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<document id="257F723A8213C83D9F880915C6EDD782" ID-DOI="10.1111/j.1096-3642.2010.00690.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5440663" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630450091946" checkinUser="valdenar" docAuthor="Krishnankutty, Sindhu M. &amp; Dietrich, Christopher H." docDate="2011" docId="D929AD541944C537FAA9FB11FB25FECE" docLanguage="en" docName="j.1096-3642.2010.00690.x.pdf" docOrigin="Zoological Journal of the Linnean Society 162 (3)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2010.00690.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Nesocerus convolutus Krishnankutty &amp; Dietrich 2011, SP. NOV." docType="treatment" docVersion="3" lastPageNumber="540" masterDocId="2510D52C196CC51EFA58FFDAFFB7FFBB" masterDocTitle="Taxonomic revision and phylogeny of an endemic leafhopper genus Nesocerus (Hemiptera: Cicadellidae: Idiocerinae) from Madagascar" masterLastPageNumber="543" masterPageNumber="499" pageNumber="539" updateTime="1699274970543" updateUser="plazi">
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<mods:title id="351CEAA3C957B0D1C261F009F566E8E0">Taxonomic revision and phylogeny of an endemic leafhopper genus Nesocerus (Hemiptera: Cicadellidae: Idiocerinae) from Madagascar</mods:title>
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<mods:namePart id="D2514DF6324AC264B25EECCC8751B084">Krishnankutty, Sindhu M.</mods:namePart>
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<mods:namePart id="BF045EEA6B4ECE27760D2277BA155D08">Dietrich, Christopher H.</mods:namePart>
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<mods:title id="23F4C0E8BF0EFE7B21EC21FD24036AED">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="9B5D85F7C87B20DC34A987495C52DDC2">2011</mods:date>
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<mods:number id="B79708A032B8D9EAE32A6FA9DD130ED0">2011-06-24</mods:number>
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<mods:number id="293DC01A89AD94D844C163A90311E990">162</mods:number>
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<mods:number id="E2DCAEAA2D8CC8536E87C557CD95A32E">3</mods:number>
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<mods:identifier id="C6A470654243210042AA084D5FF3A1CC" type="DOI">10.1111/j.1096-3642.2010.00690.x</mods:identifier>
<mods:identifier id="3BE4EE18048D8E2B24C30C5C27169287" type="ISSN">0024-4082</mods:identifier>
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<treatment id="D929AD541944C537FAA9FB11FB25FECE" LSID="urn:lsid:plazi:treatment:D929AD541944C537FAA9FB11FB25FECE" httpUri="http://treatment.plazi.org/id/D929AD541944C537FAA9FB11FB25FECE" lastPageId="41" lastPageNumber="540" pageId="40" pageNumber="539">
<subSubSection id="199A4FC91944C536FAA9FB11FD22FB64" box="[241,661,1226,1248]" pageId="40" pageNumber="539" type="nomenclature">
<paragraph id="513F1C421944C536FAA9FB11FD22FB64" blockId="40.[241,661,1226,1280]" box="[241,661,1226,1248]" pageId="40" pageNumber="539">
<heading id="0A77AB2E1944C536FAA9FB11FD22FB64" box="[241,661,1226,1248]" centered="true" fontSize="9" level="2" pageId="40" pageNumber="539" reason="4">
<taxonomicName id="968067C11944C536FAA9FB11FD91FB64" ID-CoL="8JWWV" authority="Krishnankutty &amp; Dietrich, 2011" authorityName="Krishnankutty &amp; Dietrich" authorityYear="2011" box="[241,550,1227,1248]" class="Insecta" family="Cicadellidae" genus="Nesocerus" kingdom="Animalia" order="Hemiptera" pageId="40" pageNumber="523" phylum="Arthropoda" rank="species" species="convolutus" status="sp. nov.">NESOCERUS CONVOLUTUS</taxonomicName>
<emphasis id="63F4C0501944C536F868FB14FD22FB64" bold="true" box="[560,661,1226,1247]" pageId="40" pageNumber="539">
<taxonomicNameLabel id="78C77D2B1944C536F868FB14FD22FB64" box="[560,661,1226,1247]" pageId="40" pageNumber="539" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="199A4FC91944C537FB10FB30FDABFD02" lastPageId="41" lastPageNumber="540" pageId="40" pageNumber="539" type="description">
<paragraph id="513F1C421944C536FB10FB30FD89FABB" blockId="40.[241,661,1226,1280]" box="[328,574,1258,1280]" pageId="40" pageNumber="539">
(
<figureCitation id="C9BB00C71944C536FB08FB30FE62FABB" box="[336,469,1258,1280]" captionStart="Figure 5" captionStartId="11.[164,243,1834,1853]" captionTargetBox="[200,1409,199,1799]" captionTargetId="figure-0@11.[198,1410,195,1805]" captionTargetPageId="11" captionText="Figure 5. Nesocerus spp. nov., dorsal habitus and face of holotype male. A, B, Nesocerus asymmetricus; C, D, Nesocerus uniprocessus; E, F, Nesocerus compressus; G, H, Nesocerus flexus; I, J, Nesocerus breviprocessus; K, L, Nesocerus latus; M, N, Nesocerus basiprocessus; O, P, Nesocerus serratus; Q, R, Nesocerus convolutus." figureDoi="http://doi.org/10.5281/zenodo.5440673" httpUri="https://zenodo.org/record/5440673/files/figure.png" pageId="40" pageNumber="539">FIGS 5Q, R</figureCitation>
,
<figureCitation id="C9BB00C71944C536FBBBFB30FD81FABB" box="[483,566,1258,1280]" captionStart="Figure 14" captionStartId="20.[144,223,1834,1853]" captionTargetBox="[185,1389,195,1801]" captionTargetId="figure-0@20.[177,1392,195,1805]" captionTargetPageId="20" captionText="Figure 14. Nesocerus spp. nov. Male pygofer in lateral view (A, H, O), male subgenital plate in ventral view (B, I, P) and lateral views (C, J, Q), aedeagus in lateral view (D, K, R) and caudal (E, L, S) views, style in lateral view (F, M, T), style and connective in dorsal view (G, N, U). A–G, Nesocerus basiprocessus; H–N, Nesocerus serratus; O–U, Nesocerus convolutus." figureDoi="http://doi.org/10.5281/zenodo.5440691" httpUri="https://zenodo.org/record/5440691/files/figure.png" pageId="40" pageNumber="539">14O–U</figureCitation>
)
</paragraph>
<paragraph id="513F1C421944C536FAC8FACBFE8FFA47" blockId="40.[144,759,1297,1533]" pageId="40" pageNumber="539">
<emphasis id="63F4C0501944C536FAC8FACBFEB1FA9D" box="[144,262,1297,1318]" italics="true" pageId="40" pageNumber="539">Diagnosis:</emphasis>
This species may be distinguished by the following unique characters: presence of median brown band along costa of forewing; pygofer with short process directed posterad along posterolateral margin and aedeagal shaft twisted near apex with single short process, in caudal view with lateral flaps extending from base to subapex and pair of dentate apical process.
