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<document ID-ISSN="2305-2562" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1683580829501" checkinUser="felipe" docAuthor="van Bruggen, A. C." docDate="2011" docId="D9379A342E24FFA2FE7EFA44FCFCFAC0" docLanguage="en" docName="AfrInvertebr.52.1.11-19.pdf" docOrigin="African Invertebrates 52 (1)" docStyle="DocumentStyle:0201127D758FBEB889983D376DC13FCB.3:AfrInvertebr.2008-2015.journal_article.1cover" docStyleId="0201127D758FBEB889983D376DC13FCB" docStyleName="AfrInvertebr.2008-2015.journal_article.1cover" docStyleVersion="3" docTitle="Gulella inobstructa van Bruggen 1965" docType="treatment" docVersion="2" lastPageNumber="18" masterDocId="250EE24C2E25FFAAFFEAFFCBFFB5FF8A" masterDocTitle="Northern Drakensberg range Gulella species (Gastropoda, Pulmonata, Streptaxidae): Gulella inobstructa van Bruggen, 1965, revisited, or types are not typical" masterLastPageNumber="19" masterPageNumber="11" pageNumber="11" updateTime="1683633140254" updateUser="juliana" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo>
<mods:title>Northern Drakensberg range Gulella species (Gastropoda, Pulmonata, Streptaxidae): Gulella inobstructa van Bruggen, 1965, revisited, or types are not typical</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>van Bruggen, A. C.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>African Invertebrates</mods:title>
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<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="pubDate">
<mods:number>2011-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>52</mods:number>
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<mods:detail type="issue">
<mods:number>1</mods:number>
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<mods:end>19</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="ISSN">2305-2562</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:D9379A342E24FFA2FE7EFA44FCFCFAC0" httpUri="http://treatment.plazi.org/id/D9379A342E24FFA2FE7EFA44FCFCFAC0" lastPageId="8" lastPageNumber="18" pageId="1" pageNumber="11">
<subSubSection box="[404,843,1421,1448]" pageId="1" pageNumber="11" type="nomenclature">
<paragraph blockId="1.[144,1106,1421,1670]" box="[404,843,1421,1448]" pageId="1" pageNumber="11">
<heading box="[404,843,1421,1448]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="11" reason="3">
<taxonomicName ID-CoL="3HLSC" authority="van Bruggen, 1965" authorityName="van Bruggen" authorityYear="1965" box="[404,843,1421,1448]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="11" phylum="Mollusca" rank="species" species="inobstructa">
<emphasis box="[404,623,1423,1447]" italics="true" pageId="1" pageNumber="11">Gulella inobstructa</emphasis>
van
<bibRefCitation author="BRUGGEN, A. C. &amp; VAN" box="[677,843,1421,1448]" pageId="1" pageNumber="11" pagination="21 - 26" refId="ref4477" refString="BRUGGEN, A. C., VAN. 1965. Descriptions of three new species of Gulella (Mollusca, Gastropoda Pulmonata: Streptaxidae) from South Africa. Annals of the Natal Museum 18: 21 - 26." type="journal article" year="1965">Bruggen, 1965</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="18" pageId="1" pageNumber="11" type="description">
<paragraph blockId="1.[144,1106,1421,1670]" pageId="1" pageNumber="11">
<figureCitation box="[493,593,1461,1487]" captionStart="Figs 13" captionStartId="2.[181,220,1526,1547]" captionTargetBox="[189,1134,244,1509]" captionTargetId="figure-7@2.[189,1134,244,1509]" captionTargetPageId="2" captionText="Figs 13. Shells of Gulella inobstructa van Bruggen, 1965, showing the range of variation in size, i.e. the smallest and largest specimens from Table 1 inclusive of an average shell: (1) Buffelskloof Nat. Res., NMSA W4474, length 4.4 mm (no. 1); (2) Mt Sheba Nat. Res., NMSA W4319, length 7.0 mm (no. 12); (3) Mariepskop Forest, NMSA W3502, length 9.4 mm (no. 24). Note absence of outer columellar process in the aperture. Photos Dr A.J. de Winter." pageId="1" pageNumber="11">Figs 13</figureCitation>
,
<figureCitation box="[605,621,1462,1488]" captionStart="Figs 6, 7" captionStartId="7.[144,183,769,790]" captionTargetBox="[406,882,204,749]" captionTargetId="figure-260@7.[406,882,204,749]" captionTargetPageId="7" captionText="Figs 6, 7. Type specimens of Gulella species: (6) G. perspicua, “Transvaal” (= Middelburg), paralectotype, BM 1937.12.30.1123, length 4.8 mm; (7) G. inobstructa, Woodbush Forest/Magoebaskloof, holotype, NMSA 4123/T1087, length 9.1 mm; note presence of outer columellar process in the aperture. Photos Dr D. Herbert, layout of plate by Ms Sancia van der Meij." pageId="1" pageNumber="11">7</figureCitation>
, Tables 1, 2
<taxonomicName authority=": YDQ" authorityName="YDQ" box="[144,360,1496,1520]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="11" phylum="Mollusca" rank="species" species="inobstructa">
<emphasis box="[144,317,1499,1518]" italics="true" pageId="1" pageNumber="11">Gulella inobstructa</emphasis>
:
<emphasis bold="true" box="[327,360,1497,1520]" pageId="1" pageNumber="11">YDQ</emphasis>
</taxonomicName>
<emphasis bold="true" box="[364,1106,1496,1520]" pageId="1" pageNumber="11">%UXJJHQ l965: ƻl, ¿JV l, 4 (7\SH ORF.: 6RXWK $IULFD,&gt;7UDQVYDDO@ /LPSRSR SURYLQFH,</emphasis>
</paragraph>
<paragraph blockId="1.[144,1106,1421,1670]" pageId="1" pageNumber="11">
Magoebaskloof, Woodbush Forest);
<bibRefCitation author="RICHARDSON, C. L." box="[570,761,1521,1542]" pageId="1" pageNumber="11" pagination="1 - 326" refId="ref4812" refString="RICHARDSON, C. L. 1988. Streptaxacea: catalog of species. Part I Streptaxidae. Tryonia 16: i, 1 - 326." type="journal article" year="1988">Richardson 1988: 91</bibRefCitation>
;
<bibRefCitation author="AIKEN, D." box="[769,913,1521,1542]" pageId="1" pageNumber="11" pagination="3" refId="ref4445" refString="AIKEN, D. 1995. The Streptaxidae of South Africa. Special Publication of the Conchological Society of South Africa 6: 3 unnumbered pp. + 1 - 23." type="journal article" year="1995">Aiken 1995: 17</bibRefCitation>
. New material examined (all
<collectingCountry box="[408,527,1553,1574]" name="South Africa" pageId="1" pageNumber="11">South Africa</collectingCountry>
,
<collectingRegion box="[538,747,1553,1574]" country="South Africa" name="Mpumalanga" pageId="1" pageNumber="11">Mpumalanga province</collectingRegion>
, enumerated from south to north): Lydenburg area,
<quantity box="[275,334,1577,1598]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="11" unit="km" value="20.0">20 km</quantity>
SSE of Lydenburg, Uitsoek Forest (
<geoCoordinate box="[671,784,1577,1598]" degrees="25" direction="south" minutes="16.366" orientation="latitude" pageId="1" pageNumber="11" precision="1" value="-25.272766">25°16.366'S</geoCoordinate>
:
<geoCoordinate box="[786,904,1577,1598]" degrees="30" direction="east" minutes="33.138" orientation="longitude" pageId="1" pageNumber="11" precision="1" value="30.5523">30°33.138'E</geoCoordinate>
),
<quantity box="[921,994,1577,1598]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="11" unit="m" value="1200.0">1200 m</quantity>
, indigenous forest, in leaf litter, leg. J.L. Horn leg., L 112,
<date box="[572,653,1601,1622]" pageId="1" pageNumber="11" value="2006-02-03">3.ii.2006</date>
(1 shell, NMSA W4561); Buffelskloof Nat. Res. (
<geoCoordinate box="[150,262,1625,1646]" degrees="25" direction="south" minutes="16.118" orientation="latitude" pageId="1" pageNumber="11" precision="1" value="-25.268633">25°16.118'S</geoCoordinate>
:
<geoCoordinate box="[265,382,1625,1646]" degrees="30" direction="east" minutes="31.139" orientation="longitude" pageId="1" pageNumber="11" precision="1" value="30.518984">30°31.139'E</geoCoordinate>
),
<emphasis box="[397,413,1627,1646]" italics="true" pageId="1" pageNumber="11">c.</emphasis>
<quantity box="[417,473,1625,1646]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="11" unit="km" value="25.0">25 km</quantity>
S of Lydenburg,
<quantity box="[625,695,1625,1646]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="1" pageNumber="11" unit="m" value="1750.0">1750 m</quantity>
, indigenous forest, leaf litter, J.L. Horn leg., L 180,
<date box="[188,287,1649,1670]" pageId="1" pageNumber="11" value="2006-03-16">16.iii.2006</date>
(2 shells, NMSA W4474,
<figureCitation box="[525,577,1649,1670]" captionStart="Figs 13" captionStartId="2.[181,220,1526,1547]" captionTargetBox="[189,1134,244,1509]" captionTargetId="figure-7@2.[189,1134,244,1509]" captionTargetPageId="2" captionText="Figs 13. Shells of Gulella inobstructa van Bruggen, 1965, showing the range of variation in size, i.e. the smallest and largest specimens from Table 1 inclusive of an average shell: (1) Buffelskloof Nat. Res., NMSA W4474, length 4.4 mm (no. 1); (2) Mt Sheba Nat. Res., NMSA W4319, length 7.0 mm (no. 12); (3) Mariepskop Forest, NMSA W3502, length 9.4 mm (no. 24). Note absence of outer columellar process in the aperture. Photos Dr A.J. de Winter." pageId="1" pageNumber="11">Fig. 1</figureCitation>
); Horseshoe Falls (
<geoCoordinate box="[753,826,1649,1670]" degrees="25.13" direction="south" orientation="latitude" pageId="1" pageNumber="11" precision="555" value="-25.13">25.13°S</geoCoordinate>
<geoCoordinate box="[832,896,1649,1670]" degrees="30.7" direction="east" orientation="longitude" pageId="1" pageNumber="11" precision="5555" value="30.7">30.7°E</geoCoordinate>
) near Sabie,
<emphasis box="[1012,1028,1651,1670]" italics="true" pageId="1" pageNumber="11">c.</emphasis>
<quantity box="[1033,1100,1649,1670]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="1" pageNumber="11" unit="m" value="1130.0">1130 m</quantity>
,
</paragraph>
<caption pageId="2" pageNumber="12" startId="2.[181,220,1526,1547]" targetBox="[189,1134,244,1509]" targetPageId="2" targetType="figure">
<paragraph blockId="2.[181,1143,1526,1643]" pageId="2" pageNumber="12">
Figs 13. Shells of
<taxonomicName authority="van Bruggen, 1965" authorityName="van Bruggen" authorityYear="1965" box="[358,716,1526,1547]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="12" phylum="Mollusca" rank="species" species="inobstructa">
<emphasis box="[358,535,1528,1547]" italics="true" pageId="2" pageNumber="12">Gulella inobstructa</emphasis>
van
<bibRefCitation author="BRUGGEN, A. C. &amp; VAN" box="[580,716,1526,1547]" pageId="2" pageNumber="12" pagination="21 - 26" refId="ref4477" refString="BRUGGEN, A. C., VAN. 1965. Descriptions of three new species of Gulella (Mollusca, Gastropoda Pulmonata: Streptaxidae) from South Africa. Annals of the Natal Museum 18: 21 - 26." type="journal article" year="1965">Bruggen, 1965</bibRefCitation>
</taxonomicName>
, showing the range of variation in size, i.e. the smallest and largest specimens from Table 1 inclusive of an average shell: (1) Buffelskloof Nat. Res., NMSA W4474, length 4.4 mm (no. 1); (2) Mt Sheba Nat. Res., NMSA W4319, length 7.0 mm (no. 12); (3) Mariepskop Forest, NMSA W3502, length 9.4 mm (no. 24). Note absence of outer columellar process in the aperture. Photos Dr A.J. de Winter.
</paragraph>
</caption>
<paragraph blockId="3.[144,1106,204,439]" pageId="3" pageNumber="13">
<materialsCitation collectingDate="1978-07-23" collectingDateMax="2006-12-12" collectingDateMin="1978-07-23" collectionCode="RMNH, NMSA, VKHOOV, KHED, DIURPRQWDQH, IRUHVW, OHDI, OLWWHU, WXDUW" collectorName="H. E. van Hoepen &amp; J. L. Horn &amp; A. Moussalli &amp; D. Stuart-Fox" elevation="1690" latitude="-24.93122" location="Mt Sheba Nat. Res." longLatPrecision="1" longitude="30.70965" municipality="Mt Sheba Nat. Res." pageId="3" pageNumber="13" specimenCode="W5530" specimenCount="1">
leaf litter,
<date box="[241,346,204,225]" pageId="3" pageNumber="13" value="1978-07-23">
<collectingDate box="[241,346,204,225]" pageId="3" pageNumber="13" value="1978-07-23">23.vii.1978</collectingDate>
</date>
,
<collectorName box="[358,518,204,225]" pageId="3" pageNumber="13">H.E. van Hoepen</collectorName>
leg./don. (5 shells,
<collectionCode box="[706,778,204,225]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="3" pageNumber="13" type="Museum">RMNH</collectionCode>
);
<location LSID="urn:lsid:plazi:treatment:D9379A342E24FFA2FE7EFA44FCFCFAC0:54417DF92E26FFA9FCF1FF07FC61FF6B" box="[795,980,204,225]" latitude="-24.93122" longLatPrecision="1" longitude="30.70965" municipality="Mt Sheba Nat. Res." name="Mt Sheba Nat. Res." pageId="3" pageNumber="13">Mt Sheba Nat. Res.</location>
(
<geoCoordinate box="[986,1101,204,225]" degrees="24" direction="south" minutes="56.340" orientation="latitude" pageId="3" pageNumber="13" precision="1" value="-24.939">24°56.340'S</geoCoordinate>
:
<geoCoordinate box="[144,259,228,249]" degrees="30" direction="east" minutes="42.634" orientation="longitude" pageId="3" pageNumber="13" precision="1" value="30.710567">30°42.634'E</geoCoordinate>
),
<emphasis box="[278,294,229,248]" italics="true" pageId="3" pageNumber="13">c.</emphasis>
<locationDeviation box="[302,541,228,249]" location="Wof Pilgrim's Rest" pageId="3" pageNumber="13">
<quantity box="[302,351,228,249]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="13" unit="km" value="5.0">5 km</quantity>
Wof Pilgrims Rest
</locationDeviation>
,
<quantity box="[552,626,228,249]" metricMagnitude="3" metricUnit="m" metricValue="1.69" pageId="3" pageNumber="13" unit="m" value="1690.0">
<elevation box="[552,626,228,249]" metricMagnitude="3" metricUnit="m" metricValue="1.69" pageId="3" pageNumber="13" unit="m" value="1690.0">1690 m</elevation>
</quantity>
, indigenous forest, leaf litter,
<collectorName box="[911,1001,228,249]" pageId="3" pageNumber="13">J.L. Horn</collectorName>
leg., MTS 14,
<date box="[181,285,252,273]" pageId="3" pageNumber="13" value="2006-12-12">
<collectingDate box="[181,285,252,273]" pageId="3" pageNumber="13" value="2006-12-12">12.xii.2006</collectingDate>
</date>
(29 shells,
<collectionCode box="[400,465,252,273]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="13" type="Museum">NMSA</collectionCode>
<specimenCode box="[473,543,252,273]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="13" type="Museum">W5530</specimenCode>
; duplicates in
<collectionCode box="[683,756,252,273]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="3" pageNumber="13" type="Museum">RMNH</collectionCode>
);
<collectingMunicipality box="[774,963,252,273]" pageId="3" pageNumber="13">Mt Sheba Nat. Res.</collectingMunicipality>
(
<geoCoordinate box="[977,1101,252,273]" degrees="24.931220" direction="south" orientation="latitude" pageId="3" pageNumber="13" precision="1" value="-24.93122">24.931220°S</geoCoordinate>
:
<geoCoordinate box="[144,266,276,297]" degrees="30.709650" direction="east" orientation="longitude" pageId="3" pageNumber="13" precision="1" value="30.70965">30.709650°E</geoCoordinate>
),
<quantity box="[282,354,276,297]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="13" unit="m" value="1900.0">
<elevation box="[282,354,276,297]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="13" unit="m" value="1900.0">1900 m</elevation>
</quantity>
, indigenous forest, on decaying log,
<collectorName box="[688,805,276,297]" pageId="3" pageNumber="13">A. Moussalli</collectorName>
&amp;
<collectorName box="[835,960,276,297]" pageId="3" pageNumber="13">D. Stuart-Fox</collectorName>
leg.,
<date box="[1012,1104,276,297]" pageId="3" pageNumber="13" value="2004-02-25">
<collectingDate box="[1012,1104,276,297]" pageId="3" pageNumber="13" value="2004-02-25">25.ii.2004</collectingDate>
</date>
<emphasis bold="true" pageId="3" pageNumber="13">
&gt;ƻƟ7ƻƟ9@ (5
<collectionCode box="[262,317,299,322]" pageId="3" pageNumber="13">VKHOOV</collectionCode>
,106$:4 l9,)LJ. ƻ); 0W 6
<collectionCode box="[577,624,298,321]" pageId="3" pageNumber="13">KHED</collectionCode>
(ƻ4.9 lƻ°6 Ɵ.7Ɵ97°(), l9ƟƟ P,
<collectionCode box="[927,1101,298,322]" pageId="3" pageNumber="13">DIURPRQWDQH IRUHVW</collectionCode>
,
<collectionCode box="[144,229,323,346]" pageId="3" pageNumber="13">OHDI OLWWHU</collectionCode>
,$. 0RXVVDOOL &amp; '. 6
<collectionCode box="[417,480,322,345]" pageId="3" pageNumber="13">WXDUW</collectionCode>
)
</emphasis>
</materialsCitation>
<materialsCitation box="[494,503,322,345]" collectingDate="1978-07-23" collectingDateMax="2006-12-12" collectingDateMin="1978-07-23" collectionCode="R" collectorName="H. E. van Hoepen &amp; J. L. Horn &amp; A. Moussalli &amp; D. Stuart-Fox" elevation="1690" isTailEnumeration="true" latitude="-24.93122" location="Mt Sheba Nat. Res." longLatPrecision="1" longitude="30.70965" municipality="Mt Sheba Nat. Res." pageId="3" pageNumber="13" specimenCount="1">
<collectionCode box="[494,503,322,345]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="13">
<emphasis bold="true" box="[494,503,322,345]" pageId="3" pageNumber="13">R</emphasis>
</collectionCode>
</materialsCitation>
<materialsCitation collectingDate="2005-03-01" collectionCode="VKHOOV, RUHVW, LQGLJHQRXV, DIURPRQWDQH, IRUHVW, LQ, OHDI, OLWWHU, RQ, ARRU, NMSA, LY" collectorName="Horn" latitude="-24.555117" location="Mariepskop Forest" longLatPrecision="1" longitude="30.89395" pageId="3" pageNumber="13" specimenCode="W3502" specimenCount="5">
<emphasis bold="true" pageId="3" pageNumber="13">
[OHJ., ƻ5.LL.ƻƟƟ4&gt;ƻƟ7ƻƟ9@ (ƻ
<collectionCode box="[776,831,323,346]" pageId="3" pageNumber="13">VKHOOV</collectionCode>
, 106$:ƻ4ƻ); 0DULHSVNRS)
<collectionCode box="[147,200,346,369]" pageId="3" pageNumber="13">RUHVW</collectionCode>
(ƻ4° 4.Ɵl56: Ɵ°
<specimenCount box="[368,389,347,370]" pageId="3" pageNumber="13" type="generic">5l.</specimenCount>
899(), l5ƻƟ P,
<collectionCode box="[538,817,346,370]" pageId="3" pageNumber="13">LQGLJHQRXV DIURPRQWDQH IRUHVW</collectionCode>
,
<collectionCode box="[824,1062,346,370]" pageId="3" pageNumber="13">LQ OHDI OLWWHU RQ IRUHVW ÀRRU</collectionCode>
, -./.
</emphasis>
<collectorName box="[144,191,372,393]" pageId="3" pageNumber="13">Horn</collectorName>
leg., L 19,
<date box="[291,378,372,393]" pageId="3" pageNumber="13" value="2005-03-01">
<collectingDate box="[291,378,372,393]" pageId="3" pageNumber="13" value="2005-03-01">1.iii.2005</collectingDate>
</date>
(1 shell,
<collectionCode box="[461,525,372,393]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="13" type="Museum">NMSA</collectionCode>
<specimenCode box="[530,599,372,393]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="13" type="Museum">W3502</specimenCode>
,
<figureCitation box="[607,661,372,393]" captionStart="Figs 13" captionStartId="2.[181,220,1526,1547]" captionTargetBox="[189,1134,244,1509]" captionTargetId="figure-7@2.[189,1134,244,1509]" captionTargetPageId="2" captionText="Figs 13. Shells of Gulella inobstructa van Bruggen, 1965, showing the range of variation in size, i.e. the smallest and largest specimens from Table 1 inclusive of an average shell: (1) Buffelskloof Nat. Res., NMSA W4474, length 4.4 mm (no. 1); (2) Mt Sheba Nat. Res., NMSA W4319, length 7.0 mm (no. 12); (3) Mariepskop Forest, NMSA W3502, length 9.4 mm (no. 24). Note absence of outer columellar process in the aperture. Photos Dr A.J. de Winter." pageId="3" pageNumber="13">Fig. 3</figureCitation>
);
<location LSID="urn:lsid:plazi:treatment:D9379A342E24FFA2FE7EFA44FCFCFAC0:54417DF92E26FFA9FD4DFEBFFCE4FE03" box="[679,849,372,393]" latitude="-24.555117" longLatPrecision="1" longitude="30.89395" name="Mariepskop Forest" pageId="3" pageNumber="13">Mariepskop Forest</location>
(
<geoCoordinate box="[860,974,372,393]" degrees="24" direction="south" minutes="33.307" orientation="latitude" pageId="3" pageNumber="13" precision="1" value="-24.555117">24°33.307'S</geoCoordinate>
:
<geoCoordinate box="[976,1093,372,393]" degrees="30" direction="east" minutes="53.637" orientation="longitude" pageId="3" pageNumber="13" precision="1" value="30.89395">30°53.637'E</geoCoordinate>
),
<emphasis bold="true" box="[144,1093,394,418]" pageId="3" pageNumber="13">
l46Ɵ P,
<collectionCode box="[227,511,394,418]" pageId="3" pageNumber="13">LQGLJHQRXV DIURPRQWDQH IRUHVW</collectionCode>
,
<collectionCode box="[521,773,394,418]" pageId="3" pageNumber="13">LQ OHDI OLWWHU RQ IRUHVW ÀRRU</collectionCode>
, -./. +RUQ OHJ., / l, ƻ.
<collectionCode box="[1007,1028,395,418]" country="France" name="Laboratoire de Mycologie associe au CNRS" pageId="3" pageNumber="13">LY</collectionCode>
.ƻƟƟ5 (
</emphasis>
</materialsCitation>
</paragraph>
<paragraph blockId="3.[144,1106,204,439]" lastBlockId="3.[144,1110,457,1026]" pageId="3" pageNumber="13">
shells, NMSA W3503).
<taxonomicName authorityName="van Bruggen" authorityYear="1965" box="[170,391,458,482]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="13" phylum="Mollusca" rank="species" species="inobstructa">
<emphasis box="[170,391,458,482]" italics="true" pageId="3" pageNumber="13">Gulella inobstructa</emphasis>
</taxonomicName>
has also been collected in only two of the forest study sites of
</paragraph>
<paragraph blockId="3.[144,1110,457,1026]" pageId="3" pageNumber="13">
<emphasis bold="true" box="[144,1104,486,515]" pageId="3" pageNumber="13">6ZD\H (ƻƟƟ4: 5 (WDEOH ƻ.l) &amp; 6 (¿J. ƻ.l); $SSHQGL[ƻ RQ S. ƻ ƻ), L.H. 1HZ $JDWKD</emphasis>
(Mmakgowa) Forest and Forest Glens. Both are situated in
<collectingRegion box="[810,1020,520,546]" country="South Africa" name="Limpopo" pageId="3" pageNumber="13">Limpopo Province</collectingRegion>
but are part and parcel of the northern Drakensberg range as opposed to the more northern Soutpansberg complex. This material has not been studied for the present paper.
</paragraph>
<paragraph blockId="3.[144,1110,457,1026]" pageId="3" pageNumber="13">
When
<taxonomicName authorityName="van Bruggen" authorityYear="1965" box="[247,412,618,642]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="13" phylum="Mollusca" rank="species" species="inobstructa">
<emphasis box="[247,412,618,642]" italics="true" pageId="3" pageNumber="13">G. inobstructa</emphasis>
</taxonomicName>
was described (van
<bibRefCitation author="BRUGGEN, A. C. &amp; VAN" box="[651,860,616,642]" pageId="3" pageNumber="13" pagination="21 - 26" refId="ref4477" refString="BRUGGEN, A. C., VAN. 1965. Descriptions of three new species of Gulella (Mollusca, Gastropoda Pulmonata: Streptaxidae) from South Africa. Annals of the Natal Museum 18: 21 - 26." type="journal article" year="1965">Bruggen 1965: 21</bibRefCitation>
) it was diagnosed as follows: “A smooth, medium-sized species of
<taxonomicName authorityName="Pfeiffer" authorityYear="1856" box="[672,756,650,674]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis box="[672,756,650,674]" italics="true" pageId="3" pageNumber="13">Gulella</emphasis>
</taxonomicName>
with only four teeth in a little obstructed aperture, viz., a parietal, labral and two columellar processes.” Only two
<emphasis bold="true" box="[144,1104,710,739]" pageId="3" pageNumber="13">DGXOW VKHOOV ZHUH DYDLODEOH DQG ZHUH FRQVLGHUHG VXI¿FLHQWO\ FKDUDFWHULVWLF WR ZDUUDQW</emphasis>
recognition as a separate taxon. In addition there was a juvenile shell without any apertural dentition.
</paragraph>
<paragraph blockId="3.[144,1110,457,1026]" pageId="3" pageNumber="13">
Recently a number of similar specimens has been collected in the northern Drakensberg escarpment (see above) exhibiting considerable variation in size, shape, sculpture and apertural dentition. Metric data of this material (inclusive of the
<typeStatus box="[899,947,872,898]" pageId="3" pageNumber="13">type</typeStatus>
material) are shown in Table 1, which may be summarized as 4.49.4×
<quantity box="[795,936,904,930]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.5" metricValueMin="2.5" pageId="3" pageNumber="13" unit="mm" value="3.5" valueMax="4.5" valueMin="2.5">2.54.5 mm</quantity>
, l/d 1.752.48, with 68+ whorls, which implies that the smallest adult specimen is less than half the size of the largest shell. This is rare among species of
<taxonomicName authorityName="Pfeiffer" authorityYear="1856" box="[752,836,970,994]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis box="[752,836,970,994]" italics="true" pageId="3" pageNumber="13">Gulella</emphasis>
</taxonomicName>
, particularly those with a somewhat restricted distribution (see below).
</paragraph>
<caption pageId="3" pageNumber="13" startId="3.[144,177,1630,1651]" targetBox="[172,1078,1068,1608]" targetPageId="3" targetType="figure">
<paragraph blockId="3.[144,1105,1630,1675]" pageId="3" pageNumber="13">
Fig. 4. Frequency diagram of the length of the shell of
<taxonomicName authorityName="van Bruggen" authorityYear="1965" box="[651,828,1632,1651]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="13" phylum="Mollusca" rank="species" species="inobstructa">
<emphasis box="[651,828,1632,1651]" italics="true" pageId="3" pageNumber="13">Gulella inobstructa</emphasis>
</taxonomicName>
with subdivisions of 1.0 mm. Diagram generated by Ms Sancia van der Meij.
</paragraph>
</caption>
<tableNote pageId="4" pageNumber="14" targetBox="[197,1114,299,1201]" targetPageId="4">
<paragraph blockId="4.[181,1142,205,282]" box="[617,705,205,226]" pageId="4" pageNumber="14">TABLE 1</paragraph>
<paragraph blockId="4.[181,1142,205,282]" box="[181,1142,236,260]" pageId="4" pageNumber="14">
Metric data of 24 adult shells considered to belong to
<taxonomicName authorityName="van Bruggen" authorityYear="1965" box="[667,844,239,258]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="14" phylum="Mollusca" rank="species" species="inobstructa">
<emphasis box="[667,844,239,258]" italics="true" pageId="4" pageNumber="14">Gulella inobstructa</emphasis>
</taxonomicName>
.
<emphasis bold="true" box="[854,1142,236,260]" pageId="4" pageNumber="14">$Q DGXOW VKHOO LV KHUH GH¿QHG DV</emphasis>
</paragraph>
<paragraph blockId="4.[181,1142,205,282]" box="[181,773,261,282]" pageId="4" pageNumber="14">a specimen exhibiting the full complement of apertural dentition.</paragraph>
</tableNote>
<paragraph pageId="4" pageNumber="14">
<table box="[197,1114,299,1201]" gridcols="6" gridrows="25" pageId="4" pageNumber="14">
<tr box="[197,1114,299,344]" gridrow="0" pageId="4" pageNumber="14">
<th box="[197,230,299,344]" gridcol="0" gridrow="0" pageId="4" pageNumber="14">no.</th>
<th box="[274,410,299,344]" gridcol="1" gridrow="0" pageId="4" pageNumber="14">Length × major diameter, mm</th>
<th box="[456,497,299,344]" gridcol="2" gridrow="0" pageId="4" pageNumber="14">l/d</th>
<th box="[541,640,299,344]" gridcol="3" gridrow="0" pageId="4" pageNumber="14">Number of whorls</th>
<th box="[678,941,299,344]" gridcol="4" gridrow="0" pageId="4" pageNumber="14">Additional data</th>
<th box="[971,1114,299,344]" gridcol="5" gridrow="0" pageId="4" pageNumber="14">Sample</th>
</tr>
<tr box="[197,1114,355,400]" gridrow="1" pageId="4" pageNumber="14">
<th box="[197,230,355,400]" gridcol="0" gridrow="1" pageId="4" pageNumber="14">1</th>
<td box="[274,410,355,400]" gridcol="1" gridrow="1" pageId="4" pageNumber="14">4.4 × 2.5</td>
<td box="[456,497,355,400]" gridcol="2" gridrow="1" pageId="4" pageNumber="14">1.75</td>
<td box="[541,640,355,400]" gridcol="3" gridrow="1" pageId="4" pageNumber="14">6</td>
<td box="[678,941,355,400]" gridcol="4" gridrow="1" pageId="4" pageNumber="14">adult shell: very small and very squat</td>
<td box="[971,1114,355,400]" gridcol="5" gridrow="1" pageId="4" pageNumber="14">NMSA W4474, Fig. 1</td>
</tr>
<tr box="[197,1114,411,432]" gridrow="2" pageId="4" pageNumber="14">
<th box="[197,230,411,432]" gridcol="0" gridrow="2" pageId="4" pageNumber="14">2</th>
<td box="[274,410,411,432]" gridcol="1" gridrow="2" pageId="4" pageNumber="14">5.6 × 2.9</td>
<td box="[456,497,411,432]" gridcol="2" gridrow="2" pageId="4" pageNumber="14">1.96</td>
<td box="[541,640,411,432]" gridcol="3" gridrow="2" pageId="4" pageNumber="14">7</td>
<td box="[678,941,411,432]" gridcol="4" gridrow="2" pageId="4" pageNumber="14">adult shell: small and squat</td>
<td box="[971,1114,411,432]" gridcol="5" gridrow="2" pageId="4" pageNumber="14">NMSA W4561</td>
</tr>
<tr box="[197,1114,442,463]" gridrow="3" pageId="4" pageNumber="14">
<th box="[197,230,442,463]" gridcol="0" gridrow="3" pageId="4" pageNumber="14">3</th>
<td box="[274,410,442,463]" gridcol="1" gridrow="3" pageId="4" pageNumber="14">5.9 × 2.6</td>
<td box="[456,497,442,463]" gridcol="2" gridrow="3" pageId="4" pageNumber="14">2.24</td>
<td box="[541,640,442,463]" gridcol="3" gridrow="3" pageId="4" pageNumber="14">7+</td>
<td box="[678,941,442,463]" gridcol="4" gridrow="3" pageId="4" pageNumber="14"></td>
<td box="[971,1114,442,463]" gridcol="5" gridrow="3" pageId="4" pageNumber="14">NMSA W4474</td>
</tr>
<tr box="[197,1114,474,495]" gridrow="4" pageId="4" pageNumber="14">
<th box="[197,230,474,495]" gridcol="0" gridrow="4" pageId="4" pageNumber="14">4</th>
<td box="[274,410,474,495]" gridcol="1" gridrow="4" pageId="4" pageNumber="14">6.2 × 3.0</td>
<td box="[456,497,474,495]" gridcol="2" gridrow="4" pageId="4" pageNumber="14">2.08</td>
<td box="[541,640,474,495]" gridcol="3" gridrow="4" pageId="4" pageNumber="14"></td>
<td box="[678,941,474,495]" gridcol="4" gridrow="4" pageId="4" pageNumber="14"></td>
<td box="[971,1114,474,495]" gridcol="5" gridrow="4" pageId="4" pageNumber="14">NMSA W5530</td>
</tr>
<tr box="[197,1114,506,527]" gridrow="5" pageId="4" pageNumber="14">
<th box="[197,230,506,527]" gridcol="0" gridrow="5" pageId="4" pageNumber="14">5</th>
<td box="[274,410,506,527]" gridcol="1" gridrow="5" pageId="4" pageNumber="14">6.2 ×3.1</td>
<td box="[456,497,506,527]" gridcol="2" gridrow="5" pageId="4" pageNumber="14">2.00</td>
<td box="[541,640,506,527]" gridcol="3" gridrow="5" pageId="4" pageNumber="14">
<emphasis box="[574,584,508,527]" italics="true" pageId="4" pageNumber="14">c</emphasis>
. 8
</td>
<td box="[678,941,506,527]" gridcol="4" gridrow="5" pageId="4" pageNumber="14">apex damaged</td>
<td box="[971,1114,506,527]" gridcol="5" gridrow="5" pageId="4" pageNumber="14">RMNH</td>
</tr>
<tr box="[197,1114,537,558]" gridrow="6" pageId="4" pageNumber="14">
<th box="[197,230,537,558]" gridcol="0" gridrow="6" pageId="4" pageNumber="14">6</th>
<td box="[274,410,537,558]" gridcol="1" gridrow="6" pageId="4" pageNumber="14">6.5 ×2.9</td>
<td box="[456,497,537,558]" gridcol="2" gridrow="6" pageId="4" pageNumber="14">2.26</td>
<td box="[541,640,537,558]" gridcol="3" gridrow="6" pageId="4" pageNumber="14">8</td>
<td box="[678,941,537,558]" gridcol="4" gridrow="6" pageId="4" pageNumber="14"></td>
<td box="[971,1114,537,558]" gridcol="5" gridrow="6" pageId="4" pageNumber="14">NMSA W5530</td>
</tr>
<tr box="[197,1114,569,590]" gridrow="7" pageId="4" pageNumber="14">
<th box="[197,230,569,590]" gridcol="0" gridrow="7" pageId="4" pageNumber="14">7</th>
<td box="[274,410,569,590]" gridcol="1" gridrow="7" pageId="4" pageNumber="14">6.5 ×3.1</td>
<td box="[456,497,569,590]" gridcol="2" gridrow="7" pageId="4" pageNumber="14">2.08</td>
<td box="[541,640,569,590]" gridcol="3" gridrow="7" pageId="4" pageNumber="14"></td>
<td box="[678,941,569,590]" gridcol="4" gridrow="7" pageId="4" pageNumber="14"></td>
<td box="[971,1114,569,590]" gridcol="5" gridrow="7" pageId="4" pageNumber="14">NMSA W5530</td>
</tr>
<tr box="[197,1114,601,622]" gridrow="8" pageId="4" pageNumber="14">
<th box="[197,230,601,622]" gridcol="0" gridrow="8" pageId="4" pageNumber="14">8</th>
<td box="[274,410,601,622]" gridcol="1" gridrow="8" pageId="4" pageNumber="14">6.6 ×3.0</td>
<td box="[456,497,601,622]" gridcol="2" gridrow="8" pageId="4" pageNumber="14">2.21</td>
<td box="[541,640,601,622]" gridcol="3" gridrow="8" pageId="4" pageNumber="14">8</td>
<td box="[678,941,601,622]" gridcol="4" gridrow="8" pageId="4" pageNumber="14"></td>
<td box="[971,1114,601,622]" gridcol="5" gridrow="8" pageId="4" pageNumber="14">NMSA W2423</td>
</tr>
<tr box="[197,1114,633,654]" gridrow="9" pageId="4" pageNumber="14">
<th box="[197,230,633,654]" gridcol="0" gridrow="9" pageId="4" pageNumber="14">9</th>
<td box="[274,410,633,654]" gridcol="1" gridrow="9" pageId="4" pageNumber="14">6.6 ×3.1</td>
<td box="[456,497,633,654]" gridcol="2" gridrow="9" pageId="4" pageNumber="14">2.12</td>
<td box="[541,640,633,654]" gridcol="3" gridrow="9" pageId="4" pageNumber="14">8-</td>
<td box="[678,941,633,654]" gridcol="4" gridrow="9" pageId="4" pageNumber="14"></td>
<td box="[971,1114,633,654]" gridcol="5" gridrow="9" pageId="4" pageNumber="14">NMSA W5530</td>
</tr>
<tr box="[197,1114,664,685]" gridrow="10" pageId="4" pageNumber="14">
<th box="[197,230,664,685]" gridcol="0" gridrow="10" pageId="4" pageNumber="14">10</th>
<td box="[274,410,664,685]" gridcol="1" gridrow="10" pageId="4" pageNumber="14">6.6 ×3.2</td>
<td box="[456,497,664,685]" gridcol="2" gridrow="10" pageId="4" pageNumber="14">2.04</td>
<td box="[541,640,664,685]" gridcol="3" gridrow="10" pageId="4" pageNumber="14">&lt;8</td>
<td box="[678,941,664,685]" gridcol="4" gridrow="10" pageId="4" pageNumber="14"></td>
<td box="[971,1114,664,685]" gridcol="5" gridrow="10" pageId="4" pageNumber="14">RMNH</td>
</tr>
<tr box="[197,1114,696,717]" gridrow="11" pageId="4" pageNumber="14">
<th box="[197,230,696,717]" gridcol="0" gridrow="11" pageId="4" pageNumber="14">11</th>
<td box="[274,410,696,717]" gridcol="1" gridrow="11" pageId="4" pageNumber="14">6.9 ×3.1</td>
<td box="[456,497,696,717]" gridcol="2" gridrow="11" pageId="4" pageNumber="14">2.20</td>
<td box="[541,640,696,717]" gridcol="3" gridrow="11" pageId="4" pageNumber="14">8+</td>
<td box="[678,941,696,717]" gridcol="4" gridrow="11" pageId="4" pageNumber="14"></td>
<td box="[971,1114,696,717]" gridcol="5" gridrow="11" pageId="4" pageNumber="14">NMSA W5530</td>
</tr>
<tr box="[197,1114,728,773]" gridrow="12" pageId="4" pageNumber="14">
<th box="[197,230,728,773]" gridcol="0" gridrow="12" pageId="4" pageNumber="14">12</th>
<td box="[274,410,728,773]" gridcol="1" gridrow="12" pageId="4" pageNumber="14">7.0 ×3.0</td>
<td box="[456,497,728,773]" gridcol="2" gridrow="12" pageId="4" pageNumber="14">2.33</td>
<td box="[541,640,728,773]" gridcol="3" gridrow="12" pageId="4" pageNumber="14">8</td>
<td box="[678,941,728,773]" gridcol="4" gridrow="12" pageId="4" pageNumber="14"></td>
<td box="[971,1114,728,773]" gridcol="5" gridrow="12" pageId="4" pageNumber="14">NMSA W4319, Fig. 2</td>
</tr>
<tr box="[197,1114,783,804]" gridrow="13" pageId="4" pageNumber="14">
<th box="[197,230,783,804]" gridcol="0" gridrow="13" pageId="4" pageNumber="14">13</th>
<td box="[274,410,783,804]" gridcol="1" gridrow="13" pageId="4" pageNumber="14">7.0 ×3.0</td>
<td box="[456,497,783,804]" gridcol="2" gridrow="13" pageId="4" pageNumber="14">2.33</td>
<td box="[541,640,783,804]" gridcol="3" gridrow="13" pageId="4" pageNumber="14">8</td>
<td box="[678,941,783,804]" gridcol="4" gridrow="13" pageId="4" pageNumber="14">some spiral sculpture present</td>
<td box="[971,1114,783,804]" gridcol="5" gridrow="13" pageId="4" pageNumber="14">NMSA W5530</td>
</tr>
<tr box="[197,1114,815,836]" gridrow="14" pageId="4" pageNumber="14">
<th box="[197,230,815,836]" gridcol="0" gridrow="14" pageId="4" pageNumber="14">14</th>
<td box="[274,410,815,836]" gridcol="1" gridrow="14" pageId="4" pageNumber="14">7.0 ×3.1</td>
<td box="[456,497,815,836]" gridcol="2" gridrow="14" pageId="4" pageNumber="14">2.24</td>
<td box="[541,640,815,836]" gridcol="3" gridrow="14" pageId="4" pageNumber="14">8</td>
<td box="[678,941,815,836]" gridcol="4" gridrow="14" pageId="4" pageNumber="14"></td>
<td box="[971,1114,815,836]" gridcol="5" gridrow="14" pageId="4" pageNumber="14">NMSA W4319</td>
</tr>
<tr box="[197,1114,847,868]" gridrow="15" pageId="4" pageNumber="14">
<th box="[197,230,847,868]" gridcol="0" gridrow="15" pageId="4" pageNumber="14">15</th>
<td box="[274,410,847,868]" gridcol="1" gridrow="15" pageId="4" pageNumber="14">7.1 ×3.1</td>
<td box="[456,497,847,868]" gridcol="2" gridrow="15" pageId="4" pageNumber="14">2.28</td>
<td box="[541,640,847,868]" gridcol="3" gridrow="15" pageId="4" pageNumber="14">8</td>
<td box="[678,941,847,868]" gridcol="4" gridrow="15" pageId="4" pageNumber="14"></td>
<td box="[971,1114,847,868]" gridcol="5" gridrow="15" pageId="4" pageNumber="14">NMSA W4319</td>
</tr>
<tr box="[197,1114,879,900]" gridrow="16" pageId="4" pageNumber="14">
<th box="[197,230,879,900]" gridcol="0" gridrow="16" pageId="4" pageNumber="14">16</th>
<td box="[274,410,879,900]" gridcol="1" gridrow="16" pageId="4" pageNumber="14">7.1 ×3.1</td>
<td box="[456,497,879,900]" gridcol="2" gridrow="16" pageId="4" pageNumber="14">2.28</td>
<td box="[541,640,879,900]" gridcol="3" gridrow="16" pageId="4" pageNumber="14">8</td>
<td box="[678,941,879,900]" gridcol="4" gridrow="16" pageId="4" pageNumber="14"></td>
<td box="[971,1114,879,900]" gridcol="5" gridrow="16" pageId="4" pageNumber="14">NMSA W5530</td>
</tr>
<tr box="[197,1114,910,931]" gridrow="17" pageId="4" pageNumber="14">
<th box="[197,230,910,931]" gridcol="0" gridrow="17" pageId="4" pageNumber="14">17</th>
<td box="[274,410,910,931]" gridcol="1" gridrow="17" pageId="4" pageNumber="14">7.1 ×3.2</td>
<td box="[456,497,910,931]" gridcol="2" gridrow="17" pageId="4" pageNumber="14">2.19</td>
<td box="[541,640,910,931]" gridcol="3" gridrow="17" pageId="4" pageNumber="14">&lt;8</td>
<td box="[678,941,910,931]" gridcol="4" gridrow="17" pageId="4" pageNumber="14"></td>
<td box="[971,1114,910,931]" gridcol="5" gridrow="17" pageId="4" pageNumber="14">RMNH</td>
</tr>
<tr box="[197,1114,942,963]" gridrow="18" pageId="4" pageNumber="14">
<th box="[197,230,942,963]" gridcol="0" gridrow="18" pageId="4" pageNumber="14">18</th>
<td box="[274,410,942,963]" gridcol="1" gridrow="18" pageId="4" pageNumber="14">7.2 ×3.2</td>
<td box="[456,497,942,963]" gridcol="2" gridrow="18" pageId="4" pageNumber="14">2.23</td>
<td box="[541,640,942,963]" gridcol="3" gridrow="18" pageId="4" pageNumber="14">8+</td>
<td box="[678,941,942,963]" gridcol="4" gridrow="18" pageId="4" pageNumber="14"></td>
<td box="[971,1114,942,963]" gridcol="5" gridrow="18" pageId="4" pageNumber="14">NMSA W5530</td>
</tr>
<tr box="[197,1114,974,995]" gridrow="19" pageId="4" pageNumber="14">
<th box="[197,230,974,995]" gridcol="0" gridrow="19" pageId="4" pageNumber="14">19</th>
<td box="[274,410,974,995]" gridcol="1" gridrow="19" pageId="4" pageNumber="14">7.5 ×3.1</td>
<td box="[456,497,974,995]" gridcol="2" gridrow="19" pageId="4" pageNumber="14">2.40</td>
<td box="[541,640,974,995]" gridcol="3" gridrow="19" pageId="4" pageNumber="14">8</td>
<td box="[678,941,974,995]" gridcol="4" gridrow="19" pageId="4" pageNumber="14"></td>
<td box="[971,1114,974,995]" gridcol="5" gridrow="19" pageId="4" pageNumber="14">NMSA W4319</td>
</tr>
<tr box="[197,1114,1005,1026]" gridrow="20" pageId="4" pageNumber="14">
<th box="[197,230,1005,1026]" gridcol="0" gridrow="20" pageId="4" pageNumber="14">20</th>
<td box="[274,410,1005,1026]" gridcol="1" gridrow="20" pageId="4" pageNumber="14">7.7 ×3.1</td>
<td box="[456,497,1005,1026]" gridcol="2" gridrow="20" pageId="4" pageNumber="14">2.48</td>
<td box="[541,640,1005,1026]" gridcol="3" gridrow="20" pageId="4" pageNumber="14">8+</td>
<td box="[678,941,1005,1026]" gridcol="4" gridrow="20" pageId="4" pageNumber="14"></td>
<td box="[971,1114,1005,1026]" gridcol="5" gridrow="20" pageId="4" pageNumber="14">NMSA W5530</td>
</tr>
<tr box="[197,1114,1037,1082]" gridrow="21" pageId="4" pageNumber="14">
<th box="[197,230,1037,1082]" gridcol="0" gridrow="21" pageId="4" pageNumber="14">21</th>
<td box="[274,410,1037,1082]" gridcol="1" gridrow="21" pageId="4" pageNumber="14">7.7 ×4.2</td>
<td box="[456,497,1037,1082]" gridcol="2" gridrow="21" pageId="4" pageNumber="14">1.82</td>
<td box="[541,640,1037,1082]" gridcol="3" gridrow="21" pageId="4" pageNumber="14">8</td>
<td box="[678,941,1037,1082]" gridcol="4" gridrow="21" pageId="4" pageNumber="14">shell squat and slightly costulate</td>
<td box="[971,1114,1037,1082]" gridcol="5" gridrow="21" pageId="4" pageNumber="14">NMSA W3503</td>
</tr>
<tr box="[197,1114,1093,1114]" gridrow="22" pageId="4" pageNumber="14">
<th box="[197,230,1093,1114]" gridcol="0" gridrow="22" pageId="4" pageNumber="14">22</th>
<td box="[274,410,1093,1114]" gridcol="1" gridrow="22" pageId="4" pageNumber="14">8.3 ×4.2</td>
<td box="[456,497,1093,1114]" gridcol="2" gridrow="22" pageId="4" pageNumber="14">1.98</td>
<td box="[541,640,1093,1114]" gridcol="3" gridrow="22" pageId="4" pageNumber="14">8</td>
<td box="[678,941,1093,1114]" gridcol="4" gridrow="22" pageId="4" pageNumber="14">paratype</td>
<td box="[971,1114,1093,1114]" gridcol="5" gridrow="22" pageId="4" pageNumber="14">NMSA</td>
</tr>
<tr box="[197,1114,1125,1146]" gridrow="23" pageId="4" pageNumber="14">
<th box="[197,230,1125,1146]" gridcol="0" gridrow="23" pageId="4" pageNumber="14">23</th>
<td box="[274,410,1125,1146]" gridcol="1" gridrow="23" pageId="4" pageNumber="14">9.1 ×4.5</td>
<td box="[456,497,1125,1146]" gridcol="2" gridrow="23" pageId="4" pageNumber="14">2.02</td>
<td box="[541,640,1125,1146]" gridcol="3" gridrow="23" pageId="4" pageNumber="14">8</td>
<td box="[678,941,1125,1146]" gridcol="4" gridrow="23" pageId="4" pageNumber="14">holotype</td>
<td box="[971,1114,1125,1146]" gridcol="5" gridrow="23" pageId="4" pageNumber="14">NMSA</td>
</tr>
<tr box="[197,1114,1156,1201]" gridrow="24" pageId="4" pageNumber="14">
<th box="[197,230,1156,1201]" gridcol="0" gridrow="24" pageId="4" pageNumber="14">24</th>
<td box="[274,410,1156,1201]" gridcol="1" gridrow="24" pageId="4" pageNumber="14">9.4 ×4.5</td>
<td box="[456,497,1156,1201]" gridcol="2" gridrow="24" pageId="4" pageNumber="14">2.08</td>
<td box="[541,640,1156,1201]" gridcol="3" gridrow="24" pageId="4" pageNumber="14">8</td>
<td box="[678,941,1156,1201]" gridcol="4" gridrow="24" pageId="4" pageNumber="14">shell large and slightly costulate</td>
<td box="[971,1114,1156,1201]" gridcol="5" gridrow="24" pageId="4" pageNumber="14">NMSA W3502, Fig. 3</td>
</tr>
</table>
</paragraph>
<paragraph blockId="4.[181,1142,1260,1672]" pageId="4" pageNumber="14">
<emphasis bold="true" pageId="4" pageNumber="14">8QIRUWXQDWHO\ WKHUH LV RQO\ RQH VDPSOH FRQWDLQLQJ D VXI¿FLHQW QXPEHU RI VKHOOV (ƻ7 DGXOW VKHOOV) WR UHÀHFW ORFDO YDULDWLRQ LQ VL]H DQG VKDSH (106$:55 Ɵ). $ IHZ RWKHU</emphasis>
samples each encompass more than
<specimenCount box="[580,684,1326,1352]" pageId="4" pageNumber="14" type="adult">two adult</specimenCount>
shells (see Table 2). The data in this table more or less imply that local variation in size and shape is fairly limited. However, of the
<specimenCount box="[221,372,1390,1416]" pageId="4" pageNumber="14" type="generic">24 specimens</specimenCount>
measured 11 are in the category 6.17.0 mm and
<quantity box="[918,960,1390,1416]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="4" pageNumber="14" unit="in" value="7.0">7 in</quantity>
7.18.0 mm, or, conversely,
<quantity box="[309,362,1422,1448]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="4" pageNumber="14" unit="in" value="18.0">18 in</quantity>
the joint category 6.18.0 mm against only
<quantity box="[830,870,1422,1448]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="4" pageNumber="14" unit="in" value="3.0">3 in</quantity>
the category 4.16.0mm and also only
<quantity box="[337,380,1454,1480]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="4" pageNumber="14" unit="in" value="3.0">3 in</quantity>
8.110.0 mm (see
<figureCitation box="[594,662,1454,1480]" captionStart="Fig" captionStartId="3.[144,177,1630,1651]" captionTargetBox="[172,1078,1068,1608]" captionTargetId="figure-410@3.[166,1085,1062,1614]" captionTargetPageId="3" captionText="Fig. 4. Frequency diagram of the length of the shell of Gulella inobstructa with subdivisions of 1.0 mm. Diagram generated by Ms Sancia van der Meij." pageId="4" pageNumber="14">Fig. 4</figureCitation>
). Incidentally, the sum total of the material is too limited for statistical analysis.
</paragraph>
<paragraph blockId="4.[181,1142,1260,1672]" pageId="4" pageNumber="14">
Looking at Table 1 it is clear that as regards shell length there are hardly any really
<emphasis bold="true" box="[181,1142,1548,1576]" pageId="4" pageNumber="14">VLJQL¿FDQW JDSV LQ WKH VHULHV ± WKH RQO\ RQH WKDW LV VHHPLQJO\ DEHUUDQW EHLQJ VKHOO QR. l,</emphasis>
<quantity box="[181,266,1582,1608]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="14" unit="mm" value="1.2">1.2 mm</quantity>
shorter than no. 2. The next gaps amount to no more than
<quantity box="[908,996,1582,1608]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="14" unit="mm" value="0.3">0.3 mm</quantity>
. However, in the greater lengths (&gt;
<quantity box="[415,500,1614,1640]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="4" pageNumber="14" unit="mm" value="7.7">7.7 mm</quantity>
) the gaps become increasingly larger, i.e.
<quantity box="[952,1035,1614,1640]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="14" unit="mm" value="0.6">0.6 mm</quantity>
(between nos. 21 and 22) and
<quantity box="[409,495,1646,1672]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="14" unit="mm" value="0.8">0.8 mm</quantity>
(between the
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, no. 22, and the
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, no. 23).
</paragraph>
<tableNote pageId="5" pageNumber="15" targetBox="[181,1071,297,461]" targetPageId="5">
<paragraph blockId="5.[144,1101,204,281]" box="[580,668,204,225]" pageId="5" pageNumber="15">TABLE 2</paragraph>
<paragraph blockId="5.[144,1101,204,281]" box="[144,1101,236,257]" pageId="5" pageNumber="15">
Shell measurements in
<taxonomicName authorityName="van Bruggen" authorityYear="1965" box="[354,531,237,256]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="15" phylum="Mollusca" rank="species" species="inobstructa">
<emphasis box="[354,531,237,256]" italics="true" pageId="5" pageNumber="15">Gulella inobstructa</emphasis>
</taxonomicName>
populations of which more than two adult shells are available.
</paragraph>
<paragraph blockId="5.[144,1101,204,281]" box="[144,939,260,281]" pageId="5" pageNumber="15">Samples W4319 and W2423 may be combined (cf. locality data and date of collecting).</paragraph>
</tableNote>
<paragraph pageId="5" pageNumber="15">
<table box="[181,1071,297,461]" gridcols="5" gridrows="4" pageId="5" pageNumber="15">
<tr box="[181,1071,297,342]" gridrow="0" pageId="5" pageNumber="15">
<th box="[181,378,297,342]" gridcol="0" gridrow="0" pageId="5" pageNumber="15">Measurements in mm</th>
<th box="[435,527,297,342]" gridcol="1" gridrow="0" pageId="5" pageNumber="15">l/d</th>
<th box="[571,670,297,342]" gridcol="2" gridrow="0" pageId="5" pageNumber="15">Number of whorls</th>
<th box="[723,872,297,342]" gridcol="3" gridrow="0" pageId="5" pageNumber="15">Number of adult shells measured</th>
<th box="[916,1071,297,342]" gridcol="4" gridrow="0" pageId="5" pageNumber="15">Sample</th>
</tr>
<tr box="[181,1071,353,374]" gridrow="1" pageId="5" pageNumber="15">
<th box="[181,378,353,374]" gridcol="0" gridrow="1" pageId="5" pageNumber="15">6.27.1 × 3.13.2</th>
<td box="[435,527,353,374]" gridcol="1" gridrow="1" pageId="5" pageNumber="15">2.002.19</td>
<td box="[571,670,353,374]" gridcol="2" gridrow="1" pageId="5" pageNumber="15">&lt;8</td>
<td box="[723,872,353,374]" gridcol="3" gridrow="1" pageId="5" pageNumber="15">3</td>
<td box="[916,1071,353,374]" gridcol="4" gridrow="1" pageId="5" pageNumber="15">RMNH</td>
</tr>
<tr box="[181,1071,385,406]" gridrow="2" pageId="5" pageNumber="15">
<th box="[181,378,385,406]" gridcol="0" gridrow="2" pageId="5" pageNumber="15">6.27.7 ×2.93.2</th>
<td box="[435,527,385,406]" gridcol="1" gridrow="2" pageId="5" pageNumber="15">2.082.48</td>
<td box="[571,670,385,406]" gridcol="2" gridrow="2" pageId="5" pageNumber="15">8+</td>
<td box="[723,872,385,406]" gridcol="3" gridrow="2" pageId="5" pageNumber="15">9</td>
<td box="[916,1071,385,406]" gridcol="4" gridrow="2" pageId="5" pageNumber="15">NMSA W5530</td>
</tr>
<tr box="[181,1071,416,461]" gridrow="3" pageId="5" pageNumber="15">
<th box="[181,378,416,461]" gridcol="0" gridrow="3" pageId="5" pageNumber="15">6.67.5 × 3.03.1</th>
<td box="[435,527,416,461]" gridcol="1" gridrow="3" pageId="5" pageNumber="15">2.212.40</td>
<td box="[571,670,416,461]" gridcol="2" gridrow="3" pageId="5" pageNumber="15">8</td>
<td box="[723,872,416,461]" gridcol="3" gridrow="3" pageId="5" pageNumber="15">4</td>
<td box="[916,1071,416,461]" gridcol="4" gridrow="3" pageId="5" pageNumber="15">NMSA W4319 + W2423</td>
</tr>
</table>
</paragraph>
<paragraph blockId="5.[144,1104,507,886]" pageId="5" pageNumber="15">
The major diameter or width of the shells varies from
<quantity box="[767,925,507,534]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.5" metricValueMin="2.5" pageId="5" pageNumber="15" unit="mm" value="3.5" valueMax="4.5" valueMin="2.5">2.5 to 4.5 mm</quantity>
. This is also unequally divided among the material, e.g. in the range
<quantity box="[747,884,539,566]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.2" metricValueMin="2.9" pageId="5" pageNumber="15" unit="mm" value="3.05" valueMax="3.2" valueMin="2.9">2.93.2 mm</quantity>
there are 18 shells, of which half (9) are
<quantity box="[387,478,571,598]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="5" pageNumber="15" unit="mm" value="3.1">3.1 mm</quantity>
. Large gaps here are 2.62.9 =
<quantity box="[839,929,571,598]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="15" unit="mm" value="0.3">0.3 mm</quantity>
,
<quantity box="[942,1080,571,598]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" metricValueMax="4.2" metricValueMin="3.2" pageId="5" pageNumber="15" unit="mm" value="3.7" valueMax="4.2" valueMin="3.2">3.24.2 mm</quantity>
= 1.0 mm, and
<quantity box="[293,430,603,630]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.5" metricValueMin="4.2" pageId="5" pageNumber="15" unit="mm" value="4.35" valueMax="4.5" valueMin="4.2">4.24.5 mm</quantity>
=
<quantity box="[459,548,603,630]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="15" unit="mm" value="0.3">0.3 mm</quantity>
.
</paragraph>
<paragraph blockId="5.[144,1104,507,886]" pageId="5" pageNumber="15">
The ratio length/major diameter (l/d) varies from 1.752.45 implying that on the one hand there are fairly squat specimens as opposed to distinctly cylindrical shells (Table 1 and
<figureCitation box="[211,278,699,725]" captionStart="Fig" captionStartId="5.[144,177,1650,1671]" captionTargetBox="[251,999,930,1631]" captionTargetId="figure-245@5.[247,1003,926,1635]" captionTargetPageId="5" captionText="Fig. 5. Scatter plot of length against major diameter of the shells of Gulella inobstructa and G. perspicua." pageId="5" pageNumber="15">Fig. 5</figureCitation>
). However, this is another series with small gaps, varying from 0.020.08 with a value of 0.14 as the single exception (between nos. 21 and 2, i.e. 1.82 and 1.96). There is a cluster here too, i.e. there are 16 shells with l/d values of between (and inclusive of) 1.95 and 2.25.
</paragraph>
<paragraph blockId="5.[144,1104,507,886]" lastBlockId="6.[181,1142,202,1672]" lastPageId="6" lastPageNumber="16" pageId="5" pageNumber="15">
<materialsCitation collectionCode="VDPH, DSSOLHV, WR, WKH, RXWHU, FROXPHOODU, SURFHVV, DSSHDUV, WKDW, VXSHU, FLDO, WKLFNHQLQJ" lastPageId="6" lastPageNumber="16" pageId="5" pageNumber="15" specimenCount="1">
The intergradation between smooth and costulate is slight; even pronounced growth striae may be considered costulae, so that no sharp delimitation is observed here. The
<emphasis bold="true" box="[181,1141,202,230]" pageId="6" pageNumber="16">
<collectionCode box="[181,682,202,230]" pageId="6" pageNumber="16">VDPH DSSOLHV WR WKH RXWHU FROXPHOODU SURFHVV</collectionCode>
.,W
<collectionCode box="[717,960,202,230]" pageId="6" pageNumber="16">DSSHDUV WKDW WKH VXSHU</collectionCode>
¿
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of the columellar lip, originally considered an apertural process, is of rare occurrence and has so far only been found in the
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material
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.
</paragraph>
<caption box="[144,1101,1650,1671]" pageId="5" pageNumber="15" startId="5.[144,177,1650,1671]" targetBox="[251,999,930,1631]" targetPageId="5" targetType="figure">
<paragraph blockId="5.[144,1101,1650,1671]" box="[144,1101,1650,1671]" pageId="5" pageNumber="15">
Fig. 5. Scatter plot of length against major diameter of the shells of
<taxonomicName authority="and G. perspicua" box="[757,1095,1650,1671]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="17" phylum="Mollusca" rank="species" species="inobstructa">
<emphasis box="[757,934,1652,1671]" italics="true" pageId="5" pageNumber="15">Gulella inobstructa</emphasis>
and
<emphasis box="[978,1095,1652,1671]" italics="true" pageId="5" pageNumber="15">G. perspicua</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph blockId="6.[181,1142,202,1672]" pageId="6" pageNumber="16">
To summarize, there are small squat, medium elongate, large squat, and large elongate shells. Would this be an indication that more than one taxon is involved? This is a matter of personal interpretation for the time being the present author prefers to consider this material only to represent
<taxonomicName box="[621,783,403,427]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="16" phylum="Mollusca" rank="species" species="inobstructa">
<emphasis box="[621,783,403,427]" italics="true" pageId="6" pageNumber="16">G. inobstructa</emphasis>
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of which the populations live in a fragmented environment resulting in a noticeable amount of individual variation in shell shape and size.
</paragraph>
<paragraph blockId="6.[181,1142,202,1672]" pageId="6" pageNumber="16">
The
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material (two shells only) seems to represent close to the maximum length for the shell (
<figureCitation box="[335,404,533,559]" captionStart="Fig" captionStartId="5.[144,177,1650,1671]" captionTargetBox="[251,999,930,1631]" captionTargetId="figure-245@5.[247,1003,926,1635]" captionTargetPageId="5" captionText="Fig. 5. Scatter plot of length against major diameter of the shells of Gulella inobstructa and G. perspicua." pageId="6" pageNumber="16">Fig. 5</figureCitation>
). As regards shape and apertural dentition, the extremely limited
<typeStatus box="[181,229,566,592]" pageId="6" pageNumber="16">type</typeStatus>
material also represents a slightly aberrant population. This leads to the conclusion that
<taxonomicName box="[232,396,599,623]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="16" phylum="Mollusca" rank="species" species="inobstructa">
<emphasis box="[232,396,599,623]" italics="true" pageId="6" pageNumber="16">G. inobstructa</emphasis>
</taxonomicName>
is a valid taxon but subject to considerable variation in shell size and shape. These data once again show that
<typeStatus box="[671,724,629,655]" pageId="6" pageNumber="16">type</typeStatus>
material is at times far from typical for the taxon involved.
</paragraph>
<paragraph blockId="6.[181,1142,202,1672]" pageId="6" pageNumber="16">
The phenomenon of great variation in size, implying that the smallest adult specimen is approximately less than half the size of the largest shell, is rare among species of
<taxonomicName authorityName="Pfeiffer" authorityYear="1856" box="[1052,1135,732,756]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="16" phylum="Mollusca" rank="genus">
<emphasis box="[1052,1135,732,756]" italics="true" pageId="6" pageNumber="16">Gulella</emphasis>
</taxonomicName>
, particularly those with a somewhat restricted distribution (vide e.g., van Bruggen 1980; see also
<bibRefCitation author="HERBERT, D. &amp; KILBURN, D." box="[281,563,794,820]" pageId="6" pageNumber="16" refId="ref4741" refString="HERBERT, D. &amp; KILBURN, D. 2004. Field guide to the land snails and slugs of eastern South Africa. Pietermaritzburg: Natal Museum." type="book" year="2004">Herbert &amp; Kilburn 2004</bibRefCitation>
). Some South African examples are
<taxonomicName authority="(Pfeiffer, 1859)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1859" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="16" phylum="Mollusca" rank="species" species="adamsiana">
<emphasis box="[982,1002,796,820]" italics="true" pageId="6" pageNumber="16">G</emphasis>
.
<emphasis box="[1019,1142,796,820]" italics="true" pageId="6" pageNumber="16">adamsiana</emphasis>
(Pfeiffer, 1859)
</taxonomicName>
with 6.511.0 mm (with a fairly large distribution in
<collectingRegion box="[921,1099,826,852]" country="South Africa" name="KwaZulu-Natal" pageId="6" pageNumber="16">KwaZulu-Natal</collectingRegion>
),
<taxonomicName authority="(Melvill &amp; Ponsonby, 1895)" baseAuthorityName="Melvill &amp; Ponsonby" baseAuthorityYear="1895" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="16" phylum="Mollusca" rank="species" species="farquhari">
<emphasis box="[1114,1134,828,852]" italics="true" pageId="6" pageNumber="16">G</emphasis>
.
<emphasis box="[181,287,860,884]" italics="true" pageId="6" pageNumber="16">farquhari</emphasis>
(Melvill &amp; Ponsonby, 1895)
</taxonomicName>
with
<quantity box="[664,799,858,885]" metricMagnitude="-3" metricUnit="m" metricValue="2.8499999999999996" metricValueMax="3.8" metricValueMin="1.9" pageId="6" pageNumber="16" unit="mm" value="2.8499999999999996" valueMax="3.8" valueMin="1.9">1.93.8 mm</quantity>
(includes data of van Bruggen, 1992, an in southern and East Africa very widely distributed species),
<taxonomicName authority="(Von Martens, 1866)" baseAuthorityName="Von Martens" baseAuthorityYear="1866" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="16" phylum="Mollusca" rank="species" species="infrendens">
<emphasis box="[936,956,892,916]" italics="true" pageId="6" pageNumber="16">G</emphasis>
.
<emphasis box="[968,1084,892,916]" italics="true" pageId="6" pageNumber="16">infrendens</emphasis>
(Von Martens, 1866)
</taxonomicName>
with
<quantity box="[414,549,922,949]" metricMagnitude="-3" metricUnit="m" metricValue="7.15" metricValueMax="9.1" metricValueMin="5.2" pageId="6" pageNumber="16" unit="mm" value="7.15" valueMax="9.1" valueMin="5.2">5.29.1 mm</quantity>
(a fairly widely distributed taxon in the
<collectingRegion box="[994,1142,922,949]" country="South Africa" name="Eastern Cape" pageId="6" pageNumber="16">Eastern Cape</collectingRegion>
and
<collectingRegion box="[231,411,954,980]" country="South Africa" name="KwaZulu-Natal" pageId="6" pageNumber="16">KwaZulu-Natal</collectingRegion>
), and
<taxonomicName authority="(Pfeiffer, 1856)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1856" box="[481,766,954,980]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="16" phylum="Mollusca" rank="species" species="planti">
<emphasis box="[481,582,956,980]" italics="true" pageId="6" pageNumber="16">G. planti</emphasis>
(Pfeiffer, 1856)
</taxonomicName>
with
<quantity box="[833,998,954,981]" metricMagnitude="-2" metricUnit="m" metricValue="1.71" metricValueMax="2.15" metricValueMin="1.27" pageId="6" pageNumber="16" unit="mm" value="17.1" valueMax="21.5" valueMin="12.7">12.721.5 mm</quantity>
(a restricted endemic in
<collectingRegion box="[312,493,986,1012]" country="South Africa" name="KwaZulu-Natal" pageId="6" pageNumber="16">KwaZulu-Natal</collectingRegion>
).
</paragraph>
<paragraph blockId="6.[181,1142,202,1672]" box="[208,794,1021,1049]" pageId="6" pageNumber="16">
<emphasis bold="true" box="[208,794,1021,1049]" pageId="6" pageNumber="16">7KH SUHVHQW WD[RQ PD\ QRZ EH UHGH¿QHG DV IROORZV:</emphasis>
</paragraph>
<paragraph blockId="6.[181,1142,202,1672]" pageId="6" pageNumber="16">
A normally smooth, medium-sized species of
<taxonomicName authorityName="Pfeiffer" authorityYear="1856" box="[720,804,1062,1086]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="16" phylum="Mollusca" rank="genus">
<emphasis box="[720,804,1062,1086]" italics="true" pageId="6" pageNumber="16">Gulella</emphasis>
</taxonomicName>
with only three processes in a little obstructed aperture, viz., a parietal (angular), a labral, and a columellar process (Verdcourts formula proposed in 1957/1962 1; 1; 0; 1); rimate or with closed umbilicus.,
<emphasis bold="true" box="[186,1141,1154,1183]" pageId="6" pageNumber="16">Q UDUH FDVHV JURZWK OLQHV PD\ GHYHORS LQWR IDLUO\ LQVLJQL¿FDQW FRVWXODWLRQ. $OVR, WKH</emphasis>
outer columellar wall of the lip of the aperture may exhibit a swelling that may be interpreted as an outer columellar process so that the apertural dentition then must be considered four-fold instead of three-fold (Verdcourts notation 1; 1; 0; 2). Anatomical and molecular data are as yet not available.
</paragraph>
<paragraph blockId="6.[181,1142,202,1672]" pageId="6" pageNumber="16">
Connollys group 3 (ii) should include this form, but all taxa are different in being smaller or having 4-fold instead of 3-fold dentition. Connollys group 3 (ii) is partly covered by groups 2A and 2B in
<bibRefCitation author="HERBERT, D. &amp; KILBURN, D." box="[545,910,1384,1410]" pageId="6" pageNumber="16" refId="ref4741" refString="HERBERT, D. &amp; KILBURN, D. 2004. Field guide to the land snails and slugs of eastern South Africa. Pietermaritzburg: Natal Museum." type="book" year="2004">Herbert and Kilburn (2004: 155)</bibRefCitation>
. Here, again, all taxa are different in being smaller or having 4-fold instead of 3-fold dentition. Also, none of the odd two dozen taxa of
<taxonomicName authorityName="Pfeiffer" authorityYear="1856" box="[516,600,1451,1475]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="16" phylum="Mollusca" rank="genus" sensu="lato">
<emphasis box="[516,600,1451,1475]" italics="true" pageId="6" pageNumber="16">Gulella</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[608,641,1448,1474]" pageId="6" pageNumber="16" sensu="lato">s.l.</taxonomicNameLabel>
described since Connollys treatise matches this diagnosis.
</paragraph>
<paragraph blockId="6.[181,1142,202,1672]" pageId="6" pageNumber="16">
<materialsCitation box="[208,1133,1517,1544]" collectorName="The" location="Limpopo" pageId="6" pageNumber="16" specimenCount="1" stateProvince="Limpopo">
<collectorName box="[208,252,1517,1543]" pageId="6" pageNumber="16">The</collectorName>
range of
<taxonomicName box="[358,520,1519,1543]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="16" phylum="Mollusca" rank="species" species="inobstructa">
<emphasis box="[358,520,1519,1543]" italics="true" pageId="6" pageNumber="16">G. inobstructa</emphasis>
</taxonomicName>
seems to be restricted to a limited area in SE
<collectingRegion box="[1031,1133,1517,1543]" country="South Africa" name="Limpopo" pageId="6" pageNumber="16">Limpopo</collectingRegion>
</materialsCitation>
/ NE
<collectingRegion box="[224,372,1549,1575]" country="South Africa" name="Mpumalanga" pageId="6" pageNumber="16">Mpumalanga</collectingRegion>
<materialsCitation latitude="-25.5" location="All" longLatPrecision="1246" longitude="31.0" pageId="6" pageNumber="16" specimenCount="1" stateProvince="Mpumalanga">
between 23°30'
<geoCoordinate box="[563,650,1549,1575]" degrees="25" direction="south" minutes="30" orientation="latitude" pageId="6" pageNumber="16" precision="925" value="-25.5">25°30'S</geoCoordinate>
and 29°45'
<geoCoordinate box="[787,876,1549,1575]" degrees="31" direction="east" minutes="00" orientation="longitude" pageId="6" pageNumber="16" precision="925" value="31.0">31°00'E</geoCoordinate>
where it lives in the leaf litter of afromontane forest situated between
<emphasis box="[688,707,1583,1607]" italics="true" pageId="6" pageNumber="16">c.</emphasis>
1100 and
<emphasis box="[825,844,1583,1607]" italics="true" pageId="6" pageNumber="16">c.</emphasis>
1900.
<location LSID="urn:lsid:plazi:treatment:D9379A342E24FFA2FE7EFA44FCFCFAC0:54417DF92E23FFACFC72F9E6FC09F9CD" box="[920,956,1581,1607]" latitude="-25.5" longLatPrecision="1246" longitude="31.0" name="All" pageId="6" pageNumber="16" stateProvince="Mpumalanga">All</location>
new records are from
<collectingRegion box="[242,393,1613,1639]" country="South Africa" name="Mpumalanga" pageId="6" pageNumber="16">Mpumalanga</collectingRegion>
</materialsCitation>
;
<materialsCitation latitude="-23.783333" location="Swaye's" longLatPrecision="1254" longitude="30.066668" pageId="6" pageNumber="16" specimenCount="1" stateProvince="Limpopo">
the
<typeStatus box="[444,492,1614,1640]" pageId="6" pageNumber="16">type</typeStatus>
locality (
<geoCoordinate box="[596,685,1613,1639]" degrees="23" direction="south" minutes="47" orientation="latitude" pageId="6" pageNumber="16" precision="925" value="-23.783333">23°47'S</geoCoordinate>
:
<geoCoordinate box="[689,783,1613,1639]" degrees="30" direction="east" minutes="04" orientation="longitude" pageId="6" pageNumber="16" precision="925" value="30.066668">30°04'E</geoCoordinate>
) and the two records in
<location LSID="urn:lsid:plazi:treatment:D9379A342E24FFA2FE7EFA44FCFCFAC0:54417DF92E23FFACFBF2F986FBC0F9ED" box="[1048,1141,1613,1639]" latitude="-23.783333" longLatPrecision="1254" longitude="30.066668" name="Swaye's" pageId="6" pageNumber="16" stateProvince="Limpopo">Swayes</location>
thesis are the only ones situated in
<collectingRegion box="[573,784,1645,1671]" country="South Africa" name="Limpopo" pageId="6" pageNumber="16">Limpopo province</collectingRegion>
</materialsCitation>
.
</paragraph>
<caption pageId="7" pageNumber="17" startId="7.[144,183,769,790]" targetBox="[406,882,204,749]" targetPageId="7" targetType="figure">
<paragraph blockId="7.[144,1104,769,862]" pageId="7" pageNumber="17">
Figs 6, 7. Type specimens of
<taxonomicName box="[404,546,769,790]" pageId="7" pageNumber="17">
<emphasis box="[404,472,770,789]" italics="true" pageId="7" pageNumber="17">Gulella</emphasis>
species
</taxonomicName>
: (6)
<taxonomicName baseAuthorityName="Melvill &amp; Ponsonby" baseAuthorityYear="1893" box="[585,701,770,789]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="17" phylum="Mollusca" rank="species" species="perspicua">
<emphasis box="[585,701,770,789]" italics="true" pageId="7" pageNumber="17">G. perspicua</emphasis>
</taxonomicName>
, “Transvaal” (= Middelburg), paralectotype, BM 1937.12.30.1123, length 4.8 mm; (7)
<taxonomicName authority=", Woodbush Forest" authorityName="Woodbush Forest" box="[639,951,793,814]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="17" phylum="Mollusca" rank="species" species="inobstructa">
<emphasis box="[639,774,794,813]" italics="true" pageId="7" pageNumber="17">G. inobstructa</emphasis>
, Woodbush Forest
</taxonomicName>
/Magoebaskloof, holotype, NMSA 4123/T1087, length 9.1 mm; note presence of outer columellar process in the aperture. Photos Dr D. Herbert, layout of plate by Ms Sancia van der Meij.
</paragraph>
</caption>
<paragraph blockId="7.[144,1105,902,1604]" box="[395,853,902,923]" pageId="7" pageNumber="17">
<taxonomicName authority="AND G. PERSPICUA" box="[395,853,902,923]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="17" phylum="Mollusca" rank="species" species="inobstructa">
<emphasis box="[395,646,904,923]" italics="true" pageId="7" pageNumber="17">GULELLA INOBSTRUCTA</emphasis>
AND
<emphasis box="[705,853,904,923]" italics="true" pageId="7" pageNumber="17">G. PERSPICUA</emphasis>
</taxonomicName>
</paragraph>
<paragraph blockId="7.[144,1105,902,1604]" pageId="7" pageNumber="17">
The shell of
<taxonomicName authority="(Melvill &amp; Ponsonby, 1893)" baseAuthorityName="Melvill &amp; Ponsonby" baseAuthorityYear="1893" box="[315,848,938,964]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="17" phylum="Mollusca" rank="species" species="perspicua">
<emphasis box="[315,518,940,964]" italics="true" pageId="7" pageNumber="17">Gulella perspicua</emphasis>
(
<bibRefCitation author="MELVILL, J. C. &amp; PONSONBY, J. H." box="[534,840,938,964]" pageId="7" pageNumber="17" pagination="19 - 24" refId="ref4770" refString="MELVILL, J. C. &amp; PONSONBY, J. H. 1893. Descriptions of thirteen new species of terrestrial Mollusca from South Africa. Annals and Magazine of Natural History (6) 11: 19 - 24." type="journal article" year="1893">Melvill &amp; Ponsonby, 1893</bibRefCitation>
)
</taxonomicName>
, partly sympatric with
<taxonomicName box="[144,305,972,996]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="17" phylum="Mollusca" rank="species" species="inobstructa">
<emphasis box="[144,305,972,996]" italics="true" pageId="7" pageNumber="17">G. inobstructa</emphasis>
</taxonomicName>
, shows the same
<typeStatus box="[500,548,971,997]" pageId="7" pageNumber="17">type</typeStatus>
of apertural dentition and comes within the range of measurements of the latter.
</paragraph>
<paragraph blockId="7.[144,1105,902,1604]" pageId="7" pageNumber="17">
<materialsCitation collectingDate="1978-07-09" collectionCode="RMNH, XVHXP, RQGRQ, LQ, FROOQ" collectorName="G. Deglon &amp; W. Falcon &amp; A. C. van Bruggen &amp; Waterval Boven &amp; H. E. van Hoepen &amp; The Barberton &amp; Connolly &amp; Material &amp; The Natural History &amp; U Herbert" country="South Africa" elevation="875" latitude="-25.633333" location="Barberton" longLatPrecision="1245" longitude="30.35" municipality="The" pageId="7" pageNumber="17" specimenCount="1" stateProvince="Mpumalanga" typeStatus="paralectotype">
<collectingMunicipality box="[170,214,1034,1060]" pageId="7" pageNumber="17">The</collectingMunicipality>
following material was examined for this study (
<collectingCountry box="[778,927,1034,1060]" name="South Africa" pageId="7" pageNumber="17">South Africa</collectingCountry>
,
<collectingRegion box="[939,1093,1034,1060]" country="South Africa" name="Mpumalanga" pageId="7" pageNumber="17">Mpumalanga</collectingRegion>
):
<location LSID="urn:lsid:plazi:treatment:D9379A342E24FFA2FE7EFA44FCFCFAC0:54417DF92E22FFADFF7AFBE1FEB6FBCE" box="[144,259,1066,1092]" country="South Africa" latitude="-25.633333" longLatPrecision="1245" longitude="30.35" municipality="The" name="Barberton" pageId="7" pageNumber="17" stateProvince="Mpumalanga">Barberton</location>
,
<geoCoordinate box="[269,359,1066,1092]" degrees="25" direction="south" minutes="47" orientation="latitude" pageId="7" pageNumber="17" precision="925" value="-25.783333">25°47'S</geoCoordinate>
:
<geoCoordinate box="[364,456,1066,1092]" degrees="31" direction="east" minutes="03" orientation="longitude" pageId="7" pageNumber="17" precision="925" value="31.05">31°03'E</geoCoordinate>
,
<emphasis box="[467,486,1068,1092]" italics="true" pageId="7" pageNumber="17">c.</emphasis>
<quantity box="[493,565,1066,1093]" metricMagnitude="2" metricUnit="m" metricValue="8.75" pageId="7" pageNumber="17" unit="m" value="875.0">
<elevation box="[493,565,1066,1093]" metricMagnitude="2" metricUnit="m" metricValue="8.75" pageId="7" pageNumber="17" unit="m" value="875.0">875 m</elevation>
</quantity>
, leg.
<collectorName box="[624,742,1066,1092]" pageId="7" pageNumber="17">G. Déglon</collectorName>
, ex
<collectorName box="[785,899,1066,1093]" pageId="7" pageNumber="17">W. Falcon</collectorName>
colln., don. Helen Boswell via
<collectorName box="[279,479,1098,1125]" pageId="7" pageNumber="17">A.C. van Bruggen</collectorName>
(2 shells,
<collectionCode box="[590,679,1098,1124]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="7" pageNumber="17" type="Museum">RMNH</collectionCode>
);
<collectorName box="[696,877,1098,1125]" pageId="7" pageNumber="17">Waterval Boven</collectorName>
,
<geoCoordinate box="[887,976,1098,1124]" degrees="25" direction="south" minutes="38" orientation="latitude" pageId="7" pageNumber="17" precision="925" value="-25.633333">25°38'S</geoCoordinate>
:
<geoCoordinate box="[980,1074,1098,1124]" degrees="30" direction="east" minutes="21" orientation="longitude" pageId="7" pageNumber="17" precision="925" value="30.35">30°21'E</geoCoordinate>
,
<emphasis box="[1084,1103,1100,1124]" italics="true" pageId="7" pageNumber="17">c.</emphasis>
<quantity box="[144,231,1130,1157]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="7" pageNumber="17" unit="m" value="1350.0">
<elevation box="[144,231,1130,1157]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="7" pageNumber="17" unit="m" value="1350.0">1350 m</elevation>
</quantity>
, leg./don.
<collectorName box="[348,544,1131,1157]" pageId="7" pageNumber="17">H.E. van Hoepen</collectorName>
,
<date box="[555,678,1130,1156]" pageId="7" pageNumber="17" value="1978-07-09">
<collectingDate box="[555,678,1130,1156]" pageId="7" pageNumber="17" value="1978-07-09">9.VII.1978</collectingDate>
</date>
(1 shell,
<collectionCode box="[783,872,1130,1156]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="7" pageNumber="17" type="Museum">RMNH</collectionCode>
).
<collectorName box="[890,1053,1130,1156]" pageId="7" pageNumber="17">The Barberton</collectorName>
material is particularly valuable as it was mentioned and probably examined by
<collectorName box="[1000,1104,1162,1188]" pageId="7" pageNumber="17">Connolly</collectorName>
himself (
<bibRefCitation author="CONNOLLY, M." box="[238,446,1194,1220]" pageId="7" pageNumber="17" pagination="1 - 660" refId="ref4662" refString="CONNOLLY, M. 1939. A monographic survey of South African non-marine Mollusca. Annals of the South African Museum 33: 1 - 660." type="journal article" year="1939">Connolly 1939: 33</bibRefCitation>
).
<collectorName box="[463,556,1194,1220]" pageId="7" pageNumber="17">Material</collectorName>
was also personally studied in
<collectorName box="[887,1105,1194,1220]" pageId="7" pageNumber="17">The Natural History</collectorName>
<emphasis bold="true" pageId="7" pageNumber="17">
0
<collectionCode box="[156,242,1225,1253]" pageId="7" pageNumber="17">XVHXP</collectionCode>
(/
<collectionCode box="[264,351,1224,1252]" pageId="7" pageNumber="17">RQGRQ</collectionCode>
)
<collectionCode box="[366,388,1224,1252]" pageId="7" pageNumber="17">LQ</collectionCode>
l967&gt;³7UDQVYDDO´, 3RQVRQE\
<collectionCode box="[741,806,1224,1252]" pageId="7" pageNumber="17">FROOQ</collectionCode>
., l9Ɵ..ll.44 (OHFWRW\SH), l9 7.lƻ. Ɵ.llƻ (SDUDOHFWRW\SH); %DUEHUWRQ (+.-. 3X]H\), l945.8.ƻ.ƻ6lƻ66@. '
<collectorName box="[1079,1103,1257,1285]" pageId="7" pageNumber="17">U</collectorName>
</emphasis>
Herbert has kindly supplied a photograph of the
<typeStatus box="[691,842,1290,1316]" pageId="7" pageNumber="17">paralectotype</typeStatus>
</materialsCitation>
<materialsCitation country="South Africa" location="London" pageId="7" pageNumber="17" specimenCount="1" stateProvince="Mpumalanga" typeStatus="lectotype">
shell from Middelburg (
<collectingRegion box="[151,302,1322,1348]" country="South Africa" name="Mpumalanga" pageId="7" pageNumber="17">Mpumalanga</collectingRegion>
)
<quantity box="[310,350,1322,1348]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="17" unit="in" value="1.0">
<superScript attach="left" box="[310,318,1322,1336]" fontSize="6" pageId="7" pageNumber="17">1</superScript>
in
</quantity>
<location LSID="urn:lsid:plazi:treatment:D9379A342E24FFA2FE7EFA44FCFCFAC0:54417DF92E22FFADFE8DFAE1FE0BFACE" box="[359,446,1322,1348]" country="South Africa" name="London" pageId="7" pageNumber="17" stateProvince="Mpumalanga">London</location>
(
<figureCitation box="[465,536,1322,1348]" captionStart="Figs 6, 7" captionStartId="7.[144,183,769,790]" captionTargetBox="[406,882,204,749]" captionTargetId="figure-260@7.[406,882,204,749]" captionTargetPageId="7" captionText="Figs 6, 7. Type specimens of Gulella species: (6) G. perspicua, “Transvaal” (= Middelburg), paralectotype, BM 1937.12.30.1123, length 4.8 mm; (7) G. inobstructa, Woodbush Forest/Magoebaskloof, holotype, NMSA 4123/T1087, length 9.1 mm; note presence of outer columellar process in the aperture. Photos Dr D. Herbert, layout of plate by Ms Sancia van der Meij." pageId="7" pageNumber="17">Fig. 6</figureCitation>
); this is in better condition than the now slightly damaged
<typeStatus box="[252,358,1354,1380]" pageId="7" pageNumber="17">lectotype</typeStatus>
</materialsCitation>
.
</paragraph>
<paragraph blockId="7.[144,1105,902,1604]" pageId="7" pageNumber="17">
Available shell measurements (see Table 3) give a preliminary range of 4.75.2 ×
<quantity box="[144,283,1418,1445]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.7" metricValueMin="2.4" pageId="7" pageNumber="17" unit="mm" value="2.55" valueMax="2.7" valueMin="2.4">2.42.7 mm</quantity>
, l/d 1.842.00, 77½ whorls, which is completely within the (lower parts of the) range of
<taxonomicName box="[331,496,1452,1476]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="17" phylum="Mollusca" rank="species" species="inobstructa">
<emphasis box="[331,496,1452,1476]" italics="true" pageId="7" pageNumber="17">G. inobstructa</emphasis>
</taxonomicName>
(
<figureCitation box="[512,583,1450,1476]" captionStart="Fig" captionStartId="5.[144,177,1650,1671]" captionTargetBox="[251,999,930,1631]" captionTargetId="figure-245@5.[247,1003,926,1635]" captionTargetPageId="5" captionText="Fig. 5. Scatter plot of length against major diameter of the shells of Gulella inobstructa and G. perspicua." pageId="7" pageNumber="17">Fig. 5</figureCitation>
). As regards apertural dentition there is some variation within populations, e.g. in the NMHUK series from Barberton. Four of the six shells exhibit a weak to very weak basal process, while the remaining two do not show any basal process at all. Otherwise this small sample is remarkably uniform in size, shape and whorls.
</paragraph>
<paragraph blockId="7.[144,958,1653,1675]" box="[144,958,1653,1675]" pageId="7" pageNumber="17">
<superScript attach="right" box="[144,151,1653,1665]" fontSize="5" pageId="7" pageNumber="17">1</superScript>
The
<typeStatus box="[193,232,1654,1675]" pageId="7" pageNumber="17">type</typeStatus>
locality “Transvaal” was later restricted to Middelburg by Connolly (1912: 83).
</paragraph>
<tableNote pageId="8" pageNumber="18" targetBox="[214,1115,273,509]" targetPageId="8">
<paragraph blockId="8.[255,1069,204,257]" box="[617,705,204,225]" pageId="8" pageNumber="18">TABLE 3</paragraph>
<paragraph blockId="8.[255,1069,204,257]" box="[255,1069,236,257]" pageId="8" pageNumber="18">
Shell measurements of
<taxonomicName baseAuthorityName="Melvill &amp; Ponsonby" baseAuthorityYear="1893" box="[467,630,237,256]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="18" phylum="Mollusca" rank="species" species="perspicua">
<emphasis box="[467,630,237,256]" italics="true" pageId="8" pageNumber="18">Gulella perspicua</emphasis>
</taxonomicName>
specimens studied in the context of this project.
</paragraph>
</tableNote>
<paragraph pageId="8" pageNumber="18">
<table box="[214,1115,273,509]" gridcols="5" gridrows="7" pageId="8" pageNumber="18">
<tr box="[214,1115,273,318]" gridrow="0" pageId="8" pageNumber="18">
<th box="[214,350,273,318]" gridcol="0" gridrow="0" pageId="8" pageNumber="18">Length × major diameter, mm</th>
<th box="[415,455,273,318]" gridcol="1" gridrow="0" pageId="8" pageNumber="18">l/d</th>
<th box="[499,762,273,318]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="8" pageNumber="18">Number of Locality whorls</th>
<th box="[797,1115,273,318]" gridcol="4" gridrow="0" pageId="8" pageNumber="18">Collection and/or additional data</th>
</tr>
<tr box="[214,1115,329,350]" gridrow="1" pageId="8" pageNumber="18">
<th box="[214,350,329,350]" gridcol="0" gridrow="1" pageId="8" pageNumber="18">4.7 ×2.5</th>
<td box="[415,455,329,350]" gridcol="1" gridrow="1" pageId="8" pageNumber="18">1.88</td>
<td box="[499,565,329,350]" gridcol="2" gridrow="1" pageId="8" pageNumber="18">&lt;7</td>
<td box="[616,762,329,350]" gridcol="3" gridrow="1" pageId="8" pageNumber="18">Waterval Boven</td>
<td box="[797,1115,329,350]" gridcol="4" gridrow="1" pageId="8" pageNumber="18">RMNH</td>
</tr>
<tr box="[214,1115,360,383]" gridrow="2" pageId="8" pageNumber="18">
<th box="[214,350,360,383]" gridcol="0" gridrow="2" pageId="8" pageNumber="18">4.8 ×2.4</th>
<td box="[415,455,360,383]" gridcol="1" gridrow="2" pageId="8" pageNumber="18">2.00</td>
<td box="[499,565,360,383]" gridcol="2" gridrow="2" pageId="8" pageNumber="18"></td>
<td box="[616,762,360,383]" gridcol="3" gridrow="2" pageId="8" pageNumber="18">Barberton</td>
<td box="[797,1115,360,383]" gridcol="4" gridrow="2" pageId="8" pageNumber="18">
<emphasis bold="true" box="[797,1022,360,383]" pageId="8" pageNumber="18">¿GH &amp;RQQROO\ (l9 9:)</emphasis>
</td>
</tr>
<tr box="[214,1115,392,413]" gridrow="3" pageId="8" pageNumber="18">
<th box="[214,350,392,413]" gridcol="0" gridrow="3" pageId="8" pageNumber="18">4.8 ×2.6</th>
<td box="[415,455,392,413]" gridcol="1" gridrow="3" pageId="8" pageNumber="18">1.84</td>
<td box="[499,565,392,413]" gridcol="2" gridrow="3" pageId="8" pageNumber="18">
<emphasis box="[532,542,394,413]" italics="true" pageId="8" pageNumber="18">c</emphasis>
. 7
</td>
<td box="[616,762,392,413]" gridcol="3" gridrow="3" pageId="8" pageNumber="18">“Transvaal”</td>
<td box="[797,1115,392,413]" gridcol="4" gridrow="3" pageId="8" pageNumber="18">NMHUK, paralectotype, Fig. 6</td>
</tr>
<tr box="[214,1115,424,445]" gridrow="4" pageId="8" pageNumber="18">
<th box="[214,350,424,445]" gridcol="0" gridrow="4" pageId="8" pageNumber="18">4.9 ×2.6</th>
<td box="[415,455,424,445]" gridcol="1" gridrow="4" pageId="8" pageNumber="18">1.86</td>
<td box="[499,565,424,445]" gridcol="2" gridrow="4" pageId="8" pageNumber="18">7</td>
<td box="[616,762,424,445]" gridcol="3" gridrow="4" pageId="8" pageNumber="18">Barberton</td>
<td box="[797,1115,424,445]" gridcol="4" gridrow="4" pageId="8" pageNumber="18">RMNH</td>
</tr>
<tr box="[214,1115,456,477]" gridrow="5" pageId="8" pageNumber="18">
<th box="[214,350,456,477]" gridcol="0" gridrow="5" pageId="8" pageNumber="18">5.0 ×2.7</th>
<td box="[415,455,456,477]" gridcol="1" gridrow="5" pageId="8" pageNumber="18">1.86</td>
<td box="[499,565,456,477]" gridcol="2" gridrow="5" pageId="8" pageNumber="18">7</td>
<td box="[616,762,456,477]" gridcol="3" gridrow="5" pageId="8" pageNumber="18">Barberton</td>
<td box="[797,1115,456,477]" gridcol="4" gridrow="5" pageId="8" pageNumber="18">RMNH</td>
</tr>
<tr box="[214,1115,488,509]" gridrow="6" pageId="8" pageNumber="18">
<th box="[214,350,488,509]" gridcol="0" gridrow="6" pageId="8" pageNumber="18">5.2 ×2.6</th>
<td box="[415,455,488,509]" gridcol="1" gridrow="6" pageId="8" pageNumber="18">2.00</td>
<td box="[499,565,488,509]" gridcol="2" gridrow="6" pageId="8" pageNumber="18">7</td>
<td box="[616,762,488,509]" gridcol="3" gridrow="6" pageId="8" pageNumber="18">Barberton</td>
<td box="[797,1115,488,509]" gridcol="4" gridrow="6" pageId="8" pageNumber="18">NMHUK</td>
</tr>
</table>
</paragraph>
<paragraph blockId="8.[181,1143,560,1354]" pageId="8" pageNumber="18">
Differences between the two taxa are fairly subtle, such as
<taxonomicName baseAuthorityName="Melvill &amp; Ponsonby" baseAuthorityYear="1893" box="[853,996,562,586]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="18" phylum="Mollusca" rank="species" species="perspicua">
<emphasis box="[853,996,562,586]" italics="true" pageId="8" pageNumber="18">G. perspicua</emphasis>
</taxonomicName>
having more convex whorls and a more prominent inner columellar process in a less ovate aperture; both taxa have rimate shells or a closed umbilicus. However, there is one consistent difference between
<taxonomicName baseAuthorityName="Melvill &amp; Ponsonby" baseAuthorityYear="1893" box="[394,535,658,682]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="18" phylum="Mollusca" rank="species" species="perspicua">
<emphasis box="[394,535,658,682]" italics="true" pageId="8" pageNumber="18">G. perspicua</emphasis>
</taxonomicName>
and
<taxonomicName box="[584,741,658,682]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="18" phylum="Mollusca" rank="species" species="inobstructa">
<emphasis box="[584,741,658,682]" italics="true" pageId="8" pageNumber="18">G. inobstructa</emphasis>
</taxonomicName>
. The suture of the former is narrowly indented, resulting in an excavated, trench-like line, which is visible under fairly high
<emphasis bold="true" box="[181,1052,718,746]" pageId="8" pageNumber="18">PDJQL¿FDWLRQ. 7KLV KDV EHHQ GHVFULEHG LQ GHWDLO E\ +HUEHUW IRU KLV QHZ VSHFLHV</emphasis>
<taxonomicName authority="Herbert, 2006" authorityName="Herbert" authorityYear="2006" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="18" phylum="Mollusca" rank="species" species="deviae">
<emphasis italics="true" pageId="8" pageNumber="18">Gulella deviae</emphasis>
<bibRefCitation author="HERBERT, D. G." box="[262,423,752,778]" pageId="8" pageNumber="18" pagination="1063 - 1081" refId="ref4689" refString="HERBERT, D. G. 2006. Rediscovery of the type species of Euonyma (Subulinidae) and observations on South African species of Gulella (Streptaxidae), with descriptions of two new species (Gastropoda: Eupulmonata). Journal of Natural History 40: 1063 - 1081." type="journal article" year="2006">Herbert, 2006</bibRefCitation>
</taxonomicName>
. This author refers to the suture of
<taxonomicName authority="(Herbert 2006: 1072)" baseAuthorityName="Herbert 2006:" baseAuthorityPageNumber="1072" baseAuthorityYear="1072" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="18" phylum="Mollusca" rank="species" species="perspicua">
<emphasis box="[821,967,754,778]" italics="true" pageId="8" pageNumber="18">G. perspicua</emphasis>
(
<bibRefCitation author="HERBERT, D. G." pageId="8" pageNumber="18" pagination="1063 - 1081" refId="ref4689" refString="HERBERT, D. G. 2006. Rediscovery of the type species of Euonyma (Subulinidae) and observations on South African species of Gulella (Streptaxidae), with descriptions of two new species (Gastropoda: Eupulmonata). Journal of Natural History 40: 1063 - 1081." type="journal article" year="2006">Herbert 2006: 1072</bibRefCitation>
)
</taxonomicName>
as an “unusual, roundly rebated suture”.
</paragraph>
<paragraph blockId="8.[181,1143,560,1354]" pageId="8" pageNumber="18">
Apart from
<bibRefCitation author="CONNOLLY, M." box="[335,516,816,842]" pageId="8" pageNumber="18" pagination="1 - 660" refId="ref4662" refString="CONNOLLY, M. 1939. A monographic survey of South African non-marine Mollusca. Annals of the South African Museum 33: 1 - 660." type="journal article" year="1939">Connolly (1939)</bibRefCitation>
only
<bibRefCitation author="AIKEN, D." box="[573,746,816,842]" pageId="8" pageNumber="18" pagination="3" refId="ref4445" refString="AIKEN, D. 1995. The Streptaxidae of South Africa. Special Publication of the Conchological Society of South Africa 6: 3 unnumbered pp. + 1 - 23." type="journal article" year="1995">Aiken (1995: 6)</bibRefCitation>
has contributed detailed distribution data for
<taxonomicName baseAuthorityName="Herbert 2006:" baseAuthorityYear="1072" box="[281,429,850,874]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="18" phylum="Mollusca" rank="species" species="perspicua">
<emphasis box="[281,429,850,874]" italics="true" pageId="8" pageNumber="18">G. perspicua</emphasis>
</taxonomicName>
:
<emphasis bold="true" box="[448,1143,846,875]" pageId="8" pageNumber="18">³0LGGHOEHUJ&gt;VLF!@, %DUEHUWRQ,:DWHUYDO %RYHQ, 2QGHUVDELH,</emphasis>
Wyliespoort, Strydom Tunnels”.His locality “Ondersabie”, undoubtedly derived from the Johannesburg psychiatrist/shell collector Dr H.E. van Hoepen, is somewhat misleading.
<emphasis bold="true" box="[181,1141,942,971]" pageId="8" pageNumber="18">2I¿FLDOO\ WKHUH LV RQO\ RQH 2QGHU 6DELH = /RZHU 6DELH LQ 6RXWK $IULFD, D ZHOONQRZQ</emphasis>
rest camp in the south-eastern districts of the Kruger National Park. This locality is in the Lowveld, characterized by a
<typeStatus box="[493,541,1008,1034]" pageId="8" pageNumber="18">type</typeStatus>
of savannah vegetation utterly different from that of the afromontane forest on the Drakensberg escarpment. Van Hoepen did extensively collect in the escarpment forests in
<collectingRegion box="[486,637,1072,1098]" country="South Africa" name="Mpumalanga" pageId="8" pageNumber="18">Mpumalanga</collectingRegion>
.As a genuine Afrikaner, van Hoepen did most of his labelling in Afrikaans. This is the reason why the present author prefers to interpret “Ondersabie” as a lower lying area of or near the town of Sabie (
<geoCoordinate box="[921,1011,1136,1162]" degrees="25" direction="south" minutes="06" orientation="latitude" pageId="8" pageNumber="18" precision="925" value="-25.1">25°06'S</geoCoordinate>
:
<geoCoordinate box="[1015,1110,1136,1162]" degrees="30" direction="east" minutes="47" orientation="longitude" pageId="8" pageNumber="18" precision="925" value="30.783333">30°47'E</geoCoordinate>
,
<emphasis box="[1122,1141,1138,1162]" italics="true" pageId="8" pageNumber="18">c.</emphasis>
<quantity box="[181,271,1168,1194]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="18" unit="m" value="1000.0">1000 m</quantity>
), nested in a deep valley on the escarpment, rather than the Kruger National Park locality. Thus the range of
<taxonomicName baseAuthorityName="Herbert 2006:" baseAuthorityYear="1072" box="[553,700,1202,1226]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="18" phylum="Mollusca" rank="species" species="perspicua">
<emphasis box="[553,700,1202,1226]" italics="true" pageId="8" pageNumber="18">G. perspicua</emphasis>
</taxonomicName>
stretches further south than that of
<emphasis italics="true" pageId="8" pageNumber="18">
<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="18" phylum="Mollusca" rank="species" species="inobstructa">G. inobstructa</taxonomicName>
.
</emphasis>
The
<typeStatus box="[376,424,1232,1258]" pageId="8" pageNumber="18">type</typeStatus>
locality of the former, Middelburg (
<geoCoordinate box="[838,928,1232,1258]" degrees="25" direction="south" minutes="46" orientation="latitude" pageId="8" pageNumber="18" precision="925" value="-25.766666">25°46'S</geoCoordinate>
:
<geoCoordinate box="[932,1026,1232,1258]" degrees="29" direction="east" minutes="28" orientation="longitude" pageId="8" pageNumber="18" precision="925" value="29.466667">29°28'E</geoCoordinate>
,
<quantity box="[1039,1128,1232,1258]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="8" pageNumber="18" unit="m" value="1450.0">1450 m</quantity>
), seems to be the southernmost limit of this taxon.
</paragraph>
<paragraph blockId="8.[181,1143,560,1354]" pageId="8" pageNumber="18">
<emphasis bold="true" box="[208,1141,1294,1323]" pageId="8" pageNumber="18">3HQGLQJ DQDWRPLFDO DQG PROHFXODU UHVHDUFK WKH DERYH LV VXI¿FLHQW HYLGHQFH WR FRQVLGHU</emphasis>
<taxonomicName box="[181,344,1330,1354]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="18" phylum="Mollusca" rank="species" species="inobstructa">
<emphasis box="[181,344,1330,1354]" italics="true" pageId="8" pageNumber="18">G. inobstructa</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Herbert 2006:" baseAuthorityYear="1072" box="[398,542,1330,1354]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="18" phylum="Mollusca" rank="species" species="perspicua">
<emphasis box="[398,542,1330,1354]" italics="true" pageId="8" pageNumber="18">G. perspicua</emphasis>
</taxonomicName>
to represent separate taxa.
</paragraph>
</subSubSection>
</treatment>
</document>