253 lines
20 KiB
XML
253 lines
20 KiB
XML
<document ID-DOI="10.11646/phytotaxa.594.1.2" ID-ISSN="1179-3163" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1682329257714" checkinUser="plazi" docAuthor="Lepší, Martin, Lepší, Petr, Hohla, Michael & Király, Gergely" docDate="2023" docId="D8448793FFA7FFD7FF450B98FE91FE0C" docLanguage="en" docName="phytotaxa.594.1.2.pdf" docOrigin="Phytotaxa 594 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.594.1.2" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.5:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="5" docTitle="Rubus (ser. Radula) Focke 1877" docType="key" docVersion="2" lastPageNumber="51" masterDocId="247DFFEBFFA0FFDFFFCD0F2AFF95FFD8" masterDocTitle="Rubus lentianus, a new species from Austria and Germany-another piece of the puzzle from a European hotspot of Rubus series Radula" masterLastPageNumber="52" masterPageNumber="43" pageNumber="50" updateTime="1682534635066" updateUser="jonas" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Rubus lentianus, a new species from Austria and Germany-another piece of the puzzle from a European hotspot of Rubus series Radula</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lepší, Martin</mods:namePart>
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<mods:affiliation>South Bohemian Museum in České Budějovice, Dukelská 1, CZ- 370 51 České Budějovice, Czech Republic. & Department of Botany, Faculty of Science, University of South Bohemia, Branišovská 31, CZ- 370 05 České Budějovice, Czech Republic.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lepší, Petr</mods:namePart>
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<mods:affiliation>Nature Conservation Agency of the Czech Republic, Administration of the Blanský les Protected Landscape Area, Vyšný 59, CZ- 381 01 Český Krumlov, Czech Republic.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hohla, Michael</mods:namePart>
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<mods:affiliation>Therese-Riggle-Strasse 16, A- 4982 Obernberg am Inn, Austria.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Király, Gergely</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0002-8439-2616</mods:nameIdentifier>
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<mods:affiliation>University of Sopron, Institute of Natural Resources and Forest Management, H- 9400 Sopron, Bajcsy-Zs. u. 4., Hungary. kiraly. gergely @ uni-sopron. hu; https: // orcid. org / 0000 - 0002 - 8439 - 2616</mods:affiliation>
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<mods:nameIdentifier type="email">kiraly.gergely@uni-sopron.hu</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Phytotaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-04-24</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>594</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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||
</mods:detail>
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<mods:extent unit="page">
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<mods:start>43</mods:start>
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<mods:end>52</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/phytotaxa.594.1.2</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/phytotaxa.594.1.2</mods:identifier>
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<mods:identifier type="ISSN">1179-3163</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:D8448793FFA7FFD7FF450B98FE91FE0C" httpUri="http://treatment.plazi.org/id/D8448793FFA7FFD7FF450B98FE91FE0C" lastPageId="8" lastPageNumber="51" pageId="7" pageNumber="50">
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<subSubSection box="[136,883,1202,1229]" pageId="7" pageNumber="50" type="nomenclature">
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<paragraph blockId="7.[136,883,1202,1229]" box="[136,883,1202,1229]" pageId="7" pageNumber="50">
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<heading bold="true" box="[136,883,1202,1229]" fontSize="11" level="1" pageId="7" pageNumber="50" reason="1">
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<emphasis bold="true" box="[136,883,1202,1229]" pageId="7" pageNumber="50">
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Key to the identification of
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<taxonomicName authority="(Hook. & Arn.) Focke (1877: 317)" authorityName="Focke" authorityPageNumber="317" authorityYear="1877" baseAuthorityName="Hook. & Arn." box="[462,883,1202,1229]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="50" phylum="Tracheophyta" rank="series" series="Radula">
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<emphasis bold="true" box="[462,651,1202,1229]" italics="true" pageId="7" pageNumber="50">Rubus lentianus</emphasis>
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and similar species
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="8" lastPageNumber="51" pageId="7" pageNumber="50" type="key">
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<keyStep pageId="7" pageNumber="50">
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<paragraph blockId="7.[136,1452,1271,2070]" box="[136,1451,1271,1293]" pageId="7" pageNumber="50">
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<keyLead box="[136,1451,1271,1293]" pageId="7" pageNumber="50">1. Indentation on terminal leaflet of primocane leaves regular, crenate-serrate....................................................................................2.</keyLead>
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</paragraph>
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<paragraph blockId="7.[136,1452,1271,2070]" box="[136,1451,1299,1321]" pageId="7" pageNumber="50">
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<keyLead box="[136,1451,1299,1321]" pageId="7" pageNumber="50">- Indentation on terminal leaflet of primocane leaves periodical, serrate............................................................................................3.</keyLead>
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</paragraph>
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</keyStep>
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<keyStep pageId="7" pageNumber="50">
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<paragraph blockId="7.[136,1452,1271,2070]" pageId="7" pageNumber="50">
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||
<keyLead pageId="7" pageNumber="50">
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||
2. Prickles 3–10 per
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<quantity box="[385,432,1328,1350]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="50" unit="cm" value="5.0">5 cm</quantity>
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of primocane length, 3–4(–5) mm long; apex of terminal leaflet of primocane leaves 5(–8) mm long ........ ..................................................................................................................................................................................
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<taxonomicName authorityName="Focke" authorityYear="1877" box="[1288,1450,1357,1379]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="50" phylum="Tracheophyta" rank="species" species="salisburgensis">
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<emphasis box="[1288,1450,1357,1379]" italics="true" pageId="7" pageNumber="50">R. salisburgensis</emphasis>
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</taxonomicName>
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</keyLead>
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||
</paragraph>
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<paragraph blockId="7.[136,1452,1271,2070]" pageId="7" pageNumber="50">
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<keyLead pageId="7" pageNumber="50">
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||
- Prickles (19–)24–29(–34) per
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<quantity box="[503,550,1386,1408]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="50" unit="cm" value="5.0">5 cm</quantity>
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||
of primocane length, (6.5–)7–8(–8.5) mm long; apex of terminal leaflet of primocane leaves (12–)15–17(–21) mm long ............................................................................................................................................
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||
<taxonomicName box="[1312,1451,1415,1436]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="50" phylum="Tracheophyta" rank="species" species="perpungens">
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<emphasis box="[1312,1451,1415,1436]" italics="true" pageId="7" pageNumber="50">R. perpungens</emphasis>
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</taxonomicName>
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</keyLead>
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||
</paragraph>
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||
</keyStep>
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<keyStep pageId="7" pageNumber="50">
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<paragraph blockId="7.[136,1452,1271,2070]" pageId="7" pageNumber="50">
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<keyLead pageId="7" pageNumber="50">
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||
3. Terminal leaflet of primocane leaves with incisions (2.5–)4.0–5.0(–6.0) mm deep, apex of terminal leaflet (16–)21–23(–26) mm long....................................................................................................................................................................................
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<taxonomicName authorityName="Hohla, Kiraly, M. Lepsi & P. Lepsi" authorityYear="2023" box="[1336,1451,1472,1494]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="50" phylum="Tracheophyta" rank="species" species="lentianus">
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<emphasis box="[1336,1451,1472,1494]" italics="true" pageId="7" pageNumber="50">R. lentianus</emphasis>
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</taxonomicName>
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||
</keyLead>
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||
</paragraph>
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||
<paragraph blockId="7.[136,1452,1271,2070]" box="[136,1451,1501,1523]" pageId="7" pageNumber="50">
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||
<keyLead box="[136,1451,1501,1523]" pageId="7" pageNumber="50">
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||
- Terminal leaflet of primocane leaves with incisions
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||
<quantity box="[708,787,1501,1522]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="7" pageNumber="50" unit="mm" value="2.5" valueMax="4.0" valueMin="1.0">1–4 mm</quantity>
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deep, apex of terminal leaflet 10–20(–25) mm long ........................4.
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</keyLead>
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||
</paragraph>
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||
</keyStep>
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<keyStep pageId="7" pageNumber="50">
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<paragraph blockId="7.[136,1452,1271,2070]" pageId="7" pageNumber="50">
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<keyLead pageId="7" pageNumber="50">
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||
4. Primocane with 150–280 hairs, 40–120 stalked glands per
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<quantity box="[781,828,1530,1551]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="50" unit="cm" value="1.0">1 cm</quantity>
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||
of stem side and very slender prickles, colour of leaves beneath noticeably variable along the length of the primocane, green at the base and white-greyish at the top ...............
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||
<taxonomicName box="[1264,1451,1559,1580]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="50" phylum="Tracheophyta" rank="species" species="silvae-bavaricae">
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||
<emphasis box="[1264,1451,1559,1580]" italics="true" pageId="7" pageNumber="50">R. silvae-bavaricae</emphasis>
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||
</taxonomicName>
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||
</keyLead>
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||
</paragraph>
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||
<paragraph blockId="7.[136,1452,1271,2070]" pageId="7" pageNumber="50">
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||
<keyLead pageId="7" pageNumber="50">
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||
- Primocane with fewer hairs (8–250) or fewer stalked glands (1–140) per
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||
<quantity box="[926,973,1588,1609]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="50" unit="cm" value="1.0">1 cm</quantity>
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||
of stem side and with thicker prickles, colour of leaves beneath less variable along the length of the primocane........................................................................................................5.
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||
</keyLead>
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||
</paragraph>
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||
</keyStep>
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||
<keyStep pageId="7" pageNumber="50">
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||
<paragraph blockId="7.[136,1452,1271,2070]" pageId="7" pageNumber="50">
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||
<keyLead pageId="7" pageNumber="50">5. Primocanes with transitions between stalked glands and prickles, i.e. unequal stalked glands, acicles, bristles and sometimes also pricklets present; leaves distinctly hairy to the touch beneath and hairy or glabrous above.............................................................6.</keyLead>
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||
</paragraph>
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||
<paragraph blockId="7.[136,1452,1271,2070]" pageId="7" pageNumber="50">
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||
<keyLead pageId="7" pageNumber="50">
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||
- Primocanes usually without transitions between stalked glands and prickles, stalked glands usually short; leaves usually not distinctly hairy to the touch beneath (except for
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<taxonomicName authorityName="Danner" authorityYear="2003" box="[672,796,1731,1753]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="50" phylum="Tracheophyta" rank="species" species="muhelicus">
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<emphasis box="[672,796,1731,1753]" italics="true" pageId="7" pageNumber="50">R. muhelicus</emphasis>
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</taxonomicName>
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||
, see 9) and glabrous above ..............................................................8.
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||
</keyLead>
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||
</paragraph>
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||
</keyStep>
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<keyStep pageId="7" pageNumber="50">
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||
<paragraph blockId="7.[136,1452,1271,2070]" pageId="7" pageNumber="50">
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<keyLead pageId="7" pageNumber="50">
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||
6. Primocanes sharply angled with (30–)40–100(–250) hairs per
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<quantity box="[806,853,1760,1782]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="50" unit="cm" value="1.0">1 cm</quantity>
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||
of stem side and with prickles ± patent, straight; pedicels with stalked glands longer than the longest simple hairs; leaves usually with scattered hairs above ...........................
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<taxonomicName box="[1267,1450,1790,1811]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="50" phylum="Tracheophyta" rank="species" species="jarae-cimrmanii">
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<emphasis box="[1267,1450,1790,1811]" italics="true" pageId="7" pageNumber="50">R. jarae-cimrmanii</emphasis>
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</taxonomicName>
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</keyLead>
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||
</paragraph>
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<paragraph blockId="7.[136,1452,1271,2070]" pageId="7" pageNumber="50">
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<keyLead pageId="7" pageNumber="50">
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||
- Primocanes angled or bluntly angled with 3–20 hairs per
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<quantity box="[778,825,1818,1839]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="50" unit="cm" value="1.0">1 cm</quantity>
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||
of stem side and with prickles declining, often slightly curved; pedicels with stalked glands shorter than the longest simple hairs; leaves glabrous or almost glabrous above ...............................7.
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</keyLead>
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||
</paragraph>
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||
</keyStep>
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<keyStep pageId="7" pageNumber="50">
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<paragraph blockId="7.[136,1452,1271,2070]" pageId="7" pageNumber="50">
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||
<keyLead pageId="7" pageNumber="50">
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||
7. Primocane leaves (very) distinctly pedate, with an ovate or obovate terminal leaflet abruptly narrowing into a (15–)20(–25) mm long apex, with a slightly cordate (rarely rounded) base and with distinctly periodical indentation; petals pale pink........................ .......................................................................................................................................................................................
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<taxonomicName authorityName="Zila & Weber" authorityYear="2005" box="[1317,1451,1933,1955]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="50" phylum="Tracheophyta" rank="species" species="perpedatus">
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<emphasis box="[1317,1451,1933,1955]" italics="true" pageId="7" pageNumber="50">R. perpedatus</emphasis>
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</taxonomicName>
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||
</keyLead>
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||
</paragraph>
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||
<paragraph blockId="7.[136,1452,1271,2070]" pageId="7" pageNumber="50">
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<keyLead pageId="7" pageNumber="50">
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||
- Primocane leaves not distinctly pedate, with a broadly ovate to broadly obovate terminal leaflet abruptly narrowing into a 10–15(– 20) mm long apex, with a distinctly cordate base and with slightly periodical indentation; petals white ........................
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<taxonomicName authorityName="Focke" authorityYear="1877" box="[1339,1450,1991,2012]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="50" phylum="Tracheophyta" rank="species" species="epipsilos">
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<emphasis box="[1339,1450,1991,2012]" italics="true" pageId="7" pageNumber="50">R. epipsilos</emphasis>
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||
</taxonomicName>
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||
</keyLead>
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||
</paragraph>
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||
</keyStep>
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||
<keyStep lastPageId="8" lastPageNumber="51" pageId="7" pageNumber="50">
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||
<paragraph blockId="7.[136,1452,1271,2070]" pageId="7" pageNumber="50">
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||
<keyLead pageId="7" pageNumber="50">
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||
8. Primocane leaves 3–4(–5)-foliolate, terminal leaflet oblong obovate to broadly obovate; primocanes bluntly angled; carpels densely hairy; petals white to slightly pinkish ................................................................................................................
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||
<taxonomicName box="[1324,1451,2049,2070]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="50" phylum="Tracheophyta" rank="species" species="vatavensis">
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||
<emphasis box="[1324,1451,2049,2070]" italics="true" pageId="7" pageNumber="50">R. vatavensis</emphasis>
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||
</taxonomicName>
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||
</keyLead>
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||
</paragraph>
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||
<paragraph blockId="8.[136,1452,159,468]" pageId="8" pageNumber="51">
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||
<keyLead pageId="8" pageNumber="51">- Primocane leaves (3–)5-foliolate, terminal leaflet of a different shape; primocanes angled to sharply angled; carpels glabrous or with few hairs; petals pale pink to pink.............................................................................................................................................9.</keyLead>
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||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="8" pageNumber="51">
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||
<paragraph blockId="8.[136,1452,159,468]" pageId="8" pageNumber="51">
|
||
<keyLead pageId="8" pageNumber="51">
|
||
9. Primocane leaves ± palmate, terminal leaflet broadly ovate to round with a distinctly undulating margin; prickles (9–)11–20(–22) per
|
||
<quantity box="[247,294,245,267]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="51" unit="cm" value="5.0">5 cm</quantity>
|
||
of primocane length..........................................................................................................................................
|
||
<taxonomicName authorityName="Danner" authorityYear="2003" box="[1327,1450,245,267]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="51" phylum="Tracheophyta" rank="species" species="muhelicus">
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||
<emphasis box="[1327,1450,245,267]" italics="true" pageId="8" pageNumber="51">R. muhelicus</emphasis>
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||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
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||
<paragraph blockId="8.[136,1452,159,468]" pageId="8" pageNumber="51">
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||
<keyLead pageId="8" pageNumber="51">
|
||
- Primocane leaves distinctly pedate, terminal leaflet elliptical to broadly elliptical, ovate or rhombic with a rather flat margin; prickles 5–12(–16) per
|
||
<quantity box="[435,482,303,325]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="51" unit="cm" value="5.0">5 cm</quantity>
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||
of primocane length...........................................................................................................................10.
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||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="8" pageNumber="51">
|
||
<paragraph blockId="8.[136,1452,159,468]" pageId="8" pageNumber="51">
|
||
<keyLead pageId="8" pageNumber="51">
|
||
10. Pedicels with stalked glands shorter than long simple hairs; primocanes sparsely to intermediately densely hairy, with slightly declining prickles; prickles on inflorescence rachis
|
||
<quantity box="[704,783,361,382]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="8" pageNumber="51" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">7–8 mm</quantity>
|
||
long; inflorescence conical, its branches rather thick ...........
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||
<taxonomicName box="[1360,1451,361,382]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="51" phylum="Tracheophyta" rank="species" species="radula">
|
||
<emphasis box="[1360,1451,361,382]" italics="true" pageId="8" pageNumber="51">R. radula</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="8.[136,1452,159,468]" pageId="8" pageNumber="51">
|
||
<keyLead pageId="8" pageNumber="51">
|
||
- Pedicels with stalked glands as long as short simple hairs; primocanes glabrous or with very few hairs, with distinctly declining prickles; prickles on the inflorescence rachis
|
||
<quantity box="[644,723,418,440]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="8" pageNumber="51" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
|
||
long; inflorescence broadly conical, diffuse, with rather thin branches ....
|
||
<taxonomicName authorityName="Weihe & Nees" authorityYear="1825" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="51" phylum="Tracheophyta" rank="species" species="rudis">
|
||
<emphasis italics="true" pageId="8" pageNumber="51">R. rudis</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |