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<document id="4CDCAA2560A58156CC8634B9EC07CD22" ID-DOI="10.1111/j.1096-3642.2010.00690.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5440663" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630450091946" checkinUser="valdenar" docAuthor="Krishnankutty, Sindhu M. &amp; Dietrich, Christopher H." docDate="2011" docId="D929AD541969C516F920FD27FE7DF965" docLanguage="en" docName="j.1096-3642.2010.00690.x.pdf" docOrigin="Zoological Journal of the Linnean Society 162 (3)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2010.00690.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Nesocerus ranomafanaensis Krishnankutty &amp; Dietrich 2011, SP. NOV." docType="treatment" docVersion="4" lastPageNumber="507" masterDocId="2510D52C196CC51EFA58FFDAFFB7FFBB" masterDocTitle="Taxonomic revision and phylogeny of an endemic leafhopper genus Nesocerus (Hemiptera: Cicadellidae: Idiocerinae) from Madagascar" masterLastPageNumber="543" masterPageNumber="499" pageNumber="504" updateTime="1699274970543" updateUser="plazi">
<mods:mods id="AF8A2DFF5ACC6AA469AEFCF62537F351" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="59F65E6E85BB10FAA10A775B109192CB">Taxonomic revision and phylogeny of an endemic leafhopper genus Nesocerus (Hemiptera: Cicadellidae: Idiocerinae) from Madagascar</mods:title>
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<mods:namePart id="088BBC9A24CE63F62F1F04CFC2AB950D">Krishnankutty, Sindhu M.</mods:namePart>
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<mods:namePart id="2B4F1A8F1206085E1C76C4678748FF60">Dietrich, Christopher H.</mods:namePart>
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<mods:title id="675C2757FE798AA18F448151438BDC8E">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="7CC588103AB5BFFE731EF731886A34C2">
<mods:date id="36533FD072A716C23E9EDC7C5535B443">2011</mods:date>
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<mods:number id="687A7B3793D813A2484D2CCE537F1BCD">2011-06-24</mods:number>
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<mods:number id="AE07BE46A45803A0A4B6295C1BAF24A7">162</mods:number>
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<mods:number id="431D1E8DBB473C22E306B8AF2C09F0C7">3</mods:number>
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<mods:identifier id="D8EA4EF83DFF1C861C882FF5ADAD4C28" type="DOI">10.1111/j.1096-3642.2010.00690.x</mods:identifier>
<mods:identifier id="CD3A81B081F05D65809B3D7CF96FDC10" type="ISSN">0024-4082</mods:identifier>
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<treatment id="D929AD541969C516F920FD27FE7DF965" ID-DOI="http://doi.org/10.5281/zenodo.5491961" ID-Zenodo-Dep="5491961" LSID="urn:lsid:plazi:treatment:D929AD541969C516F920FD27FE7DF965" httpUri="http://treatment.plazi.org/id/D929AD541969C516F920FD27FE7DF965" lastPageId="8" lastPageNumber="507" pageId="5" pageNumber="504">
<subSubSection id="199A4FC91969C51BF920FD27FAD4FCA9" box="[888,1379,765,786]" pageId="5" pageNumber="504" type="nomenclature">
<paragraph id="513F1C421969C51BF920FD27FAD4FCA9" blockId="5.[888,1379,765,818]" box="[888,1379,765,786]" pageId="5" pageNumber="504">
<heading id="0A77AB2E1969C51BF920FD27FAD4FCA9" box="[888,1379,765,786]" centered="true" fontSize="9" level="2" pageId="5" pageNumber="504" reason="4">
<taxonomicName id="968067C11969C51BF920FD27FB43FCAA" ID-CoL="8JWXB" authority="Krishnankutty &amp; Dietrich, 2011" authorityName="Krishnankutty &amp; Dietrich" authorityYear="2011" box="[888,1268,765,786]" class="Insecta" family="Cicadellidae" genus="Nesocerus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="511" phylum="Arthropoda" rank="species" species="ranomafanaensis" status="sp. nov.">NESOCERUS RANOMAFANAENSIS</taxonomicName>
<emphasis id="63F4C0501969C51BFEA6FCDAFAD4FCA9" bold="true" box="[1278,1379,765,786]" pageId="5" pageNumber="504">
<taxonomicNameLabel id="78C77D2B1969C51BFEA6FCDAFAD4FCA9" box="[1278,1379,765,786]" pageId="5" pageNumber="504" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="199A4FC91969C516F9A7FCC7FE7DF965" lastPageId="8" lastPageNumber="507" pageId="5" pageNumber="504" type="description">
<paragraph id="513F1C421969C51BF9A7FCC7FB6BFC89" blockId="5.[888,1379,765,818]" box="[1023,1244,796,818]" pageId="5" pageNumber="504">
(
<figureCitation id="C9BB00C71969C51BFE5FFCC7FB3CFC89" box="[1031,1163,797,818]" captionStart="Figure 3" captionStartId="9.[164,243,1834,1853]" captionTargetBox="[236,1371,204,1801]" captionTargetId="figure-0@9.[236,1371,195,1805]" captionTargetPageId="9" captionText="Figure 3. Nesocerus spp. nov., dorsal habitus and face of holotype male. A, B, Nesocerus trilineatus.; C, D, Nesocerus ranomafanaensis; E, F, Nesocerus angulatus; G, H, Nesocerus orbiculatus; I, J, Nesocerus affinis; K, L, Nesocerus sclerophyllus; M, N, Nesocerus bifurcatus; O, P, Nesocerus littoralis; Q, R, Nesocerus fasika; S, T, Nesocerus tulearensis." figureDoi="http://doi.org/10.5281/zenodo.5440669" httpUri="https://zenodo.org/record/5440669/files/figure.png" pageId="5" pageNumber="504">FIGS 3C, D</figureCitation>
,
<figureCitation id="C9BB00C71969C51BFEC2FCC6FB63FC89" box="[1178,1236,796,818]" captionStart="Figure 6" captionStartId="12.[144,223,1803,1822]" captionTargetBox="[200,1374,199,1770]" captionTargetId="figure-0@12.[192,1376,195,1773]" captionTargetPageId="12" captionText="Figure 6. Nesocerus spp. nov. Male pygofer in lateral view (A, I, R), male subgenital plate in ventral (B, J, T) and lateral views (C, K, U), aedeagus in lateral (D, L, P, V, Z) and caudal views (E, M, Q, W, A1), style in lateral view (F, N, X), style and connective in dorsal view (G, O, Y), apex of male pygofer in caudal view (H, S). A–H, Nesocerus trilineatus; I–O, Nesocerus ranomafanaensis; P–Q, Nesocerus affinis; R–Y, Nesocerus angulatus; Z–A1, Nesocerus sclerophyllus." figureDoi="http://doi.org/10.5281/zenodo.5440675" httpUri="https://zenodo.org/record/5440675/files/figure.png" pageId="5" pageNumber="504">6I–O</figureCitation>
)
</paragraph>
<paragraph id="513F1C421969C51BF962FC9EFB11FC68" blockId="5.[826,1441,835,980]" pageId="5" pageNumber="504">
<emphasis id="63F4C0501969C51BF962FC9EFC06FCE2" box="[826,945,836,857]" italics="true" pageId="5" pageNumber="504">Diagnosis:</emphasis>
This species is closely similar in habitus and male genitalia to
<taxonomicName id="968067C11969C51BFE11FCB8FB01FCCC" authorityName="Krishnankutty &amp; Dietrich" authorityYear="2011" box="[1097,1206,866,887]" class="Insecta" family="Cicadellidae" genus="Nesocerus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="504" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="63F4C0501969C51BFE11FCB8FB01FCCC" box="[1097,1206,866,887]" italics="true" pageId="5" pageNumber="504">N. affinis</emphasis>
</taxonomicName>
, but can be distinguished by the presence of a pair of long lateral processes on the aedeagal shaft and the presence of crossvein r-m
<subScript id="CD041E071969C51BF98EFC12FC69FC6F" attach="left" box="[982,990,968,980]" fontSize="5" pageId="5" pageNumber="504">1</subScript>
on the forewing.
</paragraph>
<paragraph id="513F1C421969C51BF962FC27FC35FB8B" blockId="5.[826,1441,1020,1072]" pageId="5" pageNumber="504">
<emphasis id="63F4C0501969C51BF962FC27FC75FBA9" box="[826,962,1021,1042]" italics="true" pageId="5" pageNumber="504">Description:</emphasis>
Length of male,
<quantity id="9678B1A71969C51BFEE8FC26FA80FBA9" box="[1200,1335,1020,1042]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="4.6" metricValueMin="4.4" pageId="5" pageNumber="504" unit="mm" value="4.5" valueMax="4.6" valueMin="4.4">4.4–4.6 mm</quantity>
; female,
<quantity id="9678B1A71969C51BF962FBC0FCC9FB8B" box="[826,894,1050,1072]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="504" unit="mm" value="5.0">5 mm</quantity>
.
</paragraph>
<paragraph id="513F1C421969C51BF962FB83FA34F981" blockId="5.[826,1441,1113,1594]" pageId="5" pageNumber="504">
<emphasis id="63F4C0501969C51BF962FB83FC0EFBD5" box="[826,953,1113,1134]" italics="true" pageId="5" pageNumber="504">Coloration:</emphasis>
Crown with three to four black markings extending to face reaching antennal ledge, median spots usually closer, forming v-shaped marking extending laterad to ocellus. Antennae with scape and pedicel light brown, flagellum dull yellow. Frontoclypeus with median broad black T-shaped stripe; clypeogenal and laterofrontal sutures bordered with black marking. Gena unmarked. Clypellus completely black or with central small yellow spot. Pronotum with black median markings arranged transversely along anterior region, a few irregular black markings close to posterior margin. Mesonotum with basal black triangle bordered by light brown line, not extending beyond scutellar suture. Forewing mostly hyaline. Legs yellow, fore and hind femur with pale brown lines along anterior and posterior margins.
</paragraph>
<paragraph id="513F1C421969C51BF962F9B9FBCBF948" blockId="5.[826,1441,1635,1779]" pageId="5" pageNumber="504">
<emphasis id="63F4C0501969C51BF962F9B9FC1BF9C3" box="[826,940,1635,1656]" italics="true" pageId="5" pageNumber="504">Structure:</emphasis>
Clypellus with sides concave, apex as wide as base; in profile flat. Rostrum extending beyond mid coxae. Forewing with r-m1 crossvein present. Metatibial setal rows PD, AD, and AV with 13–14, nine, and 11 macrosetae, respectively.
</paragraph>
<paragraph id="513F1C421969C516F962F8C6FE96FCD8" blockId="5.[826,1442,1820,1903]" lastBlockId="8.[144,759,478,867]" lastPageId="8" lastPageNumber="507" pageId="5" pageNumber="504">
<emphasis id="63F4C0501969C51BF962F8C6FC51F88A" box="[826,998,1820,1841]" italics="true" pageId="5" pageNumber="504">Male genitalia:</emphasis>
Pygofer, style, and connective as in
<taxonomicName id="968067C11969C51BFFDEF8C6FC06F8F4" authorityName="Krishnankutty &amp; Dietrich" authorityYear="2011" class="Insecta" family="Cicadellidae" genus="Nesocerus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="504" phylum="Arthropoda" rank="species" species="trilineatus">
<emphasis id="63F4C0501969C51BFFDEF8C6FC06F8F4" italics="true" pageId="5" pageNumber="504">N. trilineatus</emphasis>
</taxonomicName>
. Subgenital plate in lateral view depressed basally, broadened near apex and short fine setae almost equal to length of macrosetae restricted to dorsum of apex; in ventral view lateral margin strongly convex towards base, evenly narrowed toward apex, with biseriate submarginal macrosetae extending along distal two thirds. Aedeagus in lateral view with atrium broad; socle well developed; shaft in lateral view directed dorsally, broad, nearly straight and uniform in width, apex in posterior view acuminate, with pair of long ventrolaterally directed subapical processes extending to about midlength of shaft, apex of processes gradually curved laterad; gonopore subapical on posterior surface of shaft basad of processes.
</paragraph>
<paragraph id="513F1C421964C516FAC8FC56FE2EF93B" blockId="8.[144,760,908,1664]" pageId="8" pageNumber="507">
<emphasis id="63F4C0501964C516FAC8FC56FEC6FC1A" box="[144,369,908,929]" italics="true" pageId="8" pageNumber="507">Material examined:</emphasis>
<materialsCitation id="E1E8161F1964C516FB24FC56FDDDFBA7" collectingDate="2002-02-12" collectingDateMax="2002-02-19" collectingDateMin="2002-02-12" collectionCode="R, CAS" country="Madagascar" county="Province" elevation="1020" latitude="-21.266388" location="Belle Vue" longLatPrecision="21" longitude="47.42389" municipality="Parc National Ranomafana" pageId="8" pageNumber="507" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="8E3BA2E01964C516FB24FC56FE55FC19" box="[380,482,908,930]" pageId="8" pageNumber="507" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="29975CD21964C516F860FC56FD43FC1A" box="[568,756,908,929]" name="Madagascar" pageId="8" pageNumber="507">MADAGASCAR</collectingCountry>
:
<collectingCounty id="B85E64CE1964C516FAC8FC71FF44FC7A" box="[144,243,939,961]" pageId="8" pageNumber="507">Province</collectingCounty>
<collectingRegion id="9344D2A01964C516FB58FC71FE2BFC7A" box="[256,412,939,961]" country="Madagascar" name="Fianarantsoa" pageId="8" pageNumber="507">Fianarantsoa</collectingRegion>
,
<collectingMunicipality id="B15B86381964C516FBF5FC71FD44FC7A" box="[429,755,939,961]" pageId="8" pageNumber="507">Parc National Ranomafana</collectingMunicipality>
,
<location id="545F4A991964C516FAC8FC13FEA6FC64" LSID="urn:lsid:plazi:treatment:D929AD541969C516F920FD27FE7DF965:545F4A991964C516FAC8FC13FEA6FC64" box="[144,273,969,991]" country="Madagascar" county="Province" latitude="-21.266388" longLatPrecision="21" longitude="47.42389" municipality="Parc National Ranomafana" name="Belle Vue" pageId="8" pageNumber="507" stateProvince="Fianarantsoa">Belle Vue</location>
at
<location id="545F4A991964C516FB3BFC13FE6BFC64" LSID="urn:lsid:plazi:treatment:D929AD541969C516F920FD27FE7DF965:545F4A991964C516FB3BFC13FE6BFC64" box="[355,476,969,991]" country="Madagascar" county="Province" latitude="-21.266388" longLatPrecision="21" longitude="47.42389" municipality="Parc National Ranomafana" name="Talatakely" pageId="8" pageNumber="507" stateProvince="Fianarantsoa">Talatakely</location>
,
<quantity id="9678B1A71964C516FBA5FC13FDE5FC64" box="[509,594,969,991]" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="8" pageNumber="507" unit="m" value="1020.0">
<elevation id="DAADFB711964C516FBA5FC13FDE5FC64" box="[509,594,969,991]" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="8" pageNumber="507" unit="m" value="1020.0">1020 m</elevation>
</quantity>
,
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,
<geoCoordinate id="34B47A851964C516FAC8FC32FEA7FC46" box="[144,272,998,1022]" degrees="47" direction="east" minutes="25" orientation="longitude" pageId="8" pageNumber="507" precision="15" seconds="26" value="47.42389">47°25′26″E</geoCoordinate>
,
<date id="253E3A821964C516FB7CFC32FE72FC45" box="[292,453,1000,1022]" pageId="8" pageNumber="507" value="2002-02-12" valueMax="2002-02-19" valueMin="2002-02-12">
<collectingDate id="357AC36A1964C516FB7CFC32FE72FC45" box="[292,453,1000,1022]" pageId="8" pageNumber="507" value="2002-02-12" valueMax="2002-02-19" valueMin="2002-02-12">12.–19.ii.2002</collectingDate>
</date>
,
<collectionCode id="379184871964C516FB81FC32FE59FC46" box="[473,494,1000,1021]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="507">R</collectionCode>
. Harin’Hala, malaise, secondary forest, MA-02-09C-16 (
<collectionCode id="379184871964C516F873FBDCFDD5FBA7" box="[555,610,1030,1052]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="8" pageNumber="507" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E1E8161F1964C516F827FBDDFD44FBC3" collectingDate="2002-04-14" collectingDateMax="2002-04-23" collectingDateMin="2002-04-14" collectionCode="R" country="Madagascar" county="Province" elevation="1020" latitude="-21.266388" location="Belle Vue" longLatPrecision="21" longitude="47.42389" municipality="Parc National Ranomafana" pageId="8" pageNumber="507" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<emphasis id="63F4C0501964C516F827FBDDFD47FBA7" box="[639,752,1031,1052]" italics="true" pageId="8" pageNumber="507">
<typeStatus id="8E3BA2E01964C516F827FBDDFD47FBA7" box="[639,752,1031,1052]" pageId="8" pageNumber="507" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="4786D7CB1964C516FAC8FBFFFEB8FB80" box="[144,271,1061,1083]" pageId="8" pageNumber="507" type="male">four males</specimenCount>
,
<collectingCountry id="29975CD21964C516FB46FBFFFE6DFB81" box="[286,474,1061,1082]" name="Madagascar" pageId="8" pageNumber="507">MADAGASCAR</collectingCountry>
:
<collectingCounty id="B85E64CE1964C516FBB0FBFFFDFCFB80" box="[488,587,1061,1083]" pageId="8" pageNumber="507">Province</collectingCounty>
<collectingRegion id="9344D2A01964C516F80EFBFFFD45FB80" box="[598,754,1061,1083]" country="Madagascar" name="Fianarantsoa" pageId="8" pageNumber="507">Fianarantsoa</collectingRegion>
,
<collectingMunicipality id="B15B86381964C516FAC8FB9EFE67FBE2" box="[144,464,1091,1113]" pageId="8" pageNumber="507">Parc National Ranomafana</collectingMunicipality>
,
<location id="545F4A991964C516FB86FB99FDFAFBE2" LSID="urn:lsid:plazi:treatment:D929AD541969C516F920FD27FE7DF965:545F4A991964C516FB86FB99FDFAFBE2" box="[478,589,1091,1113]" country="Madagascar" county="Province" latitude="-21.266388" longLatPrecision="21" longitude="47.42389" municipality="Parc National Ranomafana" name="Belle Vue" pageId="8" pageNumber="507" stateProvince="Fianarantsoa">Belle Vue</location>
at
<location id="545F4A991964C516F821FB99FD45FBE2" LSID="urn:lsid:plazi:treatment:D929AD541969C516F920FD27FE7DF965:545F4A991964C516F821FB99FD45FBE2" box="[633,754,1091,1113]" country="Madagascar" county="Province" latitude="-21.266388" longLatPrecision="21" longitude="47.42389" municipality="Parc National Ranomafana" name="Talatakely" pageId="8" pageNumber="507" stateProvince="Fianarantsoa">Talatakely</location>
,
<quantity id="9678B1A71964C516FAC8FBB8FF52FBC3" box="[144,229,1122,1144]" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="8" pageNumber="507" unit="m" value="1020.0">
<elevation id="DAADFB711964C516FAC8FBB8FF52FBC3" box="[144,229,1122,1144]" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="8" pageNumber="507" unit="m" value="1020.0">1020 m</elevation>
</quantity>
,
<geoCoordinate id="34B47A851964C516FAA3FBB8FECFFBCC" box="[251,376,1121,1144]" degrees="21" direction="south" minutes="15" orientation="latitude" pageId="8" pageNumber="507" precision="15" seconds="59" value="-21.266388">21°15′59″S</geoCoordinate>
,
<geoCoordinate id="34B47A851964C516FBD6FBB8FDB9FBC3" box="[398,526,1121,1144]" degrees="47" direction="east" minutes="25" orientation="longitude" pageId="8" pageNumber="507" precision="15" seconds="26" value="47.42389">47°25′26″E</geoCoordinate>
,
<date id="253E3A821964C516F87CFBB8FD7EFBC3" box="[548,713,1122,1144]" pageId="8" pageNumber="507" value="2002-04-14" valueMax="2002-04-23" valueMin="2002-04-14">
<collectingDate id="357AC36A1964C516F87CFBB8FD7EFBC3" box="[548,713,1122,1144]" pageId="8" pageNumber="507" value="2002-04-14" valueMax="2002-04-23" valueMin="2002-04-14">14.–23.iv.2002</collectingDate>
</date>
,
<collectionCode id="379184871964C516F886FBB9FD44FBC3" box="[734,755,1123,1144]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="507">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="E1E8161F1964C516FAC8FB5BFE92FB0E" collectionCode="CAS, INHS" country="Madagascar" location="Harin'Hala" pageId="8" pageNumber="507" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<location id="545F4A991964C516FAC8FB5BFEA4FB2C" LSID="urn:lsid:plazi:treatment:D929AD541969C516F920FD27FE7DF965:545F4A991964C516FAC8FB5BFEA4FB2C" box="[144,275,1153,1175]" country="Madagascar" name="Harin'Hala" pageId="8" pageNumber="507" stateProvince="Fianarantsoa">Harin’Hala</location>
, malaise, secondary forest, MA-02-09C-25 (
<collectionCode id="379184871964C516FACFFB45FF7AFB0F" box="[151,205,1183,1204]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="8" pageNumber="507" type="Museum">CAS</collectionCode>
,
<collectionCode id="379184871964C516FA80FB45FEAAFB0E" box="[216,285,1183,1205]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" lsid="urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="8" pageNumber="507" type="Other type of research institution/biorepository">INHS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E1E8161F1964C516FB68FB7AFE49FAAA" collectingDate="2001-11-28" collectingDateMax="2001-12-06" collectingDateMin="2001-11-28" collectionCode="R" country="Madagascar" county="Province" elevation="1110" latitude="-21.226112" location="Vohiparara" longLatPrecision="21" longitude="47.36972" municipality="Parc National Ranomafana" pageId="8" pageNumber="507" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<specimenCount id="4786D7CB1964C516FB68FB7AFE11FB0E" box="[304,422,1183,1205]" pageId="8" pageNumber="507" type="male">two males</specimenCount>
,
<specimenCount id="4786D7CB1964C516FBE8FB7AFD9AFB0E" box="[432,557,1183,1205]" pageId="8" pageNumber="507" type="female">one female</specimenCount>
,
<collectingCountry id="29975CD21964C516F860FB45FD43FB0F" box="[568,756,1183,1204]" name="Madagascar" pageId="8" pageNumber="507">MADAGASCAR</collectingCountry>
:
<collectingCounty id="B85E64CE1964C516FAC8FB64FF44FB6F" box="[144,243,1214,1236]" pageId="8" pageNumber="507">Province</collectingCounty>
<collectingRegion id="9344D2A01964C516FB58FB64FE2BFB6F" box="[256,412,1214,1236]" country="Madagascar" name="Fianarantsoa" pageId="8" pageNumber="507">Fianarantsoa</collectingRegion>
,
<collectingMunicipality id="B15B86381964C516FBF5FB64FD44FB6F" box="[429,755,1214,1236]" pageId="8" pageNumber="507">Parc National Ranomafana</collectingMunicipality>
,
<location id="545F4A991964C516FAC8FB06FEA7FB49" LSID="urn:lsid:plazi:treatment:D929AD541969C516F920FD27FE7DF965:545F4A991964C516FAC8FB06FEA7FB49" box="[144,272,1244,1266]" country="Madagascar" county="Province" latitude="-21.226112" longLatPrecision="21" longitude="47.36972" municipality="Parc National Ranomafana" name="Vohiparara" pageId="8" pageNumber="507" stateProvince="Fianarantsoa">Vohiparara</location>
at broken bridge,
<quantity id="9678B1A71964C516F853FB07FDE9FB49" box="[523,606,1245,1266]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="8" pageNumber="507" unit="m" value="1110.0">
<elevation id="DAADFB711964C516F853FB07FDE9FB49" box="[523,606,1245,1266]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="8" pageNumber="507" unit="m" value="1110.0">1110 m</elevation>
</quantity>
,
<geoCoordinate id="34B47A851964C516F82CFB07FD46FB49" box="[628,753,1243,1266]" degrees="21" direction="south" minutes="13" orientation="latitude" pageId="8" pageNumber="507" precision="15" seconds="34" value="-21.226112">21°13′34″S</geoCoordinate>
,
<geoCoordinate id="34B47A851964C516FAC8FB21FEB9FAAA" box="[144,270,1274,1297]" degrees="47" direction="east" minutes="22" orientation="longitude" pageId="8" pageNumber="507" precision="15" seconds="11" value="47.36972">47°22′11″E</geoCoordinate>
,
<date id="253E3A821964C516FB45FB21FE6CFAAA" box="[285,475,1275,1297]" pageId="8" pageNumber="507" value="2001-11-28" valueMax="2001-12-06" valueMin="2001-11-28">
<collectingDate id="357AC36A1964C516FB45FB21FE6CFAAA" box="[285,475,1275,1297]" pageId="8" pageNumber="507" value="2001-11-28" valueMax="2001-12-06" valueMin="2001-11-28">28.xi.–6.xii.2001</collectingDate>
</date>
,
<collectionCode id="379184871964C516FBB1FB26FE49FAAA" box="[489,510,1276,1297]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="507">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="E1E8161F1964C516F854FB21FF6AFAF5" collectionCode="CAS, INHS" country="Madagascar" location="Harin'Hala" pageId="8" pageNumber="507" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<location id="545F4A991964C516F854FB21FD38FAAA" LSID="urn:lsid:plazi:treatment:D929AD541969C516F920FD27FE7DF965:545F4A991964C516F854FB21FD38FAAA" box="[524,655,1275,1297]" country="Madagascar" name="Harin'Hala" pageId="8" pageNumber="507" stateProvince="Fianarantsoa">Harin’Hala</location>
,
<collectingMethod id="88C164551964C516F8C6FB21FF77FA94" pageId="8" pageNumber="507">malaise trap</collectingMethod>
in high altitude rainforest, MA-02-09A-05 (
<collectionCode id="379184871964C516F8E4FAC0FD45FA94" box="[700,754,1306,1327]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="8" pageNumber="507" type="Museum">CAS</collectionCode>
,
<collectionCode id="379184871964C516FAC8FAE2FF62FAF5" box="[144,213,1336,1358]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" lsid="urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="8" pageNumber="507" type="Other type of research institution/biorepository">INHS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E1E8161F1964C516FAADFAE3FD74FA5C" collectingDate="1998-04-18" collectionCode="CAS" collectorName="C. E. Griswold &amp; D. H. Kavanaugh &amp; N. D. Penny &amp; M. J. Raherilalao &amp; J. S. Ranorianarisoa &amp; Jere Schweikert. &amp; Ubick" country="Madagascar" elevation="1100" latitude="-21.22389" location="Vohiparara" longLatPrecision="21" longitude="47.380833" municipality="Parc National Ranomafana" pageId="8" pageNumber="507" specimenCode="CASENT 303332" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<specimenCount id="4786D7CB1964C516FAADFAE3FEDCFAF5" box="[245,363,1336,1358]" pageId="8" pageNumber="507" type="male">one male</specimenCount>
,
<collectingCountry id="29975CD21964C516FBDCFAE2FDF7FAF6" box="[388,576,1336,1357]" name="Madagascar" pageId="8" pageNumber="507">MADAGASCAR</collectingCountry>
:
<collectingRegion id="9344D2A01964C516F80EFAE2FD45FAF5" box="[598,754,1336,1358]" country="Madagascar" name="Fianarantsoa" pageId="8" pageNumber="507">Fianarantsoa</collectingRegion>
,
<collectingMunicipality id="B15B86381964C516FAC8FA8DFE63FAD6" box="[144,468,1367,1389]" pageId="8" pageNumber="507">Parc National Ranomafana</collectingMunicipality>
,
<quantity id="9678B1A71964C516FBBDFA8DFD85FAD6" box="[485,562,1367,1389]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="8" pageNumber="507" unit="km" value="2.3">2.3 km</quantity>
north of village
<location id="545F4A991964C516FAC8FAAFFEA5FA30" LSID="urn:lsid:plazi:treatment:D929AD541969C516F920FD27FE7DF965:545F4A991964C516FAC8FAAFFEA5FA30" box="[144,274,1397,1419]" country="Madagascar" latitude="-21.22389" longLatPrecision="21" longitude="47.380833" municipality="Parc National Ranomafana" name="Vohiparara" pageId="8" pageNumber="507" stateProvince="Fianarantsoa">Vohiparara</location>
,
<quantity id="9678B1A71964C516FB19FAACFE22FA30" box="[321,405,1398,1419]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="507" unit="m" value="1100.0">
<elevation id="DAADFB711964C516FB19FAACFE22FA30" box="[321,405,1398,1419]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="507" unit="m" value="1100.0">1100 m</elevation>
</quantity>
,
<geoCoordinate id="34B47A851964C516FB9CFAACFDF5FA30" box="[452,578,1396,1419]" degrees="21" direction="south" minutes="13" orientation="latitude" pageId="8" pageNumber="507" precision="15" seconds="26" value="-21.22389">21°13′26″S</geoCoordinate>
,
<geoCoordinate id="34B47A851964C516F82AFAACFD45FA30" box="[626,754,1396,1419]" degrees="47" direction="east" minutes="22" orientation="longitude" pageId="8" pageNumber="507" precision="15" seconds="51" value="47.380833">47°22′51″E</geoCoordinate>
,
<date id="253E3A821964C516FAC8FA4EFEB3FA11" box="[144,260,1428,1450]" pageId="8" pageNumber="507" value="1998-04-18">
<collectingDate id="357AC36A1964C516FAC8FA4EFEB3FA11" box="[144,260,1428,1450]" pageId="8" pageNumber="507" value="1998-04-18">18.iv.1998</collectingDate>
</date>
,
<collectorName id="FC7579941964C516FB4CFA4EFE72FA11" box="[276,453,1428,1450]" pageId="8" pageNumber="507">C. E. Griswold</collectorName>
,
<collectorName id="FC7579941964C516FB8CFA4FFD11FA11" box="[468,678,1428,1450]" pageId="8" pageNumber="507">D. H. Kavanaugh</collectorName>
,
<collectorName id="FC7579941964C516F8EEFA4FFF6FFA73" pageId="8" pageNumber="507">N. D. Penny</collectorName>
,
<collectorName id="FC7579941964C516FABEFA69FE02FA72" box="[230,437,1459,1481]" pageId="8" pageNumber="507">M. J. Raherilalao</collectorName>
,
<collectorName id="FC7579941964C516FB9CFA69FD00FA72" box="[452,695,1459,1481]" pageId="8" pageNumber="507">J. S. Ranorianarisoa</collectorName>
,
<collectorName id="FC7579941964C516F89DFA69FEA0FA5C" pageId="8" pageNumber="507">Jere Schweikért.</collectorName>
&amp; D.
<collectorName id="FC7579941964C516FB06FA0BFE13FA5C" box="[350,420,1489,1511]" pageId="8" pageNumber="507">Ubick</collectorName>
,
<specimenCode id="0126B4391964C516FBE8FA0BFDC3FA5D" box="[432,628,1489,1510]" collectionCode="CASENT" pageId="8" pageNumber="507">CASENT 303332</specimenCode>
(
<collectionCode id="379184871964C516F8DCFA0BFD0CFA5C" box="[644,699,1489,1511]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="8" pageNumber="507" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E1E8161F1964C516F897FA08FF42F9DA" collectingDate="2002-01-21" collectingDateMax="2002-01-28" collectingDateMin="2002-01-21" collectionCode="R" collectorName="M. Irwin" country="Madagascar" elevation="1130" latitude="-22.250834" location="Parc National Ranomafana" longLatPrecision="21" longitude="47.406944" municipality="Province" pageId="8" pageNumber="507" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<specimenCount id="4786D7CB1964C516F897FA08FF7DF9BD" pageId="8" pageNumber="507" type="male">one male</specimenCount>
,
<collectingCountry id="29975CD21964C516FA83FA2AFE20F9BE" box="[219,407,1520,1541]" name="Madagascar" pageId="8" pageNumber="507">MADAGASCAR</collectingCountry>
:
<collectingMunicipality id="B15B86381964C516FBFEFA2AFDBEF9BD" box="[422,521,1520,1542]" pageId="8" pageNumber="507">Province</collectingMunicipality>
<collectingRegion id="9344D2A01964C516F84EFA2AFD05F9BD" box="[534,690,1520,1542]" country="Madagascar" name="Fianarantsoa" pageId="8" pageNumber="507">Fianarantsoa</collectingRegion>
,
<location id="545F4A991964C516F89AFA2AFE20F99F" LSID="urn:lsid:plazi:treatment:D929AD541969C516F920FD27FE7DF965:545F4A991964C516F89AFA2AFE20F99F" country="Madagascar" latitude="-22.250834" longLatPrecision="21" longitude="47.406944" municipality="Province" name="Parc National Ranomafana" pageId="8" pageNumber="507" stateProvince="Fianarantsoa">Parc National Ranomafana</location>
, radio tower at forest edge,
<quantity id="9678B1A71964C516FAC8F9F7FF53F9F8" box="[144,228,1581,1603]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="8" pageNumber="507" unit="m" value="1130.0">
<elevation id="DAADFB711964C516FAC8F9F7FF53F9F8" box="[144,228,1581,1603]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="8" pageNumber="507" unit="m" value="1130.0">1130 m</elevation>
</quantity>
,
<geoCoordinate id="34B47A851964C516FAA7F9F7FED9F9F9" box="[255,366,1580,1603]" degrees="22" direction="south" minutes="15" orientation="latitude" pageId="8" pageNumber="507" precision="15" seconds="3" value="-22.250834">22°15′3″S</geoCoordinate>
,
<geoCoordinate id="34B47A851964C516FBD1F9F7FDBEF9F8" box="[393,521,1580,1603]" degrees="47" direction="east" minutes="24" orientation="longitude" pageId="8" pageNumber="507" precision="15" seconds="25" value="47.406944">47°24′25″E</geoCoordinate>
,
<date id="253E3A821964C516F87CF9F7FD09F9F8" box="[548,702,1581,1603]" pageId="8" pageNumber="507" value="2002-01-21" valueMax="2002-01-28" valueMin="2002-01-21">
<collectingDate id="357AC36A1964C516F87CF9F7FD09F9F8" box="[548,702,1581,1603]" pageId="8" pageNumber="507" value="2002-01-21" valueMax="2002-01-28" valueMin="2002-01-21">21.–28.i.2002</collectingDate>
</date>
,
<collectorName id="FC7579941964C516F880F9F4FF66F9D9" pageId="8" pageNumber="507">M. Irwin</collectorName>
,
<collectionCode id="379184871964C516FAB8F996FF42F9DA" box="[224,245,1612,1633]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="507">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="E1E8161F1964C516FB5BF996FE22F93B" collectionCode="INHS" country="Madagascar" location="Harin'Hala" pageId="8" pageNumber="507" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<location id="545F4A991964C516FB5BF996FE31F9D9" LSID="urn:lsid:plazi:treatment:D929AD541969C516F920FD27FE7DF965:545F4A991964C516FB5BF996FE31F9D9" box="[259,390,1612,1634]" country="Madagascar" name="Harin'Hala" pageId="8" pageNumber="507" stateProvince="Fianarantsoa">Harin’Hala</location>
, malaise, mixed tropical forest, MA-02-09B-13 (
<collectionCode id="379184871964C516FB1FF9B0FE3AF93B" box="[327,397,1642,1664]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" lsid="urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="8" pageNumber="507" type="Other type of research institution/biorepository">INHS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="513F1C421964C516FAC8F973FE7DF965" blockId="8.[144,758,1705,1758]" pageId="8" pageNumber="507">
<emphasis id="63F4C0501964C516FAC8F973FEBAF905" box="[144,269,1705,1726]" italics="true" pageId="8" pageNumber="507">Etymology:</emphasis>
The specific epithet is derived from the
<typeStatus id="8E3BA2E01964C516FAC8F912FF76F966" box="[144,193,1736,1757]" pageId="8" pageNumber="507">type</typeStatus>
locality ‘Ranomafana’.
</paragraph>
</subSubSection>
</treatment>
</document>
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