381 lines
46 KiB
XML
381 lines
46 KiB
XML
<document id="3D1DA5C2D53574694A7296651687411A" ID-DOI="10.11646/zootaxa.3835.3.10" ID-GBIF-Dataset="c0146806-b336-4a86-878d-5420ef4c20a8" ID-ISSN="1175-5326" ID-Zenodo-Dep="224747" ID-ZooBank="1ECD1A7F-DE88-4BC6-9190-ABBA6ED9F6EE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460457203439" checkinUser="plazi" docAuthor="Bilton, David T. & Gentili, Elio" docDate="2014" docId="D95C87C36478AA3FF9CAF9A7FB5DFD46" docLanguage="en" docName="zt03835p400.pdf" docOrigin="Zootaxa 3835 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laccobius leopardus Bilton & Gentili, 2014, sp. n." docType="treatment" docVersion="7" lastPageNumber="400" masterDocId="2565FFBB6478AA3CF95DFFF1FFFEFF9F" masterDocTitle="Laccobius leopardus sp. nov. from the Western Cape of South Africa (Coleoptera: Hydrophilidae)" masterLastPageNumber="400" masterPageNumber="397" pageNumber="397" updateTime="1698302428745" updateUser="plazi">
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<mods:mods id="151552E90AB1AC3BEB5F6D895F0C9148" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="8D934E3ED820962CC73170B58AF84C63">
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<mods:title id="3128B9789375C89CFBAFE9BF1975FE2B">Laccobius leopardus sp. nov. from the Western Cape of South Africa (Coleoptera: Hydrophilidae)</mods:title>
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<mods:name id="88B4613683A2A2B3B6CE498FA36D318F" type="personal">
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<mods:roleTerm id="54179D1D51221AA3AEE848F1BF1D92EE">Author</mods:roleTerm>
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<mods:namePart id="B6DC41EB2D2A15BEB8FADB68416B2BDE">Bilton, David T.</mods:namePart>
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</mods:name>
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<mods:name id="5F8F2DB4028E4727640EFB7A202B21F7" type="personal">
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<mods:roleTerm id="3E758E08848230441E9E8444C333CFA0">Author</mods:roleTerm>
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<mods:namePart id="31CFCBF843281CDD487D16BD2F2454D5">Gentili, Elio</mods:namePart>
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<mods:typeOfResource id="A03D3943F1EA098C703C45CF7B90F54B">text</mods:typeOfResource>
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<mods:title id="2FE9814383EE05071AF84C1497526D33">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="0DCC31B04A2E45A73A80F8741DF4A2AF">
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<mods:date id="D58BCAFC5BD1F5E80E002BE6F7D5168D">2014</mods:date>
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<mods:number id="48ED282CDC1220F751E168C82EB04E89">3835</mods:number>
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<mods:detail id="52EAF2A3A60B249B3DDE56BDE38442F4" type="issue">
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<mods:number id="E5F137FAFFE52024957E9AC10B7B2B3F">3</mods:number>
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<mods:identifier id="9F44C5450E28594B9118861ADE9EAE9D" type="DOI">10.11646/zootaxa.3835.3.10</mods:identifier>
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<treatment id="D95C87C36478AA3FF9CAF9A7FB5DFD46" ID-DOI="http://doi.org/10.5281/zenodo.6143317" ID-GBIF-Taxon="119511317" ID-Zenodo-Dep="6143317" LSID="urn:lsid:plazi:treatment:D95C87C36478AA3FF9CAF9A7FB5DFD46" httpUri="http://treatment.plazi.org/id/D95C87C36478AA3FF9CAF9A7FB5DFD46" lastPageId="3" lastPageNumber="400" pageId="0" pageNumber="397">
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<subSubSection id="19EF655E6478AA3CF9CAF9A7FF18F90E" pageId="0" pageNumber="397" type="nomenclature">
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<paragraph id="514A36D56478AA3CF9CAF9A7FE43F9F1" blockId="0.[151,445,1622,1681]" box="[151,445,1622,1646]" pageId="0" pageNumber="397">
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<heading id="0A0281B96478AA3CF9CAF9A7FE43F9F1" bold="true" box="[151,445,1622,1646]" fontSize="10" level="2" pageId="0" pageNumber="397" reason="2">
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<emphasis id="6381EAC76478AA3CF9CAF9A7FE43F9F1" bold="true" box="[151,445,1622,1646]" pageId="0" pageNumber="397">
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<taxonomicName id="96F54D566478AA3CF9CAF9A7FE84F9F1" ID-CoL="3RLVQ" box="[151,378,1622,1646]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="397" phylum="Arthropoda" rank="species" species="leopardus" status="sp. nov.">
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<emphasis id="6381EAC76478AA3CF9CAF9A7FE84F9F1" bold="true" box="[151,378,1622,1646]" italics="true" pageId="0" pageNumber="397">Laccobius leopardus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="78B257BC6478AA3CF8DCF9A6FE43F9F1" box="[385,445,1623,1646]" pageId="0" pageNumber="397" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="514A36D56478AA3CF9CAF98BFF18F90E" blockId="0.[151,445,1622,1681]" box="[151,230,1658,1681]" pageId="0" pageNumber="397">
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(
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<figureCitation id="C9CE2A506478AA3CF9C2F98BFF23F90E" box="[159,221,1658,1681]" captionStart="FIGURE 1" captionStartId="1.[151,255,1855,1878]" captionTargetBox="[351,1224,205,1825]" captionTargetId="figure@1.[345,1230,194,1830]" captionTargetPageId="1" captionText="FIGURE 1. Morphology and habitat of Laccobius species. a) L. leopardus sp. nov. holotype dorsal habitus. b) L. leopardus sp. nov. paratype head and pronotum. c) L. caffer dorsal habitus. d) L. leopardus sp. nov. aedeagus ventral and lateral views. e) L. caffer aedeagus ventral and lateral views. f) L. leopardus sp. nov. type locality: South Africa, Western Cape, Cederberg, Matjies River at Matjiesrivier Reserve ca. 3 km N of Cederberg Oasis (photo: D. T. Bilton). Scale bars a – c = 1 mm; d – e = 500 Μm." httpUri="https://zenodo.org/record/224748/files/figure.png" pageId="0" pageNumber="397">Fig. 1</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="19EF655E6478AA3CF9CAF94EFDD0F839" pageId="0" pageNumber="397" type="materials_examined">
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<paragraph id="514A36D56478AA3CF9CAF94EFC38F966" blockId="0.[151,1436,1727,1958]" pageId="0" pageNumber="397">
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<emphasis id="6381EAC76478AA3CF9CAF94EFED8F949" bold="true" box="[151,294,1727,1750]" pageId="0" pageNumber="397">
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<typeStatus id="8E4E88776478AA3CF9CAF94EFF31F949" box="[151,207,1727,1750]" pageId="0" pageNumber="397">Type</typeStatus>
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locality
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</emphasis>
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.
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<emphasis id="6381EAC76478AA3CF869F94EFE38F949" bold="true" box="[308,454,1727,1750]" pageId="0" pageNumber="397">
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<collectingCountry id="29E276456478AA3CF869F94EFE38F949" box="[308,454,1727,1750]" name="South Africa" pageId="0" pageNumber="397">South Africa</collectingCountry>
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</emphasis>
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: Western Cape: Cederberg, Matjies River at Matjiesrivier Reserve ca.
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<quantity id="960D9B306478AA3CFDE5F94EFB10F949" box="[1208,1262,1727,1750]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="0" pageNumber="397" unit="km" value="3.0">3 km</quantity>
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N of Cederberg Oasis,
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<geoCoordinate id="34C150126478AA3CF982F913FE72F966" box="[223,396,1762,1785]" direction="south" orientation="latitude" pageId="0" pageNumber="397" precision="1" value="-32.519592">32° 31’ 10.53”S</geoCoordinate>
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<geoCoordinate id="34C150126478AA3CF8CEF913FDBDF966" box="[403,579,1762,1785]" direction="east" orientation="longitude" pageId="0" pageNumber="397" precision="1" value="19.35073">19° 21’ 02.63”E</geoCoordinate>
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,
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<quantity id="960D9B306478AA3CFB13F913FD6AF966" box="[590,660,1762,1785]" metricMagnitude="2" metricUnit="m" metricValue="7.26" pageId="0" pageNumber="397" unit="m" value="726.0">726 m</quantity>
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, D.T. Bilton leg. (
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<figureCitation id="C9CE2A506478AA3CFA0EF913FC52F966" box="[851,940,1762,1785]" captionStart="FIGURE 1" captionStartId="1.[151,255,1855,1878]" captionTargetBox="[351,1224,205,1825]" captionTargetId="figure@1.[345,1230,194,1830]" captionTargetPageId="1" captionText="FIGURE 1. Morphology and habitat of Laccobius species. a) L. leopardus sp. nov. holotype dorsal habitus. b) L. leopardus sp. nov. paratype head and pronotum. c) L. caffer dorsal habitus. d) L. leopardus sp. nov. aedeagus ventral and lateral views. e) L. caffer aedeagus ventral and lateral views. f) L. leopardus sp. nov. type locality: South Africa, Western Cape, Cederberg, Matjies River at Matjiesrivier Reserve ca. 3 km N of Cederberg Oasis (photo: D. T. Bilton). Scale bars a – c = 1 mm; d – e = 500 Μm." httpUri="https://zenodo.org/record/224748/files/figure.png" pageId="0" pageNumber="397">Figure 1</figureCitation>
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f).
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</paragraph>
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<paragraph id="514A36D56478AA3CF99AF8F5FDD0F839" blockId="0.[151,1436,1727,1958]" pageId="0" pageNumber="397">
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<emphasis id="6381EAC76478AA3CF99AF8F5FE8FF884" bold="true" box="[199,369,1796,1819]" pageId="0" pageNumber="397">
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<typeStatus id="8E4E88776478AA3CF99AF8F5FF01F884" box="[199,255,1796,1819]" pageId="0" pageNumber="397">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="8E4E88776478AA3CF826F8F5FE20F884" box="[379,478,1796,1819]" pageId="0" pageNumber="397" type="holotype">Holotype</typeStatus>
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(male): “
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||
<date id="254B10156478AA3CFB17F8F5FD19F884" box="[586,743,1796,1819]" pageId="0" pageNumber="397" value="2011-09-22" valueMax="2011-09-25" valueMin="2011-09-22">22–25/ix/2011</date>
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<collectingCountry id="29E276456478AA3CFBAFF8F5FC81F884" box="[754,895,1796,1819]" name="South Africa" pageId="0" pageNumber="397">South Africa</collectingCountry>
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||
WC, Cederberg—Matjiesrivier Reserve, Matjies River ca.
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<quantity id="960D9B306478AA3CF85DF8D6FEC9F8A1" box="[256,311,1831,1854]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="0" pageNumber="397" unit="km" value="3.0">3 km</quantity>
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||
N of, Cederberg Oasis D. T. Bilton leg.” (genitalia extracted and mounted in
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<collectionCode id="37E4AE106478AA3CFDDCF8D6FB30F8A1" box="[1153,1230,1831,1854]" pageId="0" pageNumber="397">DMHF</collectionCode>
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on same card) and red
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||
<typeStatus id="8E4E88776478AA3CF99FF8BBFEE1F8FE" box="[194,287,1866,1889]" pageId="0" pageNumber="397" type="holotype">holotype</typeStatus>
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label (
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<collectionCode id="37E4AE106478AA3CF833F8BBFE49F8FE" box="[366,439,1866,1889]" pageId="0" pageNumber="397">ISAM</collectionCode>
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).
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<emphasis id="6381EAC76478AA3CF896F8BBFDC3F8FE" bold="true" box="[459,573,1866,1889]" pageId="0" pageNumber="397">
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||
<typeStatus id="8E4E88776478AA3CF896F8BBFDC3F8FE" box="[459,573,1866,1889]" pageId="0" pageNumber="397" type="paratype">Paratypes</typeStatus>
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</emphasis>
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||
(9):
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<specimenCount id="47F3FD5C6478AA3CFB2BF8BBFD5CF8FE" box="[630,674,1866,1889]" pageId="0" pageNumber="397" type="generic">3 ♂</specimenCount>
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, 4 ♀ Same data as
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<typeStatus id="8E4E88776478AA3CFA22F8BBFC22F8FE" box="[895,988,1866,1889]" pageId="0" pageNumber="397" type="holotype">holotype</typeStatus>
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(
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<collectionCode id="37E4AE106478AA3CFAB0F8BBFBC6F8FE" box="[1005,1080,1866,1889]" pageId="0" pageNumber="397">CDTB</collectionCode>
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,
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<collectionCode id="37E4AE106478AA3CFD1BF8BBFB80F8FE" box="[1094,1150,1866,1889]" pageId="0" pageNumber="397">CGV</collectionCode>
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,
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<collectionCode id="37E4AE106478AA3CFDD7F8BBFB2DF8FE" box="[1162,1235,1866,1889]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="0" pageNumber="397">NMW</collectionCode>
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); 2 ♀ “
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<date id="254B10156478AA3CFC7BF8BBFA65F8FE" box="[1318,1435,1866,1889]" pageId="0" pageNumber="397" value="2010-09-21">21/ix/2010</date>
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<collectingCountry id="29E276456478AA3CF9CAF89DFEDCF81C" box="[151,290,1900,1923]" name="South Africa" pageId="0" pageNumber="397">South Africa</collectingCountry>
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WC, Cederberg—Matjiesrivier Reserve, Matjies River ca.
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<quantity id="960D9B306478AA3CFAF0F89DFC18F81C" box="[941,998,1900,1923]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="0" pageNumber="397" unit="km" value="3.0">3 km</quantity>
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N of, Cederberg Oasis D. T. Bilton leg (
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<collectionCode id="37E4AE106478AA3CF9C2F87EFF13F839" box="[159,237,1935,1958]" pageId="0" pageNumber="397">CDTB</collectionCode>
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).” All with red
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<typeStatus id="8E4E88776478AA3CF8CDF87EFE12F839" box="[400,492,1935,1958]" pageId="0" pageNumber="397" type="paratype">paratype</typeStatus>
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label.
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</paragraph>
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</subSubSection>
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<caption id="058A665D6479AA3DF9CAF8CEFE3EF87E" httpUri="https://zenodo.org/record/224748/files/figure.png" pageId="1" pageNumber="398" targetBox="[351,1224,205,1825]" targetPageId="1">
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<paragraph id="514A36D56479AA3DF9CAF8CEFE3EF87E" blockId="1.[151,1436,1855,2017]" pageId="1" pageNumber="398">
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||
<emphasis id="6381EAC76479AA3DF9CAF8CEFEE0F8C9" bold="true" box="[151,286,1855,1878]" pageId="1" pageNumber="398">FIGURE 1.</emphasis>
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Morphology and habitat of
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<taxonomicName id="96F54D566479AA3DFB3DF8CEFD33F8C9" box="[608,717,1855,1878]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="genus">
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||
<emphasis id="6381EAC76479AA3DFB3DF8CEFD33F8C9" box="[608,717,1855,1878]" italics="true" pageId="1" pageNumber="398">Laccobius</emphasis>
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</taxonomicName>
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species. a)
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<taxonomicName id="96F54D566479AA3DFA0AF8CEFC1FF8C9" box="[855,993,1855,1878]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="species" species="leopardus" status="sp. nov.">
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<emphasis id="6381EAC76479AA3DFA0AF8CEFC1FF8C9" box="[855,993,1855,1878]" italics="true" pageId="1" pageNumber="398">L. leopardus</emphasis>
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</taxonomicName>
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||
<emphasis id="6381EAC76479AA3DFAB1F8CEFBB8F8C9" bold="true" box="[1004,1094,1855,1878]" pageId="1" pageNumber="398">
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||
<taxonomicNameLabel id="78B257BC6479AA3DFAB1F8CEFBB8F8C9" box="[1004,1094,1855,1878]" pageId="1" pageNumber="398" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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holotype dorsal habitus. b)
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<taxonomicName id="96F54D566479AA3DFCDBF8CEFEFFF8E6" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="species" species="leopardus" status="sp. nov.">
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<emphasis id="6381EAC76479AA3DFCDBF8CEFEFFF8E6" italics="true" pageId="1" pageNumber="398">L. leopardus</emphasis>
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</taxonomicName>
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||
<emphasis id="6381EAC76479AA3DF85AF893FEA2F8E6" bold="true" box="[263,348,1890,1913]" pageId="1" pageNumber="398">
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||
<taxonomicNameLabel id="78B257BC6479AA3DF85AF893FEA2F8E6" box="[263,348,1890,1913]" pageId="1" pageNumber="398" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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paratype head and pronotum. c)
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<taxonomicName id="96F54D566479AA3DFBE4F893FCECF8E6" box="[697,786,1890,1913]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="species" species="caffer">
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||
<emphasis id="6381EAC76479AA3DFBE4F893FCECF8E6" box="[697,786,1890,1913]" italics="true" pageId="1" pageNumber="398">L. caffer</emphasis>
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||
</taxonomicName>
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||
dorsal habitus. d)
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<taxonomicName id="96F54D566479AA3DFA8BF893FBA2F8E6" box="[982,1116,1890,1913]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="species" species="leopardus" status="sp. nov.">
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||
<emphasis id="6381EAC76479AA3DFA8BF893FBA2F8E6" box="[982,1116,1890,1913]" italics="true" pageId="1" pageNumber="398">L. leopardus</emphasis>
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||
</taxonomicName>
|
||
<emphasis id="6381EAC76479AA3DFD3FF893FB49F8E6" bold="true" box="[1122,1207,1890,1913]" pageId="1" pageNumber="398">
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||
<taxonomicNameLabel id="78B257BC6479AA3DFD3FF893FB49F8E6" box="[1122,1207,1890,1913]" pageId="1" pageNumber="398" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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aedeagus ventral and lateral views. e)
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<taxonomicName id="96F54D566479AA3DF81BF875FE5EF804" box="[326,416,1924,1947]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="species" species="caffer">
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||
<emphasis id="6381EAC76479AA3DF81BF875FE5EF804" box="[326,416,1924,1947]" italics="true" pageId="1" pageNumber="398">L. caffer</emphasis>
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||
</taxonomicName>
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||
aedeagus ventral and lateral views. f)
|
||
<taxonomicName id="96F54D566479AA3DFA6AF875FC43F804" box="[823,957,1924,1947]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="species" species="leopardus" status="sp. nov.">
|
||
<emphasis id="6381EAC76479AA3DFA6AF875FC43F804" box="[823,957,1924,1947]" italics="true" pageId="1" pageNumber="398">L. leopardus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6381EAC76479AA3DFA9EF875FBE6F804" bold="true" box="[963,1048,1924,1947]" pageId="1" pageNumber="398">
|
||
<taxonomicNameLabel id="78B257BC6479AA3DFA9EF875FBE6F804" box="[963,1048,1924,1947]" pageId="1" pageNumber="398" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
type locality: South Africa, Western Cape, Cederberg, Matjies River at Matjiesrivier Reserve ca. 3 km N of Cederberg Oasis (photo: D.T. Bilton). Scale bars a–c = 1 mm; d–e = 500 Μm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="19EF655E647AAA3FF99AFF69FE32FF4E" lastPageId="3" lastPageNumber="400" pageId="2" pageNumber="399" type="description">
|
||
<paragraph id="514A36D5647AAA3EF99AFF69FB9FFE3E" blockId="2.[151,1436,152,2012]" pageId="2" pageNumber="399">
|
||
<emphasis id="6381EAC7647AAA3EF99AFF69FEB5FF30" bold="true" box="[199,331,152,175]" pageId="2" pageNumber="399">Description</emphasis>
|
||
. Size:
|
||
<typeStatus id="8E4E8877647AAA3EF8C9FF69FE05FF30" box="[404,507,152,175]" pageId="2" pageNumber="399" type="holotype">Holotype</typeStatus>
|
||
: TL =
|
||
<quantity id="960D9B30647AAA3EFB1FFF69FD68FF30" box="[578,662,152,175]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="399" unit="mm" value="3.5">3.5 mm</quantity>
|
||
, MW =
|
||
<quantity id="960D9B30647AAA3EFBB3FF69FCAEFF30" box="[750,848,152,175]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="2" pageNumber="399" unit="mm" value="1.65">1.65 mm</quantity>
|
||
;
|
||
<typeStatus id="8E4E8877647AAA3EFA06FF69FC39FF30" box="[859,967,152,175]" pageId="2" pageNumber="399" type="paratype">Paratypes</typeStatus>
|
||
: TL =
|
||
<quantity id="960D9B30647AAA3EFD50FF69FB6EFF30" box="[1037,1168,152,175]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.7" metricValueMin="3.4" pageId="2" pageNumber="399" unit="mm" value="3.55" valueMax="3.7" valueMin="3.4">3.4–3.7 mm</quantity>
|
||
, MW =
|
||
<quantity id="960D9B30647AAA3EFDB5FF69FA78FF30" box="[1256,1414,152,175]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.75" metricValueMin="1.65" pageId="2" pageNumber="399" unit="mm" value="1.7" valueMax="1.75" valueMin="1.65">1.65–1.75 mm</quantity>
|
||
. Colour: Head, including labrum, dark brown with aeneous purplish-green reflections. Distinct pale preocular spots extending from front margins of eyes to frontal angles of clypeus. Pronotum pale straw yellow-yellowish brown, with approximately parallel-sided dark central patch occupying discal 1/3, and broadly reaching front and hind margins. Elytra straw yellow, with diffuse darker patches on disc, and darker rings of pigment around punctures. Venter black, with exception of preocular region of head, pronotal hypomeron and elytral epipleurs and pseudepipleurs, which are straw yellow as upperside, and lateral areas of abdominal ventrites 2–5 which are paler brown. Legs, antennae and palpi straw yellow; trace of infuscation visible on last segment of maxillary palp (see
|
||
<figureCitation id="C9CE2A50647AAA3EFAB1FE7BFBD8FE3E" box="[1004,1062,394,417]" captionStart="FIGURE 1" captionStartId="1.[151,255,1855,1878]" captionTargetBox="[351,1224,205,1825]" captionTargetId="figure@1.[345,1230,194,1830]" captionTargetPageId="1" captionText="FIGURE 1. Morphology and habitat of Laccobius species. a) L. leopardus sp. nov. holotype dorsal habitus. b) L. leopardus sp. nov. paratype head and pronotum. c) L. caffer dorsal habitus. d) L. leopardus sp. nov. aedeagus ventral and lateral views. e) L. caffer aedeagus ventral and lateral views. f) L. leopardus sp. nov. type locality: South Africa, Western Cape, Cederberg, Matjies River at Matjiesrivier Reserve ca. 3 km N of Cederberg Oasis (photo: D. T. Bilton). Scale bars a – c = 1 mm; d – e = 500 Μm." httpUri="https://zenodo.org/record/224748/files/figure.png" pageId="2" pageNumber="399">Fig. 1</figureCitation>
|
||
a–b).
|
||
</paragraph>
|
||
<paragraph id="514A36D5647AAA3EF99AFE5CFA95FDB3" blockId="2.[151,1436,152,2012]" pageId="2" pageNumber="399">Head: Labrum curved at lateral margins, but broadly truncately rounded at front edge. Upper surface of head, including labrum, shining and distinctly shagreened, with sparse, coarse shallow punctures. Frontoclypeal suture distinct, with coarse punctures posterior to suture and inside each eye, each puncture bearing a long, white decumbent hair-like seta. Eyes oval, moderately large, occupying 1/3 side margin of head. Inter-ocular distance ca. 3.2x diameter of eye.</paragraph>
|
||
<paragraph id="514A36D5647AAA3EF99AFDC9FB48FD46" blockId="2.[151,1436,152,2012]" pageId="2" pageNumber="399">
|
||
Pronotum: Transverse; approximately
|
||
<date id="254B1015647AAA3EFB3AFDC9FD7CFDD0" box="[615,642,568,591]" pageId="2" pageNumber="399">3x</date>
|
||
broader than long. Broadly rounded at sides, with acutely rounded front angles, and broadly rounded hind angles. Front margin broadly bisinuate around centre; hind margin weakly curved. Upper surface shagreened, this microsculpture being slightly less impressed than on head, and most evident over central dark patch. With sparse, coarse shallow punctures throughout; these punctures being shallower than those on head. Larger punctures bearing long decumbent hair-like setae, visible particularly towards side margins.
|
||
</paragraph>
|
||
<paragraph id="514A36D5647AAA3EF99AFD14FA62FCFB" blockId="2.[151,1436,152,2012]" pageId="2" pageNumber="399">
|
||
Elytra: Elongate oval, subparallel in basal half and evenly rounded to apex; widest at middle. Elytral punctures rather irregular, some traces of rows on elytral disc. Each puncture bearing a seta. On disc setae of punctures short; not reaching beyond hind margin of puncture. Setae much longer at sides and in apical ½, where many are 5–
|
||
<date id="254B1015647AAA3EFC46FCDBFAD6FCDE" box="[1307,1320,810,833]" pageId="2" pageNumber="399">6</date>
|
||
x puncture diameter in length. Elytral margin with stout bristles on shoulder, and similar, slightly longer, stout bristles in apical half.
|
||
</paragraph>
|
||
<paragraph id="514A36D5647AAA3EF99AFC81FC11FAF3" blockId="2.[151,1436,152,2012]" pageId="2" pageNumber="399">Ventral side: Labrum without speculae, front margin broadly curved, and overhanging oral cavity, especially in the centre. Distinct circular apicomedian emargination visible on front margin of labrum, this occupying the downcurved upper surface and front labral edge. Ventral surface of labrum somewhat excavated either side of apicomedian emargination. Apicomedian emargination with a cluster of short, golden setae. Mentum rectangular, shining, lacking microsculpture, and with fine, scattered punctures. Submentum with similar surface properties. Gula and genae microreticulate, dull, with hydrofuge pubescence. Genae rugosely punctate towards lateral margins of head, with distinct field of long, golden, curved, hair-like erect postocular setae. Prosternum roof-like, with a central longitudinal keel. Surface punctate, with hydrofuge pubescence. Mesoventral keel with a distinct hooked anterior tooth. Tooth surrounded by broad lateral expansions, furnished with long white hairs. Keel narrowed and linear between mesocoxae. Metaventrite pubescent, with exception of a glabrous central section. Hairs thinner towards posterior margin. Glabrous central section with distinct longitudinal furrow occupying posterior ¼ of metaventrite. Pubescence overlying distinct micoreticulation; imbricate and isodiametric anteriorly, transverse posteriorly. Abdominal ventrites 1–5 shining, with sparse, coarse punctures, some bearing long golden setae arranged in loose transverse rows. Setae 1/3–3/4 of segment length. Abdominal ventrites broadly excavated laterally. Ventrite 5 with denser punctation; some punctures situated in short transverse furrows. Ventrite 6 dull, pubescent, with transverse microreticulation.</paragraph>
|
||
<paragraph id="514A36D5647AAA3EF99AFA89FBC7F986" blockId="2.[151,1436,152,2012]" pageId="2" pageNumber="399">
|
||
Legs: Moderately long and slender. Segments 2–3 of male protarsi expanded, with smooth sucker plate and modified bristle fields, typical of the genus.
|
||
<collectingCountry id="29E27645647AAA3EFB58FA6BFDD4FA2E" box="[517,554,1434,1457]" name="Palau" pageId="2" pageNumber="399">Pro</collectingCountry>
|
||
and mesotibiae straight; posterior tibiae evenly curved. All tibiae strongly spinose. Procoxae, protrochanters and base of profemora with hydrofuge pubescence. Mesocoxae with hydrofuge pubescence, mesotrochanters and anterobasal portion of mesofemora with scattered long setae. Metacoxae glabrous. Base and posterior margins of metatrochanters and base of metafemora with stout hair-like setae.
|
||
</paragraph>
|
||
<paragraph id="514A36D5647AAA3EF99AF9D4FC30F91E" blockId="2.[151,1436,152,2012]" pageId="2" pageNumber="399">Aedeagus: Parameres 2.3x length of phallobase. Paramere apices elongate, finger-like, extending beyond apex of median lobe. Median lobe with broadly rounded apex, and distinct, broad semicircular excavation on ventral face. Minutely dentate internal sac structures visible at base of this excavated area.</paragraph>
|
||
<paragraph id="514A36D5647AAA3EF99AF97CFE02F958" blockId="2.[151,1436,152,2012]" pageId="2" pageNumber="399">Female: Concavity of abdominal ventrites weaker than in males. Foretarsi unmodified. Metatrochanters with much fewer, smaller setae than in male.</paragraph>
|
||
<paragraph id="514A36D5647AAA3EF99AF923FF02F8B0" blockId="2.[151,1436,152,2012]" pageId="2" pageNumber="399">
|
||
Variability: In addition to size differences, specimens vary somewhat in the extent of the dark central patch on the pronotum (
|
||
<figureCitation id="C9CE2A50647AAA3EF850F904FEB6F893" box="[269,328,1781,1804]" captionStart="FIGURE 1" captionStartId="1.[151,255,1855,1878]" captionTargetBox="[351,1224,205,1825]" captionTargetId="figure@1.[345,1230,194,1830]" captionTargetPageId="1" captionText="FIGURE 1. Morphology and habitat of Laccobius species. a) L. leopardus sp. nov. holotype dorsal habitus. b) L. leopardus sp. nov. paratype head and pronotum. c) L. caffer dorsal habitus. d) L. leopardus sp. nov. aedeagus ventral and lateral views. e) L. caffer aedeagus ventral and lateral views. f) L. leopardus sp. nov. type locality: South Africa, Western Cape, Cederberg, Matjies River at Matjiesrivier Reserve ca. 3 km N of Cederberg Oasis (photo: D. T. Bilton). Scale bars a – c = 1 mm; d – e = 500 Μm." httpUri="https://zenodo.org/record/224748/files/figure.png" pageId="2" pageNumber="399">Fig. 1</figureCitation>
|
||
a–b) and the extent of dark mottling on the elytra, which can be slightly more extensive than seen in the
|
||
<typeStatus id="8E4E8877647AAA3EF9CAF8E9FF06F8B0" box="[151,248,1816,1839]" pageId="2" pageNumber="399" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="514A36D5647AAA3FF99AF8CBFE32FF4E" blockId="2.[151,1436,152,2012]" lastBlockId="3.[151,1437,152,729]" lastPageId="3" lastPageNumber="400" pageId="2" pageNumber="399">
|
||
<emphasis id="6381EAC7647AAA3EF99AF8CBFE3CF8CE" bold="true" box="[199,450,1850,1873]" pageId="2" pageNumber="399">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="96F54D56647AAA3EF897F8CBFDAFF8CE" box="[458,593,1850,1873]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="leopardus" status="sp. nov.">
|
||
<emphasis id="6381EAC7647AAA3EF897F8CBFDAFF8CE" box="[458,593,1850,1873]" italics="true" pageId="2" pageNumber="399">L. leopardus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6381EAC7647AAA3EFB04F8CBFD4EF8CE" bold="true" box="[601,688,1850,1873]" pageId="2" pageNumber="399">
|
||
<taxonomicNameLabel id="78B257BC647AAA3EFB04F8CBFD4EF8CE" box="[601,688,1850,1873]" pageId="2" pageNumber="399" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is morphologically closest to
|
||
<taxonomicName id="96F54D56647AAA3EFAA5F8CBFB55F8CE" box="[1016,1195,1850,1873]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="caffer">
|
||
<emphasis id="6381EAC7647AAA3EFAA5F8CBFB55F8CE" box="[1016,1195,1850,1873]" italics="true" pageId="2" pageNumber="399">Laccobius caffer</emphasis>
|
||
</taxonomicName>
|
||
, from which it can be distinguished by its larger size (TL
|
||
<quantity id="960D9B30647AAA3EFB44F8ACFD65F8EB" box="[537,667,1885,1908]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.7" metricValueMin="3.4" pageId="2" pageNumber="399" unit="mm" value="3.55" valueMax="3.7" valueMin="3.4">3.4–3.7 mm</quantity>
|
||
vs.
|
||
<quantity id="960D9B30647AAA3EFB95F8ACFCB2F8EB" box="[712,844,1885,1908]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="3.1" metricValueMin="2.5" pageId="2" pageNumber="399" unit="mm" value="2.8" valueMax="3.1" valueMin="2.5">2.5–3.1 mm</quantity>
|
||
, see
|
||
<bibRefCitation id="35644B24647AAA3EFADEF8ACFBEDF8EB" author="Gentili" box="[899,1043,1885,1908]" pageId="2" pageNumber="399" refString="Gentili, E. (1981) Il genere Laccobius nella regione Etiopica (Coleoptera, Hydrophilidae). Annuario Ossevatorio Fisica Terrestre Museo Antonio Stoppani del Seminario Arcivescovile di Milano (N. S.), 3, 31 - 42." type="journal article" year="1981">Gentili, 1981</bibRefCitation>
|
||
), darker dorsal pigmentation, with a broader and typically more parallel-sided central dark patch on the pronotum (see
|
||
<figureCitation id="C9CE2A50647AAA3EFD78F871FBA2F808" box="[1061,1116,1920,1943]" captionStart="FIGURE 1" captionStartId="1.[151,255,1855,1878]" captionTargetBox="[351,1224,205,1825]" captionTargetId="figure@1.[345,1230,194,1830]" captionTargetPageId="1" captionText="FIGURE 1. Morphology and habitat of Laccobius species. a) L. leopardus sp. nov. holotype dorsal habitus. b) L. leopardus sp. nov. paratype head and pronotum. c) L. caffer dorsal habitus. d) L. leopardus sp. nov. aedeagus ventral and lateral views. e) L. caffer aedeagus ventral and lateral views. f) L. leopardus sp. nov. type locality: South Africa, Western Cape, Cederberg, Matjies River at Matjiesrivier Reserve ca. 3 km N of Cederberg Oasis (photo: D. T. Bilton). Scale bars a – c = 1 mm; d – e = 500 Μm." httpUri="https://zenodo.org/record/224748/files/figure.png" pageId="2" pageNumber="399">Fig 1</figureCitation>
|
||
a–c). In addition the anterior labral margin is more evenly curved and less truncate dorsally in examined specimens of
|
||
<taxonomicName id="96F54D56647AAA3EFD34F853FB3AF826" box="[1129,1220,1954,1977]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="caffer">
|
||
<emphasis id="6381EAC7647AAA3EFD34F853FB3AF826" box="[1129,1220,1954,1977]" italics="true" pageId="2" pageNumber="399">L. caffer</emphasis>
|
||
</taxonomicName>
|
||
, and the abdominal ventrites have more well developed paler patches in this species. In
|
||
<taxonomicName id="96F54D56647AAA3EFA36F834FC3BF843" box="[875,965,1989,2012]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="caffer">
|
||
<emphasis id="6381EAC7647AAA3EFA36F834FC3BF843" box="[875,965,1989,2012]" italics="true" pageId="2" pageNumber="399">L. caffer</emphasis>
|
||
</taxonomicName>
|
||
the margins of abdominal ventrites 2–4, the posterior margin of ventrite 4 and much of abdominal ventrite 5 are pale yellow. The two species also differ significantly in male genitalia (
|
||
<figureCitation id="C9CE2A50647BAA3FF80AFF4BFE6CFF4E" box="[343,402,186,209]" captionStart="FIGURE 1" captionStartId="1.[151,255,1855,1878]" captionTargetBox="[351,1224,205,1825]" captionTargetId="figure@1.[345,1230,194,1830]" captionTargetPageId="1" captionText="FIGURE 1. Morphology and habitat of Laccobius species. a) L. leopardus sp. nov. holotype dorsal habitus. b) L. leopardus sp. nov. paratype head and pronotum. c) L. caffer dorsal habitus. d) L. leopardus sp. nov. aedeagus ventral and lateral views. e) L. caffer aedeagus ventral and lateral views. f) L. leopardus sp. nov. type locality: South Africa, Western Cape, Cederberg, Matjies River at Matjiesrivier Reserve ca. 3 km N of Cederberg Oasis (photo: D. T. Bilton). Scale bars a – c = 1 mm; d – e = 500 Μm." httpUri="https://zenodo.org/record/224748/files/figure.png" pageId="3" pageNumber="400">Fig. 1</figureCitation>
|
||
d–e).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="19EF655E647BAA3FF99AFF2CFB35FEC3" pageId="3" pageNumber="400" type="distribution">
|
||
<paragraph id="514A36D5647BAA3FF99AFF2CFB35FEC3" blockId="3.[151,1437,152,729]" pageId="3" pageNumber="400">
|
||
<emphasis id="6381EAC7647BAA3FF99AFF2CFEACFF6B" bold="true" box="[199,338,221,244]" pageId="3" pageNumber="400">Distribution</emphasis>
|
||
. To date only known from the
|
||
<typeStatus id="8E4E8877647BAA3FFBF7FF2CFD26FF6B" box="[682,728,221,244]" pageId="3" pageNumber="400">type</typeStatus>
|
||
locality, the sandy gravelly margins of the Matjies River, on the edge of the Cederberg range in the Western Cape Province,
|
||
<collectingCountry id="29E27645647BAA3FFA79FEF1FC4DFE88" box="[804,947,256,279]" name="South Africa" pageId="3" pageNumber="400">South Africa</collectingCountry>
|
||
. This locality is situated in a region which is somewhat transitional between the fynbos and succulent karoo biomes (Murcinia &
|
||
<bibRefCitation id="35644B24647BAA3FFD45FED3FB2AFEA6" author="Rutherford" box="[1048,1236,290,313]" pageId="3" pageNumber="400" refString="Murcina, L. & Rutherford, M. C. (2006) The vegetation of South Africa, Lesotho and Swaziland. Strelitzia, 19, 1 - 807." type="journal article" year="2006">Rutherford, 2006</bibRefCitation>
|
||
); fynbos occurring at higher altitudes, lower altitude sites such as the
|
||
<typeStatus id="8E4E8877647BAA3FFBF2FEB4FD23FEC3" box="[687,733,325,348]" pageId="3" pageNumber="400">type</typeStatus>
|
||
locality being surrounded by succulent karoo.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="19EF655E647BAA3FF99AFE99FB5DFD46" pageId="3" pageNumber="400" type="etymology">
|
||
<paragraph id="514A36D5647BAA3FF99AFE99FEF3FE5B" blockId="3.[151,1437,152,729]" pageId="3" pageNumber="400">
|
||
<emphasis id="6381EAC7647BAA3FF99AFE99FEBEFEE0" bold="true" box="[199,320,360,383]" pageId="3" pageNumber="400">Etymology</emphasis>
|
||
. Named in reference to the leopards (
|
||
<taxonomicName id="96F54D56647BAA3FFB8EFE99FBA7FEE0" authority="Linnaeus" authorityName="Linnaeus" box="[723,1113,360,383]" class="Mammalia" family="Felidae" genus="Panthera" kingdom="Animalia" order="Carnivora" pageId="3" pageNumber="400" phylum="Chordata" rank="subSpecies" species="pardus" subSpecies="pardus">
|
||
<emphasis id="6381EAC7647BAA3FFB8EFE99FC2BFEE0" box="[723,981,360,383]" italics="true" pageId="3" pageNumber="400">Panthera pardus pardus</emphasis>
|
||
(Linnaeus))
|
||
</taxonomicName>
|
||
which frequent the Cederberg Conservancy in which the
|
||
<typeStatus id="8E4E8877647BAA3FF89CFE7BFE11FE3E" box="[449,495,394,417]" pageId="3" pageNumber="400">type</typeStatus>
|
||
locality is situated, and the spotted appearance of the elytra. The name is a noun in apposition.
|
||
</paragraph>
|
||
<paragraph id="514A36D5647BAA3FF99AFE21FB5DFD46" blockId="3.[151,1437,152,729]" pageId="3" pageNumber="400">
|
||
<emphasis id="6381EAC7647BAA3FF99AFE21FEDEFE78" bold="true" box="[199,288,464,487]" pageId="3" pageNumber="400">Ecology</emphasis>
|
||
. Specimens were puddled by hand from marginal sediment in shallow river backwaters (
|
||
<figureCitation id="C9CE2A50647BAA3FFDAAFE21FACBFE78" box="[1271,1333,464,487]" captionStart="FIGURE 1" captionStartId="1.[151,255,1855,1878]" captionTargetBox="[351,1224,205,1825]" captionTargetId="figure@1.[345,1230,194,1830]" captionTargetPageId="1" captionText="FIGURE 1. Morphology and habitat of Laccobius species. a) L. leopardus sp. nov. holotype dorsal habitus. b) L. leopardus sp. nov. paratype head and pronotum. c) L. caffer dorsal habitus. d) L. leopardus sp. nov. aedeagus ventral and lateral views. e) L. caffer aedeagus ventral and lateral views. f) L. leopardus sp. nov. type locality: South Africa, Western Cape, Cederberg, Matjies River at Matjiesrivier Reserve ca. 3 km N of Cederberg Oasis (photo: D. T. Bilton). Scale bars a – c = 1 mm; d – e = 500 Μm." httpUri="https://zenodo.org/record/224748/files/figure.png" pageId="3" pageNumber="400">Fig. 1</figureCitation>
|
||
f), which were mostly without appreciable surface flow at the time of sampling, but would clearly be connected to the main river at times of higher flow. They occurred together with the water beetles
|
||
<taxonomicName id="96F54D56647BAA3FFA2EFDE4FAFDFDB3" authority="Omer-Cooper" authorityName="Omer-Cooper" box="[883,1283,533,556]" class="Insecta" family="Dytiscidae" genus="Hydropeplus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="montanus">
|
||
<emphasis id="6381EAC7647BAA3FFA2EFDE4FB97FDB3" box="[883,1129,533,556]" italics="true" pageId="3" pageNumber="400">Hydropeplus montanus</emphasis>
|
||
Omer-Cooper
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="96F54D56647BAA3FFC4DFDE4FE72FDD0" authority="Boheman" authorityName="Boheman" class="Insecta" family="Dytiscidae" genus="Herophydrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="inquinatus">
|
||
<emphasis id="6381EAC7647BAA3FFC4DFDE4FEF6FDD0" italics="true" pageId="3" pageNumber="400">Herophydrus inquinatus</emphasis>
|
||
(Boheman)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="96F54D56647BAA3FF8C6FDC9FD36FDD0" authority="Bilton" authorityName="Bilton" box="[411,712,568,591]" class="Insecta" family="Dytiscidae" genus="Sharphydrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="brincki">
|
||
<emphasis id="6381EAC7647BAA3FF8C6FDC9FD86FDD0" box="[411,632,568,591]" italics="true" pageId="3" pageNumber="400">Sharphydrus brincki</emphasis>
|
||
Bilton
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="96F54D56647BAA3FFB8AFDC9FC19FDD0" authority="Regimbart" authorityName="Regimbart" box="[727,999,568,591]" class="Insecta" family="Dytiscidae" genus="Yola" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="frontalis">
|
||
<emphasis id="6381EAC7647BAA3FFB8AFDC9FC97FDD0" box="[727,873,568,591]" italics="true" pageId="3" pageNumber="400">Yola frontalis</emphasis>
|
||
Régimbart
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="96F54D56647BAA3FFAA8FDC9FA8FFDD0" authority="Boheman" authorityName="Boheman" box="[1013,1393,568,591]" class="Insecta" family="Dytiscidae" genus="Hydroglyphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="infirmus">
|
||
<emphasis id="6381EAC7647BAA3FFAA8FDC9FB11FDD0" box="[1013,1263,568,591]" italics="true" pageId="3" pageNumber="400">Hydroglyphus infirmus</emphasis>
|
||
(Boheman)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="96F54D56647BAA3FFCDFFDC9FE72FDEE" authority="Boheman" authorityName="Boheman" class="Insecta" family="Dytiscidae" genus="Hydroglyphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="lineolatus">
|
||
<emphasis id="6381EAC7647BAA3FFCDFFDC9FEFEFDEE" italics="true" pageId="3" pageNumber="400">H. lineolatus</emphasis>
|
||
(Boheman)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="96F54D56647BAA3FF8FBFDABFCB8FDEE" authority="Omer-Cooper" authorityName="Omer-Cooper" box="[422,838,602,625]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="guignoti">
|
||
<emphasis id="6381EAC7647BAA3FF8FBFDABFD63FDEE" box="[422,669,602,625]" italics="true" pageId="3" pageNumber="400">Canthyporus guignoti</emphasis>
|
||
Omer-Cooper
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="96F54D56647BAA3FFA03FDABFBACFDEE" authority="Guignot" authorityName="Guignot" box="[862,1106,602,625]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="petulans">
|
||
<emphasis id="6381EAC7647BAA3FFA03FDABFC1BFDEE" box="[862,997,602,625]" italics="true" pageId="3" pageNumber="400">C. petulans</emphasis>
|
||
Guignot
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="96F54D56647BAA3FFD37FDABFEE5FD0B" authority="Regimbart" authorityName="Regimbart" class="Insecta" family="Dytiscidae" genus="Philaccolus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="lineatoguttatus">
|
||
<emphasis id="6381EAC7647BAA3FFD37FDABFA65FDEE" box="[1130,1435,602,625]" italics="true" pageId="3" pageNumber="400">Philaccolus lineatoguttatus</emphasis>
|
||
(Régimbart)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="96F54D56647BAA3FF875FD8CFDB5FD0B" authority="Aube" authorityName="Aube" box="[296,587,637,660]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="lineatus">
|
||
<emphasis id="6381EAC7647BAA3FF875FD8CFDF9FD0B" box="[296,519,637,660]" italics="true" pageId="3" pageNumber="400">Laccophilus lineatus</emphasis>
|
||
Aubé
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="96F54D56647BAA3FFB05FD8CFD39FD0B" box="[600,711,637,660]" class="Insecta" family="Georyssidae" genus="Georyssus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="400" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6381EAC7647BAA3FFB05FD8CFD39FD0B" box="[600,711,637,660]" italics="true" pageId="3" pageNumber="400">Georyssus</emphasis>
|
||
</taxonomicName>
|
||
spp.,
|
||
<taxonomicName id="96F54D56647BAA3FFA57FD8CFBABFD0B" authority="Boheman" authorityName="Boheman" box="[778,1109,637,660]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="punctulatus">
|
||
<emphasis id="6381EAC7647BAA3FFA57FD8CFC1AFD0B" box="[778,996,637,660]" italics="true" pageId="3" pageNumber="400">Berosus punctulatus</emphasis>
|
||
Boheman
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="96F54D56647BAA3FFD3FFD8CFA84FD0B" authority="Perris" authorityName="Perris" box="[1122,1402,637,660]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="revelieri">
|
||
<emphasis id="6381EAC7647BAA3FFD3FFD8CFACEFD0B" box="[1122,1328,637,660]" italics="true" pageId="3" pageNumber="400">Laccobius revelieri</emphasis>
|
||
Perris
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="96F54D56647BAA3FFCDBFD8CFEB8FD28" authority="Gentili" authorityName="Gentili" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="venustus">
|
||
<emphasis id="6381EAC7647BAA3FFCDBFD8CFF0DFD28" italics="true" pageId="3" pageNumber="400">L. venustus</emphasis>
|
||
Gentili
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="96F54D56647BAA3FF80CFD51FE35FD28" box="[337,459,672,695]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="400" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6381EAC7647BAA3FF80CFD51FE35FD28" box="[337,459,672,695]" italics="true" pageId="3" pageNumber="400">Helochares</emphasis>
|
||
</taxonomicName>
|
||
spp.,
|
||
<taxonomicName id="96F54D56647BAA3FFB57FD51FCB5FD28" authority="Knisch" authorityName="Knisch" box="[522,843,672,695]" class="Insecta" family="Hydrophilidae" genus="Crenitis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="zimmermanni">
|
||
<emphasis id="6381EAC7647BAA3FFB57FD51FD09FD28" box="[522,759,672,695]" italics="true" pageId="3" pageNumber="400">Crenitis zimmermanni</emphasis>
|
||
Knisch
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="96F54D56647BAA3FFA0BFD51FA80FD28" authority="Perkins & J. Balfour-Browne" authorityName="Perkins & J. Balfour-Browne" box="[854,1406,672,695]" class="Insecta" family="Hydraenidae" genus="Parasthetops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="nigritus">
|
||
<emphasis id="6381EAC7647BAA3FFA0BFD51FBC3FD28" box="[854,1085,672,695]" italics="true" pageId="3" pageNumber="400">Parasthetops nigritus</emphasis>
|
||
Perkins & J. Balfour-Browne
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="96F54D56647BAA3FFCD5FD51FEA6FD46" authority="Perkins" authorityName="Perkins" class="Insecta" family="Hydraenidae" genus="Parasthetops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="pampinus">
|
||
<emphasis id="6381EAC7647BAA3FFCD5FD51FF01FD46" italics="true" pageId="3" pageNumber="400">P. pampinus</emphasis>
|
||
Perkins
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="96F54D56647BAA3FF83FFD33FCF8FD46" authority="Perkins & J. Balfour-Browne" authorityName="Perkins & J. Balfour-Browne" box="[354,774,706,729]" class="Insecta" family="Hydraenidae" genus="Parasthetops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="rufulus">
|
||
<emphasis id="6381EAC7647BAA3FF83FFD33FE39FD46" box="[354,455,706,729]" italics="true" pageId="3" pageNumber="400">P. rufulus</emphasis>
|
||
Perkins & J. Balfour-Browne
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="96F54D56647BAA3FFA67FD33FB5DFD46" authority="Bilton." authorityName="Bilton." box="[826,1187,706,729]" class="Insecta" family="Hydraenidae" genus="Prosthetops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="wolfbergensis">
|
||
<emphasis id="6381EAC7647BAA3FFA67FD33FBACFD46" box="[826,1106,706,729]" italics="true" pageId="3" pageNumber="400">Prosthetops wolfbergensis</emphasis>
|
||
Bilton.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |