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<document ID-CLB-Dataset="229646" ID-DOI="http://dx.doi.org/10.3897/zse.99.95923" ID-GBIF-Dataset="b0059ad9-fa19-4539-821a-0bec19b167c9" ID-Pensoft-Pub="1860-0743-1-77" ID-Pensoft-UUID="F3CD023B35EE51769693A41205F7F028" ID-ZooBank="4D1F8139D4204E6E9B8E2401EF6232DA" ModsDocID="1860-0743-99-1-77" checkinTime="1674309841633" checkinUser="pensoft" docAuthor="Makino, Tomohisa, Nakano, Takafumi, Okamoto, Taku &amp; Hikida, Tsutomu" docDate="2023" docId="D886647200ED5D7DB6403CE7A01E97B0" docLanguage="en" docName="ZoosystEvol 99(1): 77-91" docOrigin="Zoosystematics and Evolution 99 (1)" docPubDate="2023-01-20" docSource="http://dx.doi.org/10.3897/zse.99.95923" docTitle="Ateuchosaurus okinavensis" docType="treatment" docVersion="6" id="F3CD023B35EE51769693A41205F7F028" lastPageNumber="77" masterDocId="F3CD023B35EE51769693A41205F7F028" masterDocTitle="Taxonomic revision and re-description of Ateuchosaurus pellopleurus (Hallowell, 1861) (Reptilia, Squamata, Scincidae) with resurrection of A. okinavensis (Thompson, 1912)" masterLastPageNumber="91" masterPageNumber="77" pageNumber="77" updateTime="1732849130218" updateUser="ExternalLinkService">
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<mods:titleInfo id="D0D9068D9B80082F73A3A1C5E34A08A7">
<mods:title id="40A580008FC2E70B0B8EEAD245EFC104">Taxonomic revision and re-description of Ateuchosaurus pellopleurus (Hallowell, 1861) (Reptilia, Squamata, Scincidae) with resurrection of A. okinavensis (Thompson, 1912)</mods:title>
</mods:titleInfo>
<mods:name id="9A502EBCC28223BD686797EC02F9FFE3" type="personal">
<mods:role id="C9DF889E3AF7FC45F3BB025CF4CEAA61">
<mods:roleTerm id="482A64F2FBDEFE5F85543AF48C668A46">Author</mods:roleTerm>
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<mods:namePart id="9150AAC1EC948C580E224037D878E73D">Makino, Tomohisa</mods:namePart>
<mods:nameIdentifier id="60B0E508C454B1A8A698DFFBC0B39F3E" type="ORCID">https://orcid.org/0000-0001-8249-7249</mods:nameIdentifier>
<mods:affiliation id="3C4FE4DBFFDC177EF78CA3477F5835E7">Department of Zoology, Graduate School of Science, Kyoto University, Sakyo-ku, Kyoto 606 - 8502, Japan</mods:affiliation>
<mods:nameIdentifier id="65E7D0CCEFFA127E04EE8E3D8DCECFB2" type="email">makino@zoo.zool.kyoto-u.ac.jp</mods:nameIdentifier>
</mods:name>
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<mods:roleTerm id="69B30B4FD812E9F0448D657D9292C1B8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A5A5487F1DC804AA9366A8524A8878F9">Nakano, Takafumi</mods:namePart>
<mods:nameIdentifier id="0085F62F7B8ED1C33ED285FC11029195" type="ORCID">https://orcid.org/0000-0001-6107-2188</mods:nameIdentifier>
<mods:affiliation id="0FD3DD1E9AC256EEFE392D3F5A85B8EE">Department of Zoology, Graduate School of Science, Kyoto University, Sakyo-ku, Kyoto 606 - 8502, Japan</mods:affiliation>
</mods:name>
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<mods:namePart id="42FB282A17F8BFF66A56AE706B5AC859">Okamoto, Taku</mods:namePart>
<mods:affiliation id="1CE7585FD6A0F1DA04944DEC9F420974">Department of Zoology, Graduate School of Science, Kyoto University, Sakyo-ku, Kyoto 606 - 8502, Japan</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="5D2B19A7FF24112DFED1D5848795FFD5">Hikida, Tsutomu</mods:namePart>
<mods:affiliation id="39C390D3CE8167F9733758D1793942C1">Department of Zoology, Graduate School of Science, Kyoto University, Sakyo-ku, Kyoto 606 - 8502, Japan</mods:affiliation>
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<mods:title id="212BD7436616B380C1E868872B6EB28B">Zoosystematics and Evolution</mods:title>
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<mods:date id="DCBCB335CC1AD6AF8187D1198615D169">2023</mods:date>
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<mods:number id="F6E16D134271D48531205FEC87F1F064">99</mods:number>
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<mods:number id="2B1A996AE9BB1AC4D9C198AD7FE6D0BA">1</mods:number>
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<subSection id="EB35FB4FC6F99DAF3C476CE9CAD4BFD5" pageId="0" pageNumber="77" type="taxonomy">
<treatment id="D886647200ED5D7DB6403CE7A01E97B0" ID-GBIF-Taxon="209013747" LSID="urn:lsid:plazi:treatment:D886647200ED5D7DB6403CE7A01E97B0" httpUri="http://treatment.plazi.org/id/D886647200ED5D7DB6403CE7A01E97B0" lastPageNumber="77" pageId="0" pageNumber="77" scope_class="Squamata" scope_family="Scincidae">
<subSubSection id="5FE359F1B3E69D21965F97B541D2CC55" pageId="0" pageNumber="77" type="nomenclature">
<paragraph id="045A44B8EB4849996491DF086B8C25C3" pageId="0" pageNumber="77">
<taxonomicName id="9E98D7AD6F39013F4E2DE7735C638417" ID-CoL="3WQT6" LSID="D8866472-00ED-5D7D-B640-3CE7A01E97B0" authority="(Thompson, 1912)" baseAuthorityName="Thompson" baseAuthorityYear="1912" family="Scincidae" genus="Ateuchosaurus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ateuchosaurus okinavensis" order="Squamata" pageId="0" pageNumber="77" rank="species" species="okinavensis">Ateuchosaurus okinavensis (Thompson, 1912)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="D9C18A52331323536054ECCED83E0AD4" pageId="0" pageNumber="77" type="description">
<paragraph id="4D86035F505F65990559A199E9D1C551" pageId="0" pageNumber="77">
<figureCitation id="D3514ABCE2112BE1FF688CF216ABCF37" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal (a) and ventral (b) views of the holotype of Ateuchosaurus okinavensis (CAS 21537)." figureDoi="10.3897/zse.99.95923.figure5" httpUri="https://binary.pensoft.net/fig/798126" pageId="0" pageNumber="77">Figs 5</figureCitation>
<figureCitation id="AD00C43B5393B3D58F18E4260F3C8237" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Dorsal, lateral, and ventral views of head scalation of Ateuchosaurus okinavensis (KUZ R 77462). See Fig. 4 legend for abbreviations of scale names." figureDoi="10.3897/zse.99.95923.figure6" httpUri="https://binary.pensoft.net/fig/798127" pageId="0" pageNumber="77">, 6</figureCitation>
<figureCitation id="2ADB2D7F3BA167ABBB938A9B2091F597" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Photograph of a live individual of Ateuchosaurus okinavensis collected at Urasoe Park in Urasoe City, Okinawajima Island (26 ° 15 ' 17.37 &quot; N, 127 ° 43 ' 41.25 &quot; E, 38 m above sea level)." figureDoi="10.3897/zse.99.95923.figure7" httpUri="https://binary.pensoft.net/fig/798128" pageId="0" pageNumber="77">, 7 Suggested Japanese name: Okinawa-Hime-Tokage</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="1607BBA083AABCBF75D9FC59D2462A6A" pageId="0" pageNumber="77" type="reference_group">
<paragraph id="CD76E914A6B30FA627AAC9CADAC30966" pageId="0" pageNumber="77">
<taxonomicName id="4F651BADD57F05B9D0E98E6D15C61E83" authorityName="Hallowell" authorityYear="1861" family="Scincidae" genus="Lygosaurus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Lygosaurus pellopleurus" order="Squamata" pageId="0" pageNumber="77" rank="species" species="pellopleurus">Lygosaurus pellopleurus</taxonomicName>
Hallowell, 1861: 496-497 (part);
<bibRefCitation id="C644C6329BFC45ADB4C25E15FC9970DD" author="Stejneger, L" journalOrPublisher="Bulletin - United States National Museum" pageId="0" pageNumber="77" pagination="1 - 577" refId="B36" refString="Stejneger, L, 1907. Herpetology of Japan and adjacent territory. Bulletin - United States National Museum 58: 1 - 577" title="Herpetology of Japan and adjacent territory." volume="58" year="1907">Stejneger 1907</bibRefCitation>
: 222-224 (part); 1927: 2;
<bibRefCitation id="49FB293DB61AF306E803F8EEF77B5D8F" author="Van Denburgh, J" journalOrPublisher="Private publication, San Francisco" pageId="0" pageNumber="77" refId="B41" refString="Van Denburgh, J, 1912a. Advance diagnoses of new reptiles and amphibians from the Loo Choo Islands and Formosa. Private publication, San Francisco, 1-8." title="Advance diagnoses of new reptiles and amphibians from the Loo Choo Islands and Formosa. Private publication, San Francisco, 1 - 8." year="1912 a">Van Denburgh 1912a</bibRefCitation>
: 7 (part); 1912b: 240-241 (part);
<bibRefCitation id="A59EC51F9B8BAA62CBD385609EF502BF" author="Sclater, JA" journalOrPublisher="BMC Evolutionary Biology" pageId="0" pageNumber="77" refId="B32" refString="Sclater, JA, 1950. Bionomic notes on some reptiles form Okinawa Shima, Ryukyu Islands, Japan. The Annals &amp; magazine of natural history. 12th series, Zoology, Botany, and Geology 3: 113-116." title="Bionomic notes on some reptiles form Okinawa Shima, Ryukyu Islands, Japan. The Annals &amp; magazine of natural history. 12 th series, Zoology, Botany, and Geology 3: 113 - 116." year="1950">Sclater 1950</bibRefCitation>
: 114.
