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<document ID-CLB-Dataset="183006" ID-DOI="http://dx.doi.org/10.3897/zookeys.1128.89751" ID-GBIF-Dataset="122b5dc5-8722-4597-858b-767dca75e0d7" ID-PMC="PMC9836590" ID-Pensoft-Pub="1313-2970-1128-129" ID-Pensoft-UUID="9462C7DFB85D5DC780F645BC75150827" ID-PubMed="36762242" ID-ZooBank="7AC1EB83EE914109892AE5714A65EFB9" ModsDocID="1313-2970-1128-129" checkinTime="1668024534632" checkinUser="pensoft" docAuthor="Saxton, Natalie A., Marinov, Milen G. &amp; Bybee, Seth M." docDate="2022" docId="D8738B6ACDCB5BC9AC8E9FE4849ED810" docLanguage="en" docName="ZooKeys 1128: 129-169" docOrigin="ZooKeys 1128" docPubDate="2022-11-09" docSource="http://dx.doi.org/10.3897/zookeys.1128.89751" docTitle="Vanuatubasis santoensis Ober &amp; Staniczek 2009" docType="treatment" docVersion="5" id="9462C7DFB85D5DC780F645BC75150827" lastPageNumber="129" masterDocId="9462C7DFB85D5DC780F645BC75150827" masterDocTitle="Revision of Vanuatubasis Ober &amp; Staniczek, 2009 (Odonata, Coenagrionidae), with description of seven new species" masterLastPageNumber="169" masterPageNumber="129" pageNumber="129" updateTime="1732743181810" updateUser="ExternalLinkService">
<mods:mods id="756385EE8517B3330B9C7AC09C422006" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A24E4B3CE490D2FF62F2615A92AA2DE9">
<mods:title id="47BA2FD5A3537846D54F8214914A93E9">Revision of Vanuatubasis Ober &amp; Staniczek, 2009 (Odonata, Coenagrionidae), with description of seven new species</mods:title>
</mods:titleInfo>
<mods:name id="4E0C3142FF382BD95C31FB9A2CF3AE60" type="personal">
<mods:role id="91AB8E765805E1E0199763C6C9EF1C12">
<mods:roleTerm id="55F0EECE51DBD5C1AB467E41DCF0179F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="78E7E9B55737B836BE0781B9A0B294C2">Saxton, Natalie A.</mods:namePart>
<mods:nameIdentifier id="885B2B8B3D4E6832568526BB2D3A09AD" type="ORCID">https://orcid.org/0000-0001-5993-9782</mods:nameIdentifier>
<mods:affiliation id="B4D2BB78A3CB2862A58B36B1D7C1D9DE">Research and Collections Division, The Cleveland Museum of Natural History, Cleveland, OH, 44106, USA &amp; Department of Biology, Case Western Reserve University, Cleveland, OH, 44106, USA &amp; Department of Biology and Monte L. Bean Museum Brigham Young University, Provo, UT 84604, USA</mods:affiliation>
<mods:nameIdentifier id="C379D557072DE47265C9F4A2C4BFCD3F" type="email">nsaxton55@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="8C2D61EF6345A263B15A84A99FB5DB82" type="personal">
<mods:role id="450C5702B19FAD94F5B26A367E8CD8F1">
<mods:roleTerm id="D94C301185C5625E2CB550A710BB3F2E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CD413FC087BCC248CA1D94C05C88619E">Marinov, Milen G.</mods:namePart>
<mods:nameIdentifier id="BDC9D146931A2245DC6470FF7717DA7C" type="ORCID">https://orcid.org/0000-0003-3284-2555</mods:nameIdentifier>
<mods:affiliation id="5D8BF052CF975588B8E741F6D60FE2C3">Biosecurity Surveillance and Incursion Investigation Plant Health Team, Ministry for Primary Industries, 14 Sir William Pickering Drive, Christchurch 8053, New Zealand</mods:affiliation>
</mods:name>
<mods:name id="EEC0354C2E00B0C72AF6130814179475" type="personal">
<mods:role id="9C7822E552E9231385F387464371CBF6">
<mods:roleTerm id="C289794E1E816A932ACE3D4634637A70">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2798629FAF691CBF096F00226754C07B">Bybee, Seth M.</mods:namePart>
<mods:affiliation id="68FB832BF478D2FA66E57310BCAF158C">Department of Biology and Monte L. Bean Museum Brigham Young University, Provo, UT 84604, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource id="5C59C7E49FBD4D9DB961683456FD15DC">text</mods:typeOfResource>
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<mods:title id="2FE2D6AF375C14D7AEDDA68C6155676C">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="76575966016CF301842F1EC9DDAD0E3C">
<mods:date id="3109F018C7B24E0D0590D3E3FF32E8A6">2022</mods:date>
<mods:detail id="DC153CE43666EDB398009D1C6B4A0295" type="pubDate">
<mods:number id="706054E2825151DC87176D7AEE3C72E8">2022-11-09</mods:number>
</mods:detail>
<mods:detail id="CF57DDE774D8BB04A42AF268AAA20152" type="volume">
<mods:number id="AF214AF156D4CC39301C6DE1FFB4DE7D">1128</mods:number>
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<mods:start id="CE5F3AE27EB18D3C48B578B8575A2468">129</mods:start>
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<mods:location id="ACEDF6BF24BEA9061890F477D0E3E3EC">
<mods:url id="9C32C797A9295AB075CD6A0C1157F016">http://dx.doi.org/10.3897/zookeys.1128.89751</mods:url>
</mods:location>
<mods:classification id="77B45AEC2CD19EFD31D86DAFE3C690A2">journal article</mods:classification>
<mods:identifier id="B139FDBCC4DD607B1DED17A0C7C4EE07" type="DOI">http://dx.doi.org/10.3897/zookeys.1128.89751</mods:identifier>
<mods:identifier id="15A43E436C8A055B8BE488BF667D9625" type="Pensoft-Pub">1313-2970-1128-129</mods:identifier>
<mods:identifier id="8DE54C11FD41B719DF0C6E4D46C6B703" type="ZooBank">7AC1EB83EE914109892AE5714A65EFB9</mods:identifier>
<mods:identifier id="9815E666E640760FE6AE06FAD21CFE53" type="Pensoft-UUID">9462C7DFB85D5DC780F645BC75150827</mods:identifier>
</mods:mods>
<treatment id="D8738B6ACDCB5BC9AC8E9FE4849ED810" ID-GBIF-Taxon="204393636" LSID="urn:lsid:plazi:treatment:D8738B6ACDCB5BC9AC8E9FE4849ED810" httpUri="http://treatment.plazi.org/id/D8738B6ACDCB5BC9AC8E9FE4849ED810" lastPageNumber="129" pageId="0" pageNumber="129" scope_family="Coenagrionidae" scope_order="Odonata" scope_subOrder="Zygoptera">
<subSubSection id="3CD7CACF22859BEE5EB07E4BB49986CD" pageId="0" pageNumber="129" type="nomenclature">
<paragraph id="FE8582900045E453EB0C557F3D3631B5" pageId="0" pageNumber="129">
<taxonomicName id="85C31BD199D03E41B324039945C23636" ID-CoL="7FGNH" LSID="D8738B6A-CDCB-5BC9-AC8E-9FE4849ED810" authority="Ober &amp; Staniczek, 2009" authorityName="Ober &amp; Staniczek" authorityYear="2009" class="Insecta" family="Coenagrionidae" genus="Vanuatubasis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vanuatubasis santoensis" order="Odonata" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="santoensis">Vanuatubasis santoensis Ober &amp; Staniczek, 2009</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="E51EBC756BF977CD9AD21DDE142C1913" pageId="0" pageNumber="129" type="description">
<paragraph id="29193BF673828A5D1888FAC891FE2362" pageId="0" pageNumber="129">
<figureCitation id="207AC74A4EE93455B1DB03C49E8352FA" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Vanuatubasis santoensis (♂ BYU) A dorsal thorax B lateral terminalia C dorsal terminalia D lateral habitus V. santoensis (♀ BYU) E dorsal thorax F lateral terminalia. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1128.89751.figure10" httpUri="https://binary.pensoft.net/fig/767336" pageId="0" pageNumber="129">Figs 10</figureCitation>
<figureCitation id="AF472B3D3FAE48EB38EDA795D5B06180" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male dorso-posterior and dorsal terminalia of A V. bidens B V. evelynae C V. insularivorum D V. kapularum E V. malekulana F V. nunggoli G V. santoensis." figureDoi="10.3897/zookeys.1128.89751.figure20" httpUri="https://binary.pensoft.net/fig/767346" pageId="0" pageNumber="129">, 20G</figureCitation>
<figureCitation id="0AC2AF7FDFEEC20BD885419F697EFA12" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Female face and dorsal terminalia of A V. bidens B V. discontinua C V. evelynae D V. insularivorum E V. kapularum F V. malekulana G V. nunggoli H V. rhomboides I V. santoensis J V. xanthochroa." figureDoi="10.3897/zookeys.1128.89751.figure21" httpUri="https://binary.pensoft.net/fig/767347" pageId="0" pageNumber="129">, 21I</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BE6FEDA9CC50DA1DB941A1C1D96994FB" pageId="0" pageNumber="129" type="reference_group">
<paragraph id="14BC9F3B8D05B5FFF1A03E929A963E49" pageId="0" pageNumber="129">
<taxonomicName id="48AD8DEE6749FC80873C93797508FB88" authorityName="Ober &amp; Staniczek" authorityYear="2009" class="Insecta" family="Coenagrionidae" genus="Vanuatubasis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vanuatubasis santoensis" order="Odonata" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="santoensis">Vanuatubasis santoensis</taxonomicName>
Ober &amp; Staniczek (2009): 487-490.
</paragraph>
</subSubSection>
<subSubSection id="9DB671188E36A373299AE7B4B6D839F6" pageId="0" pageNumber="129" type="remarks">
<paragraph id="BEFFA64292C09BCD1E2E20A20BADC5ED" pageId="0" pageNumber="129">Remark.</paragraph>
<paragraph id="60D167BB5F6B8E002A159309F09F47FE" pageId="0" pageNumber="129">
<taxonomicName id="069F962453791702A897762AFF2CD6A1" authorityName="Ober &amp; Staniczek" authorityYear="2009" class="Insecta" family="Coenagrionidae" genus="Vanuatubasis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vanuatubasis santoensis" order="Odonata" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="santoensis">
<emphasis id="894910B4ED1E1F13D1ED89C462065F45" italics="true" pageId="0" pageNumber="129">Vanuatubasis santoensis</emphasis>
</taxonomicName>
was known by males only. Here we describe the female.
</paragraph>
</subSubSection>
<subSubSection id="ACD184D3ED55AFC715AE810E8B8005B7" pageId="0" pageNumber="129" type="materials_examined">
<paragraph id="E784F65330BA0B7D7C42E6A5EA41F640" pageId="0" pageNumber="129">Material examined.</paragraph>
<paragraph id="FB19188293B1765871F6CA15AAE2AB07" pageId="0" pageNumber="129">
<emphasis id="E885B8BBA9B76850EFA35CF2BF0E78ED" bold="true" pageId="0" pageNumber="129">
<emphasis id="7179F0982963DDB21952D86E762C151B" italics="true" pageId="0" pageNumber="129">
<typeStatus id="665043A1994CE9D26241A78FDBCCF8E7">Holotype</typeStatus>
</emphasis>
(
<specimenCount id="4BDF1F275BE1352EA5892767C6D8796E" type="male">1 ♂</specimenCount>
</emphasis>
<emphasis id="A2F8790E4ED2830CD7F62EF4B9BA4694" bold="true" pageId="0" pageNumber="129">SMNS).</emphasis>
&quot;Vanuatubasis santoensis
<specimenCount id="20FF4712D5D49BF45BA95AA17B85288B">♂</specimenCount>
<materialsCitation id="C1AF9FDCDD36D94C8CD09502C201A006" collectingDate="2006-11-13" collectingDateMax="2018-06-06" collectingDateMin="2006-11-13" collectorName="A. H. Staniczek, M. Pallmann, S. Bybee, G. Powell" country="VANUATU" county="Penaoru River" determinerName="S. V. Ober" elevation="90" latitude="-14.96105" location="Espiritu Santo" longLatPrecision="1" longitude="166.63316" municipality="Penaoru" specimenCount="28" specimenCount-female="9" specimenCount-male="19" stateProvince="Sanma Province" typeStatus="HOLOTYPE">
<typeStatus id="F555489977B9696D8A0F6ECAA84E71E6">HOLOTYPE</typeStatus>
|
<bibRefCitation id="5212D6FA211C63AD25C90D488512AE46" DOI="https://doi.org/10.5252/z2009n3a6" author="Ober, S" journalOrPublisher="Zoosystema" pageId="0" pageNumber="129" pagination="485 - 497" refId="B9" refString="Ober, S, Staniczek, A, 2009. A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485 - 497, DOI: https://doi.org/10.5252/z2009n3a6" title="A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu." url="https://doi.org/10.5252/z2009n3a6" volume="31" year="2009">Ober and Staniczek 2009</bibRefCitation>
| Vanuatu,
<collectingRegion id="2A2B9F984BE20D5BE3BBB3F25C0DADC9" country="Vanuatu" name="Sanma">Sanma Province</collectingRegion>
|
<location id="A4A51B84D109514573F29CF1772FE0A2" LSID="urn:lsid:plazi:treatment:D8738B6ACDCB5BC9AC8E9FE4849ED810:A4A51B84D109514573F29CF1772FE0A2" country="VANUATU" county="Penaoru River" latitude="-14.96105" longLatPrecision="1" longitude="166.63316" municipality="Penaoru" name="Espiritu Santo" stateProvince="Sanma Province">Espiritu Santo</location>
, surroundings of |
<collectingMunicipality id="5BF2FF846B6E657BBB7D277EECD504DE">Penaoru</collectingMunicipality>
,
<collectingCounty id="8BA570F75DE8EBAC7B79CE3C26FC7C91">Penaoru River</collectingCounty>
|leg.
<collectorName id="8087FB619105514DF9CA4C32469B0675">A.H. Staniczek</collectorName>
and
<collectorName id="5BA6FC6C1E2ACD9249C9907A857B9D42">M. Pallmann</collectorName>
,
<collectingDate id="89B271DCD443A119F1CB782224F44B67" value="2006-11-13">13.XI.2006</collectingDate>
| det.
<determinerName id="47A098C341A19CF69B6937BF53A83037">S.V. Ober</determinerName>
,
<collectingDate id="043F9119DD408A586635EC8470B29FD6" value="2008-07-21">21.VII.2008</collectingDate>
| coll.- |
<bibRefCitation id="01BF7227D81A7BCFC15102E227F0BE4D" DOI="https://doi.org/10.5252/z2009n3a6" author="Ober, S" journalOrPublisher="Zoosystema" pageId="0" pageNumber="129" pagination="485 - 497" refId="B9" refString="Ober, S, Staniczek, A, 2009. A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485 - 497, DOI: https://doi.org/10.5252/z2009n3a6" title="A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu." url="https://doi.org/10.5252/z2009n3a6" volume="31" year="2009">Ober and Staniczek 2009</bibRefCitation>
|
<geoCoordinate id="2AE029E49FA74E088DACB0D8656BA8D8" degrees="14.96105" direction="south" orientation="latitude" precision="1" value="-14.96105">14.96105°S</geoCoordinate>
,
<geoCoordinate id="DEFD263B27D9BD554BCBD003BF74E976" degrees="166.63316" direction="east" orientation="longitude" precision="1" value="166.63316">166.63316°E</geoCoordinate>
<quantity id="025D6151331545CB57D45154C8EFF2F2" metricMagnitude="1" metricUnit="m" metricValue="9.0" unit="m" value="90.0">
<elevation id="567C14105396C3D272E32C48E53DE0F6" metricMagnitude="1" metricUnit="m" metricValue="9.0" unit="m" value="90.0">90 m</elevation>
</quantity>
| ODO 000242 K.&quot;
<emphasis id="A4C7A307568C230C4C8EAE6A934A80A7" bold="true" pageId="0" pageNumber="129">
(
<specimenCount id="7CE1A3CC536071FB1204BC0E2DF4D916" type="male">4 ♂♂</specimenCount>
</emphasis>
<emphasis id="0D3FE1406BC6861C5F6149E4D7943768" bold="true" pageId="0" pageNumber="129">BYU)</emphasis>
&quot;
<collectingCountry id="2B0FCCDD0B50DCB221FD7FB536156C8C" name="Vanuatu">VANUATU</collectingCountry>
: Santo Is: | Wailapa, -15.5781 | 167.0024,
<collectingDate id="A82D9CF17EAA1B4C8E03C7DD686C654E" value="2018-06-06">6.vi.2018</collectingDate>
, coll, |
<collectorName id="75C5A510B55FF6E22160549E89788576">S. Bybee</collectorName>
and
<collectorName id="ED51DD08C72AED589EF9E190465F85BB">G. Powell</collectorName>
&quot;
<emphasis id="4A8DAF3D9EA5EDD2C3781707453BF161" bold="true" pageId="0" pageNumber="129">
(
<specimenCount id="E1EACA92BB6E0BDE140BE56B2071597B" type="male">1 ♂</specimenCount>
</emphasis>
<emphasis id="090C0572AFFF77022FCD6E5CD2046FA3" bold="true" pageId="0" pageNumber="129">BYU)</emphasis>
&quot;
<collectingCountry id="AD30CDF3C2CA8AE11B801E22E4E35434" name="Vanuatu">VANUATU</collectingCountry>
: Santo Is: | Narango, -15.6274 | 166.8535,
<collectingDate id="7CBA4DBA57F211E365B5AD5AA0A8A5D4" value="2018-06-04">4.vi.2018</collectingDate>
, | coll,
<collectorName id="ACF304F602B9ED99AE41065F33684740">S. Bybee</collectorName>
and
<collectorName id="0B6283350A44AE88D12F0A8F6B917A75">G. Powell</collectorName>
&quot;
<emphasis id="E1A6E0FAC4A3C4E551381B4FD2DAED46" bold="true" pageId="0" pageNumber="129">
(
<specimenCount id="48402BDB60BE85D097FFD6A0F71A9A46" type="male">2 ♂♂</specimenCount>
</emphasis>
<emphasis id="16D3F1FBDD86A4DB430F8E9AC591DE64" bold="true" pageId="0" pageNumber="129">BYU)</emphasis>
&quot;
<collectingCountry id="9B93AB7EA910BD23A3D8EA9E413E0A15" name="Vanuatu">VANUATU</collectingCountry>
: Santo Is: | Narango, -15.5538 | 166.9814,
<collectingDate id="368D7207EA7798B1B572B47F95FD8CA6" value="2018-06-04">4.vi.2018</collectingDate>
, | coll,
<collectorName id="8F4D42E795DDF4908F51AC733634A491">S. Bybee</collectorName>
and
<collectorName id="02D7A84A1764EE529E0832496D0030B6">G. Powell</collectorName>
&quot;
<emphasis id="4B4D21DE5B5D0BA9780EAA36718EE113" bold="true" pageId="0" pageNumber="129">
(
<specimenCount id="1C3C1DFA3ABF2678FFA68E8B7D9DC4BB" type="male">12 ♂♂</specimenCount>
</emphasis>
<specimenCount id="0F50456F7C9C6C775B23DF5B9B4683FE" type="female">
<emphasis id="579AB395C30C009419CD83B52ADC205F" bold="true" pageId="0" pageNumber="129">4</emphasis>
♀♀
</specimenCount>
<emphasis id="17EBFF7B66C87BE69D486D4115A7AE78" bold="true" pageId="0" pageNumber="129">BYU)</emphasis>
&quot;
<collectingCountry id="77CD7B11CC340846EE2010F43A3B0E95" name="Vanuatu">VANUATU</collectingCountry>
: Santo Is: | Ipayato, -15.6296| 166.8426,
<collectingDate id="D8C13C970CD148872D047649AC061C42" value="2018-06-04">4 vi.2018</collectingDate>
. | coll.
<collectorName id="51938FE41706AA6DBCD145C664E1970F">S. Bybee</collectorName>
and
<collectorName id="4D5E701582B18E81B179568550821C2D">G. Powell</collectorName>
&quot;
<emphasis id="07F3B3A08DD097C0CB65B098623A0890" bold="true" pageId="0" pageNumber="129">
(
<specimenCount id="0F103D62F4BCFD6986FEDFA6AF372C09" type="female">5 ♀♀</specimenCount>
</emphasis>
<emphasis id="C6B41085331DB358C47AAAB39BD00C27" bold="true" pageId="0" pageNumber="129">BYU)</emphasis>
&quot;
<collectingCountry id="378C365BAC847AD4044A765E309030D0" name="Vanuatu">VANUATU</collectingCountry>
: Santo Is: Felea |
<geoCoordinate id="BFDFBFA91FAA871AAC7D3DBB2C390381" degrees="15.3839" direction="south" orientation="latitude" precision="5" value="-15.3839">-15.3839</geoCoordinate>
<geoCoordinate id="15B4534C7775E63AB9C09919BC67803B" degrees="166.8426" direction="east" orientation="longitude" precision="5" value="166.8426">166.8426</geoCoordinate>
,
<collectingDate id="1BA5792900A1361EBDE8CF629F469C5E" value="2018-06-04">4.vi.2018</collectingDate>
, | coll,
<collectorName id="EE801EF47B2CF80C4E4BBAE99E8C4A74">S. Bybee</collectorName>
and
<collectorName id="A1847D664E56DAEE247C99690833AB1B">G. Powell</collectorName>
&quot;
</materialsCitation>
</paragraph>
<paragraph id="51374FC081A2901ED30744A67E6C4DFB" pageId="0" pageNumber="129">
<materialsCitation id="628D414A1BDA114FA1C5C9861B0849AA" collectingDate="2006-11-09" collectingDateMax="2006-11-11" collectingDateMin="2006-11-09" collectorName="A. H. Staniczek, M. Pallmann" country="Vanuatu" county="Espiritu Santo" determinerName="S. V. Ober" elevation="139" latitude="-15.2175" location="Pae River" longLatPrecision="5" longitude="166.68706" municipality="Tasmate" specimenCount="7" specimenCount-male="7" stateProvince="Sanma Province" typeStatus="Paratypes">
<emphasis id="6EA6F48665F703643C4C763297270C62" bold="true" pageId="0" pageNumber="129">
<emphasis id="0BFE4196BC1AB8F0F4DFFA1FD7D3C233" italics="true" pageId="0" pageNumber="129">
<typeStatus id="460AD59D568FF495D1599DA96FD079D0">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="456FB1AE3B2E92F1F89028E866AD2E54" type="male">2 ♂♂</specimenCount>
</emphasis>
<emphasis id="A7978A3524DFD220E6D9105F686D719E" bold="true" pageId="0" pageNumber="129">SMNS).</emphasis>
same label data as Holotype with the following barcode numbers: ODO 000246 K, ODO 000247 K.
<emphasis id="252DF96B1FD2EE37FB8A4F12B27910BD" bold="true" pageId="0" pageNumber="129">
(
<specimenCount id="EEA562FDFDE2BA741A6C1A8E463B2075" type="male">1 ♂</specimenCount>
</emphasis>
<emphasis id="F4C9A7E48E702A37870E5E8EE3BDBA22" bold="true" pageId="0" pageNumber="129">SMNS).</emphasis>
&quot;
<collectingCountry id="19759213D1A8E47D49869DC81B02D58A" name="Vanuatu">Vanuatu</collectingCountry>
,
<collectingRegion id="D69D589AA8E41177172AE7A24B75C769" country="Vanuatu" name="Sanma">Sanma Province</collectingRegion>
|
<collectingCounty id="358D490F03276117CDDE450906CE8AF0">Espiritu Santo</collectingCounty>
, surroundings of
<collectingMunicipality id="72CF17E49D60E2904819C2B847D1996E">Tasmate</collectingMunicipality>
,
<location id="833467CD654822EFF1A36CD7892DC3D8" LSID="urn:lsid:plazi:treatment:D8738B6ACDCB5BC9AC8E9FE4849ED810:833467CD654822EFF1A36CD7892DC3D8" country="Vanuatu" county="Espiritu Santo" latitude="-15.2175" longLatPrecision="5" longitude="166.68706" municipality="Tasmate" name="Pae River" stateProvince="Sanma Province">
<normalizedToken id="CFC24C98C5DEB382EDD801AEFAB29639" originalValue="Paé">Pae</normalizedToken>
River
</location>
,
<geoCoordinate id="757687A96B6F694244E3E8373B9733E5" degrees="15.2175" direction="south" orientation="latitude" precision="5" value="-15.2175">15.2175°S</geoCoordinate>
,
<geoCoordinate id="C680B6170419C50CC91264F59CDA1444" degrees="166.68706" direction="east" orientation="longitude" precision="1" value="166.68706">166.68706°E</geoCoordinate>
,
<quantity id="70D56DB846FA8A4247C294E8E3181E9A" metricMagnitude="2" metricUnit="m" metricValue="1.39" unit="m" value="139.0">
<elevation id="0C47FAB72858B346AAB26A5009A1A5FD" metricMagnitude="2" metricUnit="m" metricValue="1.39" unit="m" value="139.0">139 m</elevation>
</quantity>
,
<collectingDate id="000CE950D8321CF3D7975F00694C8899" value="2006-11-11">11.XI.2006</collectingDate>
, leg.
<collectorName id="08D4EA8EFD92F162A79D772FDC6CAD98">A.H. Staniczek</collectorName>
and
<collectorName id="D16B5ABEE9CB5BDFADB2669AA99AFA72">M. Pallmann</collectorName>
&quot;, with the following barcode number: ODO 000245 K.
<emphasis id="8CBDC2A9B01CBE82EFC2B5279009B423" bold="true" pageId="0" pageNumber="129">
(
<specimenCount id="2D6754C00F917551FA8B5404991E261F" type="male">2 ♂♂</specimenCount>
</emphasis>
<emphasis id="3D23F95109F08C51537E28B10FDE8F54" bold="true" pageId="0" pageNumber="129">
SNMS,
<specimenCount id="AA6ED419BCB7E8BDB455BB65FC0E854A" type="male">2 ♂♂</specimenCount>
</emphasis>
<emphasis id="E80033E5A510529E1F72864D758DC9AD" bold="true" pageId="0" pageNumber="129">MNHN).</emphasis>
&quot;
<collectingCountry id="2EE71A11A38E51D5DE04435E92FC625D" name="Vanuatu">Vanuatu</collectingCountry>
,
<collectingRegion id="5401403A043F39438F0A3E3EB7C3C049" country="Vanuatu" name="Sanma">Sanma Province</collectingRegion>
, Espiritu Santo, surroundings of Tasmate,
<location id="740EE93AF4D4F70F53B7D30E5BF5B098" LSID="urn:lsid:plazi:treatment:D8738B6ACDCB5BC9AC8E9FE4849ED810:740EE93AF4D4F70F53B7D30E5BF5B098" country="Vanuatu" county="Espiritu Santo" latitude="-15.2175" longLatPrecision="5" longitude="166.68706" municipality="Tasmate" name="Mamasa River" stateProvince="Sanma Province">Mamasa River</location>
,
<geoCoordinate id="B95568C5AE551E48369F4501D577E8BB" degrees="15.20976" direction="south" orientation="latitude" precision="1" value="-15.20976">15.20976°S</geoCoordinate>
,
<geoCoordinate id="34162FA444D820FB3FE26F87AE198793" degrees="166.67705" direction="east" orientation="longitude" precision="1" value="166.67705">166.67705°E</geoCoordinate>
,
<quantity id="2FF6A838EAC5E0861ADD9B99DB1AF63F" metricMagnitude="1" metricUnit="m" metricValue="2.0" unit="m" value="20.0">
<elevation id="21232D48A29E711373FE741C7CB1627A" metricMagnitude="1" metricUnit="m" metricValue="2.0" unit="m" value="20.0">20 m</elevation>
</quantity>
,
<collectingDate id="227061847F7001A4DA39556D60C865D4" value="2006-11-09">9.XI.2006</collectingDate>
, leg.
<collectorName id="0C0FB070DA2ADE071763909FE83B2FFD">A. H. Staniczek</collectorName>
and
<collectorName id="AC6287CC533E33EC2EE6D4A01584B811">M. Pallmann</collectorName>
&quot;, SNMS specimens with the following barcodes: ODO 000243 K, ODO 000244 K
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="83B5FEE0E9DD4B4A7417F994C26D8D9E" pageId="0" pageNumber="129" type="description">
<paragraph id="98D34D4AE79243D8E8FF3AB29C88137D" pageId="0" pageNumber="129">Description of the female.</paragraph>
<paragraph id="A2A98CAC3C534209C1C3293519B8AD28" pageId="0" pageNumber="129">
<emphasis id="C59CDAA4793DCD0FFE4D4CA3FF7EBEE3" bold="true" italics="true" pageId="0" pageNumber="129">Head</emphasis>
: Labium overall pale beige; labrum pale green with brown latero-posterior edges and small, slightly indented medially brown spot at posterior edge; anteclypeus, genae, and mandibles (expect for reddish tips) pale green; postclypeus translucent brown with green lateral edges; frons pale green, turning black posteriorly; scapes, pedicels, and flagella pale brown; vertex and rear of head black with line of setae and a bronze shimmer; three pale ocelli with small beige patch apical of the median ocellus; two white post-ocular spots; eyes pale green.
</paragraph>
<paragraph id="140ECCF5094FA0628A6816B02E673A8F" pageId="0" pageNumber="129">
<emphasis id="55CD9C60239D4D11A9EA40DC875048BC" bold="true" italics="true" pageId="0" pageNumber="129">Thorax</emphasis>
: Prothorax dorsally black; laterally green; pronotum black medially with brown edges, latero-posterior corners rounded to acute angles and weakly explanate, mid-line obviously indented across the pronotum, hind lobe with raised ridge that is shorter than the width of the pronotum, is curved outward medially, and extends to a sharp, rounded point that protrudes posteriorly; mesostigmal plate black with green lateral edges and not significantly raised. Pterothorax with black carina; laterally mesepisternum with black stripe reaching the dorsal carina and extending more than 3/4 of the mesepisternum, pale green on latter 1/4 of mesepisternum; mesepimeron overall pale green with hints of yellow; metepisternum overall green with short, brown line located on metapleural suture near the base of the wings, extending ~ 1/6 the
<normalizedToken id="81795E54974E0E4BB35A151886D2C4DA" originalValue="suture’s">suture's</normalizedToken>
length; mesinfraepisternum pale green with apical brown spot; metepimeron green, with beige anterior portion; coxae and trochanters beige, femora dorsally dark brown and ventrally beige with black spines; tibiae pale brown with slightly darker, and smaller, spines than that of the femora; tarsi beige with dark brown edges and small, dense spines; pale brown tarsal claws that darken apically to reddish tips, claws with a small tooth located on the basal 1/4 of their length.
</paragraph>
<caption id="AD6F2F855BB923878623CF1E4C8A5CFC" doi="10.3897/zookeys.1128.89751.figure10" httpUri="https://binary.pensoft.net/fig/767336" pageId="0" pageNumber="129" start="Figure 10" startId="F10">
<paragraph id="F4B7E2CF16AA7FE7A1277184990564B8" pageId="0" pageNumber="129">
<emphasis id="B4127839F8787C8EBB7FC05182BA0F50" bold="true" pageId="0" pageNumber="129">Figure 10.</emphasis>
<taxonomicName id="05086B75395E2DF40555174EA31B6E95" authorityName="Ober &amp; Staniczek" authorityYear="2009" class="Insecta" family="Coenagrionidae" genus="Vanuatubasis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vanuatubasis santoensis" order="Odonata" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="santoensis">
<emphasis id="C3C16BE86064F80058505DA34D4DC538" italics="true" pageId="0" pageNumber="129">Vanuatubasis santoensis</emphasis>
</taxonomicName>
(♂ BYU)
<emphasis id="72A18651761177C6F7EA197BF2C91993" bold="true" pageId="0" pageNumber="129">A</emphasis>
dorsal thorax
<emphasis id="8BA25DCC3E679D0EADF592A19C83CB59" bold="true" pageId="0" pageNumber="129">B</emphasis>
lateral terminalia
<emphasis id="D22335BC21FD3C307C80A0C2155B6CFC" bold="true" pageId="0" pageNumber="129">C</emphasis>
dorsal terminalia
<emphasis id="ED5499AA4E68632EB71A53383A84222E" bold="true" pageId="0" pageNumber="129">D</emphasis>
lateral habitus
<taxonomicName id="1AA603090449DBD189ADA09338CFCE43" lsidName="V. santoensis" pageId="0" pageNumber="129" rank="species" species="santoensis">
<emphasis id="3F4DEB50D296C6F4D6652833F572E91B" italics="true" pageId="0" pageNumber="129">V. santoensis</emphasis>
</taxonomicName>
(♀ BYU)
<emphasis id="E4478EE2C92882B1E56FB31E9F2C0523" bold="true" pageId="0" pageNumber="129">E</emphasis>
dorsal thorax
<emphasis id="E0012F71CAF52C585CF274E45998B141" bold="true" pageId="0" pageNumber="129">F</emphasis>
lateral terminalia. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="B0D8800BF858B02328F2B6B4F8467BFC" pageId="0" pageNumber="129">
<emphasis id="C2D54A40CC7EFEA52C8176141646C27B" bold="true" italics="true" pageId="0" pageNumber="129">Wings</emphasis>
: Hyaline with dark brown; pterostigma dark brown and rhombus-shaped, darkest on the edges; CuP halfway between antenodals in both wings; arculus just distal of second antenodal crossvein in both wings; discoidal cells unequal with FW dorsal edge being 1/2 as long as HW. Nodal index: 13/2-2/12 in FW and 12/2-2/11 in HW.
</paragraph>
<paragraph id="74E201E4BD8FF7D1D694A82D3269E81E" pageId="0" pageNumber="129">
<emphasis id="E01B59760F8152375455225E92C88B68" bold="true" italics="true" pageId="0" pageNumber="129">Abdomen</emphasis>
: Overall beige yellow with black dorsal stripe from S1-8 and lightening towards the edges, with pale brown setae; S1 and S2 green laterally; S3-S8 yellow laterally; S9 with pale dorsal stripe; S10 pale brown; ovipositor overall beige, with darker serrated ventral edge; stylus with rounded edges, pale brown and lightening apically; gonapophysis reddish brown with slightly serrated dorsal edge. Cerci roughly triangular, dark brown, and narrowing to a rounded apex, dorsal edge straight a ventral edge slightly rounded ventrally.
</paragraph>
<paragraph id="8FD0B7848E625FDF115D12B0D81A8012" pageId="0" pageNumber="129">
<emphasis id="4E2430668531971D55588305059698C7" bold="true" pageId="0" pageNumber="129">
<emphasis id="74882240054EC05E05E547F4B93C9A78" italics="true" pageId="0" pageNumber="129">Measurements</emphasis>
(mm)
</emphasis>
: total length 36-39 mm, abdomen 29-33 mm, HW 22-24 mm (
<emphasis id="274F5E971C21AA3D3AFB472100E62F55" italics="true" pageId="0" pageNumber="129">n</emphasis>
= 9).
</paragraph>
</subSubSection>
<subSubSection id="8F9DEF39F70C021A600C8E8A1CAD179B" pageId="0" pageNumber="129" type="diagnosis">
<paragraph id="4DC761AC64A67065DE812630FA25B87F" pageId="0" pageNumber="129">Diagnosis.</paragraph>
<paragraph id="6FAE14BE1290061C9B7B6F5E1E5E149C" pageId="0" pageNumber="129">
<emphasis id="8DC0BA0836C64B5D24B556095BC36151" bold="true" pageId="0" pageNumber="129">Male.</emphasis>
<taxonomicName id="F55D13A35D37E9B8C5690FC28AB04E19" authorityName="Ober &amp; Staniczek" authorityYear="2009" class="Insecta" family="Coenagrionidae" genus="Vanuatubasis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vanuatubasis santoensis" order="Odonata" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="santoensis">
<emphasis id="9597EC6271CEA02D5085FBD47351A1DD" italics="true" pageId="0" pageNumber="129">Vanuatubasis santoensis</emphasis>
</taxonomicName>
can be distinguished from other
<taxonomicName id="B9818EFADC61B0A284127D530D415935" authorityName="Ober &amp; Staniczek" authorityYear="2009" class="Insecta" family="Coenagrionidae" genus="Vanuatubasis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vanuatubasis" order="Odonata" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis id="92A6D2621182C6D64BD34CE8484B2500" italics="true" pageId="0" pageNumber="129">Vanuatubasis</emphasis>
</taxonomicName>
by its black pterostigma, pale blue postclypeus without any bars or maculations, dorso-posterior corner of mesostigmal plate not raised in an auricle (or if elevated, only slightly so).
<emphasis id="1EF6E4BF787B2B1387E9A70884D75D2E" bold="true" pageId="0" pageNumber="129">Female.</emphasis>
<taxonomicName id="F64467AE72A2D6C644B22B1DD43DDE78" authorityName="Ober &amp; Staniczek" authorityYear="2009" class="Insecta" family="Coenagrionidae" genus="Vanuatubasis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vanuatubasis santoensis" order="Odonata" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="santoensis">
<emphasis id="6BF457913E7440D0C93B1F48B6A6D2AD" italics="true" pageId="0" pageNumber="129">Vanuatubasis santoensis</emphasis>
</taxonomicName>
can be distinguished from other females in this genus, by its dark brown to black pterostigma, uniformly colored postclypeus (sometimes with pale spots), frons with a dark triangle extending from the base of the median ocelli, and the dark area on the mesepisternum reaching the black spot at the posterior end of the mesopleural suture.
</paragraph>
</subSubSection>
<subSubSection id="DC23938107E79729D093E4842FD2F404" pageId="0" pageNumber="129" type="variation">
<paragraph id="04A20A931332437A9C4DF030683D98C5" pageId="0" pageNumber="129">Variation.</paragraph>
<paragraph id="0F745DE0DFCA48D2EB6B3F9239073F58" pageId="0" pageNumber="129">
<emphasis id="D1AB6B39F642C236E5C50DD879D594F2" bold="true" pageId="0" pageNumber="129">Male.</emphasis>
Postclypeal maculation more prominent than in some individuals. Pterothorax sometimes with variable short black stripes.
<emphasis id="D57ABA4B5E92C6990D90C8269C4B5A99" bold="true" pageId="0" pageNumber="129">Female.</emphasis>
Thoracic coloring varies from yellow to green, presumably due to the age of the individual.
</paragraph>
</subSubSection>
<subSubSection id="F0F3D7DE5CCBFA62F643B7975AF6CF7A" pageId="0" pageNumber="129" type="distribution">
<paragraph id="1E18C875AA04FCBDFAAB5A9BE91B2FC6" pageId="0" pageNumber="129">Distribution.</paragraph>
<paragraph id="7C06252D2579C985630223311D97D493" pageId="0" pageNumber="129">Espiritu Santo, Vanuatu.</paragraph>
</subSubSection>
</treatment>
</document>
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