treatments-xml/data/DA/3F/78/DA3F784A5E2C55B2ACA461EC4E615FCB.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1147.93824" ID-Pensoft-Pub="1313-2970-1147-1" ID-Pensoft-UUID="FF2B6B317B505F9EA0E1000BDCD16CE7" ID-ZooBank="9BB9881B0076473D8E53155D37CA1F50" ModsDocID="1313-2970-1147-1" checkinTime="1676378625502" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Hunyadi, Andras, Vermeulen, Jaap J., Grego, Jozef, Sutcharit, Chirasak, Reischuetz, Alexander, Dumrongrojwattana, Pongrat, Botta-Dukat, Zoltan, Oerstan, Aydin, Fekete, Judit &amp; Jochum, Adrienne" docDate="2023" docId="DA3F784A5E2C55B2ACA461EC4E615FCB" docLanguage="en" docName="ZooKeys 1147: 1-177" docOrigin="ZooKeys 1147" docPubDate="2023-02-13" docSource="http://dx.doi.org/10.3897/zookeys.1147.93824" docTitle="Angustopila steffeki Pall-Gergely &amp; Grego 2023, sp. nov." docType="treatment" docUuid="76FA631F-18B7-4743-993F-B6D626BB982D" docUuidSource="ZooBank" docVersion="1" id="FF2B6B317B505F9EA0E1000BDCD16CE7" lastPageNumber="1" masterDocId="FF2B6B317B505F9EA0E1000BDCD16CE7" masterDocTitle="Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)" masterLastPageNumber="177" masterPageNumber="1" pageNumber="1" updateTime="1676378625502" updateUser="pensoft">
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<mods:title>Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
<mods:affiliation>Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hunyadi, Andras</mods:namePart>
<mods:affiliation>Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
<mods:affiliation>JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grego, Jozef</mods:namePart>
<mods:affiliation>Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reischuetz, Alexander</mods:namePart>
<mods:affiliation>Puechhaimgasse 52, A- 3580 Horn, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dumrongrojwattana, Pongrat</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Botta-Dukat, Zoltan</mods:namePart>
<mods:affiliation>Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Oerstan, Aydin</mods:namePart>
<mods:affiliation>12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fekete, Judit</mods:namePart>
<mods:affiliation>University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary &amp; Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary</mods:affiliation>
</mods:name>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Jochum, Adrienne</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6624-6412</mods:nameIdentifier>
<mods:affiliation>Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland &amp; Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland &amp; Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany</mods:affiliation>
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<mods:date>2023</mods:date>
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<mods:number>2023-02-13</mods:number>
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<mods:detail type="volume">
<mods:number>1147</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1147.93824</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1147.93824</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:76FA631F-18B7-4743-993F-B6D626BB982D" httpUri="http://treatment.plazi.org/id/DA3F784A5E2C55B2ACA461EC4E615FCB" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="https://zoobank.org/76FA631F-18B7-4743-993F-B6D626BB982D" authority="Páll-Gergely &amp; Grego" authorityName="Pall-Gergely &amp; Grego" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila steffeki" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="steffeki" status="sp. nov.">
Angustopila steffeki 
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Grego
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 81" captionStartId="F81" captionText="Figure 81. Angustopila steffeki Pall-Gergely &amp; Grego, sp. nov. (holotype, NHMUK 20170296). Apertural (A), ventral (B), lateral (C) and apical (D) sides of the shell; sculpture of the protoconch (E); aperture (F); ventral (G) and frontal (H) surface of the body whorl." figureDoi="10.3897/zookeys.1147.93824.figure81" httpUri="https://binary.pensoft.net/fig/808890" pageId="0" pageNumber="1">Fig. 81</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: Laos • 1 empty shell (H: 0.64 mm, D: 0.74 mm); Bolikhamsay Province, large spring lake at S foot of limestone massif 2 km NNE of Na Pavan village, ca. 5 km from Phontan crossing at road 8 (locality code: JG16); 
<geoCoordinate degrees="18" direction="north" minutes="13.30" orientation="latitude" precision="9" value="18.221666">18°13.30'N</geoCoordinate>
, 
<geoCoordinate degrees="104" direction="east" minutes="44.81" orientation="longitude" precision="9" value="104.74683">104°44.81'E</geoCoordinate>
; 18 Feb. 2017; J. Grego leg.; NHMUK 20170296.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
A small, depressed-globular 
<taxonomicName authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
</taxonomicName>
species with the aperture oblique to shell axis, a strongly protruding aperture, and relatively weak parietal and rather strong upper palatal tooth.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Shell small, wider than high; off-white, depressed-globular with domed spire; body whorl widest in standard apertural view; protoconch consists of 1.25 whorls, microstructure finely pitted and granular with a powdery superficial texture and with weak spiral striae (although holotype corroded; ca. 19 on body whorl from standard apertural view); teleoconch ornamented by some fine irregular, radial growth lines and stronger, equidistantly-arranged spiral striae; whorls 4, shouldered; aperture not oblique to shell axis from lateral view (although the peristome might be broken); umbilicus very wide; aperture pear-shaped with strongly distinct, relatively narrow sinulus and wider main section; peristome slightly expanded, not reflected; parietal callus strongly protruding, detached from penultimate whorl; upper parietal tooth relatively weak, low, but quite long, reaching peristome, perpendicular to parietal side; palatal tooth relatively strong, situated directly opposite of parietal tooth just at peristome edge.</paragraph>
<caption doi="10.3897/zookeys.1147.93824.figure81" httpUri="https://binary.pensoft.net/fig/808890" pageId="0" pageNumber="1" start="Figure 81" startId="F81">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 81.</emphasis>
<taxonomicName authorityName="Pall-Gergely &amp; Grego" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila steffeki" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="steffeki">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila steffeki</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Grego, sp. nov. (holotype, NHMUK 20170296). Apertural (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
), ventral (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
), lateral (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
) and apical (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
) sides of the shell; sculpture of the protoconch (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
); aperture (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
); ventral (
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
) and frontal (
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
) surface of the body whorl.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements (in mm).</paragraph>
<paragraph pageId="0" pageNumber="1">H = 0.64, D = 0.74, H/D*100 = 86.5, RUD = 36.3 (holotype).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
In comparison to other 
<taxonomicName authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
</taxonomicName>
species possessing two apertural denticles, 
<taxonomicName authorityName="Pall-Gergely &amp; Grego" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila steffeki" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="steffeki">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila steffeki</emphasis>
</taxonomicName>
sp. nov. shows a more depressed shell with a dome-like appearance. 
<taxonomicName authorityName="Dumrongrojwattana, Chuenit &amp; Wongkamhaeng" authorityYear="2021" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila pallgergelyi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pallgergelyi">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila pallgergelyi</emphasis>
</taxonomicName>
has a more conical shell, a lower spire, and denser spiral striation. 
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila somsaki" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="somsaki">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila somsaki</emphasis>
</taxonomicName>
sp. nov. has a less globular shell, less cylindrical (more funnel-shaped) umbilicus, and a less pointed palatal tooth.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is dedicated to and named after Jozef 
<normalizedToken originalValue="Šteffek">Steffek</normalizedToken>
(1952-2013), Slovak malacologist.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is known from the type locality only in Bolikhamsay Province of Laos (Fig. 
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Distribution of Angustopila species." figureDoi="10.3897/zookeys.1147.93824.figure48" httpUri="https://binary.pensoft.net/fig/808857" pageId="0" pageNumber="1">48</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
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