plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/D9/29/87/D92987FD6767FFC9BCC7FD3CFF07F816.xml
maintenance 2337d110b5 added D9
2024-06-21 12:53:51 +02:00

392 lines
58 KiB
XML
Raw Permalink Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="E20CCD4A5A8B9C8D207EBD69AC79B08D" ID-CLB-Dataset="198867" ID-DOI="10.11646/zootaxa.5213.4.3" ID-GBIF-Dataset="1076c5fd-90bf-495b-83d2-579e6ea3b56a" ID-ISSN="1175-5326" ID-Zenodo-Dep="7381394" ID-ZooBank="B5C1E409-DF40-4BDE-9702-DE79C3A252A9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669809047837" checkinUser="plazi" docAuthor="Ballarin, Francesco &amp; Eguchi, Katsuyuki" docDate="2022" docId="D92987FD6767FFC9BCC7FD3CFF07F816" docLanguage="en" docName="zootaxa.5213.4.3.pdf" docOrigin="Zootaxa 5213 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Masirana suzukii Ballarin &amp; Eguchi 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="379" masterDocId="2510FF856761FFC1BC50FFC5FFF6FFD7" masterDocTitle="Taxonomic notes on leptonetid spiders from the Ryukyu Archipelago with the description of two new species and the first record of the genus Longileptoneta from Japan (Araneae: Leptonetidae)" masterLastPageNumber="387" masterPageNumber="371" pageNumber="377" updateTime="1699412544694" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="953E92C68A2705627BDE8F31FFE1E7C3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AB946CDECF5CA4804848DEDC5EA112FF">
<mods:title id="020D82A92DB7391211745FD466FC6C35">Taxonomic notes on leptonetid spiders from the Ryukyu Archipelago with the description of two new species and the first record of the genus Longileptoneta from Japan (Araneae: Leptonetidae)</mods:title>
</mods:titleInfo>
<mods:name id="C4A0291FA059E91265FECA4177CC4DB2" type="personal">
<mods:role id="605B1EAEBCD23077BF3F06DF75D32564">
<mods:roleTerm id="E3B112E86D8587067FD1186F7C25E047">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8C79D08E63E5AB9CFE5AE4A98B418175">Ballarin, Francesco</mods:namePart>
<mods:nameIdentifier id="F16411A5DB26B726E48C8401866BBFBE" type="ORCID">0000-0002-1054-1295</mods:nameIdentifier>
<mods:affiliation id="6AEA9E5D042A74A6846FACA5535553B2">Systematic Zoology Laboratory, Department of Biological Sciences, Tokyo Metropolitan University, 1 - 1 Minami-Osawa, Hachioji-shi, 192 - 0397, Tokyo, Japan. https: // orcid. org / 0000 - 0002 - 1054 - 1295 &amp; Department of Zoology, Museo di Storia Naturale of Verona, Lungadige Porta Vittoria, 9, I- 37129 Verona, Italy.</mods:affiliation>
</mods:name>
<mods:name id="1DEBD8F8FBCAFF01285C2277BC331779" type="personal">
<mods:role id="508E41F67CB65067CC50D2F9C8B93907">
<mods:roleTerm id="8021D131969B42982175696C9EB28666">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="04EB39BB94497A6136F8B4959F1E4B2E">Eguchi, Katsuyuki</mods:namePart>
<mods:nameIdentifier id="EF4F4F98E2F5DBFDA5A82CE1610D4D6E" type="ORCID">0000-0002-1054-1295</mods:nameIdentifier>
<mods:affiliation id="2FA0CFB13CEB7149A5E546EC5FAA378F">Systematic Zoology Laboratory, Department of Biological Sciences, Tokyo Metropolitan University, 1 - 1 Minami-Osawa, Hachioji-shi, 192 - 0397, Tokyo, Japan. https: // orcid. org / 0000 - 0002 - 1054 - 1295</mods:affiliation>
</mods:name>
<mods:typeOfResource id="088ABB94923D1DEC09D20399AD262408">text</mods:typeOfResource>
<mods:relatedItem id="41F653E82A4218EEAFE45C3BC0700ED3" type="host">
<mods:titleInfo id="90BA7D9B40FBE650A434BE8C7FF7A80C">
<mods:title id="2D99073AF3AE76A4D136C963FFE1622C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A444D4FBA84025C3D2AFE06C06689DB5">
<mods:date id="7977735E3B338D14381EFA594D4F8355">2022</mods:date>
<mods:detail id="6558F17A284680FF45E14240A3AC74D4" type="pubDate">
<mods:number id="85BA2D5092191C8D0ED2C22D8765FC3F">2022-11-28</mods:number>
</mods:detail>
<mods:detail id="19E3EDC3855A7143922A52118A25C83D" type="volume">
<mods:number id="0FF43F0814594CC063D3739DD9B09375">5213</mods:number>
</mods:detail>
<mods:detail id="EA2687AEF00988AC479298AC8FC9CC66" type="issue">
<mods:number id="4860B58FF3CD84F7119F8F4702257236">4</mods:number>
</mods:detail>
<mods:extent id="B0EDB356B192BF8411957377270BE238" unit="page">
<mods:start id="0950105C6C20F7B646D6946047AB4143">371</mods:start>
<mods:end id="8E7366091597EF809FD02F5CA548C315">387</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="01C08F0DD885A6DE8FB6C1116AAF5159">journal article</mods:classification>
<mods:identifier id="2EDD36066A17E1F85899F1C63FB8FB11" type="CLB-Dataset">198867</mods:identifier>
<mods:identifier id="E4089A73E435E5368202CF5A18B0246D" type="DOI">10.11646/zootaxa.5213.4.3</mods:identifier>
<mods:identifier id="F565AC122EAC9610166E1CB567E9A664" type="GBIF-Dataset">1076c5fd-90bf-495b-83d2-579e6ea3b56a</mods:identifier>
<mods:identifier id="E9B2C5F00EA79A706C4AFB8EABDFEC8E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="7D2363E3CC0CA3D571581ADF70AFA70B" type="Zenodo-Dep">7381394</mods:identifier>
<mods:identifier id="3D792628FC99CEA4A99AE1633FB5D587" type="ZooBank">B5C1E409-DF40-4BDE-9702-DE79C3A252A9</mods:identifier>
</mods:mods>
<treatment id="D92987FD6767FFC9BCC7FD3CFF07F816" ID-DOI="http://doi.org/10.5281/zenodo.7383527" ID-GBIF-Taxon="205334650" ID-Zenodo-Dep="7383527" LSID="urn:lsid:plazi:treatment:D92987FD6767FFC9BCC7FD3CFF07F816" httpUri="http://treatment.plazi.org/id/D92987FD6767FFC9BCC7FD3CFF07F816" lastPageId="8" lastPageNumber="379" pageId="6" pageNumber="377">
<subSubSection id="199A65606767FFC7BCC7FD3CFC2CFCE0" pageId="6" pageNumber="377" type="nomenclature">
<paragraph id="513F36EB6767FFC7BCC7FD3CFE39FCC4" blockId="6.[151,986,761,860]" box="[151,463,761,788]" pageId="6" pageNumber="377">
<heading id="0A7781876767FFC7BCC7FD3CFE39FCC4" bold="true" box="[151,463,761,788]" fontSize="11" level="1" pageId="6" pageNumber="377" reason="1">
<emphasis id="63F4EAF96767FFC7BCC7FD3CFE39FCC4" bold="true" box="[151,463,761,788]" pageId="6" pageNumber="377">
<taxonomicName id="96804D686767FFC7BCC7FD3CFE9CFCC3" authority="Ballarin &amp; Eguchi, 2022" authorityName="Ballarin &amp; Eguchi" authorityYear="2022" box="[151,362,761,788]" class="Arachnida" family="Leptonetidae" genus="Masirana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="suzukii" status="sp. nov.">
<emphasis id="63F4EAF96767FFC7BCC7FD3CFE9CFCC3" bold="true" box="[151,362,761,788]" italics="true" pageId="6" pageNumber="377">Masirana suzukii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="78C757826767FFC7BD22FD3CFE39FCC4" box="[370,463,761,787]" pageId="6" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="513F36EB6767FFC7BCC7FCD8FC2CFCE0" blockId="6.[151,986,761,860]" box="[151,986,797,824]" pageId="6" pageNumber="377">(Japanese name: tiragamahinamashiragumoティラガマヒナマシラグモ)</paragraph>
</subSubSection>
<subSubSection id="199A65606767FFC7BCC7FC84FE4FFC8C" box="[151,441,833,860]" pageId="6" pageNumber="377" type="description">
<paragraph id="513F36EB6767FFC7BCC7FC84FE4FFC8C" blockId="6.[151,986,761,860]" box="[151,441,833,860]" pageId="6" pageNumber="377">
<figureCitation id="C9BB2A6E6767FFC7BCC7FC84FEE0FC8C" box="[151,278,833,859]" captionStart="FIGURE 4" captionStartId="7.[152,254,1948,1972]" captionTargetBox="[187,1402,561,1925]" captionTargetId="figure-206@7.[185,1402,561,1925]" captionTargetPageId="7" captionText="FIGURE 4. Male palp of Masirana suzukii sp. nov. from Okinawa. A palp, prolateral view B same, retrolateral view; C tip of palp, prolateral view; D same, retrolateral view; E same, dorsal view; F same, ventral view. Scale bars: A, B = 0.2 mm; C–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7381412" httpUri="https://zenodo.org/record/7381412/files/figure.png" pageId="6" pageNumber="377">Figs. 4A–F</figureCitation>
,
<figureCitation id="C9BB2A6E6767FFC7BD71FC84FE93FC8B" box="[289,357,833,860]" captionStart="FIGURE 5" captionStartId="8.[152,255,1312,1336]" captionTargetBox="[151,1436,181,1288]" captionTargetId="figure-25@8.[151,1436,181,1288]" captionTargetPageId="8" captionText="FIGURE 5. Female and male of Masirana suzukii sp. nov. A internal genitalia of female, ventral view; B same, dorsal view; C habitus of male, dorsal view; D habitus of female, dorsal view E same, frontal view. Scale bars: A, B, E = 0.2 mm, C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7381414" httpUri="https://zenodo.org/record/7381414/files/figure.png" pageId="6" pageNumber="377">5A–E</figureCitation>
,
<figureCitation id="C9BB2A6E6767FFC7BD20FC84FE43FC8C" box="[368,437,833,859]" captionStart="FIGURE 6" captionStartId="9.[152,255,1807,1831]" captionTargetBox="[151,1436,181,1771]" captionTargetId="figure-19@9.[151,1436,181,1783]" captionTargetPageId="9" captionText="FIGURE 6. male and female of Masirana suzukii sp. nov. A male palp, prolateral view; B same, retrolateral view C bulb, ventral view; D male chelicera, internal view; E internal genitalia of female. Abbreviations: AT = atrium; E = embolus; MS = median sclerite; PL = prolateral lobe; PS = prolateral sclerite; S = spermatheca; SS = spermatheca stalk; TA = tibial apophyses; TS = tarsal spur. Scale bars: A–D = 0.1 mm, E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7381416" httpUri="https://zenodo.org/record/7381416/files/figure.png" pageId="6" pageNumber="377">6A–E</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="199A65606767FFC7BCC7FC41FB4CFBDD" pageId="6" pageNumber="377" type="materials_examined">
<paragraph id="513F36EB6767FFC7BCC7FC41FC2AFC31" blockId="6.[151,1437,900,2043]" pageId="6" pageNumber="377">
<emphasis id="63F4EAF96767FFC7BCC7FC41FE73FC49" bold="true" box="[151,389,900,926]" pageId="6" pageNumber="377">Material examined.</emphasis>
<materialsCitation id="E1E83CB66767FFC7BDC0FC40FC2AFC31" ID-GBIF-Occurrence="3979374306" collectingDate="2020-11-17" collectorName="Okinawa-honto Is. &amp; F. Ballarin" country="Japan" county="Kunigami-gun" elevation="55" latitude="26.684" location="Tiragama cave" longLatPrecision="74" longitude="128.005" municipality="Nakijin-son" pageId="6" pageNumber="377" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="holotype">
<emphasis id="63F4EAF96767FFC7BDC0FC40FBBEFC49" bold="true" box="[400,1096,900,929]" pageId="6" pageNumber="377">
<specimenCount id="4786FD626767FFC7BDC0FC40FE53FC76" box="[400,421,901,929]" pageId="6" pageNumber="377" type="male">♂</specimenCount>
<typeStatus id="8E3B88496767FFC7BDE1FC41FDD6FC49" box="[433,544,900,926]" pageId="6" pageNumber="377" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="2997767B6767FFC7BE60FC41FD78FC49" box="[560,654,900,926]" name="Japan" pageId="6" pageNumber="377">JAPAN</collectingCountry>
:
<collectingRegion id="9344F8096767FFC7BECFFC41FCA3FC49" box="[671,853,900,926]" country="Japan" name="Okinawa" pageId="6" pageNumber="377">Okinawa Pref.</collectingRegion>
:
<collectorName id="FC75533D6767FFC7BF36FC41FBBEFC49" box="[870,1096,900,926]" pageId="6" pageNumber="377">Okinawa-honto Is.</collectorName>
</emphasis>
,
<collectingCounty id="B85E4E676767FFC7B80BFC41FAF7FC49" box="[1115,1281,900,926]" pageId="6" pageNumber="377">Kunigami-gun</collectingCounty>
,
<collectingMunicipality id="B15BAC916767FFC7B941FC41FA6EFC49" box="[1297,1432,900,926]" pageId="6" pageNumber="377">Nakijin-son</collectingMunicipality>
,
<location id="545F60306767FFC7BCC7FC6DFECBFC14" LSID="urn:lsid:plazi:treatment:D92987FD6767FFC9BCC7FD3CFF07F816:545F60306767FFC7BCC7FC6DFECBFC14" box="[151,317,936,963]" country="Japan" county="Kunigami-gun" latitude="26.684" longLatPrecision="74" longitude="128.005" municipality="Nakijin-son" name="Tiragama cave" pageId="6" pageNumber="377" stateProvince="Okinawa">Tiragama cave</location>
(ティラガマ = Tametomo no horaana, Ẫ朝の洞Ẉ),
<geoCoordinate id="34B4502C6767FFC7BFFBFC6DFBECFC15" box="[939,1050,936,962]" degrees="26.684" direction="north" orientation="latitude" pageId="6" pageNumber="377" precision="55" value="26.684">26.684°N</geoCoordinate>
,
<geoCoordinate id="34B4502C6767FFC7B877FC6DFB57FC14" box="[1063,1185,936,963]" degrees="128.005" direction="east" orientation="longitude" pageId="6" pageNumber="377" precision="55" value="128.005">128.005°E</geoCoordinate>
,
<elevation id="DAADD1D86767FFC7B8FEFC6DFAD0FC15" box="[1198,1318,936,963]" metricMagnitude="1" metricUnit="m" metricValue="5.5" pageId="6" pageNumber="377" unit="m" value="55.0">
<quantity id="96789B0E6767FFC7B8FEFC6DFB1EFC14" box="[1198,1256,936,963]" metricMagnitude="1" metricUnit="m" metricValue="5.5" pageId="6" pageNumber="377" unit="m" value="55.0">55 m</quantity>
a.s.l.
</elevation>
, short and rather humid cave,
<date id="253E102B6767FFC7BD3FFC09FDBDFC31" box="[367,587,972,998]" pageId="6" pageNumber="377" value="2020-11-17">
<collectingDate id="357AE9C36767FFC7BD3FFC09FDBDFC31" box="[367,587,972,998]" pageId="6" pageNumber="377" value="2020-11-17">17 November 2020</collectingDate>
</date>
,
<collectorName id="FC75533D6767FFC7BE06FC08FD3DFC31" box="[598,715,972,999]" pageId="6" pageNumber="377">F. Ballarin</collectorName>
leg. (NSMT-Ar 22246).
</materialsCitation>
</paragraph>
<paragraph id="513F36EB6767FFC7BC97FC35FB4CFBDD" blockId="6.[151,1437,900,2043]" box="[199,1210,1008,1035]" pageId="6" pageNumber="377">
<materialsCitation id="E1E83CB66767FFC7BC97FC35FCB6FBDD" ID-GBIF-Occurrence="3979374311" box="[199,832,1008,1035]" collectingDate="2020-11-17" collectorName="Okinawa-honto Is. &amp; F. Ballarin" country="Japan" county="Kunigami-gun" elevation="55" latitude="26.684" location="Tiragama cave" longLatPrecision="74" longitude="128.005" municipality="Nakijin-son" pageId="6" pageNumber="377" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Okinawa" typeStatus="paratype">
<emphasis id="63F4EAF96767FFC7BC97FC35FE41FBDD" bold="true" box="[199,439,1008,1034]" pageId="6" pageNumber="377">
<typeStatus id="8E3B88496767FFC7BC97FC35FEB7FBDD" box="[199,321,1008,1034]" pageId="6" pageNumber="377" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="2997767B6767FFC7BD1DFC35FE51FBDD" box="[333,423,1008,1034]" name="Japan" pageId="6" pageNumber="377">JAPAN</collectingCountry>
-
</emphasis>
same data as the holotype,
<specimenCount id="4786FD626767FFC7BEBCFC35FCE7FBDD" box="[748,785,1008,1034]" pageId="6" pageNumber="377" type="male">2♂</specimenCount>
,
<specimenCount id="4786FD626767FFC7BF4DFC35FCB6FBDD" box="[797,832,1008,1034]" pageId="6" pageNumber="377" type="female">1♀</specimenCount>
</materialsCitation>
<materialsCitation id="E1E83CB66767FFC7BF17FC35FB40FBDD" ID-GBIF-Occurrence="3979374305" box="[839,1206,1008,1035]" collectionCode="RMUF, FBPC" country="Japan" pageId="6" pageNumber="377" specimenCount="5" specimenCount-female="2" specimenCount-male="3" typeStatus="paratype">
(
<specimenCount id="4786FD626767FFC7BF00FC35FC83FBDD" box="[848,885,1008,1034]" pageId="6" pageNumber="377" type="male">2♂</specimenCount>
,
<specimenCount id="4786FD626767FFC7BFD1FC35FC52FBDD" box="[897,932,1008,1034]" pageId="6" pageNumber="377" type="female">1♀</specimenCount>
<collectionCode id="3791AE2E6767FFC7BFFBFC34FC08FBDC" box="[939,1022,1009,1035]" pageId="6" pageNumber="377">RMUF</collectionCode>
;
<specimenCount id="4786FD626767FFC7B859FC35FBD8FBDD" box="[1033,1070,1008,1034]" pageId="6" pageNumber="377" type="male">1♂</specimenCount>
,
<specimenCount id="4786FD626767FFC7B86AFC35FBABFBDD" box="[1082,1117,1008,1034]" pageId="6" pageNumber="377" type="female">1♀</specimenCount>
<collectionCode id="3791AE2E6767FFC7B834FC35FB58FBDD" box="[1124,1198,1008,1034]" pageId="6" pageNumber="377">FBPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="199A65606767FFC7BC97FBD1FEF5FB4D" pageId="6" pageNumber="377" type="etymology">
<paragraph id="513F36EB6767FFC7BC97FBD1FEF5FB4D" blockId="6.[151,1437,900,2043]" pageId="6" pageNumber="377">
<emphasis id="63F4EAF96767FFC7BC97FBD1FEA6FBF9" bold="true" box="[199,336,1044,1070]" pageId="6" pageNumber="377">Etymology.</emphasis>
The new species is a patronym in honor to our colleague and friend Yuya Suzuki (United Graduate School of Agricultural Sciences,
<collectingRegion id="9344F8096767FFC7BE4BFBFDFD6BFB85" box="[539,669,1080,1106]" country="Japan" name="Kagoshima" pageId="6" pageNumber="377">Kagoshima</collectingRegion>
University,
<collectingCountry id="2997767B6767FFC7BF7FFBFDFC83FB85" box="[815,885,1080,1106]" name="Japan" pageId="6" pageNumber="377">Japan</collectingCountry>
). Yuya is a young and promising arachnologist actively working on ethology, ecology and taxonomy of Japanese spiders, including cave species from the Ryukyus.
</paragraph>
</subSubSection>
<subSubSection id="199A65606767FFC7BC97FB61FDBDF99D" pageId="6" pageNumber="377" type="diagnosis">
<paragraph id="513F36EB6767FFC7BC97FB61FDBDF99D" blockId="6.[151,1437,900,2043]" pageId="6" pageNumber="377">
<emphasis id="63F4EAF96767FFC7BC97FB61FEB4FB69" bold="true" box="[199,322,1188,1214]" pageId="6" pageNumber="377">Diagnosis.</emphasis>
Species closely related to
<taxonomicName id="96804D686767FFC7BE29FB60FBCAFB69" authority="(Zhu &amp; Tso, 2002)" baseAuthorityName="Zhu &amp; Tso" baseAuthorityYear="2002" box="[633,1084,1188,1215]" class="Arachnida" family="Leptonetidae" genus="Masirana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="changlini">
<emphasis id="63F4EAF96767FFC7BE29FB60FCAFFB69" box="[633,857,1188,1214]" italics="true" pageId="6" pageNumber="377">Masirana changlini</emphasis>
(Zhu &amp; Tso, 2002)
</taxonomicName>
from
<collectingCountry id="2997767B6767FFC7B8D5FB61FB2DFB69" box="[1157,1243,1188,1214]" name="Taiwan" pageId="6" pageNumber="377">Taiwan</collectingCountry>
. The male of
<taxonomicName id="96804D686767FFC7B92EFB60FF13FB35" authority="Ballarin &amp; Eguchi, 2022" authorityName="Ballarin &amp; Eguchi" authorityYear="2022" class="Arachnida" family="Leptonetidae" genus="Masirana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="suzukii" status="sp. nov.">
<emphasis id="63F4EAF96767FFC7B92EFB60FF13FB35" italics="true" pageId="6" pageNumber="377">M. suzukii</emphasis>
</taxonomicName>
<emphasis id="63F4EAF96767FFC7BCBEFB0DFEBCFB35" bold="true" box="[238,330,1224,1250]" pageId="6" pageNumber="377">
<taxonomicNameLabel id="78C757826767FFC7BCBEFB0DFEBCFB35" box="[238,330,1224,1250]" pageId="6" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from the male of
<taxonomicName id="96804D686767FFC7BF43FB0CFC52FB35" baseAuthorityName="Zhu &amp; Tso" baseAuthorityYear="2002" box="[787,932,1224,1250]" class="Arachnida" family="Leptonetidae" genus="Masirana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="changlini">
<emphasis id="63F4EAF96767FFC7BF43FB0CFC52FB35" box="[787,932,1224,1250]" italics="true" pageId="6" pageNumber="377">M. changlini</emphasis>
</taxonomicName>
by the different number of denticles on the chelicera (a single row of 9 denticles on the promargin in
<taxonomicName id="96804D686767FFC7BF60FB28FC53FAD1" authority="Ballarin &amp; Eguchi, 2022" authorityName="Ballarin &amp; Eguchi" authorityYear="2022" box="[816,933,1260,1286]" class="Arachnida" family="Leptonetidae" genus="Masirana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="suzukii" status="sp. nov.">
<emphasis id="63F4EAF96767FFC7BF60FB28FC53FAD1" box="[816,933,1260,1286]" italics="true" pageId="6" pageNumber="377">M. suzukii</emphasis>
</taxonomicName>
<emphasis id="63F4EAF96767FFC7BFFEFB29FBFFFAD1" bold="true" box="[942,1033,1260,1286]" pageId="6" pageNumber="377">
<taxonomicNameLabel id="78C757826767FFC7BFFEFB29FBFFFAD1" box="[942,1033,1260,1286]" pageId="6" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
vs. two rows of denticles, 6 on the promargin and 7 on the retromargin, in
<taxonomicName id="96804D686767FFC7BE04FAD4FD15FAFD" baseAuthorityName="Zhu &amp; Tso" baseAuthorityYear="2002" box="[596,739,1296,1322]" class="Arachnida" family="Leptonetidae" genus="Masirana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="changlini">
<emphasis id="63F4EAF96767FFC7BE04FAD4FD15FAFD" box="[596,739,1296,1322]" italics="true" pageId="6" pageNumber="377">M. changlini</emphasis>
</taxonomicName>
; cf.
<figureCitation id="C9BB2A6E6767FFC7BF46FAD5FC9BFAFD" box="[790,877,1296,1322]" captionStart="FIGURE 6" captionStartId="9.[152,255,1807,1831]" captionTargetBox="[151,1436,181,1771]" captionTargetId="figure-19@9.[151,1436,181,1783]" captionTargetPageId="9" captionText="FIGURE 6. male and female of Masirana suzukii sp. nov. A male palp, prolateral view; B same, retrolateral view C bulb, ventral view; D male chelicera, internal view; E internal genitalia of female. Abbreviations: AT = atrium; E = embolus; MS = median sclerite; PL = prolateral lobe; PS = prolateral sclerite; S = spermatheca; SS = spermatheca stalk; TA = tibial apophyses; TS = tarsal spur. Scale bars: A–D = 0.1 mm, E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7381416" httpUri="https://zenodo.org/record/7381416/files/figure.png" pageId="6" pageNumber="377">Fig. 6D</figureCitation>
vs. fig.
