177 lines
20 KiB
XML
177 lines
20 KiB
XML
<document id="2A9115BD98426F8B4A6374B61DFA34D5" ID-DOI="10.1111/j.1096-3642.2010.00690.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5440663" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630450091946" checkinUser="valdenar" docAuthor="Krishnankutty, Sindhu M. & Dietrich, Christopher H." docDate="2011" docId="D929AD541970C502FAC7FBB5FBE2F9BD" docLanguage="en" docName="j.1096-3642.2010.00690.x.pdf" docOrigin="Zoological Journal of the Linnean Society 162 (3)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2010.00690.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Nesocerus fasika Krishnankutty & Dietrich 2011, SP. NOV." docType="treatment" docVersion="4" lastPageNumber="527" masterDocId="2510D52C196CC51EFA58FFDAFFB7FFBB" masterDocTitle="Taxonomic revision and phylogeny of an endemic leafhopper genus Nesocerus (Hemiptera: Cicadellidae: Idiocerinae) from Madagascar" masterLastPageNumber="543" masterPageNumber="499" pageNumber="527" updateTime="1699274970543" updateUser="plazi">
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<mods:titleInfo id="C4DB998720D0FD4DD4095DFBC4186416">
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<mods:title id="F15F90F4006280C55D9BD3DEC60379B9">Taxonomic revision and phylogeny of an endemic leafhopper genus Nesocerus (Hemiptera: Cicadellidae: Idiocerinae) from Madagascar</mods:title>
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<mods:roleTerm id="AD4A457AFA4A5DC5699EF0221BDFCC8E">Author</mods:roleTerm>
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<mods:namePart id="FE4ADFFDD3598CEB0CD1B46078E77F60">Krishnankutty, Sindhu M.</mods:namePart>
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<mods:roleTerm id="0C5B72BE796A92D31F38721E6CD94FBA">Author</mods:roleTerm>
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<mods:namePart id="420292DC89B3BA2FCD15D329E02E4489">Dietrich, Christopher H.</mods:namePart>
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<mods:typeOfResource id="FCAE224EC7C4902AC0785B0D06CA251E">text</mods:typeOfResource>
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<mods:title id="557159D03F680C1FCDB8F138157FBF8E">Zoological Journal of the Linnean Society</mods:title>
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</mods:titleInfo>
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<mods:part id="70EA6B3F9EC4E6CECC0C9E612D1D76E4">
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<mods:date id="73DD354E7BA4A39B7CAFBECFDD9146A8">2011</mods:date>
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<mods:detail id="69CE4F2931564CDD7B57046BF88D7ACD" type="pubDate">
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<mods:number id="F2F8F26603ECA75383B0385CA1D572B6">2011-06-24</mods:number>
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<mods:detail id="A4B609C1F64D48348E6EDAF520F5D4E7" type="volume">
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<mods:number id="C7C85354A888FD0205531CB2A0771732">162</mods:number>
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<mods:number id="38B1A9979B4532198DF584EDFDADED76">3</mods:number>
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<mods:url id="57BF423E3BCE5E392E6276AC8E2B3371">http://dx.doi.org/10.1111/j.1096-3642.2010.00690.x</mods:url>
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<mods:classification id="EFB4C1AFA535D3501CC713E43ACCD659">journal article</mods:classification>
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<mods:identifier id="C5B9AEC7AF955CA5284AFB33CC89FAC9" type="DOI">10.1111/j.1096-3642.2010.00690.x</mods:identifier>
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<mods:identifier id="3461946776C14705B6B419B93B993C56" type="ISSN">0024-4082</mods:identifier>
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<treatment id="D929AD541970C502FAC7FBB5FBE2F9BD" ID-DOI="http://doi.org/10.5281/zenodo.5491969" ID-Zenodo-Dep="5491969" LSID="urn:lsid:plazi:treatment:D929AD541970C502FAC7FBB5FBE2F9BD" httpUri="http://treatment.plazi.org/id/D929AD541970C502FAC7FBB5FBE2F9BD" lastPageNumber="527" pageId="28" pageNumber="527">
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<subSubSection id="199A4FC91970C502FAC7FBB5FD5FFB3E" box="[159,744,1135,1157]" pageId="28" pageNumber="527" type="nomenclature">
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<paragraph id="513F1C421970C502FAC7FBB5FD5FFB3E" blockId="28.[159,744,1135,1157]" box="[159,744,1135,1157]" pageId="28" pageNumber="527">
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<heading id="0A77AB2E1970C502FAC7FBB5FD5FFB3E" box="[159,744,1135,1157]" centered="true" fontSize="9" level="2" pageId="28" pageNumber="527" reason="4">
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<taxonomicName id="968067C11970C502FAC7FBB5FE3EFB38" ID-CoL="8JWWY" authority="Krishnankutty & Dietrich, 2011" authorityName="Krishnankutty & Dietrich" authorityYear="2011" box="[159,393,1135,1156]" class="Insecta" family="Cicadellidae" genus="Nesocerus" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="524" phylum="Arthropoda" rank="species" species="fasika" status="sp. nov.">NESOCERUS FASIKA</taxonomicName>
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<emphasis id="63F4C0501970C502FBCBFBA8FE4FFB3F" bold="true" box="[403,504,1135,1156]" pageId="28" pageNumber="527">
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<taxonomicNameLabel id="78C77D2B1970C502FBCBFBA8FE4FFB3F" box="[403,504,1135,1156]" pageId="28" pageNumber="527" rank="species">SP. NOV.</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="C9BB00C71970C502F853FBB5FD38FB3F" box="[523,655,1135,1156]" captionStart="Figure 3" captionStartId="9.[164,243,1834,1853]" captionTargetBox="[236,1371,204,1801]" captionTargetId="figure-0@9.[236,1371,195,1805]" captionTargetPageId="9" captionText="Figure 3. Nesocerus spp. nov., dorsal habitus and face of holotype male. A, B, Nesocerus trilineatus.; C, D, Nesocerus ranomafanaensis; E, F, Nesocerus angulatus; G, H, Nesocerus orbiculatus; I, J, Nesocerus affinis; K, L, Nesocerus sclerophyllus; M, N, Nesocerus bifurcatus; O, P, Nesocerus littoralis; Q, R, Nesocerus fasika; S, T, Nesocerus tulearensis." figureDoi="http://doi.org/10.5281/zenodo.5440669" httpUri="https://zenodo.org/record/5440669/files/figure.png" pageId="28" pageNumber="527">FIGS 3Q, R</figureCitation>
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,
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<figureCitation id="C9BB00C71970C502F8C5FBB5FD57FB3E" box="[669,736,1135,1157]" captionStart="Figure 8" captionStartId="14.[144,223,1834,1853]" captionTargetBox="[178,1398,202,1799]" captionTargetId="figure-0@14.[169,1398,195,1805]" captionTargetPageId="14" captionText="Figure 8. Nesocerus spp. nov. Male pygofer in lateral view (A, H, O), male subgenital plate in ventral (B, I, P) and lateral views (C, J, Q), aedeagus in lateral (D, K, R) and caudal views (E, L, S), style in lateral view (F, M, T), style and connective in dorsal view (G, N, U). A–G, Nesocerus fasika; H–N, Nesocerus tulearensis; O–U, Nesocerus recurvus." figureDoi="http://doi.org/10.5281/zenodo.5440679" httpUri="https://zenodo.org/record/5440679/files/figure.png" pageId="28" pageNumber="527">8A–G</figureCitation>
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)
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="199A4FC91970C502FAC8FB4CFBE2F9BD" pageId="28" pageNumber="527" type="diagnosis">
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<paragraph id="513F1C421970C502FAC8FB4CFDA0FA05" blockId="28.[144,759,1173,1471]" pageId="28" pageNumber="527">
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<emphasis id="63F4C0501970C502FAC8FB4CFEB1FB10" box="[144,262,1174,1195]" italics="true" pageId="28" pageNumber="527">Diagnosis:</emphasis>
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This species may be distinguished by the following unique combination of characters: frontoclypeus with pair of tiny circular ivory spots near ocelli and remainder covered with dark brown mottled markings, pygofer process forked in posterior view; style abruptly curved anterodorsad beyond middle with broad laterally compressed apex; connective with dorsal margin shallowly concave and aedeagal shaft slender, sinuate near apex with laterally directed short apical process.
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</paragraph>
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<paragraph id="513F1C421970C502FAC8FA32FDC7FA46" blockId="28.[144,624,1512,1534]" box="[144,624,1512,1534]" pageId="28" pageNumber="527">
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<emphasis id="63F4C0501970C502FAC8FA32FEAFFA46" box="[144,280,1512,1533]" italics="true" pageId="28" pageNumber="527">Description:</emphasis>
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Length of male,
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<quantity id="9678B1A71970C502FBBCFA32FDDBFA46" box="[484,620,1512,1533]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.4" metricValueMin="4.3" pageId="28" pageNumber="527" unit="mm" value="4.35" valueMax="4.4" valueMin="4.3">4.3–4.4 mm</quantity>
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.
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</paragraph>
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<paragraph id="513F1C421970C502FAC8F9FDFB1DFEED" blockId="28.[144,759,1575,1903]" lastBlockId="28.[806,1421,198,342]" pageId="28" pageNumber="527">
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<emphasis id="63F4C0501970C502FAC8F9FDFEB9F987" box="[144,270,1575,1596]" italics="true" pageId="28" pageNumber="527">Coloration:</emphasis>
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Crown mottled with light brown markings. Antennae with scape and flagellum yellow, pedicel light brown towards base. Frontoclypeus mottled with black or brown markings varying in darkness from crown to base of clypellus, two oval or circular yellow spots below ocelli. Gena with black marking. Clypellus completely black or with central small yellow spot. Pronotum with irregular black marking, lighter in posterior half. Mesonotum with basal black triangle without any border, median irregular pattern not extending beyond scutellar suture. Scutellum with light brown longitudinal lines. Forewing with claval vein white with alternate brown coloration. Legs yellow, fore and hind femur with black stripe near base and apex, hind femur with black stripe along dorsal margin.
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</paragraph>
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<paragraph id="513F1C421970C502F97EFEA5FBDFFDB4" blockId="28.[806,1421,383,527]" pageId="28" pageNumber="527">
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<emphasis id="63F4C0501970C502F97EFEA5FC2FFE2F" box="[806,920,383,404]" italics="true" pageId="28" pageNumber="527">Structure:</emphasis>
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Clypellus with sides concave, apex as wide as base, in profile flat. Rostrum extending beyond mid coxae. Forewing with r-m1 crossvein present. Metatibial setal rows PD, AD, and AV with 13–14, nine, and 11 macrosetae, respectively.
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</paragraph>
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<paragraph id="513F1C421970C502F97EFDE2FABEFB83" blockId="28.[806,1421,568,1080]" pageId="28" pageNumber="527">
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<emphasis id="63F4C0501970C502F97EFDE2FC63FDF6" box="[806,980,568,589]" italics="true" pageId="28" pageNumber="527">Male genitalia:</emphasis>
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Pygofer broad with acute posterodorsal process; posterolateral margin with short setae. Subgenital plate in lateral view depressed basally, broadened near apex, with short fine setae restricted to dorsum of apex; in ventral view with lateral margin strongly convex towards base, evenly narrowed toward apex, with submarginal row of macrosetae along apical half. Style in lateral view with broad apodeme; apophysis gradually narrowed along middle, then abruptly curved anterodorsad with broad laterally compressed apex. Connective in dorsal view semicircular, broadly flattened, fused to preatrium of aedeagus. Aedeagus in lateral view with atrium narrow; socle well developed; shaft in lateral view slender, S-shaped, tapered towards apex; apex in posterior view narrow, rounded, with pair of laterally directed short processes; gonopore apical.
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</paragraph>
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<paragraph id="513F1C421970C502F97EFBBBFB06FA31" blockId="28.[806,1422,1121,1418]" pageId="28" pageNumber="527">
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<emphasis id="63F4C0501970C502F97EFBBBFBB0FBCD" box="[806,1031,1121,1142]" italics="true" pageId="28" pageNumber="527">Material examined:</emphasis>
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<materialsCitation id="E1E8161F1970C502FE4BFBBBFCDAFAAB" collectingDate="2001-10-13" collectingDateMax="2001-11-12" collectingDateMin="2001-10-13" collectionCode="R, CAS" collectorName="M. E. Irwin & F. D. Parker & Harin'Hala" country="Madagascar" county="Hotel Paradisia" elevation="9" latitude="-23.179443" location="Province Tulear" longLatPrecision="21" longitude="43.616665" municipality="Ifaty" pageId="28" pageNumber="527" specimenCount="1" typeStatus="holotype">
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<typeStatus id="8E3BA2E01970C502FE4BFBBBFBCEFBCC" box="[1043,1145,1121,1143]" pageId="28" pageNumber="527" type="holotype">Holotype</typeStatus>
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male,
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<collectingCountry id="29975CD21970C502FE97FBBBFA3CFBCD" box="[1231,1419,1121,1142]" name="Madagascar" pageId="28" pageNumber="527">MADAGASCAR</collectingCountry>
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:
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<location id="545F4A991970C502F97EFBA5FC5EFB2E" LSID="urn:lsid:plazi:treatment:D929AD541970C502FAC7FBB5FBE2F9BD:545F4A991970C502F97EFBA5FC5EFB2E" box="[806,1001,1151,1173]" country="Madagascar" county="Hotel Paradisia" latitude="-23.179443" longLatPrecision="21" longitude="43.616665" municipality="Ifaty" name="Province Tulear" pageId="28" pageNumber="527">Province Tulear</location>
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,
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<collectingMunicipality id="B15B86381970C502FE5AFBA5FB8EFB2E" box="[1026,1081,1151,1173]" pageId="28" pageNumber="527">Ifaty</collectingMunicipality>
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, near
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<collectingCounty id="B85E64CE1970C502FEC2FBA5FAEAFB2E" box="[1178,1373,1151,1173]" pageId="28" pageNumber="527">Hotel Paradisia</collectingCounty>
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, in coastal dunes,
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,
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,
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<geoCoordinate id="34B47A851970C502FEECFB44FA83FB08" box="[1204,1332,1180,1203]" degrees="43" direction="east" minutes="37" orientation="longitude" pageId="28" pageNumber="527" precision="15" seconds="00" value="43.616665">43°37′00″E</geoCoordinate>
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,
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<date id="253E3A821970C502FF11FB44FC2AFB69" pageId="28" pageNumber="527" value="2001-10-13" valueMax="2001-11-12" valueMin="2001-10-13">
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<collectingDate id="357AC36A1970C502FF11FB44FC2AFB69" pageId="28" pageNumber="527" value="2001-10-13" valueMax="2001-11-12" valueMin="2001-10-13">13.x.– 12.xi.2001</collectingDate>
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</date>
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,
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<collectorName id="FC7579941970C502F9F3FB67FB81FB69" box="[939,1078,1212,1234]" pageId="28" pageNumber="527">M. E. Irwin</collectorName>
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,
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<collectorName id="FC7579941970C502FE1CFB67FB61FB69" box="[1092,1238,1212,1234]" pageId="28" pageNumber="527">F. D. Parker</collectorName>
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,
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<collectionCode id="379184871970C502FEBBFB67FB4FFB69" box="[1251,1272,1213,1234]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="28" pageNumber="527">R</collectionCode>
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.
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<collectorName id="FC7579941970C502FF5EFB66FA3EFB69" box="[1286,1417,1212,1234]" pageId="28" pageNumber="527">Harin’Hala</collectorName>
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,
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<collectingMethod id="88C164551970C502F97EFB01FC0CFB4B" box="[806,955,1243,1265]" pageId="28" pageNumber="527">malaise trap</collectingMethod>
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, vegetation in sandy area, MA-02-16-01 (
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||
<collectionCode id="379184871970C502F976FB20FCD2FAAB" box="[814,869,1274,1296]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="28" pageNumber="527" type="Museum">CAS</collectionCode>
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)
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||
</materialsCitation>
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.
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||
<materialsCitation id="E1E8161F1970C502F926FB20FC04FAD0" collectingDate="2002-01-18" collectingDateMax="2002-01-25" collectingDateMin="2002-01-18" collectionCode="R" collectorName="Parcelle I" country="Madagascar" elevation="165" latitude="-23.68861" location="Tulear Province" longLatPrecision="21" longitude="44.59083" municipality="Beza Mahafaly Reserve" pageId="28" pageNumber="527" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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<emphasis id="63F4C0501970C502F926FB20FC53FAB4" box="[894,996,1274,1295]" italics="true" pageId="28" pageNumber="527">
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<typeStatus id="8E3BA2E01970C502F926FB20FC53FAB4" box="[894,996,1274,1295]" pageId="28" pageNumber="527" type="paratype">Paratype</typeStatus>
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</emphasis>
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:
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||
<specimenCount id="4786D7CB1970C502F9AFFB20FBD1FAAB" box="[1015,1126,1274,1296]" pageId="28" pageNumber="527" type="male">one male</specimenCount>
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,
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<collectingCountry id="29975CD21970C502FE2FFB20FA84FAB4" box="[1143,1331,1274,1295]" name="Madagascar" pageId="28" pageNumber="527">MADAGASCAR</collectingCountry>
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:
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<location id="545F4A991970C502FF1BFB20FC3BFA95" LSID="urn:lsid:plazi:treatment:D929AD541970C502FAC7FBB5FBE2F9BD:545F4A991970C502FF1BFB20FC3BFA95" country="Madagascar" latitude="-23.68861" longLatPrecision="21" longitude="44.59083" municipality="Beza Mahafaly Reserve" name="Tulear Province" pageId="28" pageNumber="527">Tulear Province</location>
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||
,
|
||
<collectingMunicipality id="B15B86381970C502F9F8FAC3FB77FA95" box="[928,1216,1304,1326]" pageId="28" pageNumber="527">Beza Mahafaly Reserve</collectingMunicipality>
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||
,
|
||
<collectorName id="FC7579941970C502FE8CFAC2FAFEFA95" box="[1236,1353,1304,1326]" pageId="28" pageNumber="527">Parcelle I</collectorName>
|
||
near research station,
|
||
<quantity id="9678B1A71970C502F9B7FAEDFB80FAF7" box="[1007,1079,1335,1357]" metricMagnitude="2" metricUnit="m" metricValue="1.65" pageId="28" pageNumber="527" unit="m" value="165.0">
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<elevation id="DAADFB711970C502F9B7FAEDFB80FAF7" box="[1007,1079,1335,1357]" metricMagnitude="2" metricUnit="m" metricValue="1.65" pageId="28" pageNumber="527" unit="m" value="165.0">165 m</elevation>
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</quantity>
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,
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<geoCoordinate id="34B47A851970C502FE1DFAEDFB75FAF7" box="[1093,1218,1333,1356]" degrees="23" direction="south" minutes="41" orientation="latitude" pageId="28" pageNumber="527" precision="15" seconds="19" value="-23.68861">23°41′19″S</geoCoordinate>
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,
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<geoCoordinate id="34B47A851970C502FE88FAEDFAE7FAF7" box="[1232,1360,1333,1357]" degrees="44" direction="east" minutes="35" orientation="longitude" pageId="28" pageNumber="527" precision="15" seconds="27" value="44.59083">44°35′27″E</geoCoordinate>
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,
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<date id="253E3A821970C502FF05FAEDFC27FAD0" pageId="28" pageNumber="527" value="2002-01-18" valueMax="2002-01-25" valueMin="2002-01-18">
|
||
<collectingDate id="357AC36A1970C502FF05FAEDFC27FAD0" pageId="28" pageNumber="527" value="2002-01-18" valueMax="2002-01-25" valueMin="2002-01-18">18.– 25.i.2002</collectingDate>
|
||
</date>
|
||
,
|
||
<collectionCode id="379184871970C502F9C6FA8CFC04FAD0" box="[926,947,1366,1387]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="28" pageNumber="527">R</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="E1E8161F1970C502F998FA8FFB1AFA31" collectionCode="INHS" country="Madagascar" pageId="28" pageNumber="527" specimenCount="1" typeStatus="paratype">
|
||
Harin’Hala,
|
||
<collectingMethod id="88C164551970C502FE08FA8FFB55FAD0" box="[1104,1250,1365,1387]" pageId="28" pageNumber="527">malaise trap</collectingMethod>
|
||
in dry deciduous forest, MA-02-14A-12 (
|
||
<collectionCode id="379184871970C502FE07FAAEFB12FA31" box="[1119,1189,1396,1418]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" lsid="urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="28" pageNumber="527" type="Other type of research institution/biorepository">INHS</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="513F1C421970C502F97EFA69FBE2F9BD" blockId="28.[806,1421,1459,1542]" pageId="28" pageNumber="527">
|
||
<emphasis id="63F4C0501970C502F97EFA69FC13FA73" box="[806,932,1459,1480]" italics="true" pageId="28" pageNumber="527">Etymology:</emphasis>
|
||
The specific epithet is derived from the Malagasy word
|
||
<taxonomicName id="968067C11970C502F987FA0BFB93FA5D" authorityName="Krishnankutty & Dietrich" authorityYear="2011" box="[991,1060,1489,1510]" class="Insecta" family="Cicadellidae" genus="Nesocerus" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="527" phylum="Arthropoda" rank="species" species="fasika">
|
||
<emphasis id="63F4C0501970C502F987FA0BFB93FA5D" box="[991,1060,1489,1510]" italics="true" pageId="28" pageNumber="527">fasika</emphasis>
|
||
</taxonomicName>
|
||
meaning sandy area, referring to the sandy
|
||
<typeStatus id="8E3BA2E01970C502F99AFA2AFC44F9BE" box="[962,1011,1520,1541]" pageId="28" pageNumber="527">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |