearly visible. With regard to shape and setation of the metatrochanter and metafemur, however, the specimen is most de®nitely not a
<taxonomicName id="95CCD4F9FF85FB2AFD2C32B3FCBFFC34" box="[758,868,993,1017]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1051" phylum="Arthropoda" rank="species" species="abditus">
<emphasis id="60B87368FF85FB2AFD2C32B3FCBFFC34" box="[758,868,993,1017]" italics="true" pageId="40" pageNumber="1051">B. abditus</emphasis>
</taxonomicName>
but most likely a
<taxonomicName id="95CCD4F9FF85FB2AFFA83553FF00FBD4" box="[114,219,1025,1049]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="1051" phylum="Arthropoda" rank="species" species="dudley">
<emphasis id="60B87368FF85FB2AFFA83553FF00FBD4" box="[114,219,1025,1049]" italics="true" pageId="40" pageNumber="1051">B. dudley</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>