<quantity id="96789B0E6767FFC7BF98FAD5FC02FAFD" box="[968,1012,1296,1322]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="6" pageNumber="377" unit="in" value="3.0">3 in</quantity>
Zhu &amp; Tso 2002) and by the slimmer and longer tip of the cymbium bearing a short, stronger spine (vs. a shorter, tougher tip of cymbium lacking any thick apical spine in
<taxonomicName id="96804D686767FFC7BDD9FA9CFDEDFAA5" baseAuthorityName="Zhu &amp; Tso" baseAuthorityYear="2002" box="[393,539,1368,1394]" class="Arachnida" family="Leptonetidae" genus="Masirana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="changlini">
<emphasis id="63F4EAF96767FFC7BDD9FA9CFDEDFAA5" box="[393,539,1368,1394]" italics="true" pageId="6" pageNumber="377">M. changlini</emphasis>
</taxonomicName>
; cf.
<figureCitation id="C9BB2A6E6767FFC7BE03FA9DFD00FAA5" box="[595,758,1368,1394]" captionStart="FIGURE 4" captionStartId="7.[152,254,1948,1972]" captionTargetBox="[187,1402,561,1925]" captionTargetId="figure-206@7.[185,1402,561,1925]" captionTargetPageId="7" captionText="FIGURE 4. Male palp of Masirana suzukii sp. nov. from Okinawa. A palp, prolateral view B same, retrolateral view; C tip of palp, prolateral view; D same, retrolateral view; E same, dorsal view; F same, ventral view. Scale bars: A, B = 0.2 mm; C–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7381412" httpUri="https://zenodo.org/record/7381412/files/figure.png" pageId="6" pageNumber="377">Figs. 4B, C–F</figureCitation>
,
<figureCitation id="C9BB2A6E6767FFC7BF54FA9DFCBFFAA4" box="[772,841,1368,1395]" captionStart="FIGURE 6" captionStartId="9.[152,255,1807,1831]" captionTargetBox="[151,1436,181,1771]" captionTargetId="figure-19@9.[151,1436,181,1783]" captionTargetPageId="9" captionText="FIGURE 6. male and female of Masirana suzukii sp. nov. A male palp, prolateral view; B same, retrolateral view C bulb, ventral view; D male chelicera, internal view; E internal genitalia of female. Abbreviations: AT = atrium; E = embolus; MS = median sclerite; PL = prolateral lobe; PS = prolateral sclerite; S = spermatheca; SS = spermatheca stalk; TA = tibial apophyses; TS = tarsal spur. Scale bars: A–D = 0.1 mm, E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7381416" httpUri="https://zenodo.org/record/7381416/files/figure.png" pageId="6" pageNumber="377">6A, B</figureCitation>
vs. figs. 4,
<quantity id="96789B0E6767FFC7BF85FA9DFBF5FAA5" box="[981,1027,1368,1395]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="6" pageNumber="377" unit="in" value="5.0">5 in</quantity>
Zhu &amp; Tso, 2002). In addition, the embolus of the new species shows a more rounded apex in contrast with a pointy apex in
<taxonomicName id="96804D686767FFC7B8CDFAB8FAD8FA41" baseAuthorityName="Zhu &amp; Tso" baseAuthorityYear="2002" box="[1181,1326,1404,1430]" class="Arachnida" family="Leptonetidae" genus="Masirana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="changlini">
<emphasis id="63F4EAF96767FFC7B8CDFAB8FAD8FA41" box="[1181,1326,1404,1430]" italics="true" pageId="6" pageNumber="377">M. changlini</emphasis>
</taxonomicName>
(cf.
<figureCitation id="C9BB2A6E6767FFC7B935FAB9FF4CFA6D" captionStart="FIGURE 4" captionStartId="7.[152,254,1948,1972]" captionTargetBox="[187,1402,561,1925]" captionTargetId="figure-206@7.[185,1402,561,1925]" captionTargetPageId="7" captionText="FIGURE 4. Male palp of Masirana suzukii sp. nov. from Okinawa. A palp, prolateral view B same, retrolateral view; C tip of palp, prolateral view; D same, retrolateral view; E same, dorsal view; F same, ventral view. Scale bars: A, B = 0.2 mm; C–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7381412" httpUri="https://zenodo.org/record/7381412/files/figure.png" pageId="6" pageNumber="377">Figs. 4B</figureCitation>
,
<figureCitation id="C9BB2A6E6767FFC7BC96FA65FF11FA6D" box="[198,231,1440,1466]" captionStart="FIGURE 6" captionStartId="9.[152,255,1807,1831]" captionTargetBox="[151,1436,181,1771]" captionTargetId="figure-19@9.[151,1436,181,1783]" captionTargetPageId="9" captionText="FIGURE 6. male and female of Masirana suzukii sp. nov. A male palp, prolateral view; B same, retrolateral view C bulb, ventral view; D male chelicera, internal view; E internal genitalia of female. Abbreviations: AT = atrium; E = embolus; MS = median sclerite; PL = prolateral lobe; PS = prolateral sclerite; S = spermatheca; SS = spermatheca stalk; TA = tibial apophyses; TS = tarsal spur. Scale bars: A–D = 0.1 mm, E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7381416" httpUri="https://zenodo.org/record/7381416/files/figure.png" pageId="6" pageNumber="377">6B</figureCitation>
vs. fig.
<quantity id="96789B0E6767FFC7BD13FA65FE98FA6D" box="[323,366,1440,1466]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="6" pageNumber="377" unit="in" value="4.0">4 in</quantity>
Zhu &amp; Tso 2002). The male of
<taxonomicName id="96804D686767FFC7BE84FA64FCBEFA6D" authority="Ballarin &amp; Eguchi, 2022" authorityName="Ballarin &amp; Eguchi" authorityYear="2022" box="[724,840,1440,1466]" class="Arachnida" family="Leptonetidae" genus="Masirana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="suzukii" status="sp. nov.">
<emphasis id="63F4EAF96767FFC7BE84FA64FCBEFA6D" box="[724,840,1440,1466]" italics="true" pageId="6" pageNumber="377">M. suzukii</emphasis>
</taxonomicName>
<emphasis id="63F4EAF96767FFC7BF1FFA65FC5FFA6D" bold="true" box="[847,937,1440,1466]" pageId="6" pageNumber="377">
<taxonomicNameLabel id="78C757826767FFC7BF1FFA65FC5FFA6D" box="[847,937,1440,1466]" pageId="6" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be easily separated from the male of
<taxonomicName id="96804D686767FFC7B92EFA64FEE3FA09" authorityName="Komatsu" authorityYear="1972" class="Arachnida" family="Leptonetidae" genus="Masirana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="longipalpis">
<emphasis id="63F4EAF96767FFC7B92EFA64FEE3FA09" italics="true" pageId="6" pageNumber="377">M. longipalpis</emphasis>
</taxonomicName>
by the different length of the palpal femur and tibia (cf.
<figureCitation id="C9BB2A6E6767FFC7BFD5FA01FBF2FA08" box="[901,1028,1476,1503]" captionStart="FIGURE 4" captionStartId="7.[152,254,1948,1972]" captionTargetBox="[187,1402,561,1925]" captionTargetId="figure-206@7.[185,1402,561,1925]" captionTargetPageId="7" captionText="FIGURE 4. Male palp of Masirana suzukii sp. nov. from Okinawa. A palp, prolateral view B same, retrolateral view; C tip of palp, prolateral view; D same, retrolateral view; E same, dorsal view; F same, ventral view. Scale bars: A, B = 0.2 mm; C–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7381412" httpUri="https://zenodo.org/record/7381412/files/figure.png" pageId="6" pageNumber="377">Figs. 4A, B</figureCitation>
vs. 7A, B). The female of
<taxonomicName id="96804D686767FFC7B97AFA00FA6AFA09" authority="Ballarin &amp; Eguchi, 2022" authorityName="Ballarin &amp; Eguchi" authorityYear="2022" box="[1322,1436,1476,1502]" class="Arachnida" family="Leptonetidae" genus="Masirana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="suzukii" status="sp. nov.">
<emphasis id="63F4EAF96767FFC7B97AFA00FA6AFA09" box="[1322,1436,1476,1502]" italics="true" pageId="6" pageNumber="377">M. suzukii</emphasis>
</taxonomicName>
<emphasis id="63F4EAF96767FFC7BCC7FA2DFF05F9D5" bold="true" box="[151,243,1512,1538]" pageId="6" pageNumber="377">
<taxonomicNameLabel id="78C757826767FFC7BCC7FA2DFF05F9D5" box="[151,243,1512,1538]" pageId="6" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinguished from the female of other congeners, including
<taxonomicName id="96804D686767FFC7BF9DFA2CFB85F9D5" authorityName="Komatsu" authorityYear="1972" box="[973,1139,1512,1538]" class="Arachnida" family="Leptonetidae" genus="Masirana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="longipalpis">
<emphasis id="63F4EAF96767FFC7BF9DFA2CFB85F9D5" box="[973,1139,1512,1538]" italics="true" pageId="6" pageNumber="377">M. longipalpis</emphasis>
</taxonomicName>
, by the different shape of internal genitalia (e.g., smaller and rounded spermathecae (S) in contrast with larger, sac-like S in
<taxonomicName id="96804D686767FFC7B8A1F9C8FA63F9F1" authorityName="Komatsu" authorityYear="1972" box="[1265,1429,1548,1574]" class="Arachnida" family="Leptonetidae" genus="Masirana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="longipalpis">
<emphasis id="63F4EAF96767FFC7B8A1F9C8FA63F9F1" box="[1265,1429,1548,1574]" italics="true" pageId="6" pageNumber="377">M. longipalpis</emphasis>
</taxonomicName>
, cf.
<figureCitation id="C9BB2A6E6767FFC7BCEBF9F5FECAF99C" box="[187,316,1584,1611]" captionStart="FIGURE 5" captionStartId="8.[152,255,1312,1336]" captionTargetBox="[151,1436,181,1288]" captionTargetId="figure-25@8.[151,1436,181,1288]" captionTargetPageId="8" captionText="FIGURE 5. Female and male of Masirana suzukii sp. nov. A internal genitalia of female, ventral view; B same, dorsal view; C habitus of male, dorsal view; D habitus of female, dorsal view E same, frontal view. Scale bars: A, B, E = 0.2 mm, C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7381414" httpUri="https://zenodo.org/record/7381414/files/figure.png" pageId="6" pageNumber="377">Figs. 5A, B</figureCitation>
and
<figureCitation id="C9BB2A6E6767FFC7BD23F9F5FE64F99D" box="[371,402,1584,1610]" captionStart="FIGURE 6" captionStartId="9.[152,255,1807,1831]" captionTargetBox="[151,1436,181,1771]" captionTargetId="figure-19@9.[151,1436,181,1783]" captionTargetPageId="9" captionText="FIGURE 6. male and female of Masirana suzukii sp. nov. A male palp, prolateral view; B same, retrolateral view C bulb, ventral view; D male chelicera, internal view; E internal genitalia of female. Abbreviations: AT = atrium; E = embolus; MS = median sclerite; PL = prolateral lobe; PS = prolateral sclerite; S = spermatheca; SS = spermatheca stalk; TA = tibial apophyses; TS = tarsal spur. Scale bars: A–D = 0.1 mm, E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7381416" httpUri="https://zenodo.org/record/7381416/files/figure.png" pageId="6" pageNumber="377">6E</figureCitation>
vs.
<figureCitation id="C9BB2A6E6767FFC7BD90F9F5FDC8F99D" box="[448,574,1584,1610]" captionStart="FIGURE 8" captionStartId="12.[152,255,1832,1856]" captionTargetBox="[218,1369,181,1808]" captionTargetId="figure-25@12.[218,1369,181,1809]" captionTargetPageId="12" captionText="FIGURE 8. male and female of Masirana longipalpis. A male from Nisshu-do cave, Okinawa-honto Is., tip of palp retrolateral view; B same, ventral view; C female from Nisshu-do cave, frontal view; D habitus of female from Fukubukuîzâ cave, Ishigakijima Is., dorsal view; E internal genitalia of female from Abuntogama cave, ventral view; F same, dorsal view; G habitus of male from Abuntogama cave, dorsal view; H same, habitus of female; I same, frontal view.Abbreviations:AT = atrium; E = embolus; MS = median sclerite; RS = retrolateral sclerite; S = spermathecae; SS = spermathecae stalk; TA = tibial apophyses. Scale bars: A–B = 0.1 mm, C, E, F, I = 0.2; D, G, H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7381424" httpUri="https://zenodo.org/record/7381424/files/figure.png" pageId="6" pageNumber="377">Figs. 8E, F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="199A65606767FFC9BC97F991FAD4F9C7" lastPageId="8" lastPageNumber="379" pageId="6" pageNumber="377" type="description">
<paragraph id="513F36EB6767FFC6BC97F991FD41FF2E" blockId="6.[151,1437,900,2043]" lastBlockId="7.[151,1437,151,537]" lastPageId="7" lastPageNumber="378" pageId="6" pageNumber="377">
<emphasis id="63F4EAF96767FFC7BC97F991FEAFF9B9" bold="true" box="[199,345,1620,1646]" pageId="6" pageNumber="377">Description.</emphasis>
Male (
<typeStatus id="8E3B88496767FFC7BDF5F991FDF8F9B9" box="[421,526,1620,1646]" pageId="6" pageNumber="377" type="holotype">holotype</typeStatus>
). Habitus as in
<figureCitation id="C9BB2A6E6767FFC7BEE2F991FCF1F9B8" box="[690,775,1620,1647]" captionStart="FIGURE 5" captionStartId="8.[152,255,1312,1336]" captionTargetBox="[151,1436,181,1288]" captionTargetId="figure-25@8.[151,1436,181,1288]" captionTargetPageId="8" captionText="FIGURE 5. Female and male of Masirana suzukii sp. nov. A internal genitalia of female, ventral view; B same, dorsal view; C habitus of male, dorsal view; D habitus of female, dorsal view E same, frontal view. Scale bars: A, B, E = 0.2 mm, C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7381414" httpUri="https://zenodo.org/record/7381414/files/figure.png" pageId="6" pageNumber="377">Fig. 5C</figureCitation>
. Total length: 1.66; prosoma 0.72 long, 0.63 wide. Carapace brownish with sightly lighter central area. Median groove, cervical grooves and radial furrows distinct. Cephalic area poorly defined, slightly raised from carapace. Sternum uniformly brownish. Six eyes all well-developed. ALE=0.05, PLE=0.05, PME=0.05, ALE-PLE=0.01, PLE-PME=0.03. Chelicera, labium and maxillae uniformly brownish. Chelicera bearing a single row of 8 denticles on promargin; denticles on retromargin missing (
<figureCitation id="C9BB2A6E6767FFC7B965F921FA66F929" box="[1333,1424,1764,1790]" captionStart="FIGURE 6" captionStartId="9.[152,255,1807,1831]" captionTargetBox="[151,1436,181,1771]" captionTargetId="figure-19@9.[151,1436,181,1783]" captionTargetPageId="9" captionText="FIGURE 6. male and female of Masirana suzukii sp. nov. A male palp, prolateral view; B same, retrolateral view C bulb, ventral view; D male chelicera, internal view; E internal genitalia of female. Abbreviations: AT = atrium; E = embolus; MS = median sclerite; PL = prolateral lobe; PS = prolateral sclerite; S = spermatheca; SS = spermatheca stalk; TA = tibial apophyses; TS = tarsal spur. Scale bars: A–D = 0.1 mm, E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7381416" httpUri="https://zenodo.org/record/7381416/files/figure.png" pageId="6" pageNumber="377">Fig. 6D</figureCitation>
). Legs uniformly light brown. Leg formula: I, IV, II, III. Leg measurements: I = 4.67 (1.42, 0.22, 1.74, 1.38, 0.91), II = 3.75 (1.06, 0.22, 1.07, 0.81, 0.59), III = 3.09 (0.87, 0.21, 0.80, 0.73, 0.48), IV = 4.03 (1.19, 0.23, 1.26, 1.03, 0.59). Opisthosoma yellowish with faint dorsal marks. Male palp as in
<figureCitation id="C9BB2A6E6767FFC7BF27F895FC0EF8BD" box="[887,1016,1872,1898]" captionStart="FIGURE 4" captionStartId="7.[152,254,1948,1972]" captionTargetBox="[187,1402,561,1925]" captionTargetId="figure-206@7.[185,1402,561,1925]" captionTargetPageId="7" captionText="FIGURE 4. Male palp of Masirana suzukii sp. nov. from Okinawa. A palp, prolateral view B same, retrolateral view; C tip of palp, prolateral view; D same, retrolateral view; E same, dorsal view; F same, ventral view. Scale bars: A, B = 0.2 mm; C–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7381412" httpUri="https://zenodo.org/record/7381412/files/figure.png" pageId="6" pageNumber="377">Figs. 4A–F</figureCitation>
,
<figureCitation id="C9BB2A6E6767FFC7B854F895FBBCF8BD" box="[1028,1098,1872,1898]" captionStart="FIGURE 6" captionStartId="9.[152,255,1807,1831]" captionTargetBox="[151,1436,181,1771]" captionTargetId="figure-19@9.[151,1436,181,1783]" captionTargetPageId="9" captionText="FIGURE 6. male and female of Masirana suzukii sp. nov. A male palp, prolateral view; B same, retrolateral view C bulb, ventral view; D male chelicera, internal view; E internal genitalia of female. Abbreviations: AT = atrium; E = embolus; MS = median sclerite; PL = prolateral lobe; PS = prolateral sclerite; S = spermatheca; SS = spermatheca stalk; TA = tibial apophyses; TS = tarsal spur. Scale bars: A–D = 0.1 mm, E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7381416" httpUri="https://zenodo.org/record/7381416/files/figure.png" pageId="6" pageNumber="377">6A–C</figureCitation>
. Femur with several long and robust spines on its ventral and dorsal margins. Patella and tibia both elongated, approximately of the same length. Two apophyses (TA) on retrodistal margin of tibia close to each other; ventral apophysis large and lanceolate; dorsal apophysis spine-like, larger at its base and ending with a sharp and long tip (
<figureCitation id="C9BB2A6E6767FFC7B86DF879FB4BF800" box="[1085,1213,1980,2007]" captionStart="FIGURE 4" captionStartId="7.[152,254,1948,1972]" captionTargetBox="[187,1402,561,1925]" captionTargetId="figure-206@7.[185,1402,561,1925]" captionTargetPageId="7" captionText="FIGURE 4. Male palp of Masirana suzukii sp. nov. from Okinawa. A palp, prolateral view B same, retrolateral view; C tip of palp, prolateral view; D same, retrolateral view; E same, dorsal view; F same, ventral view. Scale bars: A, B = 0.2 mm; C–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7381412" httpUri="https://zenodo.org/record/7381412/files/figure.png" pageId="6" pageNumber="377">Figs. 4D, E</figureCitation>
and
<figureCitation id="C9BB2A6E6767FFC7B8A4F879FAEEF801" box="[1268,1304,1980,2006]" captionStart="FIGURE 6" captionStartId="9.[152,255,1807,1831]" captionTargetBox="[151,1436,181,1771]" captionTargetId="figure-19@9.[151,1436,181,1783]" captionTargetPageId="9" captionText="FIGURE 6. male and female of Masirana suzukii sp. nov. A male palp, prolateral view; B same, retrolateral view C bulb, ventral view; D male chelicera, internal view; E internal genitalia of female. Abbreviations: AT = atrium; E = embolus; MS = median sclerite; PL = prolateral lobe; PS = prolateral sclerite; S = spermatheca; SS = spermatheca stalk; TA = tibial apophyses; TS = tarsal spur. Scale bars: A–D = 0.1 mm, E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7381416" httpUri="https://zenodo.org/record/7381416/files/figure.png" pageId="6" pageNumber="377">6B</figureCitation>
). Cymbium bearing several long and thin spines, ending pointy and with a robust spine at its apex (= tarsal spur). Bulb with two sclerite: median sclerite (MS) spine-like, long and thin; prolateral sclerite (PS) wide and transparent, ribbon-like and wrapped around MS. Embolus (E) long and robust, thread-like and laterally flattened, ending with a rounded tip slightly curved dorsally (
<figureCitation id="C9BB2A6E6766FFC6BDFFFF1AFDDBFF2E" box="[431,557,223,249]" captionStart="FIGURE 4" captionStartId="7.[152,254,1948,1972]" captionTargetBox="[187,1402,561,1925]" captionTargetId="figure-206@7.[185,1402,561,1925]" captionTargetPageId="7" captionText="FIGURE 4. Male palp of Masirana suzukii sp. nov. from Okinawa. A palp, prolateral view B same, retrolateral view; C tip of palp, prolateral view; D same, retrolateral view; E same, dorsal view; F same, ventral view. Scale bars: A, B = 0.2 mm; C–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7381412" httpUri="https://zenodo.org/record/7381412/files/figure.png" pageId="7" pageNumber="378">Figs. 4C–F</figureCitation>
and
<figureCitation id="C9BB2A6E6766FFC6BE33FF1AFD5CFF2E" box="[611,682,223,249]" captionStart="FIGURE 6" captionStartId="9.[152,255,1807,1831]" captionTargetBox="[151,1436,181,1771]" captionTargetId="figure-19@9.[151,1436,181,1783]" captionTargetPageId="9" captionText="FIGURE 6. male and female of Masirana suzukii sp. nov. A male palp, prolateral view; B same, retrolateral view C bulb, ventral view; D male chelicera, internal view; E internal genitalia of female. Abbreviations: AT = atrium; E = embolus; MS = median sclerite; PL = prolateral lobe; PS = prolateral sclerite; S = spermatheca; SS = spermatheca stalk; TA = tibial apophyses; TS = tarsal spur. Scale bars: A–D = 0.1 mm, E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7381416" httpUri="https://zenodo.org/record/7381416/files/figure.png" pageId="7" pageNumber="378">6A–C</figureCitation>
).
</paragraph>
<paragraph id="513F36EB6766FFC6BC97FEC6FAB3FDCE" blockId="7.[151,1437,151,537]" pageId="7" pageNumber="378">
Female (one of the
<typeStatus id="8E3B88496766FFC6BDFCFEC6FDEBFECA" box="[428,541,259,285]" pageId="7" pageNumber="378" type="paratype">paratypes</typeStatus>
). Habitus as in
<figureCitation id="C9BB2A6E6766FFC6BE84FEC6FCA7FECA" box="[724,849,258,285]" captionStart="FIGURE 5" captionStartId="8.[152,255,1312,1336]" captionTargetBox="[151,1436,181,1288]" captionTargetId="figure-25@8.[151,1436,181,1288]" captionTargetPageId="8" captionText="FIGURE 5. Female and male of Masirana suzukii sp. nov. A internal genitalia of female, ventral view; B same, dorsal view; C habitus of male, dorsal view; D habitus of female, dorsal view E same, frontal view. Scale bars: A, B, E = 0.2 mm, C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7381414" httpUri="https://zenodo.org/record/7381414/files/figure.png" pageId="7" pageNumber="378">Fig. 5D, E</figureCitation>
. Total length: 1.59; prosoma 0.69 long, 0.63 wide. General coloration and pattern as in male. Frontal view of cephalic area as in
<figureCitation id="C9BB2A6E6766FFC6B863FEE2FB7AFE96" box="[1075,1164,294,321]" captionStart="FIGURE 5" captionStartId="8.[152,255,1312,1336]" captionTargetBox="[151,1436,181,1288]" captionTargetId="figure-25@8.[151,1436,181,1288]" captionTargetPageId="8" captionText="FIGURE 5. Female and male of Masirana suzukii sp. nov. A internal genitalia of female, ventral view; B same, dorsal view; C habitus of male, dorsal view; D habitus of female, dorsal view E same, frontal view. Scale bars: A, B, E = 0.2 mm, C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7381414" httpUri="https://zenodo.org/record/7381414/files/figure.png" pageId="7" pageNumber="378">Fig. 5E</figureCitation>
. ALE=0.05, PLE=0.05, PME=0.05, ALE-PLE=0.01, PLE-PME=0.03. Leg formula: I, IV, II, III. Leg measurements: I = 4.69 (1.28, 0.23, 1.42, 1.09, 0.67), II = 3.51 (1.01, 0.21, 0.95, 0.77, 0.57), III = 3.03 (0.85, 0.23, 0.76, 0.69, 0.50), IV = 4.09 (1.12, 0.22, 1.19, 0.95, 0.61). Opisthosoma brownish or yellowish, strongly wrinkled in the frontal part (
<figureCitation id="C9BB2A6E6766FFC6B8BFFE56FAB1FE7A" box="[1263,1351,402,429]" captionStart="FIGURE 5" captionStartId="8.[152,255,1312,1336]" captionTargetBox="[151,1436,181,1288]" captionTargetId="figure-25@8.[151,1436,181,1288]" captionTargetPageId="8" captionText="FIGURE 5. Female and male of Masirana suzukii sp. nov. A internal genitalia of female, ventral view; B same, dorsal view; C habitus of male, dorsal view; D habitus of female, dorsal view E same, frontal view. Scale bars: A, B, E = 0.2 mm, C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7381414" httpUri="https://zenodo.org/record/7381414/files/figure.png" pageId="7" pageNumber="378">Fig. 5E</figureCitation>
). Other characters as in male. Internal genitalia as in
<figureCitation id="C9BB2A6E6766FFC6BECEFE72FCECFE06" box="[670,794,438,465]" captionStart="FIGURE 5" captionStartId="8.[152,255,1312,1336]" captionTargetBox="[151,1436,181,1288]" captionTargetId="figure-25@8.[151,1436,181,1288]" captionTargetPageId="8" captionText="FIGURE 5. Female and male of Masirana suzukii sp. nov. A internal genitalia of female, ventral view; B same, dorsal view; C habitus of male, dorsal view; D habitus of female, dorsal view E same, frontal view. Scale bars: A, B, E = 0.2 mm, C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7381414" httpUri="https://zenodo.org/record/7381414/files/figure.png" pageId="7" pageNumber="378">Fig. 5A, B</figureCitation>
,
<figureCitation id="C9BB2A6E6766FFC6BF77FE72FCBCFE06" box="[807,842,439,465]" captionStart="FIGURE 6" captionStartId="9.[152,255,1807,1831]" captionTargetBox="[151,1436,181,1771]" captionTargetId="figure-19@9.[151,1436,181,1783]" captionTargetPageId="9" captionText="FIGURE 6. male and female of Masirana suzukii sp. nov. A male palp, prolateral view; B same, retrolateral view C bulb, ventral view; D male chelicera, internal view; E internal genitalia of female. Abbreviations: AT = atrium; E = embolus; MS = median sclerite; PL = prolateral lobe; PS = prolateral sclerite; S = spermatheca; SS = spermatheca stalk; TA = tibial apophyses; TS = tarsal spur. Scale bars: A–D = 0.1 mm, E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7381416" httpUri="https://zenodo.org/record/7381416/files/figure.png" pageId="7" pageNumber="378">6E</figureCitation>
. Atrium (AT) wide, cup-shaped. Spermathecae stalk (SS) starting at sides of atrium, first bending inward with a comma course, then reaching spermathecae (S) after one convolution. Spermathecae small and round, separated from each other by two and a half of their diameter.