</paragraph>
<paragraph id="99DE4E6505962640C1F9441F3AC00571" pageId="0" pageNumber="77">
<taxonomicName id="F7707FB02F73CB2A5967D151C8653760" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Lygosoma pellopleurum" order="Squamata" pageId="0" pageNumber="77" rank="species" species="pellopleurum">Lygosoma pellopleurum</taxonomicName>
:
<bibRefCitation id="30B851A0B4953214BFBB4EF15A5B0E30" author="Boulenger, GA" journalOrPublisher="British Museum (Natural history), London" pageId="0" pageNumber="77" refId="B3" refString="Boulenger, GA, 1887. Catalogue of the lizards in the British Museum (Natural History). 2nd edn. Volume III. British Museum (Natural history), London" title="Catalogue of the lizards in the British Museum (Natural History). 2 nd edn. Volume III." year="1887">Boulenger 1887</bibRefCitation>
: 319;
<bibRefCitation id="5AC3E420ED51B505A9D7BF18D769452D" author="Okada, S" journalOrPublisher="Kinkōdō, Tokyo" pageId="0" pageNumber="77" refId="B23" refString="Okada, S, 1891. Catalogue of Vertebrated Animals of Japan. Kinkōdō, Tokyo" title="Catalogue of Vertebrated Animals of Japan." year="1891">Okada 1891</bibRefCitation>
: 70.
</paragraph>
<paragraph id="0245F9C46A5267BA08C0CE0DE46E3AFA" pageId="0" pageNumber="77">
<taxonomicName id="FED0A18E0643DE2B69E6774010CE6C0A" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Lygosoma (Homolepida) pellopleurus" order="Squamata" pageId="0" pageNumber="77" rank="species" species="pellopleurus" subGenus="Homolepida">Lygosoma (Homolepida) pellopleurus</taxonomicName>
:
<bibRefCitation id="B37C3C638C39A721EBA8458B2465DF9F" author="Boettger, O" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="77" refId="B2" refString="Boettger, O, 1895. Neue Froesche und Schlangen von den Liukiu-Inseln. Offenbacker Vereins fur Naturkunde. Bericht 33-36: 101-117." title="Neue Froesche und Schlangen von den Liukiu-Inseln. Offenbacker Vereins fur Naturkunde. Bericht 33 - 36: 101 - 117." year="1895">Boettger 1895</bibRefCitation>
: 107.
</paragraph>
<paragraph id="C881F089A38733C99B6114868541C993" pageId="0" pageNumber="77">
<taxonomicName id="8782B1F7976E0053238067FA6D02F427" authorityName="Thompson" authorityYear="1912" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Lygosoma okinavensis" order="Squamata" pageId="0" pageNumber="77" rank="species" species="okinavensis">Lygosoma okinavensis</taxonomicName>
Thompson, 1912: 4.
</paragraph>
<paragraph id="65384944BACF6AD1B689F638D3F8D148" pageId="0" pageNumber="77">
<taxonomicName id="307E6669C864983AEF8CA08C465275FB" baseAuthorityName="Hallowell" baseAuthorityYear="1861" family="Scincidae" genus="Ateuchosaurus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ateuchosaurus pellopleurus" order="Squamata" pageId="0" pageNumber="77" rank="species" species="pellopleurus">Ateuchosaurus pellopleurus</taxonomicName>
:
<bibRefCitation id="BF5225A6211D37F4152D8A38C2F74B8C" author="Smith, MA" journalOrPublisher="Records of the Indian Museum" pageId="0" pageNumber="77" pagination="213 - 234" refId="B34" refString="Smith, MA, 1937. A review of the genus Lygosoma (Scincidae: Reptilia) and its allies. Records of the Indian Museum 39: 213 - 234" title="A review of the genus Lygosoma (Scincidae: Reptilia) and its allies." volume="39" year="1937">Smith 1937</bibRefCitation>
: 231;
<bibRefCitation id="6CC9EF9B48D09E3BEB4EE9883A31167A" author="Tanaka, S" journalOrPublisher="Ienohikari Kyokai, Tokyo" pageId="0" pageNumber="77" refId="B38" refString="Tanaka, S, 1979. Heriguro Himetokage. In: Sengoku S (Ed.) Gensyoku Ryosei Hachurui. Ienohikari Kyokai, Tokyo" title="Heriguro Himetokage. In: Sengoku S (Ed.) Gensyoku Ryosei Hachurui." year="1979">Tanaka 1979</bibRefCitation>
: 37 (part); Hikida, 1996: 81 (part);
<bibRefCitation id="F832E58736C566A8C0069E4DCED0E631" DOI="https://doi.org/10.2307/1565549" author="Ota, H" journalOrPublisher="Journal of Herpetology" pageId="0" pageNumber="77" pagination="106 - 118" refId="B30" refString="Ota, H, Miyaguni, H, Hikida, T, 1999. Geographic variation in the endemic skink, Ateuchosaurus pellopleurus from the Ryukyu Archipelago. Journal of Herpetology 33 (1): 106 - 118, DOI: https://doi.org/10.2307/1565549" title="Geographic variation in the endemic skink, Ateuchosaurus pellopleurus from the Ryukyu Archipelago." url="https://doi.org/10.2307/1565549" volume="33" year="1999">Ota et al. 1999</bibRefCitation>
: 106 (part);
<bibRefCitation id="5E6EA41B0C8A216A97D423FDD8837040" author="Goris, RC" journalOrPublisher="Krieger, Malabar" pageId="0" pageNumber="77" refId="B4" refString="Goris, RC, Maeda, N, 2004. Guide to the Amphibians and Reptiles of Japan. Krieger, Malabar" title="Guide to the Amphibians and Reptiles of Japan." year="2004">Goris and Maeda 2004</bibRefCitation>
: 155-156 (part);
<bibRefCitation id="4C106103CB9E351803A79AB59876B817" DOI="https://doi.org/10.1016/j.ympev.2005.12.011" author="Austin, JJ" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="77" pagination="503 - 511" refId="B1" refString="Austin, JJ, Arnold, EN, 2006. Using ancient and recent DNA to explore relationships of extinct and endangered Leiolopisma skinks (Reptilia: Scincidae) in the Mascarene islands. Molecular Phylogenetics and Evolution 39 (2): 503 - 511, DOI: https://doi.org/10.1016/j.ympev.2005.12.011" title="Using ancient and recent DNA to explore relationships of extinct and endangered Leiolopisma skinks (Reptilia: Scincidae) in the Mascarene islands." url="https://doi.org/10.1016/j.ympev.2005.12.011" volume="39" year="2006">Austin and Arnold 2006</bibRefCitation>
: fig. 2, table 1;
<bibRefCitation id="08358525A41E0A60E029F5A897C4104A" DOI="https://doi.org/10.1186/1471-2148-13-93" author="Pyron, RA" journalOrPublisher="BMC Evolutionary Biology" pageId="0" pageNumber="77" pagination=": 93" refId="B31" refString="Pyron, RA, Burbrink, FT, Wiens, JJ, 2013. A phylogeny and revised classification of Squamata, including 4161 species of lizards and snakes. BMC Evolutionary Biology 13 (1): 93, DOI: https://doi.org/10.1186/1471-2148-13-93" title="A phylogeny and revised classification of Squamata, including 4161 species of lizards and snakes." url="https://doi.org/10.1186/1471-2148-13-93" volume="13" year="2013">Pyron et al. 2013</bibRefCitation>
: fig. 7;
<bibRefCitation id="2C7F052419B20C2D56A90306A8E3846E" DOI="https://doi.org/10.11646/zootaxa.3765.4.2" author="Hedges, SB" journalOrPublisher="Zootaxa" pageId="0" pageNumber="77" pagination="317 - 338" refId="B12" refString="Hedges, SB, 2014. The high-level classification of skinks (Reptilia, Squamata, Scincomorpha). Zootaxa 3765 (4): 317 - 338, DOI: https://doi.org/10.11646/zootaxa.3765.4.2" title="The high-level classification of skinks (Reptilia, Squamata, Scincomorpha)." url="https://doi.org/10.11646/zootaxa.3765.4.2" volume="3765" year="2014">Hedges 2014</bibRefCitation>
: 322;
<bibRefCitation id="23DA1BCC03479A42B3A1507B07BD4240" DOI="https://doi.org/10.1016/j.ympev.2015.10.009" author="Zheng, Y" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="77" pagination="537 - 547" refId="B44" refString="Zheng, Y, Wiens, JJ, 2016. Combining phylogenomic and supermatrix approaches, and a time-calibrated phylogeny for squamate reptiles (lizards and snakes) based on 52 genes and 4162 species. Molecular Phylogenetics and Evolution 94: 537 - 547, DOI: https://doi.org/10.1016/j.ympev.2015.10.009" title="Combining phylogenomic and supermatrix approaches, and a time-calibrated phylogeny for squamate reptiles (lizards and snakes) based on 52 genes and 4162 species." url="https://doi.org/10.1016/j.ympev.2015.10.009" volume="94" year="2016">Zheng and Wiens 2016</bibRefCitation>
: figs 1-2;
<bibRefCitation id="201C2332AA6AE7DC80E15CDA1871E83A" DOI="https://doi.org/10.1007/978-4-431-56432-4_5" author="Okamoto, T" editor="Motokawa, M" journalOrPublisher="Springer Japan KK, Tokyo" pageId="0" pageNumber="77" pagination="135 - 163" refId="B26" refString="Okamoto, T, 2017. Historical biogeography of the terrestrial reptiles of Japan: a comparative analysis of geographic ranges and molecular phylogenies. In: Motokawa, M, Kajihara, H, Eds., Species Diversity of Animals in Japan. Springer Japan KK, Tokyo: 135 - 163, DOI: https://doi.org/10.1007/978-4-431-56432-4_5" title="Historical biogeography of the terrestrial reptiles of Japan: a comparative analysis of geographic ranges and molecular phylogenies." url="https://doi.org/10.1007/978-4-431-56432-4_5" volumeTitle="Species Diversity of Animals in Japan." year="2017">Okamoto 2017</bibRefCitation>
: table 5.2;
<bibRefCitation id="74B74AAE5753C2DF33D05C9C90E596C4" DOI="https://doi.org/10.1016/j.jcz.2020.06.008" author="Makino, T" journalOrPublisher="Zoologischer Anzeiger" pageId="0" pageNumber="77" pagination="1 - 10" refId="B17" refString="Makino, T, Okamoto, T, Kurita, K, Nakano, T, Hikida, T, 2020. Origin and intraspecific diversification of the scincid lizard Ateuchosaurus pellopleurus with implications for historical island biogeography of the Central Ryukyus of Japan. Zoologischer Anzeiger 288: 1 - 10, DOI: https://doi.org/10.1016/j.jcz.2020.06.008" title="Origin and intraspecific diversification of the scincid lizard Ateuchosaurus pellopleurus with implications for historical island biogeography of the Central Ryukyus of Japan." url="https://doi.org/10.1016/j.jcz.2020.06.008" volume="288" year="2020">Makino et al. 2020</bibRefCitation>
: 7 (&quot;southern lineage&quot;);
<bibRefCitation id="0B11D10466FB6B9AAA5E14366D9E40AA" author="Okamoto, T" journalOrPublisher="Sunrise Publishing, Hikone" pageId="0" pageNumber="77" pagination="142 - 143" refId="B27" refString="Okamoto, T, Kurita, K, 2021. Ateuchosaurus pellopleurus (Hallowell, 1861). In: Amphibians and Reptiles of Japan. Sunrise Publishing, Hikone: 142 - 143" title="Ateuchosaurus pellopleurus (Hallowell, 1861)." volumeTitle="Amphibians and Reptiles of Japan." year="2021">Okamoto and Kurita 2021</bibRefCitation>
: 142-143 (part).
</paragraph>
<paragraph id="F3323B51A0C7F837A38FF07CEE90FE83" pageId="0" pageNumber="77">
<taxonomicName id="E9CABAA3E19A3A435ABD928DFCD81828" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Lygosoma pellopleurus" order="Squamata" pageId="0" pageNumber="77" rank="species" species="pellopleurus">Lygosoma pellopleurus</taxonomicName>
:
<bibRefCitation id="DC5F8E41064064264D0FC5C2C78C2280" author="Okada, Y" journalOrPublisher="Section B" pageId="0" pageNumber="77" pagination="159 - 214" refId="B24" refString="Okada, Y, 1939. Studies on the lizards of Japan. Contribution III. Scincidae. Science Reports of the Tokyo Bunrika Daigaku. Section B 4: 159 - 214" title="Studies on the lizards of Japan. Contribution III. Scincidae. Science Reports of the Tokyo Bunrika Daigaku." volume="4" year="1939">Okada 1939</bibRefCitation>
: 206-209 (part).
</paragraph>
<paragraph id="345D8B7BF357905C237111236EC07543" pageId="0" pageNumber="77">
<taxonomicName id="B381D9CBBA746D18BCCB515CF67452D3" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Lygosoma (Ateuchosaurus) pellopleurum" order="Squamata" pageId="0" pageNumber="77" rank="species" species="pellopleurum" subGenus="Ateuchosaurus">Lygosoma (Ateuchosaurus) pellopleurum</taxonomicName>
:
<bibRefCitation id="79EBEE25C54FC654E3DEF71949278872" author="Nakamura, K" journalOrPublisher="Hoikusha Publishing, Osaka" pageId="0" pageNumber="77" refId="B19" refString="Nakamura, K, Ueno, S, 1963. Japanese reptiles and amphibians in colour. Hoikusha Publishing, Osaka" title="Japanese reptiles and amphibians in colour." year="1963">
Nakamura and
<normalizedToken id="62166DFBCADB30701A96151D58046351" originalValue="Uéno">Ueno</normalizedToken>
1963
</bibRefCitation>
: 123-125 (part).
</paragraph>
</subSubSection>
<subSubSection id="DB0BE968EE0D8C708CF2BAD2600786FF" pageId="0" pageNumber="77" type="holotype">
<paragraph id="7845A3B3C5A9417FAEA145816FBA80AF" pageId="0" pageNumber="77">Holotype.</paragraph>
<paragraph id="5C64042018E607224F438AF1A9514314" pageId="0" pageNumber="77">CAS 21537 collected from Nago, Okinawajima Island, Okinawa Group, Ryukyu Archipelago, Japan.</paragraph>
</subSubSection>
<subSubSection id="87B3D7C24592B3BE3781FE7DC1D5FCE4" pageId="0" pageNumber="77" type="materials_examined">
<paragraph id="EF0452997080630D4C893B78840AE6FD" pageId="0" pageNumber="77">Material examined.</paragraph>
<paragraph id="D53AC0DFB631FEF6E60AB946B7558714" pageId="0" pageNumber="77">
<materialsCitation id="4B42A896D4B6E80D7EA2484347BE2B1E" accessionNumber="R55183" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Japan" location="Iheyajima Island" municipality="Iheya Village" specimenCount="1" stateProvince="Okinawa Prefecture">
<collectingCountry id="43B383B9F7DF2EF8F242792B65B8A6D9" name="Japan">Japan</collectingCountry>
•
<collectingRegion id="EFAC2D99DCCC08CD28D63AA6AB3A7356" country="Japan" name="Okinawa">Okinawa Prefecture</collectingRegion>
,
<collectingMunicipality id="4AD1F0E627F90D8C19112E69429009ED">Iheya Village</collectingMunicipality>
,
<location id="666BBE02474E993BC11E185847FFA96F" LSID="urn:lsid:plazi:treatment:D886647200ED5D7DB6403CE7A01E97B0:666BBE02474E993BC11E185847FFA96F" country="Japan" municipality="Iheya Village" name="Iheyajima Island" stateProvince="Okinawa Prefecture">Iheyajima Island</location>
; KUZ
<accessionNumber id="B2BD3DD72B8820DDF0DB18AD92E525E1" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R55183">R55183</accessionNumber>
</materialsCitation>
to 55185, 77521 to 77529, 77531 to 77533 •
<materialsCitation id="C80BAED14994C99B8BCDA3682BFD95C4" accessionNumber="R65611" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Japan" location="Okinawajima Island" specimenCount="1" stateProvince="Okinawa Prefecture">
<collectingRegion id="E3D788DC56259C380D17BFA0ED853E35" country="Japan" name="Okinawa">Okinawa Prefecture</collectingRegion>
,
<location id="8C6F4829FB759287935DBDBB191E3910" LSID="urn:lsid:plazi:treatment:D886647200ED5D7DB6403CE7A01E97B0:8C6F4829FB759287935DBDBB191E3910" country="Japan" name="Okinawajima Island" stateProvince="Okinawa Prefecture">Okinawajima Island</location>
; KUZ
<accessionNumber id="72CDEEC90AB7041252AAEFE3CBD6E55C" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R65611">R65611</accessionNumber>
</materialsCitation>
, 66980 to 66982, 68039 to 68040, 69900, 70122 to 70123, 70125, 70130 to 70131, 70673 to 70674, 77442, 77445 to 77446, 77450 to 77454, 77458 to 77462, 77464, 77466 to 77467, 77470 to 77471, 77587, OMNH R745, 1143, 3061 to 3064 •
<materialsCitation id="E61583C5772C8322119CF0E3F4626592" accessionNumber="R77438" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Japan" location="Minnajima Island" municipality="Motobu Town" specimenCount="1" stateProvince="Okinawa Prefecture">
<collectingRegion id="A318FC42683AEE3352E4460E164C219C" country="Japan" name="Okinawa">Okinawa Prefecture</collectingRegion>
,
<collectingMunicipality id="E4BCEE7CE3704364F9AC15E0D3DA9555">Motobu Town</collectingMunicipality>
,
<location id="1EB891512BCECA46A1ED40BDFC3A66EA" LSID="urn:lsid:plazi:treatment:D886647200ED5D7DB6403CE7A01E97B0:1EB891512BCECA46A1ED40BDFC3A66EA" country="Japan" municipality="Motobu Town" name="Minnajima Island" stateProvince="Okinawa Prefecture">Minnajima Island</location>
; KUZ
<accessionNumber id="007890AB37693AD1B1B310087DB4C7AB" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R77438">R77438</accessionNumber>
</materialsCitation>
to 77440, 77574 to 77584 •
<materialsCitation id="79448240F6B8425D32DB885479D5F7A1" accessionNumber="R70132" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Japan" location="Miyagijima Island" municipality="Uruma City" specimenCount="1" stateProvince="Okinawa Prefecture">
<collectingRegion id="E0471FDBB9590800B0A59EB46A3FA6A0" country="Japan" name="Okinawa">Okinawa Prefecture</collectingRegion>
,
<collectingMunicipality id="720C6C75E3B85C8DE185AC2E9EA6041D">Uruma City</collectingMunicipality>
,
<location id="48BEB4A5796B84D65C098E6629B91E39" LSID="urn:lsid:plazi:treatment:D886647200ED5D7DB6403CE7A01E97B0:48BEB4A5796B84D65C098E6629B91E39" country="Japan" municipality="Uruma City" name="Miyagijima Island" stateProvince="Okinawa Prefecture">Miyagijima Island</location>
; KUZ
<accessionNumber id="B9D7983EE73C908DE992906082F2C7A5" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R70132">R70132</accessionNumber>
</materialsCitation>
to 70133 •
<materialsCitation id="C11898C0010A0738A1EA12C307CDFF20" accessionNumber="R70126" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Japan" location="Hamahigajima Island" municipality="Uruma City" specimenCount="1" stateProvince="Okinawa Prefecture">
<collectingRegion id="04BE39800B3C982708CF276AEB038E07" country="Japan" name="Okinawa">Okinawa Prefecture</collectingRegion>
,
<collectingMunicipality id="11A107A3184085CBFF6358F63F6190C0">Uruma City</collectingMunicipality>
,
<location id="93464FDAE895699C61B6303CE318A63F" LSID="urn:lsid:plazi:treatment:D886647200ED5D7DB6403CE7A01E97B0:93464FDAE895699C61B6303CE318A63F" country="Japan" municipality="Uruma City" name="Hamahigajima Island" stateProvince="Okinawa Prefecture">Hamahigajima Island</location>
; KUZ
<accessionNumber id="560611398D15917E11DC8169A2B27987" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R70126">R70126</accessionNumber>
</materialsCitation>
to 70128 •
<materialsCitation id="1F680109A581E99D8F0530F207B8189F" accessionNumber="R47122" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Japan" location="Tokashikijima Island" municipality="Tokashiki Village" specimenCount="1" stateProvince="Okinawa Prefecture">
<collectingRegion id="A29A1B06F72AB6B22B0CCC5537E7F3BF" country="Japan" name="Okinawa">Okinawa Prefecture</collectingRegion>
,
<collectingMunicipality id="5E4ED22E6D0EC1C61C85F75A608862F7">Tokashiki Village</collectingMunicipality>
,
<location id="13EDBDC6B0B085AF5189D4501437E8AC" LSID="urn:lsid:plazi:treatment:D886647200ED5D7DB6403CE7A01E97B0:13EDBDC6B0B085AF5189D4501437E8AC" country="Japan" municipality="Tokashiki Village" name="Tokashikijima Island" stateProvince="Okinawa Prefecture">Tokashikijima Island</location>
; KUZ
<accessionNumber id="9500E7EBF56EABA2185AA84C26B708E8" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R47122">R47122</accessionNumber>
</materialsCitation>
to 47141, 47143, 70501 to 70503, 70505 to 70507, OMNH R1102, 1107 to 1108 •
<materialsCitation id="31C24F91DE3AAD36FF73A2C999E7169C" accessionNumber="R77623" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Japan" location="Agunijima Island" municipality="Aguni Village" specimenCount="1" stateProvince="Okinawa Prefecture">
<collectingRegion id="54CC7D1A9BBDFFEDCB432E626C8B6688" country="Japan" name="Okinawa">Okinawa Prefecture</collectingRegion>
,
<collectingMunicipality id="AEA7D3ADC08F92398E0B92B8B85EE983">Aguni Village</collectingMunicipality>
,
<location id="3E03A124EB14FE3D10AD41F945D3CC9B" LSID="urn:lsid:plazi:treatment:D886647200ED5D7DB6403CE7A01E97B0:3E03A124EB14FE3D10AD41F945D3CC9B" country="Japan" municipality="Aguni Village" name="Agunijima Island" stateProvince="Okinawa Prefecture">Agunijima Island</location>
; KUZ
<accessionNumber id="B687C937312F244103F722710EAF4702" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R77623">R77623</accessionNumber>
</materialsCitation>
to 77625, 77628 to 77632 •
<materialsCitation id="3E6A77C35A8808208E9D5CD58501BC3A" accessionNumber="R66983" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Japan" location="Tonakijima Island" municipality="Tonaki Village" specimenCount="1" stateProvince="Okinawa Prefecture">
<collectingRegion id="65B4ED5137F74EBB91669544E033657B" country="Japan" name="Okinawa">Okinawa Prefecture</collectingRegion>
,
<collectingMunicipality id="3F58555488B0E1A3F67C18BA9B242D7D">Tonaki Village</collectingMunicipality>
,
<location id="D0B4A37E93A622E6FF1433EF051ED6F1" LSID="urn:lsid:plazi:treatment:D886647200ED5D7DB6403CE7A01E97B0:D0B4A37E93A622E6FF1433EF051ED6F1" country="Japan" municipality="Tonaki Village" name="Tonakijima Island" stateProvince="Okinawa Prefecture">Tonakijima Island</location>
; KUZ
<accessionNumber id="3E1473B359D12950849B5F9786D4DD3E" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R66983">R66983</accessionNumber>
</materialsCitation>
•
<materialsCitation id="B195C576523635895720DBF32D51CDD4" accessionNumber="R70480" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Japan" location="Kumejima Island" municipality="Kumejima Town" specimenCount="1" stateProvince="Okinawa Prefecture">
<collectingRegion id="2B4BDF49CF729B03DF2920648CB56499" country="Japan" name="Okinawa">Okinawa Prefecture</collectingRegion>
,
<collectingMunicipality id="DA9CCE6D33C1F76FB2AFDCEAC151E35E">Kumejima Town</collectingMunicipality>
,
<location id="D1C7EDDB20B10000DE90357D92D4D5E9" LSID="urn:lsid:plazi:treatment:D886647200ED5D7DB6403CE7A01E97B0:D1C7EDDB20B10000DE90357D92D4D5E9" country="Japan" municipality="Kumejima Town" name="Kumejima Island" stateProvince="Okinawa Prefecture">Kumejima Island</location>
; KUZ
<accessionNumber id="8802161301E30961163C764054116F63" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R70480">R70480</accessionNumber>
</materialsCitation>
, 70483 to 70488, 70490 to 70491, 70496 to 70497, 70499 to 70500, OMNH R3065.
</paragraph>
<caption id="0912ECDF5012211E02896CA64E989315" doi="10.3897/zse.99.95923.figure5" httpUri="https://binary.pensoft.net/fig/798126" pageId="0" pageNumber="77" start="Figure 5" startId="F5">
<paragraph id="FC4E6EABB005AB2A2AF6D49C40A37303" pageId="0" pageNumber="77">
<emphasis id="DB40E3DD85D8555FB128B002965AA0BE" bold="true" pageId="0" pageNumber="77">Figure 5.</emphasis>
Dorsal (
<emphasis id="8983958156833253CAFC7ADA22D306AE" bold="true" pageId="0" pageNumber="77">a</emphasis>
) and ventral (
<emphasis id="278C094F1372CFEEC070D486FC2D5623" bold="true" pageId="0" pageNumber="77">b</emphasis>
) views of the
<typeStatus id="AD31820CCE76824C717D9BBD56304B1E">holotype</typeStatus>
of
<taxonomicName id="45CB6BFB58FC8958AA7A139865D1464F" baseAuthorityName="Thompson" baseAuthorityYear="1912" family="Scincidae" genus="Ateuchosaurus" kingdom="Animalia" lsidName="Ateuchosaurus okinavensis" order="Squamata" pageId="0" pageNumber="77" rank="species" species="okinavensis">
<emphasis id="7A6F9EE2222A52E2EAEDB081F9E73CE2" italics="true" pageId="0" pageNumber="77">Ateuchosaurus okinavensis</emphasis>
</taxonomicName>
(
<accessionNumber id="3A98002FB54A8811A7D8D35E3C1AD7E6" httpUri="http://www.ncbi.nlm.nih.gov/protein/CAS21537">CAS 21537</accessionNumber>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="DEE07BB9910EF80ACE621FC7BD1101A1" pageId="0" pageNumber="77" type="emended diagnosis">
<paragraph id="946AC1DC6629648150FFBF3134E091DF" pageId="0" pageNumber="77">Emended diagnosis.</paragraph>
<paragraph id="2ED5AAF4499C07D27FB38C2983FA4EFF" pageId="0" pageNumber="77">
An
<taxonomicName id="5FADE4308A1B7911980478F573A911A9" authorityName="Gray" authorityYear="1845" family="Scincidae" genus="Ateuchosaurus" kingdom="Animalia" lsidName="Ateuchosaurus" order="Squamata" pageId="0" pageNumber="77" rank="genus">
<emphasis id="AC7E010A68EC81D4390325CA8238E02A" italics="true" pageId="0" pageNumber="77">Ateuchosaurus</emphasis>
</taxonomicName>
species characterized by the following: usually fused FrNa and frontal; a pair of frontoparietals that do not contact each other; eight SuCis; anteroposteriorly reduced parietals separated from SuO and pretemporal by EcP; no distinct nuchals; usually six InLas; body size medium (SVL ca. 42-70 mm); widely separated forelimb and hindlimb when appressed; usually 26 or 28 MSRs (mode = 28, 25-28); 54-64 (mode = 59) and 55-67 DMSs (mode = 58) in male and female, respectively; 10-16 TIVs (mode = 13); 10 preanals not enlarged; usually no black stripe on dorsal scale row or one pair of DSRBS; a karyotype of 2
<emphasis id="F28F6E354973F3E5B756C09E83E77689" italics="true" pageId="0" pageNumber="77">n</emphasis>
= 28 (
<bibRefCitation id="205F8099FFA50B4BDCB924390261713E" DOI="https://doi.org/10.2307/1565622" author="Ota, H" journalOrPublisher="Journal of Herpetology" pageId="0" pageNumber="77" pagination="604 - 607" refId="B29" refString="Ota, H, Lin, JT, Bogadek, A, Lau, MW, 1998. Karyotype of the lygosomine genus Ateuchosaurus from East Asia. Journal of Herpetology 31 (4): 604 - 607, DOI: https://doi.org/10.2307/1565622" title="Karyotype of the lygosomine genus Ateuchosaurus from East Asia." url="https://doi.org/10.2307/1565622" volume="31" year="1998">Ota et al. 1998</bibRefCitation>
).
</paragraph>
<caption id="0F189DE63A9C4E5C5C0177C6FCBC5A25" doi="10.3897/zse.99.95923.figure6" httpUri="https://binary.pensoft.net/fig/798127" pageId="0" pageNumber="77" start="Figure 6" startId="F6">
<paragraph id="D4D1F74C242A03D98EF3EA496FFD66E1" pageId="0" pageNumber="77">
<emphasis id="E3FE647F95D7A1972981BA4A0525B603" bold="true" pageId="0" pageNumber="77">Figure 6.</emphasis>
Dorsal, lateral, and ventral views of head scalation of
<taxonomicName id="B67C32A9C1E2D4939DF71DB630827244" baseAuthorityName="Thompson" baseAuthorityYear="1912" family="Scincidae" genus="Ateuchosaurus" kingdom="Animalia" lsidName="Ateuchosaurus okinavensis" order="Squamata" pageId="0" pageNumber="77" rank="species" species="okinavensis">
<emphasis id="21738E641F216C7C5D382B881375F96E" italics="true" pageId="0" pageNumber="77">Ateuchosaurus okinavensis</emphasis>
</taxonomicName>
(KUZ R77462). See Fig.
<figureCitation id="5EE86281D253D3D0B30007C539A67CAB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal, lateral, and ventral views of head scalation of Ateuchosaurus pellopleurus (KUZ R 77703). CS, chinshield; EcP, ectoparietal; F, frontal; FrNa, frontonasal; FrP, frontoparietal; InLa, infralabial; IP, interparietal; Lo, loreal; M, mental; Na, nasal; P, parietal; PE, parietal eye; PoG, postgenial; PoLa, postlabial; PoM, postmental; PoO, postocular; PoSO, postsubocular; PrO, preocular; PrSO, presubocular; PrT, pretemporal; R, rostral; SuCi, supraciliary; SuLa, supralabial; SuLo, supraloreal; SuNa, supranasal; SuO, supraocular; T, temporal. The dashed line on the fused scale of Na and first SuLa in the lateral view indicates the line of a shallow groove on the scale." figureDoi="10.3897/zse.99.95923.figure4" httpUri="https://binary.pensoft.net/fig/798125" pageId="0" pageNumber="77">4</figureCitation>
legend for abbreviations of scale names.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="F79547CDC50B3BAA1422C9E7B64C566F" pageId="0" pageNumber="77" type="comparisons">
<paragraph id="DF96FFFB6E5A9DF8A31BA57AF6E00CCA" pageId="0" pageNumber="77">Comparison.</paragraph>
<paragraph id="9208E6FDC2626AE9788B3FD3755F6B3A" pageId="0" pageNumber="77">
<taxonomicName id="493FA68752DEE0AD5E70482EFC7C2F2B" baseAuthorityName="Thompson" baseAuthorityYear="1912" family="Scincidae" genus="Ateuchosaurus" kingdom="Animalia" lsidName="Ateuchosaurus okinavensis" order="Squamata" pageId="0" pageNumber="77" rank="species" species="okinavensis">
<emphasis id="B47124BEEDF37D60CFBF8EB45B4099F8" italics="true" pageId="0" pageNumber="77">Ateuchosaurus okinavensis</emphasis>
</taxonomicName>
, together with
<taxonomicName id="47563C48A959C2363A6DAF8810B2E5F5" lsidName="A. pellopleurus" pageId="0" pageNumber="77" rank="species" species="pellopleurus">
<emphasis id="907FC2BE958D87A1E6AF7D8A44DEC0B2" italics="true" pageId="0" pageNumber="77">A. pellopleurus</emphasis>
</taxonomicName>
, is distinguished from
<taxonomicName id="0DF3E87B858F72DBE1D3F4924C83138A" lsidName="A. chinensis" pageId="0" pageNumber="77" rank="species" species="chinensis">
<emphasis id="A38CDB0EB19C650E8DFABBA52E95D2DA" italics="true" pageId="0" pageNumber="77">A. chinensis</emphasis>
</taxonomicName>
by having the following characters (
<bibRefCitation id="58573D46D6936ACF39198249C22BB461" author="Nguyen, QT" journalOrPublisher="Herpetology Notes" pageId="0" pageNumber="77" pagination="17 - 21" refId="B21" refString="Nguyen, QT, Tung, TT, Ngoc, HV, Boehme, W, Ziegler, T, 2008. Rediscovery and redescription of Ateuchosaurus chinensis Gray, 1845 (Squamata: Sauria: Scincidae) from northeastern Vietnam. Herpetology Notes 1: 17 - 21" title="Rediscovery and redescription of Ateuchosaurus chinensis Gray, 1845 (Squamata: Sauria: Scincidae) from northeastern Vietnam." volume="1" year="2008">Nguyen et al. 2008</bibRefCitation>
): usually six InLas (vs. seven InLas); smaller body size (SVL ca. 42-70 mm vs. 70.0-83.8 mm; see also
<bibRefCitation id="B7EF25E21AFB1311AF2E8E832FE76E6C" author="Okamoto, T" journalOrPublisher="Sunrise Publishing, Hikone" pageId="0" pageNumber="77" pagination="142 - 143" refId="B27" refString="Okamoto, T, Kurita, K, 2021. Ateuchosaurus pellopleurus (Hallowell, 1861). In: Amphibians and Reptiles of Japan. Sunrise Publishing, Hikone: 142 - 143" title="Ateuchosaurus pellopleurus (Hallowell, 1861)." volumeTitle="Amphibians and Reptiles of Japan." year="2021">Okamoto and Kurita 2021</bibRefCitation>
); usually 26 or 28 MSRs (vs. 30 scale rows); usually 10-14 TIVs (vs. usually 16-18 TIVs); 10 preanals (vs. usually six preanals); blackish line on dorsolateral surface from snout to midbody (vs. no such line). This species also is distinguished from
<taxonomicName id="C0C053D1683FA327217C17B44610C55D" lsidName="A. chinensis" pageId="0" pageNumber="77" rank="species" species="chinensis">
<emphasis id="FF6FED52EB7BAA9F99B6C857F23E96F3" italics="true" pageId="0" pageNumber="77">A. chinensis</emphasis>
</taxonomicName>
by the karyotype of 2
<emphasis id="F9E8D054FF47A0E36AB688D73E7B3634" italics="true" pageId="0" pageNumber="77">n</emphasis>
= 28 (vs. 2
<emphasis id="F0D8761997909F77E96D3322D5862130" italics="true" pageId="0" pageNumber="77">n</emphasis>
= 26; no information about the karyotype of
<taxonomicName id="7137B31C52576FEBF4D9ABD932E78D5C" lsidName="A. pellopleurus" pageId="0" pageNumber="77" rank="species" species="pellopleurus">
<emphasis id="4612267F6A5BD45C08A4590A72A855AC" italics="true" pageId="0" pageNumber="77">A. pellopleurus</emphasis>
</taxonomicName>
;
<bibRefCitation id="3036E8B9AB9FE9E0920C60AD614DE692" DOI="https://doi.org/10.2307/1565622" author="Ota, H" journalOrPublisher="Journal of Herpetology" pageId="0" pageNumber="77" pagination="604 - 607" refId="B29" refString="Ota, H, Lin, JT, Bogadek, A, Lau, MW, 1998. Karyotype of the lygosomine genus Ateuchosaurus from East Asia. Journal of Herpetology 31 (4): 604 - 607, DOI: https://doi.org/10.2307/1565622" title="Karyotype of the lygosomine genus Ateuchosaurus from East Asia." url="https://doi.org/10.2307/1565622" volume="31" year="1998">Ota et al. 1998</bibRefCitation>
).
<taxonomicName id="B1758A9311658C4FB80D9360867678A3" baseAuthorityName="Thompson" baseAuthorityYear="1912" family="Scincidae" genus="Ateuchosaurus" kingdom="Animalia" lsidName="Ateuchosaurus okinavensis" order="Squamata" pageId="0" pageNumber="77" rank="species" species="okinavensis">
<emphasis id="E083DB474B28521D2E0CBD7D8E6AEC39" italics="true" pageId="0" pageNumber="77">Ateuchosaurus okinavensis</emphasis>
</taxonomicName>
resembles
<taxonomicName id="FAB8138102E70C75E9622FC9B7F22F00" lsidName="A. pellopleurus" pageId="0" pageNumber="77" rank="species" species="pellopleurus">
<emphasis id="4F370B0C90958DCFA73C2D7A2533661B" italics="true" pageId="0" pageNumber="77">A. pellopleurus</emphasis>
</taxonomicName>
in MSR and body size but differs in almost having a fused FrNa and frontal (vs. separated ones), 54-64 and 55-67 DMSs in male and female (vs. 53-70 and 56-69 DMSs in male and female), 10-16 TIVs (vs. 8-14 TIVs), usually 0-1 pair of DSRBS (vs. usually 2-3 pairs).
</paragraph>
<caption id="2A3394A5F707AEFAB8B99354BA863261" doi="10.3897/zse.99.95923.figure7" httpUri="https://binary.pensoft.net/fig/798128" pageId="0" pageNumber="77" start="Figure 7" startId="F7">
<paragraph id="1686F11FF6F084958F8DD9B306DB0789" pageId="0" pageNumber="77">
<emphasis id="7AA5250C8AFA556CA15810F4E144E5C9" bold="true" pageId="0" pageNumber="77">Figure 7.</emphasis>
Photograph of a live individual of
<taxonomicName id="A27028A679DE8262FFBFE982BD0F2310" baseAuthorityName="Thompson" baseAuthorityYear="1912" family="Scincidae" genus="Ateuchosaurus" kingdom="Animalia" lsidName="Ateuchosaurus okinavensis" order="Squamata" pageId="0" pageNumber="77" rank="species" species="okinavensis">
<emphasis id="01612455F22AD4DDD38A9387261492ED" italics="true" pageId="0" pageNumber="77">Ateuchosaurus okinavensis</emphasis>
</taxonomicName>
collected at Urasoe Park in Urasoe City, Okinawajima Island (
<geoCoordinate id="1AE1FDA092373706AE57CBCEA7F7241D" degrees="26" direction="north" minutes="15" orientation="latitude" precision="1" seconds="17.37" value="26.254826">26°15'17.37&quot;N</geoCoordinate>
,
<geoCoordinate id="48B9A91D29CD5CAC9C54F6F348878A11" degrees="127" direction="east" minutes="43" orientation="longitude" precision="1" seconds="41.25" value="127.72813">127°43'41.25&quot;E</geoCoordinate>
, 38 m above sea level).
</paragraph>
</caption>
<paragraph id="4F42558F2AAAE26EEDD115B48CFD9ACC" pageId="0" pageNumber="77">
The mean K2P distance
<normalizedToken id="34C8C212F2DA663EC0E25D9EB9E8333C" originalValue="±">+/-</normalizedToken>
standard deviation (range in parenthesis) based on cytochrome
<emphasis id="D07CDEEA48FA17966C3CF7DBCA39BF11" italics="true" pageId="0" pageNumber="77">b</emphasis>
sequences between
<taxonomicName id="46A04935DAFB27D7BB493DEC42693090" lsidName="A. okinavensis" pageId="0" pageNumber="77" rank="species" species="okinavensis">
<emphasis id="A093D58B7E22A964681E2B1C081A70FF" italics="true" pageId="0" pageNumber="77">A. okinavensis</emphasis>
</taxonomicName>
and
<taxonomicName id="0507A0B2A456D1720D3C3E00EEA5B755" lsidName="A. chinensis" pageId="0" pageNumber="77" rank="species" species="chinensis">
<emphasis id="523680C9E5680A22E6398C0E5444DD11" italics="true" pageId="0" pageNumber="77">A. chinensis</emphasis>
</taxonomicName>
was 24.7%
<normalizedToken id="38E1B8D7145A4623BFA65FF38B82F4D8" originalValue="±">+/-</normalizedToken>
0.3% (23.9%-25.5%); the value between
<taxonomicName id="519923CC649F43BB31745624F13971AA" lsidName="A. okinavensis" pageId="0" pageNumber="77" rank="species" species="okinavensis">
<emphasis id="A170748CFD5F4837A656E2A28E357416" italics="true" pageId="0" pageNumber="77">A. okinavensis</emphasis>
</taxonomicName>
and
<taxonomicName id="A723BA1C64BD3F34992CD85D0D5D8FD1" lsidName="A. pellopleurus" pageId="0" pageNumber="77" rank="species" species="pellopleurus">
<emphasis id="FAECF701804EEEBAF96172AFAE36D46D" italics="true" pageId="0" pageNumber="77">A. pellopleurus</emphasis>
</taxonomicName>
, see Comparison of
<taxonomicName id="1BD4E379DC68ED0C6D890B5010503174" lsidName="A. pellopleurus" pageId="0" pageNumber="77" rank="species" species="pellopleurus">
<emphasis id="5BCC5E9788F23907E73BFB41EEF1BFD1" italics="true" pageId="0" pageNumber="77">A. pellopleurus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="60AB41D7CE523729EBE3CA15B466DC61" pageId="0" pageNumber="77" type="description">
<paragraph id="8E7A4910164266BD0EFA72F5E48B7DEC" pageId="0" pageNumber="77">Description.</paragraph>
<paragraph id="102743A4C9A5182DBC5954F2EF1BF1C8" pageId="0" pageNumber="77">
Mainly based on a specimen (KUZ R77462), an adult male collected from Ogimi Village in Okinawajima Island (
<geoCoordinate id="61BD111E11E903577A6A5023C1A4C7E8" degrees="26" direction="north" minutes="41" orientation="latitude" precision="1" seconds="44.88" value="26.695799">26°41'44.88&quot;N</geoCoordinate>
,
<geoCoordinate id="0D0ACD93B1246B8B862AA0059104E18F" degrees="128" direction="east" minutes="07" orientation="longitude" precision="1" seconds="31.14" value="128.12532">128°07'31.14&quot;E</geoCoordinate>
, 111 m above sea level) (followed by ranges of several morphometric and meristic characters in parenthesis;
<emphasis id="FD9E9C3E30906F42ED911411D3DDC2AF" italics="true" pageId="0" pageNumber="77">N</emphasis>
= 13 collected from Ogimi Village, KUZ R70673-70674, 77446, 77451-77454, 77458-77462, 77464); SVL 54.4 mm (55.0
<normalizedToken id="766E139D95C6749FA289442129B78385" originalValue="±">+/-</normalizedToken>
4.4, 49.8-64.8); HL 6.7 mm (6.3
<normalizedToken id="80A1D9FB73A77D4DEE6DF6766A2464E5" originalValue="±">+/-</normalizedToken>
0.4, 5.7-7.0); HW 5.4 mm (5.5
<normalizedToken id="658A472FE71941EF9CECFCF947B0345B" originalValue="±">+/-</normalizedToken>
0.4, 5.1-6.1); HH 3.8 mm (3.8
<normalizedToken id="EF9D7B7AD5371991EA44C80C90DE4ECC" originalValue="±">+/-</normalizedToken>
0.3, 3.3-4.4); SEyL 3.2 mm (3.1
<normalizedToken id="0AD68D2EA42076036184F96A2B8F8D7B" originalValue="±">+/-</normalizedToken>
0.2, 2.8-3.4); EyL 2.3 mm (2.3
<normalizedToken id="D567AE2B5256C0FBED388A1486292E07" originalValue="±">+/-</normalizedToken>
0.2, 2.0-2.6); EyEaL 3.3 mm (3.0
<normalizedToken id="224A1BF4B591259994D98CC61A4BFCCD" originalValue="±">+/-</normalizedToken>
0.3, 2.6-3.3); EaD 1.0 mm (1.1
<normalizedToken id="FA2579BB965A263679BD8813798F7649" originalValue="±">+/-</normalizedToken>
0.1, 0.9-1.4); SAL 19.3 mm (19.2
<normalizedToken id="F55204E7F5F931A09850083F3900DB30" originalValue="±">+/-</normalizedToken>
1.5, 16.7-21.8); AGL 31.1 mm (31.8
<normalizedToken id="1FFECE102C91E633CCCF166F34372250" originalValue="±">+/-</normalizedToken>
2.8, 27.5-37.5); TaL 38.8 mm (the tip of tail lost; the remaining tail original; 61.4
<normalizedToken id="77AEC166B572547D9CD7B7D30D5D4A8A" originalValue="±">+/-</normalizedToken>
3.5, 57.6-64.5,
<emphasis id="9CC9E38704E05AC0ED9CFE5725F3A98D" italics="true" pageId="0" pageNumber="77">N</emphasis>
= 3 with undamaged original tail); FlL 8.5 mm (8.9
<normalizedToken id="F7A6D2FE6FFB5E46517174CB8371CE60" originalValue="±">+/-</normalizedToken>
0.9, 7.5-10.2,
<emphasis id="C7C9DF3AE78683BED5484822C85E8E18" italics="true" pageId="0" pageNumber="77">N</emphasis>
= 12); HlL 14.3 mm (14.2
<normalizedToken id="97F21489D39B4240FF7B2D4A3495E1F7" originalValue="±">+/-</normalizedToken>
1.0, 12.3-15.5); FIIIL 2.0 mm (1.9
<normalizedToken id="9917D35615C27FB036922CE3A64CB379" originalValue="±">+/-</normalizedToken>
0.2, 1.6-2.3,
<emphasis id="0DCA3F0F587225C3FE9552615B75B088" italics="true" pageId="0" pageNumber="77">N</emphasis>
= 12); TIVL 4.4 mm (4.3
<normalizedToken id="F4AF1EBA88D680CB30DC8646F9D61D41" originalValue="±">+/-</normalizedToken>
0.4, 3.5-5.1).
</paragraph>
<paragraph id="5F35681A16B87BFBEBE012BFBCD5D8C6" pageId="0" pageNumber="77">
Snout obtusely pointed; rostral visible from above, overlapping supranasal and nasal; nasal fused with first supralabial, with no groove on boundary between nasal and labial parts; nostril at center of nasal part of nasal; supranasal unpaired with posterior edge concaved, overlapped by nasal; supraloreal overlapped by supranasal and anterior and posterior loreals, overlapping frontonasal part and first supraciliary; frontonasal and frontal fused, frontonasal part overlapped by supranasal, overlapping first supraocular, frontal part overlapping frontoparietals and interparietal; no prefrontal; four supraoculars (no variation,
<emphasis id="E295360CC8E24E323A9A9A49B70B67F1" italics="true" pageId="0" pageNumber="77">N</emphasis>
= 12), anterior ones overlapping posterior ones, first one overlapped by first, second and third supraciliaries, first and second ones overlapping frontal part, third and fourth ones overlapping frontoparietal; a pair of frontoparietals as long as width, separated from each other by interparietal, overlapping interparietal, parietal and ectoparietal; interparietal with short arrowhead-like tetragonal shape, a little longer and narrower than frontoparietal, overlapping parietals; ectoparietal pentagonal, shorter than and as wide as fourth supraocular, overlapped by fourth supraocular and upper pretemporal, overlapping parietal; parietals shorter and almost 1.2 times wider than interparietal, separated from each other by interparietal; no distinct nuchal.
</paragraph>
<paragraph id="7C8E6889696A412066F8C11B7A8A0245" pageId="0" pageNumber="77">Two loreals with a half length and similar height of second supralabial, anterior one overlapped by nasal, overlapping supranasal, posterior one and second supralabial, posterior one overlapped by second supralabial, overlapping first supraciliary and upper and lower preoculars; eight supraciliaries (mode = 8, 8-9), anterior ones overlapping posterior ones, first one overlapped by upper preocular, third and fourth ones overlapping second supraocular, fifth and sixth ones overlapping third supraocular, seventh one overlapping fourth supraocular, eighth one overlapping upper pretemporal and upper postocular; two preoculars, upper one similar size to first supraciliary, overlapped by lower one, lower one larger than upper one, overlapped by second supralabial, overlapping third supralabial; two presuboculars, anterior one overlapped by lower preocular in left side (overlapped by third supralabial and lower preocular in right side), overlapping third supralabial and posterior one in left side (overlapping posterior one in right side), posterior one overlapped by third supralabial, overlapping fourth supralabial; three postsuboculars, lower ones overlapping upper ones, lowermost one largest, overlapped by fourth supralabial, lowermost and middle ones overlapping fifth supralabial, middle and uppermost ones overlapping primary temporal, uppermost one overlapping anterior and lower postoculars; three postoculars, anterior one overlapping upper and lower ones, upper one overlapping lower one; two pretemporals, upper one overlapped by fourth supraocular and upper postocular, overlapping lower one and uppermost secondary temporal, lower one smaller than upper one, overlapped by upper and lower postoculars, overlapping first to third secondary temporals; temporals with similar shape and size of body scale; single primary temporal overlapped by lower postocular and fifth supralabial, overlapping sixth supralabial; four secondary temporals, uppermost one overlapped by ectoparietal, second one overlapped by uppermost one, third one overlapped by primary temporal and second one on left side (overlapped by primary temporal, overlapping second one on right side), lowermost one overlapped by primary temporal, third one and sixth supralabial, overlapping upper postlabial; six supralabials (no variation), anterior ones overlapping posterior ones, fourth and fifth ones largely interrupted by lower-anterior postsubocular, fifth one longest, third to fifth ones under eye; two postlabials overlapped by sixth supralabial, upper one overlapping lower one.</paragraph>
<paragraph id="8D5519CE129FB7E3494E90159E5153DF" pageId="0" pageNumber="77">Mental as wide as and lower than rostral, overlapping postmental and first infralabial; single postmental as long as and narrower than mental, overlapped by first infralabial, overlapping chinshields; six infralabials on left side (five on right side, mode = 6, 4-7), anterior ones overlapping posterior ones, first one overlapping chinshield, sixth one smallest; a pair of chinshields, separated from each other, smaller than postmental, overlapped by second infralabial, overlapping postgenial; a pair of postgenials, smaller than chinshields, overlapped by second and third infralabials.</paragraph>
<paragraph id="74822D5E1D9927E3E7E866C802FA7A33" pageId="0" pageNumber="77">
Ear opening with oval shape, no ear lobule; forelimbs with five short but distinct fingers, third one longest; third finger with supradigital scales of single row at tip and two rows at base and eight subdigital scales (mode = 7, 7-8,
<emphasis id="021E1E77A806FB81F290A925D7244B3E" italics="true" pageId="0" pageNumber="77">N</emphasis>
= 12); 25 scale rows around midbody (mode = 26, 25-28); 59 pairs of dorsal median scales from posterior side of parietal to position of posterior margin of preanal (mode = 56, 55-61); usually three keels on a dorsal scale; hindlimbs with five distinct toes, fourth one longest; fourth toe with three rows of supradigital scales at base and 14 subdigital scales (mode = 14, 13-16); 10 preanals, central ones with shorter length and similar width of midbody scales, overlapped by outer ones, right-central one overlapped by left-central one, outermost ones small; subcaudal not enlarged; 18 scale rows around tail at 10th subcaudal position.
</paragraph>
<paragraph id="1A55F77C7CDD8335FF1EF592286C0AB2" pageId="0" pageNumber="77">In alcohol, dorsal surface of head to tail light brown, first scale row from dorsal median line on midbody marginally darker with small blackish spots; no distinct blackish stripe on dorsal scale row (mode = 0, 0-1); first scale row from dorsal median line on original tail with a pale blackish stripe; upper and lower eyelids whitish with blackish edge; blackish line on dorsolateral surface from snout to midbody interrupting dorsal and lateral sides, with narrower width of second supralabial on snout, interrupted by eye, with similar width of dorsolateral scale from eye to neck, 1.0-1.5 times wider than dorsolateral scale at position above forelimb, obscure from that position to midbody; upper margin of blackish line distinct from posterior corner of eye to midbody, located on middle position of scales on fourth scale row from dorsal median line; lateral surface of head to neck brownish white with many black spots; lateral surface of upper and lower jaws with several black spots; lateral surface around forelimb to tail dark brown; ventral surface pale yellow from head to tail, with many short stripes and spots irregularly scattered in head, black spots between hindlimb, and a pale black spot on each scale in tail; dorsal surface of forelimb dark brown with a rough blackish line on each scale row; ventral surface of forelimb pale yellow without blackish line; dorsal surface of hindlimb dark brown with a blackish line on each scale row; ventral surface of hindlimb pale yellow with a blackish dotted line on each scale row.</paragraph>
</subSubSection>
<subSubSection id="772C84A30C50001668AD8177E05C1CBC" pageId="0" pageNumber="77" type="variation">
<paragraph id="A2120582979185E785653F6BC916ED14" pageId="0" pageNumber="77">Variation.</paragraph>
<paragraph id="F549BA60F5344E11C0F0DF8039949073" pageId="0" pageNumber="77">For SuNa, this species usually shows a single scale with a concave posterior edge (84/104 specimens), and sometimes two laterally separated scales (1/104 specimens) or a single scale with an almost straight posterior edge (19/104 specimens).</paragraph>
<paragraph id="B360AB4F5B4418480C068F7055C59DB8" pageId="0" pageNumber="77">
This species usually has a long fused FrNa and frontal scale, but sometimes FrNa and frontal are distinct (Table
<tableCitation id="0952F0806C5D60AFFBA19D9562EE5B39" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Variation of the condition of frontonasal-frontal and the number of subdigital scales of the fourth toe (TIV) in Ateuchosaurus pellopleurus and A. okinavensis." httpUri="http://table.plazi.org/id/4D51506529DC5DA604A7AEAEFFC59096" pageId="0" pageNumber="77" tableUuid="4D51506529DC5DA604A7AEAEFFC59096">1</tableCitation>
;
<bibRefCitation id="1DFA945B5D6286AEE4D8C6FA8D01A41D" DOI="https://doi.org/10.2307/1565549" author="Ota, H" journalOrPublisher="Journal of Herpetology" pageId="0" pageNumber="77" pagination="106 - 118" refId="B30" refString="Ota, H, Miyaguni, H, Hikida, T, 1999. Geographic variation in the endemic skink, Ateuchosaurus pellopleurus from the Ryukyu Archipelago. Journal of Herpetology 33 (1): 106 - 118, DOI: https://doi.org/10.2307/1565549" title="Geographic variation in the endemic skink, Ateuchosaurus pellopleurus from the Ryukyu Archipelago." url="https://doi.org/10.2307/1565549" volume="33" year="1999">Ota et al. 1999</bibRefCitation>
).
</paragraph>
<paragraph id="27E7E13A69967AEB3C5E05EB9DE1D8B3" pageId="0" pageNumber="77">In the Tokashikijima population, males have smaller DMSs (mode = 57, 54-61) than females (mode = 60, 56-62).</paragraph>
<paragraph id="E0D28DE4F3312DFC832D2BCD93C0C429" pageId="0" pageNumber="77">
Mitochondrial DNA sequences differ among the four groups of Iheyajima and Izenajima Islands (site 14 and the adjacent island), the northern part of Okinawajima Island (site 15) and the adjacent islets (including site 16), the southern part of Okinawajima Island and the islands of the Kerama Group (site 19 and the adjacent islets), and Agunijima, Tonakijima, and Kumejima Islands (sites 20-22) (
<bibRefCitation id="9437C5209EC5C4D613DE6E2EB80AA113" DOI="https://doi.org/10.1016/j.jcz.2020.06.008" author="Makino, T" journalOrPublisher="Zoologischer Anzeiger" pageId="0" pageNumber="77" pagination="1 - 10" refId="B17" refString="Makino, T, Okamoto, T, Kurita, K, Nakano, T, Hikida, T, 2020. Origin and intraspecific diversification of the scincid lizard Ateuchosaurus pellopleurus with implications for historical island biogeography of the Central Ryukyus of Japan. Zoologischer Anzeiger 288: 1 - 10, DOI: https://doi.org/10.1016/j.jcz.2020.06.008" title="Origin and intraspecific diversification of the scincid lizard Ateuchosaurus pellopleurus with implications for historical island biogeography of the Central Ryukyus of Japan." url="https://doi.org/10.1016/j.jcz.2020.06.008" volume="288" year="2020">Makino et al. 2020</bibRefCitation>
). The populations in the northern and southern parts of Okinawajima Island have different
<emphasis id="74CF75F270028B1036A85452D4944DA4" italics="true" pageId="0" pageNumber="77">RAG1</emphasis>
allele compositions (
<bibRefCitation id="DAF7B36C6EA5B666FC161FA668FB4F8D" DOI="https://doi.org/10.1016/j.jcz.2020.06.008" author="Makino, T" journalOrPublisher="Zoologischer Anzeiger" pageId="0" pageNumber="77" pagination="1 - 10" refId="B17" refString="Makino, T, Okamoto, T, Kurita, K, Nakano, T, Hikida, T, 2020. Origin and intraspecific diversification of the scincid lizard Ateuchosaurus pellopleurus with implications for historical island biogeography of the Central Ryukyus of Japan. Zoologischer Anzeiger 288: 1 - 10, DOI: https://doi.org/10.1016/j.jcz.2020.06.008" title="Origin and intraspecific diversification of the scincid lizard Ateuchosaurus pellopleurus with implications for historical island biogeography of the Central Ryukyus of Japan." url="https://doi.org/10.1016/j.jcz.2020.06.008" volume="288" year="2020">Makino et al. 2020</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8C362C24A2B4523380EDDEEA4E637EE2" pageId="0" pageNumber="77" type="distribution">
<paragraph id="3278EA9F474314B967C1D8F40C150E7C" pageId="0" pageNumber="77">Distribution.</paragraph>
<paragraph id="B7B0A162EB6F73748744E1563B9D2FAB" pageId="0" pageNumber="77">
Okinawa Group (sites 14-22 and surrounding islets in Fig.
<figureCitation id="50E6BE70D94106CF064E7660BF66BE44" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map of the geographic range of Ateuchosaurus pellopleurus and A. okinavensis in the Ryukyu Archipelago, and the sampling sites of the specimens examined in this study. A. The geographic location of the Ryukyu Archipelago; B. Sampling sites in the northern and central parts of the Ryukyu Archipelago. Each solid line indicates the geographic range of the Ateuchosaurus species revealed by Makino et al. (2020). Site numbers correspond to the locality numbers in Tables 1 - 2 and Suppl. materials 1 - 3." figureDoi="10.3897/zse.99.95923.figure1" httpUri="https://binary.pensoft.net/fig/798122" pageId="0" pageNumber="77">1</figureCitation>
) of the Ryukyu Archipelago.
</paragraph>
</subSubSection>
<subSubSection id="A1FC635482AC30F861153D2C3AFE394E" pageId="0" pageNumber="77" type="natural history">
<paragraph id="89A32816E4DD06B01D318DC03083CBDE" pageId="0" pageNumber="77">Natural history.</paragraph>
<paragraph id="D34140B2CA7EAC434449466A62CD50EB" pageId="0" pageNumber="77">
<taxonomicName id="2D00E370C31B8F6CD0936914E7A248F4" baseAuthorityName="Thompson" baseAuthorityYear="1912" family="Scincidae" genus="Ateuchosaurus" kingdom="Animalia" lsidName="Ateuchosaurus okinavensis" order="Squamata" pageId="0" pageNumber="77" rank="species" species="okinavensis">
<emphasis id="42D135E495BCB41417960AECA0710C1F" italics="true" pageId="0" pageNumber="77">Ateuchosaurus okinavensis</emphasis>
</taxonomicName>
exhibits similar microhabitat use to
<taxonomicName id="5B06E686987803AA5810A637EB8DFA01" lsidName="A. pellopleurus" pageId="0" pageNumber="77" rank="species" species="pellopleurus">
<emphasis id="1ECBC1ADC4F1D46F433CEAFF8267ABAD" italics="true" pageId="0" pageNumber="77">A. pellopleurus</emphasis>
</taxonomicName>
. On Iheyajima, Okinawajima, Minnajima, Tokashikijima, Agunijima and Kumejima Islands (sites 14-16, 19, 20, and 22 in Fig.
<figureCitation id="8F18610BCA55760F6CACEBE8EFC27513" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map of the geographic range of Ateuchosaurus pellopleurus and A. okinavensis in the Ryukyu Archipelago, and the sampling sites of the specimens examined in this study. A. The geographic location of the Ryukyu Archipelago; B. Sampling sites in the northern and central parts of the Ryukyu Archipelago. Each solid line indicates the geographic range of the Ateuchosaurus species revealed by Makino et al. (2020). Site numbers correspond to the locality numbers in Tables 1 - 2 and Suppl. materials 1 - 3." figureDoi="10.3897/zse.99.95923.figure1" httpUri="https://binary.pensoft.net/fig/798122" pageId="0" pageNumber="77">1</figureCitation>
),
<taxonomicName id="4C1B56A913073190EB6E35F65739DD27" lsidName="A. okinavensis" pageId="0" pageNumber="77" rank="species" species="okinavensis">
<emphasis id="EA439D1139FCE9B8DF97F14138A2A8CF" italics="true" pageId="0" pageNumber="77">A. okinavensis</emphasis>
</taxonomicName>
is usually seen in the daytime (6:53-17:54, 62/72 specimens) in spring to autumn (T.M., personal observation); however, it is sometimes also seen at night (18:06-23:18, 10/72 specimens). This species is found in warm microhabitats (28.6
<normalizedToken id="D168403D8B7BE5A888573B15872C1841" originalValue="±">+/-</normalizedToken>
2.1 °C, 25.1-33.2 °C, 72 specimens), and is more frequently seen on sunny days (47/72 specimens) than on cloudy (15/72 specimens) or rainy days (10/72 specimens).
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>
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