</paragraph>
<paragraph id="513F1C421944C536FAC8F9FCFDF9F980" blockId="40.[144,590,1573,1595]" box="[144,590,1573,1595]" pageId="40" pageNumber="539">
<emphasis id="63F4C0501944C536FAC8F9FCFEAFF980" box="[144,280,1574,1595]" italics="true" pageId="40" pageNumber="539">Description:</emphasis>
Length of male,
<quantity id="9678B1A71944C536FBBCF9FFFDFEF980" box="[484,585,1573,1595]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" pageId="40" pageNumber="539" unit="mm" value="3.75">3.75 mm</quantity>
.
</paragraph>
<paragraph id="513F1C421944C536FAC8F9BEFA39FE28" blockId="40.[144,759,1636,1903]" lastBlockId="40.[806,1422,198,403]" pageId="40" pageNumber="539">
<emphasis id="63F4C0501944C536FAC8F9BEFEB9F9C2" box="[144,270,1636,1657]" italics="true" pageId="40" pageNumber="539">Coloration:</emphasis>
Crown yellow with three black markings; median one V-shaped extending to face, lateral ones comma-shaped with yellow spot at the apex. Antennae with pedicel and flagellum black, scape yellow. Frontoclypeus with short, median black stripe not extending near base of clypellus. Laterofrontal suture and clypeogenal suture with black marking close to ocelli and base of clypellus, respectively. Gena with black elongate marking. Clypellus with U-shaped black marking. Lorum and legs yellow without any markings. Pronotum with numerous irregular black spots along middle and near lateral margins. Mesonotum with dull orange basal triangles, median longitudinal marking not extending beyond scutellar suture. Forewings mostly hyaline with dark brown veins, light brown band at middle along costal region.
</paragraph>
<paragraph id="513F1C421944C536F97EFE67FB1FFDF6" blockId="40.[806,1421,444,589]" pageId="40" pageNumber="539">
<emphasis id="63F4C0501944C536F97EFE67FC2FFE69" box="[806,920,445,466]" italics="true" pageId="40" pageNumber="539">Structure:</emphasis>
Clypellus with sides concave, apex wider than base, in profile flat. Rostrum extending beyond mid coxae. Forewing with r-m1 crossvein absent. Metatibial setal rows PD, AD, and AV with 12, ten, and ten macrosetae, respectively.
</paragraph>
<paragraph id="513F1C421944C536F97EFDAFFB56FB2F" blockId="40.[806,1422,629,1172]" pageId="40" pageNumber="539">
<emphasis id="63F4C0501944C536F97EFDAFFC6BFD31" box="[806,988,629,650]" italics="true" pageId="40" pageNumber="539">Male genitalia:</emphasis>
Pygofer broad with short process directed posterad in lateral view, posterolateral margin with short setae. Subgenital plate in lateral view slightly depressed basally, slightly broadened near apex, with short fine setae confined to dorsum of apex; in ventral view lateral margin gradually narrowed beyond middle towards apex, with macrosetae along mesal and lateral margin. Style in lateral view with narrow apodeme; apophysis slender, uniform in width throughout length, curved dorsad near apex, apex hooked. Connective Y-shaped, articulated with preatrium of aedeagus. Aedeagus in lateral view with atrium broad, socle well developed, shaft in lateral view slender, twisted near apex, in caudal view with lateral flaps extending from base to subapex, gradually narrowed towards apex, with single lateral process arising near the bend, pair of short dentate processes near apex. Gonopore apical.
</paragraph>
<paragraph id="513F1C421944C536F97EFB66FCC6FAF6" blockId="40.[806,1422,1212,1357]" pageId="40" pageNumber="539">
<emphasis id="63F4C0501944C536F97EFB66FBB0FB6A" box="[806,1031,1212,1233]" italics="true" pageId="40" pageNumber="539">Material examined:</emphasis>
<materialsCitation id="E1E8161F1944C536FE4BFB66FCC6FAF6" accessionNumber="BLF15448" collectingDate="2006-12-02" collectingDateMax="2006-12-04" collectingDateMin="2006-12-02" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" county="Tolagnaro" elevation="200" latitude="-24.56889" location="Tulear Province" longLatPrecision="21" longitude="47.203888" municipality="Foret Ivohibe" pageId="40" pageNumber="539" specimenCount="1" typeStatus="holotype">
<typeStatus id="8E3BA2E01944C536FE4BFB66FBCEFB69" box="[1043,1145,1212,1234]" pageId="40" pageNumber="539" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="29975CD21944C536FE97FB66FA3CFB6A" box="[1231,1419,1212,1233]" name="Madagascar" pageId="40" pageNumber="539">MADAGASCAR</collectingCountry>
:
<location id="545F4A991944C536F97EFB01FC69FB4A" LSID="urn:lsid:plazi:treatment:D929AD541944C537FAA9FB11FB25FECE:545F4A991944C536F97EFB01FC69FB4A" box="[806,990,1243,1265]" country="Madagascar" county="Tolagnaro" latitude="-24.56889" longLatPrecision="21" longitude="47.203888" municipality="Foret Ivohibe" name="Tulear Province" pageId="40" pageNumber="539">Tulear Province</location>
,
<collectingMunicipality id="B15B86381944C536F9B1FB01FB32FB4A" box="[1001,1157,1243,1265]" pageId="40" pageNumber="539">Forêt Ivohibe</collectingMunicipality>
, 55.0 km north of
<collectingCounty id="B85E64CE1944C536FF0DFB01FCDDFAB4" pageId="40" pageNumber="539">Tolagnaro</collectingCounty>
,
<quantity id="9678B1A71944C536F923FB20FC74FAB4" box="[891,963,1274,1295]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="40" pageNumber="539" unit="m" value="200.0">
<elevation id="DAADFB711944C536F923FB20FC74FAB4" box="[891,963,1274,1295]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="40" pageNumber="539" unit="m" value="200.0">200 m</elevation>
</quantity>
,
<geoCoordinate id="34B47A851944C536F98CFB20FBE5FAB4" box="[980,1106,1272,1295]" degrees="24" direction="south" minutes="34" orientation="latitude" pageId="40" pageNumber="539" precision="15" seconds="08" value="-24.56889">24°34′08″S</geoCoordinate>
,
<geoCoordinate id="34B47A851944C536FE3CFB20FB53FAB4" box="[1124,1252,1272,1295]" degrees="47" direction="east" minutes="12" orientation="longitude" pageId="40" pageNumber="539" precision="15" seconds="14" value="47.203888">47°12′14″E</geoCoordinate>
,
<date id="253E3A821944C536FEADFB23FA3EFAB4" box="[1269,1417,1273,1295]" pageId="40" pageNumber="539" value="2006-12-02" valueMax="2006-12-04" valueMin="2006-12-02">
<collectingDate id="357AC36A1944C536FEADFB23FA3EFAB4" box="[1269,1417,1273,1295]" pageId="40" pageNumber="539" value="2006-12-02" valueMax="2006-12-04" valueMin="2006-12-02">2.–4.xii.2006</collectingDate>
</date>
,
<collectorName id="FC7579941944C536F97EFAC3FC19FA95" box="[806,942,1304,1326]" pageId="40" pageNumber="539">B. L. Fisher</collectorName>
<emphasis id="63F4C0501944C536F9EDFAC3FC52FA96" box="[949,997,1304,1326]" italics="true" pageId="40" pageNumber="539">et al</emphasis>
.,
<collectingMethod id="88C164551944C536F9A2FAC2FB3AFA95" box="[1018,1165,1304,1326]" pageId="40" pageNumber="539">malaise trap</collectingMethod>
, rainforest, BLF15448 (
<collectionCode id="379184871944C536F976FAEDFCD2FAF6" box="[814,869,1335,1357]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="40" pageNumber="539" type="Museum">CAS</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="513F1C421944C536F97EFAACFBD7FA11" blockId="40.[806,1421,1398,1450]" pageId="40" pageNumber="539">
<emphasis id="63F4C0501944C536F97EFAACFC13FA30" box="[806,932,1398,1419]" italics="true" pageId="40" pageNumber="539">Etymology:</emphasis>
The specific epithet refers to the unique twisted shape of aedeagus.
</paragraph>
<paragraph id="513F1C421944C536F919FA3BFAC5FA4E" blockId="40.[833,1394,1504,1558]" box="[833,1394,1504,1526]" pageId="40" pageNumber="539">
<taxonomicName id="968067C11944C536F919FA3BFAC5FA4E" authority="FREYTAG &amp; CWIKLA" authorityName="FREYTAG &amp; CWIKLA" box="[833,1394,1504,1526]" class="Insecta" family="Cicadellidae" genus="Nesocerus" kingdom="Animalia" order="Hemiptera" pageId="40" pageNumber="539" phylum="Arthropoda" rank="species" species="trimaculatus">
<emphasis id="63F4C0501944C536F919FA3BFBC8FA4F" box="[833,1151,1505,1526]" italics="true" pageId="40" pageNumber="539">NESOCERUS TRIMACULATUS</emphasis>
FREYTAG &amp; CWIKLA
</taxonomicName>
</paragraph>
<paragraph id="513F1C421944C536FE56F9DAFB11F9AD" blockId="40.[833,1394,1504,1558]" box="[1038,1190,1536,1558]" pageId="40" pageNumber="539">
(
<figureCitation id="C9BB00C71944C536FE4EF9DAFB29F9AD" box="[1046,1182,1536,1558]" captionStart="Figure 15" captionStartId="21.[164,243,842,861]" captionTargetBox="[164,1445,196,812]" captionTargetId="figure-378@21.[164,1445,196,812]" captionTargetPageId="21" captionText="Figure 15. Nesocerus trimaculatus Freytag &amp; Cwikla. A, dorsal habitus; B, face; C–D, aedeagus in lateral (C) and caudal (D) views." figureDoi="http://doi.org/10.5281/zenodo.5440693" httpUri="https://zenodo.org/record/5440693/files/figure.png" pageId="40" pageNumber="539">FIG. 15A–D</figureCitation>
)
</paragraph>
<paragraph id="513F1C421944C537F97EF9FDFEC0FECE" blockId="40.[806,1422,1575,1903]" lastBlockId="41.[164,779,198,373]" lastPageId="41" lastPageNumber="540" pageId="40" pageNumber="539">
<materialsCitation id="E1E8161F1944C537F97EF9FDFEC4FECE" country="Madagascar" county="We" lastPageId="41" lastPageNumber="540" location="This" municipality="Ankarafantsika forest" pageId="40" pageNumber="539" specimenCount="1" stateProvince="Mahajanga" typeStatus="holotype">
<location id="545F4A991944C536F97EF9FDFCEEF986" LSID="urn:lsid:plazi:treatment:D929AD541944C537FAA9FB11FB25FECE:545F4A991944C536F97EF9FDFCEEF986" box="[806,857,1575,1597]" country="Madagascar" county="We" municipality="Ankarafantsika forest" name="This" pageId="40" pageNumber="539" stateProvince="Mahajanga">This</location>
species was described by
<bibRefCitation id="351161B31944C536FEF6F9FDFCD9F9E7" author="Freytag PH &amp; Cwikla PS" pageId="40" pageNumber="539" pagination="341 - 344" refId="ref27098" refString="Freytag PH, Cwikla PS. 1984. Two new species of idiocerine leafhoppers from Malagasy Republic (Homoptera: Cicadellidae). The Pan-Pacific Entomologist 60: 341 - 344." type="journal article" year="1984">Freytag &amp; Cwikla (1984)</bibRefCitation>
based on a single male specimen from
<collectingMunicipality id="B15B86381944C536F97EF9BEFB95F9C1" box="[806,1058,1636,1658]" pageId="40" pageNumber="539">Ankarafantsika forest</collectingMunicipality>
reserve in the
<collectingRegion id="9344D2A01944C536FE93F9BEFCE3F922" country="Madagascar" name="Mahajanga" pageId="40" pageNumber="539">Mahajanga province</collectingRegion>
of
<collectingCountry id="29975CD21944C536F926F959FBBDF922" box="[894,1034,1667,1689]" name="Madagascar" pageId="40" pageNumber="539">Madagascar</collectingCountry>
.
<collectingCounty id="B85E64CE1944C536FE40F959FB8CF923" box="[1048,1083,1667,1688]" pageId="40" pageNumber="539">We</collectingCounty>
could not examine the
<typeStatus id="8E3BA2E01944C536FF0DF959FCE0F90C" pageId="40" pageNumber="539" type="holotype">holotype</typeStatus>
as it is apparently lost. We identified this species in the present samples based on the similarity of both external morphology and genitalia characters. Pygofer, subgenital plate, style, connective, and shape of aedeagus are the same as in original description. The only difference we noticed is in the number of aedeagal processes. Aedeagus in lateral view has two pairs of lateral apical processes and a single process arising on ventral surface of shaft subapically. In the original description, the presence of this single subapical process is not mentioned or illustrated and we assume that this process might have broken off in the original specimen
</materialsCitation>
.
</paragraph>
<paragraph id="513F1C421945C537FAFCFE44FE02FDF6" blockId="41.[164,779,414,589]" pageId="41" pageNumber="540">
<emphasis id="63F4C0501945C537FAFCFE44FE36FE08" box="[164,385,414,435]" italics="true" pageId="41" pageNumber="540">Material examined:</emphasis>
<materialsCitation id="E1E8161F1945C537FBD1FE44FE06FDF6" collectingDate="2001-11-06" collectingDateMax="2001-11-10" collectingDateMin="2001-11-06" collectionCode="CAS, INHS" country="Madagascar" county="Six" elevation="50" latitude="-19.141945" location="Parc National Tsingy de Bemaraha" longLatPrecision="21" longitude="44.828056" municipality="Province" pageId="41" pageNumber="540" specimenCount="1" stateProvince="Mahajanga">
<collectingCounty id="B85E64CE1945C537FBD1FE44FE1AFE0F" box="[393,429,414,436]" pageId="41" pageNumber="540">Six</collectingCounty>
males,
<collectingCountry id="29975CD21945C537F85FFE44FD74FE08" box="[519,707,414,435]" name="Madagascar" pageId="41" pageNumber="540">MADAGASCAR</collectingCountry>
:
<collectingMunicipality id="B15B86381945C537F896FE45FF65FE68" pageId="41" pageNumber="540">Province</collectingMunicipality>
<collectingRegion id="9344D2A01945C537FA83FE67FEE8FE68" box="[219,351,445,467]" country="Madagascar" name="Mahajanga" pageId="41" pageNumber="540">Mahajanga</collectingRegion>
:
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,
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of
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,
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,
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,
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,
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,
<date id="253E3A821945C537F856FE20FD10FDAB" box="[526,679,506,528]" pageId="41" pageNumber="540" value="2001-11-06" valueMax="2001-11-10" valueMin="2001-11-06">
<collectingDate id="357AC36A1945C537F856FE20FD10FDAB" box="[526,679,506,528]" pageId="41" pageNumber="540" value="2001-11-06" valueMax="2001-11-10" valueMin="2001-11-06">6.–10.xi.2001</collectingDate>
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, Fisher, Griswold
<emphasis id="63F4C0501945C537FB4FFDC3FEF0FD96" box="[279,327,536,558]" italics="true" pageId="41" pageNumber="540">et al</emphasis>
.,
<collectingMethod id="88C164551945C537FB3AFDC2FE4EFD95" box="[354,505,536,558]" pageId="41" pageNumber="540">malaise trap</collectingMethod>
, in tropical dry forest, BLF4233 (
<collectionCode id="379184871945C537FB47FDEDFEE2FDF7" box="[287,341,567,588]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="41" pageNumber="540" type="Museum">CAS</collectionCode>
,
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)
</materialsCitation>
.
</paragraph>
<paragraph id="513F1C421945C537FAADFD5EFD0EFD22" blockId="41.[245,697,644,697]" box="[245,697,644,665]" pageId="41" pageNumber="540">
<taxonomicName id="968067C11945C537FAADFD5EFD0EFD22" authority="(EVANS)" baseAuthorityName="EVANS" box="[245,697,644,665]" class="Insecta" family="Cicadellidae" genus="Nesocerus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="540" phylum="Arthropoda" rank="species" species="clavopunctatus">
<emphasis id="63F4C0501945C537FAADFD5EFDE4FD2C" box="[245,595,644,665]" italics="true" pageId="41" pageNumber="540">NESOCERUS CLAVOPUNCTATUS</emphasis>
(EVANS)
</taxonomicName>
</paragraph>
<paragraph id="513F1C421945C537FBCAFD7EFDABFD02" blockId="41.[245,697,644,697]" box="[402,540,676,697]" pageId="41" pageNumber="540">
(
<figureCitation id="C9BB00C71945C537FBC2FD7EFDA3FD02" box="[410,532,676,697]" captionStart="Figure 1" captionStartId="2.[144,223,1819,1838]" captionTargetBox="[201,1368,194,1786]" captionTargetId="figure-0@2.[200,1368,193,1789]" captionTargetPageId="2" captionText="Figure 1. Nesocerus spp., dorsal habitus &amp; face. Holotypes, A–J; A, B, Nesocerus acuminatus Freytag &amp; Knight.; C, D, Nesocerus chelatus Freytag &amp; Knight; E, F, Nesocerus clavopunctatus (Evans); G, H, Nesocerus madagascariensis (Evans); I, J, Nesocerus spurus Freytag &amp; Knight; K, L, female of N. spurus Freytag &amp; Knight." figureDoi="http://doi.org/10.5281/zenodo.5440665" httpUri="https://zenodo.org/record/5440665/files/figure.png" pageId="41" pageNumber="540">FIG. 1E, F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="199A4FC91945C537FAFCFD10FE30FCA7" pageId="41" pageNumber="540" type="reference_group">
<paragraph id="513F1C421945C537FAFCFD10FD24FD5B" blockId="41.[164,779,714,797]" box="[164,659,714,736]" pageId="41" pageNumber="540">
<treatmentCitationGroup id="71903B6C1945C537FAFCFD10FD24FD5B" box="[164,659,714,736]" pageId="41" pageNumber="540">
<treatmentCitation id="D0213A531945C537FAFCFD10FD24FD5B" author="Evans JW" box="[164,659,714,736]" page="106" pageId="41" pageNumber="540" year="1953">
<taxonomicName id="968067C11945C537FAFCFD10FD24FD5B" ID-CoL="8JTQW" authority="Evans, 1953: 106" authorityName="Evans" authorityPageNumber="106" authorityYear="1953" box="[164,659,714,736]" class="Insecta" family="Cicadellidae" genus="Idiocerus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="540" phylum="Arthropoda" rank="species" species="clavopunctatus">
<emphasis id="63F4C0501945C537FAFCFD10FE76FD64" box="[164,449,714,735]" italics="true" pageId="41" pageNumber="540">Idiocerus clavopunctatus</emphasis>
<bibRefCitation id="351161B31945C537FB93FD11FD24FD5B" author="Evans JW" box="[459,659,714,736]" pageId="41" pageNumber="540" pagination="87 - 137" refId="ref27040" refString="Evans JW. 1953. Les Cicadellidae de Madagascar. Memoires De L' Institut Scientifique Madagascar 4: 87 - 137." type="journal article" year="1953">Evans, 1953: 106</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="513F1C421945C537FAFCFD33FE30FCA7" blockId="41.[164,779,714,797]" pageId="41" pageNumber="540">
<treatmentCitationGroup id="71903B6C1945C537FAFCFD33FE30FCA7" pageId="41" pageNumber="540">
<taxonomicName id="968067C11945C537FAFCFD33FDF0FD44" ID-CoL="477HV" authority="(Evans)" baseAuthorityName="Evans" box="[164,583,745,767]" class="Insecta" family="Cicadellidae" genus="Nesocerus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="540" phylum="Arthropoda" rank="species" species="clavopunctatus">
<emphasis id="63F4C0501945C537FAFCFD33FE6FFD45" box="[164,472,745,766]" italics="true" pageId="41" pageNumber="540">Nesocerus clavopunctatus</emphasis>
(Evans)
</taxonomicName>
–
<treatmentCitation id="D0213A531945C537F8DDFD33FE30FCA7" author="Freytag PH &amp; Knight WJ" page="83" pageId="41" pageNumber="540" year="1966">
<bibRefCitation id="351161B31945C537F8DDFD33FE30FCA7" author="Freytag PH &amp; Knight WJ" pageId="41" pageNumber="540" pagination="75 - 103" refId="ref27130" refString="Freytag PH, Knight WJ. 1966. The Idiocerinae of Madagascar (Homoptera: Cicadellidae). Annales De La Societe Entomologique De France 2: 75 - 103." type="journal article" year="1966">Freytag &amp; Knight, 1966: 83</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="199A4FC91945C537FAFCFC9DFB25FECE" pageId="41" pageNumber="540" type="discussion">
<paragraph id="513F1C421945C537FAFCFC9DFE71F92C" blockId="41.[164,779,838,1687]" pageId="41" pageNumber="540">
<emphasis id="63F4C0501945C537FAFCFC9DFE95FCE7" box="[164,290,839,860]" italics="true" pageId="41" pageNumber="540">Comments:</emphasis>
This species and
<taxonomicName id="968067C11945C537F852FC9DFF43FCC0" authority="Freytag &amp; Knight" authorityName="Freytag &amp; Knight" class="Insecta" family="Cicadellidae" genus="Nesocerus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="540" phylum="Arthropoda" rank="species" species="spurus">
<emphasis id="63F4C0501945C537F852FC9DFD37FCE7" box="[522,640,839,860]" italics="true" pageId="41" pageNumber="540">N. spurus</emphasis>
Freytag &amp; Knight
</taxonomicName>
are very similar to each other, but different from other known
<taxonomicName id="968067C11945C537FBCFFC5EFDDAFC21" box="[407,621,900,922]" class="Insecta" family="Cicadellidae" genus="Nesocerus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="540" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="63F4C0501945C537FBCFFC5EFDBFFC22" box="[407,520,900,921]" italics="true" pageId="41" pageNumber="540">Nesocerus</emphasis>
species
</taxonomicName>
, in external colour pattern (
<figureCitation id="C9BB00C71945C537FB0EFC78FE7DFC03" box="[342,458,930,952]" captionStart="Figure 1" captionStartId="2.[144,223,1819,1838]" captionTargetBox="[201,1368,194,1786]" captionTargetId="figure-0@2.[200,1368,193,1789]" captionTargetPageId="2" captionText="Figure 1. Nesocerus spp., dorsal habitus &amp; face. Holotypes, A–J; A, B, Nesocerus acuminatus Freytag &amp; Knight.; C, D, Nesocerus chelatus Freytag &amp; Knight; E, F, Nesocerus clavopunctatus (Evans); G, H, Nesocerus madagascariensis (Evans); I, J, Nesocerus spurus Freytag &amp; Knight; K, L, female of N. spurus Freytag &amp; Knight." figureDoi="http://doi.org/10.5281/zenodo.5440665" httpUri="https://zenodo.org/record/5440665/files/figure.png" pageId="41" pageNumber="540">Fig. 1E, F</figureCitation>
).
<bibRefCitation id="351161B31945C537FB86FC79FDC2FC03" author="Evans JW" box="[478,629,930,952]" pageId="41" pageNumber="540" pagination="87 - 137" refId="ref27040" refString="Evans JW. 1953. Les Cicadellidae de Madagascar. Memoires De L' Institut Scientifique Madagascar 4: 87 - 137." type="journal article" year="1953">Evans (1953)</bibRefCitation>
described
<taxonomicName id="968067C11945C537F8A8FC79FEF8FC6D" class="Insecta" family="Cicadellidae" genus="Nesocerus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="540" phylum="Arthropoda" rank="species" species="clavopunctatus">
<emphasis id="63F4C0501945C537F8A8FC79FEF8FC6D" italics="true" pageId="41" pageNumber="540">N. clavopunctatus</emphasis>
</taxonomicName>
from a single female specimen based on external coloration and did not describe the genitalia.
<bibRefCitation id="351161B31945C537FAB0FC24FDB0FBAF" author="Freytag PH &amp; Knight WJ" box="[232,519,1022,1044]" pageId="41" pageNumber="540" pagination="75 - 103" refId="ref27130" refString="Freytag PH, Knight WJ. 1966. The Idiocerinae of Madagascar (Homoptera: Cicadellidae). Annales De La Societe Entomologique De France 2: 75 - 103." type="journal article" year="1966">Freytag &amp; Knight (1966)</bibRefCitation>
illustrated the female seventh sternum of
<taxonomicName id="968067C11945C537FBC8FBC7FDD7FB89" box="[400,608,1053,1074]" class="Insecta" family="Cicadellidae" genus="Nesocerus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="540" phylum="Arthropoda" rank="species" species="clavopunctatus">
<emphasis id="63F4C0501945C537FBC8FBC7FDD7FB89" box="[400,608,1053,1074]" italics="true" pageId="41" pageNumber="540">N. clavopunctatus</emphasis>
</taxonomicName>
and described
<taxonomicName id="968067C11945C537FAFCFBE6FEABFBEA" box="[164,284,1084,1105]" class="Insecta" family="Cicadellidae" genus="Nesocerus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="540" phylum="Arthropoda" rank="species" species="spurus">
<emphasis id="63F4C0501945C537FAFCFBE6FEABFBEA" box="[164,284,1084,1105]" italics="true" pageId="41" pageNumber="540">N. spurus</emphasis>
</taxonomicName>
based on a series of male and female specimens, distinguishing the latter species based on its smaller size and more truncate female seventh sternite. Comparison of the
<typeStatus id="8E3BA2E01945C537F858FB4DFD86FB17" box="[512,561,1175,1196]" pageId="41" pageNumber="540">type</typeStatus>
and
<specimenCount id="4786D7CB1945C537F821FB4DFEB9FB77" pageId="41" pageNumber="540" type="generic">one nontype specimen</specimenCount>
of
<taxonomicName id="968067C11945C537FB64FB6CFDB8FB70" box="[316,527,1206,1227]" class="Insecta" family="Cicadellidae" genus="Nesocerus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="540" phylum="Arthropoda" rank="species" species="clavopunctatus">
<emphasis id="63F4C0501945C537FB64FB6CFDB8FB70" box="[316,527,1206,1227]" italics="true" pageId="41" pageNumber="540">N. clavopunctatus</emphasis>
</taxonomicName>
to a series of specimens of
<taxonomicName id="968067C11945C537FB40FB0FFE25FB51" box="[280,402,1237,1258]" class="Insecta" family="Cicadellidae" genus="Nesocerus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="540" phylum="Arthropoda" rank="species" species="spurus">
<emphasis id="63F4C0501945C537FB40FB0FFE25FB51" box="[280,402,1237,1258]" italics="true" pageId="41" pageNumber="540">N. spurus</emphasis>
</taxonomicName>
in the MNHN revealed that
<bibRefCitation id="351161B31945C537FAFCFB29FE6EFAB2" author="Freytag PH &amp; Knight WJ" box="[164,473,1267,1289]" pageId="41" pageNumber="540" pagination="75 - 103" refId="ref27130" refString="Freytag PH, Knight WJ. 1966. The Idiocerinae of Madagascar (Homoptera: Cicadellidae). Annales De La Societe Entomologique De France 2: 75 - 103." type="journal article" year="1966">Freytag &amp; Knight’s (1966)</bibRefCitation>
illustration of the female seventh sternite of
<taxonomicName id="968067C11945C537FBCAFAC8FDD2FA9D" box="[402,613,1297,1319]" class="Insecta" family="Cicadellidae" genus="Nesocerus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="540" phylum="Arthropoda" rank="species" species="clavopunctatus">
<emphasis id="63F4C0501945C537FBCAFAC8FDD2FA9D" box="[402,613,1297,1319]" italics="true" pageId="41" pageNumber="540">N. clavopunctatus</emphasis>
</taxonomicName>
is inaccurate and that its shape is very similar to that of
<taxonomicName id="968067C11945C537F8CDFAEBFCB3FAFD" box="[661,772,1329,1350]" class="Insecta" family="Cicadellidae" genus="Nesocerus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="540" phylum="Arthropoda" rank="species" species="spurus">
<emphasis id="63F4C0501945C537F8CDFAEBFCB3FAFD" box="[661,772,1329,1350]" italics="true" pageId="41" pageNumber="540">N. spurus</emphasis>
</taxonomicName>
. Nevertheless,
<taxonomicName id="968067C11945C537FB0DFA95FD86FADF" box="[341,561,1359,1380]" class="Insecta" family="Cicadellidae" genus="Nesocerus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="540" phylum="Arthropoda" rank="species" species="clavopunctatus">
<emphasis id="63F4C0501945C537FB0DFA95FD86FADF" box="[341,561,1359,1380]" italics="true" pageId="41" pageNumber="540">N. clavopunctatus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="968067C11945C537F8A8FA95FF46FA38" class="Insecta" family="Cicadellidae" genus="Nesocerus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="540" phylum="Arthropoda" rank="species" species="spurus">
<emphasis id="63F4C0501945C537F8A8FA95FF46FA38" italics="true" pageId="41" pageNumber="540">N. spurus</emphasis>
</taxonomicName>
as follows: body size larger, forewing with lateral yellow spot near base of clavus shorter and broader, median yellow spot on costal margin absent, and frontoclypeus with median black stripe. Although discovery of males from the
<typeStatus id="8E3BA2E01945C537F846FA32FDF8FA46" box="[542,591,1512,1533]" pageId="41" pageNumber="540">type</typeStatus>
locality of
<taxonomicName id="968067C11945C537F8A8FA32FEF8F9A0" class="Insecta" family="Cicadellidae" genus="Nesocerus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="540" phylum="Arthropoda" rank="species" species="clavopunctatus">
<emphasis id="63F4C0501945C537F8A8FA32FEF8F9A0" italics="true" pageId="41" pageNumber="540">N. clavopunctatus</emphasis>
</taxonomicName>
– Massif du Tsaratanana – may yet reveal that this taxon and
<taxonomicName id="968067C11945C537FB86F9FFFDF8F981" box="[478,591,1573,1594]" class="Insecta" family="Cicadellidae" genus="Nesocerus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="540" phylum="Arthropoda" rank="species" species="spurus">
<emphasis id="63F4C0501945C537FB86F9FFFDF8F981" box="[478,591,1573,1594]" italics="true" pageId="41" pageNumber="540">N. spurus</emphasis>
</taxonomicName>
are colour variants of the same species, the evidence available at present is sufficient to continue treating these two taxa as separate species.
</paragraph>
<paragraph id="513F1C421945C537FAFCF91AFED4F8D5" blockId="41.[164,779,1728,1903]" pageId="41" pageNumber="540">
<emphasis id="63F4C0501945C537FAFCF91AFD9AF96E" box="[164,557,1728,1749]" italics="true" pageId="41" pageNumber="540">
Specimens of
<taxonomicName id="968067C11945C537FB19F91AFE06F96E" box="[321,433,1728,1749]" class="Insecta" family="Cicadellidae" genus="Nesocerus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="540" phylum="Arthropoda" rank="species" species="spurus">N. spurus</taxonomicName>
examined:
</emphasis>
Six males, MADA- GASCAR: Massif du Marojejy Andasy II,
<quantity id="9678B1A71945C537F8E8F905FCB1F94F" box="[688,774,1759,1780]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="41" pageNumber="540" unit="m" value="1300.0">1300 m</quantity>
,
<date id="253E3A821945C537FAFCF927FF41F8A8" box="[164,246,1789,1811]" pageId="41" pageNumber="540" value="1961-04">iv.1961</date>
, P. Soja (MNHN);
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male,
<materialsCitation id="E1E8161F1945C537F8D8F927FEE8F8D5" collectingDate="1959-11" collectionCode="MADAGAS-" collectorName="Est &amp; de Sambava &amp; Reserve &amp; du Marojejy-Ouest &amp; P. Soga" elevation="1140" pageId="41" pageNumber="540" specimenCount="1">
<collectionCode id="379184871945C537F8D8F927FCBCF8A9" box="[640,779,1789,1810]" pageId="41" pageNumber="540">MADAGAS-</collectionCode>
CAR
<collectorName id="FC7579941945C537FAB2F8C6FEA3F88A" box="[234,276,1820,1841]" pageId="41" pageNumber="540">Est</collectorName>
: district
<collectorName id="FC7579941945C537FBD3F8C6FD95F889" box="[395,546,1820,1842]" pageId="41" pageNumber="540">de Sambava</collectorName>
,
<collectorName id="FC7579941945C537F86FF8C6FD26F889" box="[567,657,1820,1842]" pageId="41" pageNumber="540">Réserve</collectorName>
naturelle intégrale XII, massif
<collectorName id="FC7579941945C537FBE1F8E0FD2CF8EB" box="[441,667,1850,1872]" pageId="41" pageNumber="540">du Marojejy-Ouest</collectorName>
,
<quantity id="9678B1A71945C537F8EAF8E0FCB1F8EB" box="[690,774,1850,1872]" metricMagnitude="3" metricUnit="m" metricValue="1.14" pageId="41" pageNumber="540" unit="m" value="1140.0">
<elevation id="DAADFB711945C537F8EAF8E0FCB1F8EB" box="[690,774,1850,1872]" metricMagnitude="3" metricUnit="m" metricValue="1.14" pageId="41" pageNumber="540" unit="m" value="1140.0">1140 m</elevation>
</quantity>
,
<date id="253E3A821945C537FAFCF883FF4FF8D4" box="[164,248,1881,1903]" pageId="41" pageNumber="540" value="1959-11">
<collectingDate id="357AC36A1945C537FAFCF883FF4FF8D4" box="[164,248,1881,1903]" pageId="41" pageNumber="540" value="1959-11">xi.1959</collectingDate>
</date>
,
<collectorName id="FC7579941945C537FB5FF883FEE8F8D5" box="[263,351,1881,1902]" pageId="41" pageNumber="540">P. Soga</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="513F1C421945C537F962FF1CFB25FECE" blockId="41.[826,1441,198,373]" pageId="41" pageNumber="540">
<emphasis id="63F4C0501945C537F962FF1CFA98FF60" box="[826,1327,198,219]" italics="true" pageId="41" pageNumber="540">
Specimens of
<taxonomicName id="968067C11945C537F986FF1CFB07FF60" box="[990,1200,198,219]" class="Insecta" family="Cicadellidae" genus="Nesocerus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="540" phylum="Arthropoda" rank="species" species="clavopunctatus">N. clavopunctatus</taxonomicName>
examined:
</emphasis>
<materialsCitation id="E1E8161F1945C537FF63FF1CFB25FECE" collectingDate="1949-10" collectingDateMax="1966-12-18" collectingDateMin="1949-10" collectionCode="R, MNHN" collectorName="du Tsaratanana &amp; Andohanambatoafa &amp; P. Soga" country="Madagascar" elevation="1900" location="Sambirano" municipality="Paulian" pageId="41" pageNumber="540" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="8E3BA2E01945C537FF63FF1CFA16FF67" box="[1339,1441,198,220]" pageId="41" pageNumber="540" type="holotype">Holotype</typeStatus>
female,
<collectingCountry id="29975CD21945C537F9CDFF3FFBFAFF41" box="[917,1101,229,250]" name="Madagascar" pageId="41" pageNumber="540">MADAGASCAR</collectingCountry>
<location id="545F4A991945C537FE0DFF3EFB63FF41" LSID="urn:lsid:plazi:treatment:D929AD541944C537FAA9FB11FB25FECE:545F4A991945C537FE0DFF3EFB63FF41" box="[1109,1236,228,250]" country="Madagascar" municipality="Paulian" name="Sambirano" pageId="41" pageNumber="540">Sambirano</location>
, massif
<collectorName id="FC7579941945C537FF6BFF3EFC3AFEA3" pageId="41" pageNumber="540">du Tsaratanana</collectorName>
,
<quantity id="9678B1A71945C537F9FCFED9FC4DFEA3" box="[932,1018,259,280]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="41" pageNumber="540" unit="m" value="1900.0">
<elevation id="DAADFB711945C537F9FCFED9FC4DFEA3" box="[932,1018,259,280]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="41" pageNumber="540" unit="m" value="1900.0">1900 m</elevation>
</quantity>
,
<date id="253E3A821945C537FE49FED9FBE8FEA3" box="[1041,1119,259,280]" pageId="41" pageNumber="540" value="1949-10">
<collectingDate id="357AC36A1945C537FE49FED9FBE8FEA3" box="[1041,1119,259,280]" pageId="41" pageNumber="540" value="1949-10">x.1949</collectingDate>
</date>
,
<collectionCode id="379184871945C537FE2EFED9FB3CFEA3" box="[1142,1163,259,280]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="41" pageNumber="540">R</collectionCode>
.
<collectingMunicipality id="B15B86381945C537FEF9FED9FB4AFEA2" box="[1185,1277,259,281]" pageId="41" pageNumber="540">Paulian</collectingMunicipality>
;
<specimenCount id="4786D7CB1945C537FF4CFED9FA2AFEA2" box="[1300,1437,259,281]" pageId="41" pageNumber="540" type="female">one female</specimenCount>
,
<collectingCountry id="29975CD21945C537F962FEF8FC45FE8C" box="[826,1010,290,311]" name="Madagascar" pageId="41" pageNumber="540">MADAGASCAR</collectingCountry>
nord, massif
<collectorName id="FC7579941945C537FE8CFEF8FA16FE8C" box="[1236,1441,290,312]" pageId="41" pageNumber="540">du Tsaratanana</collectorName>
(versant Sud),
<quantity id="9678B1A71945C537FE5BFE9AFBEEFEED" box="[1027,1113,320,342]" metricMagnitude="3" metricUnit="m" metricValue="2.03" pageId="41" pageNumber="540" unit="m" value="2030.0">
<elevation id="DAADFB711945C537FE5BFE9AFBEEFEED" box="[1027,1113,320,342]" metricMagnitude="3" metricUnit="m" metricValue="2.03" pageId="41" pageNumber="540" unit="m" value="2030.0">2030 m</elevation>
</quantity>
,
<collectorName id="FC7579941945C537FE2BFE9AFAE2FEED" box="[1139,1365,320,342]" pageId="41" pageNumber="540">Andohanambatoafa</collectorName>
,
<date id="253E3A821945C537FF29FE9AFC0EFECE" pageId="41" pageNumber="540" value="1966-12-16" valueMax="1966-12-18" valueMin="1966-12-16">
<collectingDate id="357AC36A1945C537FF29FE9AFC0EFECE" pageId="41" pageNumber="540" value="1966-12-16" valueMax="1966-12-18" valueMin="1966-12-16">16.– 18.xii.1966</collectingDate>
</date>
,
<collectorName id="FC7579941945C537F99FFE85FBABFECF" box="[967,1052,351,372]" pageId="41" pageNumber="540">P. Soga</collectorName>
(
<collectionCode id="379184871945C537FE75FE85FB31FECE" box="[1069,1158,351,373]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="41" pageNumber="540">MNHN</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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