</paragraph>
<caption id="05FF66636766FFC6BCC8F859FA6DF80F" ID-DOI="http://doi.org/10.5281/zenodo.7381412" ID-Zenodo-Dep="7381412" httpUri="https://zenodo.org/record/7381412/files/figure.png" pageId="7" pageNumber="378" startId="7.[152,254,1948,1972]" targetBox="[187,1402,561,1925]" targetPageId="7">
<paragraph id="513F36EB6766FFC6BCC8F859FA6DF80F" blockId="7.[151,1436,1948,2008]" pageId="7" pageNumber="378">
<emphasis id="63F4EAF96766FFC6BCC8F859FEE3F863" bold="true" box="[152,277,1948,1972]" pageId="7" pageNumber="378">FIGURE 4.</emphasis>
Male palp of
<taxonomicName id="96804D686766FFC6BDCBF858FDB3F863" authority="Ballarin &amp; Eguchi, 2022" authorityName="Ballarin &amp; Eguchi" authorityYear="2022" box="[411,581,1948,1972]" class="Arachnida" family="Leptonetidae" genus="Masirana" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="378" phylum="Arthropoda" rank="species" species="suzukii" status="sp. nov.">
<emphasis id="63F4EAF96766FFC6BDCBF858FDB3F863" box="[411,581,1948,1972]" italics="true" pageId="7" pageNumber="378">Masirana suzukii</emphasis>
</taxonomicName>
<emphasis id="63F4EAF96766FFC6BE1AF859FD61F863" bold="true" box="[586,663,1948,1972]" pageId="7" pageNumber="378">
<taxonomicNameLabel id="78C757826766FFC6BE1AF859FD61F863" box="[586,663,1948,1972]" pageId="7" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from Okinawa.
<emphasis id="63F4EAF96766FFC6BF66F859FCBEF863" bold="true" box="[822,840,1948,1972]" pageId="7" pageNumber="378">A</emphasis>
palp, prolateral view
<emphasis id="63F4EAF96766FFC6B84BF859FBDAF863" bold="true" box="[1051,1068,1948,1972]" pageId="7" pageNumber="378">B</emphasis>
same, retrolateral view;
<emphasis id="63F4EAF96766FFC6B94CF859FAD8F863" bold="true" box="[1308,1326,1948,1972]" pageId="7" pageNumber="378">C</emphasis>
tip of palp, prolateral view;
<emphasis id="63F4EAF96766FFC6BD69F805FEBDF80F" bold="true" box="[313,331,1984,2008]" pageId="7" pageNumber="378">D</emphasis>
same, retrolateral view;
<emphasis id="63F4EAF96766FFC6BE6FF805FDA6F80F" bold="true" box="[575,592,1984,2008]" pageId="7" pageNumber="378">E</emphasis>
same, dorsal view;
<emphasis id="63F4EAF96766FFC6BF45F805FCD2F80F" bold="true" box="[789,804,1984,2008]" pageId="7" pageNumber="378">F</emphasis>
same, ventral view. Scale bars: A, B = 0.2 mm; C–F = 0.1 mm.
</paragraph>
</caption>
<caption id="05FF66636769FFC9BCC8FAE5FF30FA57" ID-DOI="http://doi.org/10.5281/zenodo.7381414" ID-Zenodo-Dep="7381414" httpUri="https://zenodo.org/record/7381414/files/figure.png" pageId="8" pageNumber="379" startId="8.[152,255,1312,1336]" targetBox="[151,1436,181,1288]" targetPageId="8">
<paragraph id="513F36EB6769FFC9BCC8FAE5FF30FA57" blockId="8.[151,1436,1312,1408]" pageId="8" pageNumber="379">
<emphasis id="63F4EAF96769FFC9BCC8FAE5FEEEFAEF" bold="true" box="[152,280,1312,1336]" pageId="8" pageNumber="379">FIGURE 5.</emphasis>
Female and male of
<taxonomicName id="96804D686769FFC9BDBAFAE4FD6EFAEF" authority="Ballarin &amp; Eguchi, 2022" authorityName="Ballarin &amp; Eguchi" authorityYear="2022" box="[490,664,1312,1336]" class="Arachnida" family="Leptonetidae" genus="Masirana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="suzukii" status="sp. nov.">
<emphasis id="63F4EAF96769FFC9BDBAFAE4FD6EFAEF" box="[490,664,1312,1336]" italics="true" pageId="8" pageNumber="379">Masirana suzukii</emphasis>
</taxonomicName>
<emphasis id="63F4EAF96769FFC9BECEFAE5FCF0FAEF" bold="true" box="[670,774,1312,1336]" pageId="8" pageNumber="379">
<taxonomicNameLabel id="78C757826769FFC9BECEFAE5FD18FAEF" box="[670,750,1312,1336]" pageId="8" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
A
</emphasis>
internal genitalia of female, ventral view;
<emphasis id="63F4EAF96769FFC9B8E1FAE5FB34FAEF" bold="true" box="[1201,1218,1312,1336]" pageId="8" pageNumber="379">B</emphasis>
same, dorsal view;
<emphasis id="63F4EAF96769FFC9B9DAFAE5FA6AFAEF" bold="true" box="[1418,1436,1312,1336]" pageId="8" pageNumber="379">C</emphasis>
habitus of male, dorsal view;
<emphasis id="63F4EAF96769FFC9BD95FA81FE21FA8B" bold="true" box="[453,471,1348,1372]" pageId="8" pageNumber="379">D</emphasis>
habitus of female, dorsal view
<emphasis id="63F4EAF96769FFC9BF49FA81FCDCFA8B" bold="true" box="[793,810,1348,1372]" pageId="8" pageNumber="379">E</emphasis>
same, frontal view. Scale bars: A, B, E = 0.2 mm, C, D = 0.5 mm.
</paragraph>
</caption>
<paragraph id="513F36EB6769FFC9BC97FA17FAD4F9C7" blockId="8.[151,1436,1490,1985]" pageId="8" pageNumber="379">
<emphasis id="63F4EAF96769FFC9BC97FA17FE8EFA3B" bold="true" box="[199,376,1490,1516]" pageId="8" pageNumber="379">Size variation:</emphasis>
Male (based on
<specimenCount id="4786FD626769FFC9BE12FA16FD23FA3B" box="[578,725,1490,1517]" pageId="8" pageNumber="379" type="generic">4 specimens</specimenCount>
): total length: 1.55–1.66, Prosoma 0.65–0.72 long, 0.63–0.66 wide. Female (based on
<specimenCount id="4786FD626769FFC9BDF8FA32FDCEF9C7" box="[424,568,1526,1553]" pageId="8" pageNumber="379" type="generic">2 specimens</specimenCount>
): total length: 1.59–1.70, prosoma 0.69–0.72 long, 0.63–0.66wide.
</paragraph>
</subSubSection>
<subSubSection id="199A65606769FFC9BC97F9DFFDADF9AA" pageId="8" pageNumber="379" type="distribution">
<paragraph id="513F36EB6769FFC9BC97F9DFFCAAF9E2" blockId="8.[151,1436,1490,1985]" box="[199,860,1562,1589]" pageId="8" pageNumber="379">
<emphasis id="63F4EAF96769FFC9BC97F9DFFE97F9E3" bold="true" box="[199,353,1562,1588]" pageId="8" pageNumber="379">Distribution.</emphasis>
Known only from the
<typeStatus id="8E3B88496769FFC9BE30F9DEFD66F9E2" box="[608,656,1563,1589]" pageId="8" pageNumber="379">type</typeStatus>
locality (
<figureCitation id="C9BB2A6E6769FFC9BEABF9DFFCB8F9E2" box="[763,846,1562,1589]" captionStart="FIGURE 10" captionStartId="14.[152,255,1636,1660]" captionTargetBox="[154,1433,453,1612]" captionTargetId="figure-143@14.[154,1433,453,1612]" captionTargetPageId="14" captionText="FIGURE 10. Known records of Leptonetidae in the Ryukyu Archipelago and location of the related caves. Legend: pink hexagon = Longileptoneta yamasakii sp. nov.; blue dot = Masirana. suzukii sp. nov.; yellow dots = M. longipalpis; red squares = Falcileptoneta okinawaensis. Type localities of the species are highlighted with an internal black dot." figureDoi="http://doi.org/10.5281/zenodo.7381428" httpUri="https://zenodo.org/record/7381428/files/figure.png" pageId="8" pageNumber="379">Fig. 10</figureCitation>
).
</paragraph>
<paragraph id="513F36EB6769FFC9BC97F9FBFDADF9AA" blockId="8.[151,1436,1490,1985]" pageId="8" pageNumber="379">
<emphasis id="63F4EAF96769FFC9BC97F9FBFEDCF98F" bold="true" box="[199,298,1598,1624]" pageId="8" pageNumber="379">Habitat.</emphasis>
Caves. The new species was found in empty spaces under stones and in recesses of the floor in the twilight zone of a short and humid cave.
</paragraph>
</subSubSection>
<subSubSection id="199A65606769FFC9BC97F942FF07F816" pageId="8" pageNumber="379" type="discussion">
<paragraph id="513F36EB6769FFC9BC97F942FF07F816" blockId="8.[151,1436,1490,1985]" pageId="8" pageNumber="379">
<emphasis id="63F4EAF96769FFC9BC97F942FECDF976" bold="true" box="[199,315,1671,1697]" pageId="8" pageNumber="379">Remarks.</emphasis>
<taxonomicName id="96804D686769FFC9BD13F942FE40F977" authority="Ballarin &amp; Eguchi, 2022" authorityName="Ballarin &amp; Eguchi" authorityYear="2022" box="[323,438,1670,1696]" class="Arachnida" family="Leptonetidae" genus="Masirana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="suzukii" status="sp. nov.">
<emphasis id="63F4EAF96769FFC9BD13F942FE40F977" box="[323,438,1670,1696]" italics="true" pageId="8" pageNumber="379">M. suzukii</emphasis>
</taxonomicName>
<emphasis id="63F4EAF96769FFC9BDEEF942FDEEF976" bold="true" box="[446,536,1671,1697]" pageId="8" pageNumber="379">
<taxonomicNameLabel id="78C757826769FFC9BDEEF942FDEEF976" box="[446,536,1671,1697]" pageId="8" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
shows reduced pigmentation and faint dorsal color pattern. Despite lacking any real troglobitic adaptation, the collecting environment suggests troglophilic habits. However, we do not exclude that this species might also inhabit screes and other shallow subterranean habitats or even external environments (e.g., forest litter). According to
<bibRefCitation id="35114B1A6769FFC9BE5BF937FD26F8DA" author="Shimojana, M." box="[523,720,1778,1805]" pageId="8" pageNumber="379" pagination="337 - 365" refId="ref8780" refString="Shimojana, M. (1977) Preliminary report on the cave spider fauna of the Ryukyu Archipelago. Acta Arachnologica, 27 (Spec. No.), 337 - 365. https: // doi. org / 10.2476 / asjaa. 27. Specialnumber _ 337" type="journal article" year="1977">Shimojana (1977</bibRefCitation>
, pg. 347)
<taxonomicName id="96804D686769FFC9BF15F937FB7EF8DB" authorityName="Komatsu" authorityYear="1972" box="[837,1160,1778,1804]" class="Arachnida" family="Leptonetidae" genus="Falcileptoneta" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="okinawaensis">
<emphasis id="63F4EAF96769FFC9BF15F937FB7EF8DB" box="[837,1160,1778,1804]" italics="true" pageId="8" pageNumber="379">Falcileptoneta okinawaensis</emphasis>
</taxonomicName>
was also recorded from the same cave where
<taxonomicName id="96804D686769FFC9BDD7F8D2FE0DF8E7" authority="Ballarin &amp; Eguchi, 2022" authorityName="Ballarin &amp; Eguchi" authorityYear="2022" box="[391,507,1814,1840]" class="Arachnida" family="Leptonetidae" genus="Masirana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="suzukii" status="sp. nov.">
<emphasis id="63F4EAF96769FFC9BDD7F8D2FE0DF8E7" box="[391,507,1814,1840]" italics="true" pageId="8" pageNumber="379">M. suzukii</emphasis>
</taxonomicName>
<emphasis id="63F4EAF96769FFC9BE52F8D2FDADF8E6" bold="true" box="[514,603,1815,1841]" pageId="8" pageNumber="379">
<taxonomicNameLabel id="78C757826769FFC9BE52F8D2FDADF8E6" box="[514,603,1815,1841]" pageId="8" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was collected. Despite extensive collections inside the cave, we could not find any other leptonetid species. In addition, the records of
<taxonomicName id="96804D686769FFC9BF6FF8FEFC04F883" authorityName="Komatsu" authorityYear="1972" box="[831,1010,1850,1876]" class="Arachnida" family="Leptonetidae" genus="Falcileptoneta" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="okinawaensis">
<emphasis id="63F4EAF96769FFC9BF6FF8FEFC04F883" box="[831,1010,1850,1876]" italics="true" pageId="8" pageNumber="379">F. okinawaensis</emphasis>
</taxonomicName>
by Shimojana in Tiragama cave were based on females only. Due to the external similarities of females in
<taxonomicName id="96804D686769FFC9BFC0F89BFBD4F8AF" authorityName="Simon" authorityYear="1890" box="[912,1058,1886,1912]" class="Arachnida" family="Leptonetidae" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="379" phylum="Arthropoda" rank="family">Leptonetidae</taxonomicName>
and the lack of information about the internal genitalia of these species, it is possible that such records refer to misidentified samples of
<taxonomicName id="96804D686769FFC9B978F846FA6AF84B" authority="Ballarin &amp; Eguchi, 2022" authorityName="Ballarin &amp; Eguchi" authorityYear="2022" box="[1320,1436,1922,1948]" class="Arachnida" family="Leptonetidae" genus="Masirana" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="suzukii" status="sp. nov.">
<emphasis id="63F4EAF96769FFC9B978F846FA6AF84B" box="[1320,1436,1922,1948]" italics="true" pageId="8" pageNumber="379">M. suzukii</emphasis>
</taxonomicName>
<emphasis id="63F4EAF96769FFC9BCC7F862FF07F816" bold="true" box="[151,241,1959,1985]" pageId="8" pageNumber="379">
<taxonomicNameLabel id="78C757826769FFC9BCC7F862FF07F816" box="[151,241,1959,1985]" pageId="8" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink