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<document ID-CLB-Dataset="191986" ID-DOI="http://dx.doi.org/10.3897/zookeys.1130.91467" ID-GBIF-Dataset="4793f979-7563-478f-9c51-0bcac41ad564" ID-PMC="PMC9836665" ID-Pensoft-Pub="1313-2970-1130-167" ID-Pensoft-UUID="DCA2FBA40D0051C18F31827BCF7D25C2" ID-PubMed="36761017" ID-ZooBank="991C76184FD04191A4119F6AB775D3AF" ModsDocID="1313-2970-1130-167" checkinTime="1669095659592" checkinUser="pensoft" docAuthor="Wang, Shuqiao, Lu, Ying, Li, Ya, Li, Shuqiang &amp; Lin, Yucheng" docDate="2022" docId="D65CF3D42C895E54AE2B17713D135C49" docLanguage="en" docName="ZooKeys 1130: 167-189" docOrigin="ZooKeys 1130" docPubDate="2022-11-21" docSource="http://dx.doi.org/10.3897/zookeys.1130.91467" docTitle="Anapistula panensis Lin, Tao &amp; S. Li 2013" docType="treatment" docVersion="5" id="DCA2FBA40D0051C18F31827BCF7D25C2" lastPageNumber="167" masterDocId="DCA2FBA40D0051C18F31827BCF7D25C2" masterDocTitle="Systematic notes on three troglobitic Anapistula (Araneae, Symphytognathidae) spiders from China, with the descriptions of two new species" masterLastPageNumber="189" masterPageNumber="167" pageNumber="167" updateTime="1685912906400" updateUser="ExternalLinkService">
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<mods:title id="36A4ACDB8F5B9F82103DF8A118990B38">Systematic notes on three troglobitic Anapistula (Araneae, Symphytognathidae) spiders from China, with the descriptions of two new species</mods:title>
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<mods:namePart id="3975E76CF191F7C26B99E7D52E067C9E">Wang, Shuqiao</mods:namePart>
<mods:affiliation id="D270F84EC2F28A939357F34EBCE0F00B">Key Laboratory of Bio-resources and Eco-environment (Ministry of Education), College of Life Sciences, Sichuan University, Chengdu 610065, China</mods:affiliation>
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<mods:namePart id="77B2151FFAEBB0453BBB104713462A31">Lu, Ying</mods:namePart>
<mods:affiliation id="55B55A1227641F890BC7E50E312B3C62">The Sichuan Key Laboratory for Conservation Biology of Endangered Wildlife, Sichuan University, Chengdu, Sichuan 610064, China</mods:affiliation>
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<mods:namePart id="A94BA3F1E345E8B6BC1A59CA977AE05F">Li, Ya</mods:namePart>
<mods:nameIdentifier id="4C1E84EEFCD2AF86A23F617BF5DF9D6D" type="ORCID">https://orcid.org/0000-0002-1097-6192</mods:nameIdentifier>
<mods:affiliation id="719EC53C8C808CBFF4A60C94BB46DB9C">Key Laboratory of Bio-resources and Eco-environment (Ministry of Education), College of Life Sciences, Sichuan University, Chengdu 610065, China</mods:affiliation>
</mods:name>
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<mods:namePart id="FDAFFF1FED29DCA5B5427F6B98B277A0">Li, Shuqiang</mods:namePart>
<mods:nameIdentifier id="A5BF09A51ABFB2042A9E1683EEBC6259" type="ORCID">https://orcid.org/0000-0002-3290-5416</mods:nameIdentifier>
<mods:affiliation id="CFE15BD0ACA0D23822001DD87F34A16B">The Sichuan Key Laboratory for Conservation Biology of Endangered Wildlife, Sichuan University, Chengdu, Sichuan 610064, China</mods:affiliation>
<mods:nameIdentifier id="E68AD19C770C5E4C9BAE9A123915F946" type="email">lisq@ioz.ac.cn</mods:nameIdentifier>
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<mods:namePart id="386BC80E1923C6B0A576E8EA0D1108CC">Lin, Yucheng</mods:namePart>
<mods:nameIdentifier id="E6923337BBB452DD0805A60D112951E9" type="ORCID">https://orcid.org/0000-0002-5054-0633</mods:nameIdentifier>
<mods:affiliation id="2F9F69F8632076F45676F38B6209C05B">Key Laboratory of Bio-resources and Eco-environment (Ministry of Education), College of Life Sciences, Sichuan University, Chengdu 610065, China</mods:affiliation>
<mods:nameIdentifier id="A066E728D51781A1E002FBDDCF6415E0" type="email">linyucheng@scu.edu.cn</mods:nameIdentifier>
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<mods:typeOfResource id="43265B0F3747220A3C080396D413A068">text</mods:typeOfResource>
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<mods:title id="25647E34727536802E78D1B3F2707381">ZooKeys</mods:title>
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<mods:date id="F5C8B60CBE15DF4E8FE82653BE285CA6">2022</mods:date>
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<mods:number id="A13972D7C584C6D9EB39BBE00FBDC32C">2022-11-21</mods:number>
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<mods:number id="FCEF0E86BDAC66889910BC107602C55C">1130</mods:number>
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<mods:start id="3344B71358A308561EF9D20AE4480F8E">167</mods:start>
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<mods:identifier id="FB433E97305B5772064145BF190FF4F8" type="Pensoft-Pub">1313-2970-1130-167</mods:identifier>
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<subSection id="B375AAE4C843F27C448F01A03906EB76" pageId="0" pageNumber="167" type="results">
<treatment id="D65CF3D42C895E54AE2B17713D135C49" LSID="urn:lsid:plazi:treatment:D65CF3D42C895E54AE2B17713D135C49" httpUri="http://treatment.plazi.org/id/D65CF3D42C895E54AE2B17713D135C49" lastPageNumber="167" pageId="0" pageNumber="167">
<subSubSection id="464DBB43BCDCB9AD5DE85FFDC73232AE" pageId="0" pageNumber="167" type="nomenclature">
<paragraph id="6C8B873227C592A469D3D8BC5E99E8CE" pageId="0" pageNumber="167">
<taxonomicName id="AE9F5B73773750893A33A7D9934A743C" ID-CoL="DFSD" ID-ENA="2969356" LSID="D65CF3D4-2C89-5E54-AE2B-17713D135C49" authority="Lin, Tao &amp; S. Li, 2013" authorityName="Lin, Tao &amp; S. Li" authorityYear="2013" class="Arachnida" family="Symphytognathidae" genus="Anapistula" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Anapistula panensis" order="Araneae" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="panensis">Anapistula panensis Lin, Tao &amp; S. Li, 2013</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C4354C0A9B00C84DCD567066D6EC23F5" pageId="0" pageNumber="167" type="description">
<paragraph id="439ACBB11825CE348BC9D523B2851A29" pageId="0" pageNumber="167">
<figureCitation id="8F7277104CEEEB3F25CA3536C0D06DB2" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Anapistula sanjiao sp. nov. (A, D, G, J), Anapistula walayaku sp. nov. (B, E, H, K), and Anapistula panensis (C, F, I, L) A, C male habitus, dorsal D, F male habitus, ventral B male prosoma, dorsal E male prosoma, ventral G-I female habitus, dorsal J-L female habitus, ventral. Scale bars: 0.20 (A-L)." figureDoi="10.3897/zookeys.1130.91467.figure3" httpUri="https://binary.pensoft.net/fig/772116" pageId="0" pageNumber="167">Figs 3C, F, I, L</figureCitation>
<figureCitation id="0BEB3CEAC6DEECE594CC3CB8D446A04B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Anapistula panensis A male palp, prolateral B male palp, retrolateral C epigyne, ventral D vulva, ventral E vulva, dorsal. Abbreviations: A = epigynal atrium; Co = conductor; C 1 = anterior projection of conductor; C 2 = posterior projection of conductor; Cy = cymbium; E = embolus; Fd = fertilization duct; Lb = lateral branch of the MD; Llb = distal lobe of lateral branch; MD = median duct of vulva; Pa = palpal patella; S = spermatheca; Sd = sperm duct; Te = palpal tibia. Scale bars: 0.10 (A-E)." figureDoi="10.3897/zookeys.1130.91467.figure6" httpUri="https://binary.pensoft.net/fig/772119" pageId="0" pageNumber="167">, 6A-E</figureCitation>
<figureCitation id="E4DC67D8823014B4538B151E4FE0D1DB" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Anapistula panensis A male palp, prolateral B male palp, retrolateral C vulva, ventral D vulva, dorsal. Abbreviations: A = epigynal atrium; Co = conductor; C 1 = anterior projection of conductor; C 2 = posterior projection of conductor; Cy = cymbium; E = embolus; Fd = fertilization duct; Lb = lateral branch of the MD; Llb = distal lobe of lateral branch; MD = median duct of vulva; Pa = palpal patella; S = spermatheca; Sd = sperm duct; Te = palpal tibia. Scale bars: 0.10 (A-D)." figureDoi="10.3897/zookeys.1130.91467.figure9" httpUri="https://binary.pensoft.net/fig/772122" pageId="0" pageNumber="167">, 9A-D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C488368EE5DDB42E0E122704A425302C" pageId="0" pageNumber="167" type="reference_group">
<paragraph id="9269F361BB8EC742653F3319572D8E76" pageId="0" pageNumber="167">
<taxonomicName id="6494859252177537B5D33FFF8830616A" authorityName="Lin, Tao &amp; S. Li" authorityYear="2013" class="Arachnida" family="Symphytognathidae" genus="Anapistula" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Anapistula panensis" order="Araneae" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="panensis">Anapistula panensis</taxonomicName>
Lin, Tao &amp; Li, 2013: 53, figs 1-5 (♂♀).
</paragraph>
</subSubSection>
<subSubSection id="6A032AD30205E8EED432B1FC907F7D38" pageId="0" pageNumber="167" type="type material">
<paragraph id="9048F47E33B51908E5AA3240480FDCA7" pageId="0" pageNumber="167">Type material.</paragraph>
<paragraph id="A18123CBC3F5893493C8C27E0FA98613" pageId="0" pageNumber="167">
<emphasis id="B68956781A5A47C25F9DD03363156752" bold="true" italics="true" pageId="0" pageNumber="167">Holotype</emphasis>
♂ and
<emphasis id="33DC1705C8700E25237DCC551C72B263" bold="true" italics="true" pageId="0" pageNumber="167">paratypes</emphasis>
1♂ 50♀ (IZCAS) from
<emphasis id="5DB514BD6E385D23EFA3C71206FFD3BE" bold="true" pageId="0" pageNumber="167">China</emphasis>
: Guizhou Province, Liupanshui City, Pan County, Zhudong Township, Shiliping Village, Shenxian Cave (
<geoCoordinate id="4F97156C2DF148F87165738E5E5BD7A0" degrees="25.62367" direction="north" orientation="latitude" precision="1" value="25.62367">25.62367°N</geoCoordinate>
,
<geoCoordinate id="4A0111DF4523F01E5DA42DA8542D61BA" degrees="104.75653" direction="east" orientation="longitude" precision="1" value="104.75653">104.75653°E</geoCoordinate>
, 1687 m), 15.IV.2007, J. Liu &amp; Y. Lin leg. Examined.
</paragraph>
</subSubSection>
<subSubSection id="0C5E1E062CAD5BBD2CC0EC3377EB2572" pageId="0" pageNumber="167" type="materials_examined">
<paragraph id="BA8C6526A42D1CDC27C753DC74D7DAC5" pageId="0" pageNumber="167">Other material examined.</paragraph>
<paragraph id="4591D89708EED57FCEFDF556DF080169" pageId="0" pageNumber="167">
<materialsCitation id="D4493E14D2C06973949C9F86A5F8F867" collectingDate="2010-04-26" collectorName="Y. Lin, Q. Zhao" country="China" location="China" specimenCount="51" specimenCount-female="51">
<specimenCount id="B45828539CFAB5B10965F4CA875F9616" type="female">51♀</specimenCount>
18 juvs (NHMSU-HA020) from
<emphasis id="C6CF5A5B1C1AD25D73E9B1111845F306" bold="true" pageId="0" pageNumber="167">
<collectingCountry id="790C3B91CFC8982A0FFC4AAB200C3CA5" name="China">China</collectingCountry>
</emphasis>
: same data as type locality,
<collectingDate id="1B4918BCD429D42A140DC16914FC349E" value="2010-04-26">26.IV.2010</collectingDate>
,
<collectorName id="1E49B68926BD17C81AAA06EB50E36E08">Y. Lin</collectorName>
&amp;
<collectorName id="74592052F514A14A19165B96B3F3EE7F">Q. Zhao</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="FF9BF51FEA55E580D9561D0F71BA1C6D" collectingDate="2020-08-24" collectorName="Y. Lin" country="China" specimenCount="68" specimenCount-female="67" specimenCount-male="1">
<specimenCount id="A522969818230EC59369335CED3C24FF" type="male">1♂</specimenCount>
<specimenCount id="33ADBCA3095AE40A4F3DC4DF7FACBFB1" type="female">67♀</specimenCount>
20 juvs (NHMSU-QX003) same data as type locality,
<collectingDate id="55EA9EC06A5F24F2EFDEF7A1F23CEC82" value="2020-08-24">24.VIII.2020</collectingDate>
,
<collectorName id="358877DDCF0FDD5C5A83B6C8B430084A">Y. Lin</collectorName>
et al. leg
</materialsCitation>
.;
<materialsCitation id="BDC056293EAF5F71F46351CD60EA1F61" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Y. Lin" country="China" specimenCount="2" specimenCount-female="1" specimenCount-juvenile="1">
<specimenCount id="59E44E6ED08DBA17103971D438C1CB20" type="female">1♀</specimenCount>
<specimenCount id="19ECF0C8E9DC0D183B209FC0C905715B" type="juvenile">1 juv</specimenCount>
(NHMSU-HA020) used for sequencing, GenBank accession numbers given in Table
<tableCitation id="FF78E3A1C8A977A8F1B3FC3E145F0306" captionStart="Table 2" captionStartId="T2" captionText="Table 2. GenBank accession numbers for new DNA sequence data from three Anapistula species." httpUri="http://table.plazi.org/id/2C65119DC82E48657BD255A2ADDAF307" pageId="0" pageNumber="167" tableUuid="2C65119DC82E48657BD255A2ADDAF307">2</tableCitation>
, same data as for preceding
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="482EE5BC28CE1D48CA176097A76EE91C" pageId="0" pageNumber="167" type="diagnosis">
<paragraph id="DD0B017A008BF2101ED1B9EF740E33BD" pageId="0" pageNumber="167">Diagnosis.</paragraph>
<paragraph id="BFE0C6083F34DAB93D9A7C723473FA02" pageId="0" pageNumber="167">
The male of
<taxonomicName id="FFCC65D77BD8FC105DE4F397CA38A8A1" lsidName="A. panensis" pageId="0" pageNumber="167" rank="species" species="panensis">
<emphasis id="04D555EEDDB250B472E20FA7E15BC799" italics="true" pageId="0" pageNumber="167">A. panensis</emphasis>
</taxonomicName>
is similar to that of
<taxonomicName id="E988BFEC22AE96A887C0EF172A8B4FAF" lsidName="A. choojaiae" pageId="0" pageNumber="167" rank="species" species="choojaiae">
<emphasis id="6882C6BD945445EB87AB33C5836180D4" italics="true" pageId="0" pageNumber="167">A. choojaiae</emphasis>
</taxonomicName>
in the shape of the palp and in having C1 and C2 roughly equal in length, but it differs by a narrower C1 and a wider C2, a longer embolus, and having three setae on the paracymbium (vs. two; cf. Figs
<figureCitation id="E64DA0490B8C8AE20DBA766CE43FEC81" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Anapistula panensis A male palp, prolateral B male palp, retrolateral C epigyne, ventral D vulva, ventral E vulva, dorsal. Abbreviations: A = epigynal atrium; Co = conductor; C 1 = anterior projection of conductor; C 2 = posterior projection of conductor; Cy = cymbium; E = embolus; Fd = fertilization duct; Lb = lateral branch of the MD; Llb = distal lobe of lateral branch; MD = median duct of vulva; Pa = palpal patella; S = spermatheca; Sd = sperm duct; Te = palpal tibia. Scale bars: 0.10 (A-E)." figureDoi="10.3897/zookeys.1130.91467.figure6" httpUri="https://binary.pensoft.net/fig/772119" pageId="0" pageNumber="167">6A, B</figureCitation>
,
<figureCitation id="DD079ED66419B427BDD8E6B190C000FD" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Anapistula panensis A male palp, prolateral B male palp, retrolateral C vulva, ventral D vulva, dorsal. Abbreviations: A = epigynal atrium; Co = conductor; C 1 = anterior projection of conductor; C 2 = posterior projection of conductor; Cy = cymbium; E = embolus; Fd = fertilization duct; Lb = lateral branch of the MD; Llb = distal lobe of lateral branch; MD = median duct of vulva; Pa = palpal patella; S = spermatheca; Sd = sperm duct; Te = palpal tibia. Scale bars: 0.10 (A-D)." figureDoi="10.3897/zookeys.1130.91467.figure9" httpUri="https://binary.pensoft.net/fig/772122" pageId="0" pageNumber="167">9A, B</figureCitation>
;
<bibRefCitation id="09BEC38AA2FA4D96002E09DE116C9165" author="Lin, Y" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="0" pageNumber="167" pagination="53 - 58" refId="B28" refString="Lin, Y, Tao, Y, Li, S, 2013. Two new species of the genus Anapistula (Araneae, Symphytognathidae) from southern China. Acta Zootaxonomica Sinica 38: 53 - 58" title="Two new species of the genus Anapistula (Araneae, Symphytognathidae) from southern China." volume="38" year="2013">Lin et al. 2013</bibRefCitation>
: figs 1, 2 with
<bibRefCitation id="08100D5896E15F950BC2E353C58A9CDB" DOI="https://doi.org/10.3897/zookeys.1012.57047" author="Rivera-Quiroz, FA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="167" pagination="21 - 53" refId="B39" refString="Rivera-Quiroz, FA, Petcharad, B, Miller, JA, 2021. First records and three new species of the family Symphytognathidae (Arachnida, Araneae) from Thailand, and the circumscription of the genus Crassignatha Wunderlich, 1995. ZooKeys 1012: 21 - 53, DOI: https://doi.org/10.3897/zookeys.1012.57047" title="First records and three new species of the family Symphytognathidae (Arachnida, Araneae) from Thailand, and the circumscription of the genus Crassignatha Wunderlich, 1995." url="https://doi.org/10.3897/zookeys.1012.57047" volume="1012" year="2021">Rivera-Quiroz et al. 2021</bibRefCitation>
: figs 7c, 9a, b). The female differs from most
<taxonomicName id="6014D4CEECB875D22EB4ABF679CC7ACD" authorityName="Gertsch" authorityYear="1941" class="Arachnida" family="Symphytognathidae" genus="Anapistula" kingdom="Animalia" lsidName="Anapistula" order="Araneae" pageId="0" pageNumber="167" phylum="Arthropoda" rank="genus">
<emphasis id="1E29FEF7E8FD78FF0192214D545F546A" italics="true" pageId="0" pageNumber="167">Anapistula</emphasis>
</taxonomicName>
species by the Y-shaped epigynal median duct; it shares this character with
<taxonomicName id="E192B00DEA1E4DC40008AA4BBD18A74D" lsidName="A. orbisterna" pageId="0" pageNumber="167" rank="species" species="orbisterna">
<emphasis id="AA0227DE82212FD791406CA057790207" italics="true" pageId="0" pageNumber="167">A. orbisterna</emphasis>
</taxonomicName>
,
<taxonomicName id="D9ED156FF66ED61CB0CBD8D564014EBF" lsidName="A. secreta" pageId="0" pageNumber="167" rank="species" species="secreta">
<emphasis id="F40138AD58EB38EB8A6A9063384E4D3F" italics="true" pageId="0" pageNumber="167">A. secreta</emphasis>
</taxonomicName>
,
<taxonomicName id="07E86E5632A7869F99E3A045BF45421A" lsidName="A. bifurcate" pageId="0" pageNumber="167" rank="species" species="bifurcate">
<emphasis id="3A4920C0B1800633F24D7F0DC8EA913F" italics="true" pageId="0" pageNumber="167">A. bifurcate</emphasis>
</taxonomicName>
,
<taxonomicName id="9BE0050C94ED80E2DC322F33DAF1D666" lsidName="A. tonga" pageId="0" pageNumber="167" rank="species" species="tonga">
<emphasis id="5B55483DB366B580A2DD5880322315C0" italics="true" pageId="0" pageNumber="167">A. tonga</emphasis>
</taxonomicName>
,
<taxonomicName id="A740C1A57A0405BE58F8B3D6CF48CB42" lsidName="A. choojaiae" pageId="0" pageNumber="167" rank="species" species="choojaiae">
<emphasis id="6EFDBBF0405FD21B9430672CEAD9991F" italics="true" pageId="0" pageNumber="167">A. choojaiae</emphasis>
</taxonomicName>
,
<taxonomicName id="26E7A1B9BB7CD49AC0545558C81714CF" lsidName="A. equatoriana" pageId="0" pageNumber="167" rank="species" species="equatoriana">
<emphasis id="AE9B8FBF7E567CD4E4C3F028AF08AF08" italics="true" pageId="0" pageNumber="167">A. equatoriana</emphasis>
</taxonomicName>
,
<taxonomicName id="E6829521C6ECF7B508C9E2321D02E299" lsidName="A. zhengi" pageId="0" pageNumber="167" rank="species" species="zhengi">
<emphasis id="162B062B425461072ABC184C67DBBC21" italics="true" pageId="0" pageNumber="167">A. zhengi</emphasis>
</taxonomicName>
,
<taxonomicName id="BCF93FA9361A7D71CFBAB38BFADC6382" lsidName="A. sanjiao" pageId="0" pageNumber="167" rank="species" species="sanjiao">
<emphasis id="3E6DFCC5DC71F6A78E2E5E4C84B90EB2" italics="true" pageId="0" pageNumber="167">A. sanjiao</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName id="A1C61FD568349E9C22F145881DF5C6AD" lsidName="A. walayaku" pageId="0" pageNumber="167" rank="species" species="walayaku">
<emphasis id="F0091DABC4C7914D2C5629D0455E9453" italics="true" pageId="0" pageNumber="167">A. walayaku</emphasis>
</taxonomicName>
sp. nov.
<taxonomicName id="1B5F9A929C00D1BD331F04AF1D6BF964" authorityName="Lin, Tao &amp; S. Li" authorityYear="2013" class="Arachnida" family="Symphytognathidae" genus="Anapistula" kingdom="Animalia" lsidName="Anapistula panensis" order="Araneae" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="panensis">
<emphasis id="EFAE5600A963BC3A43E1EF8F74E7AC91" italics="true" pageId="0" pageNumber="167">Anapistula panensis</emphasis>
</taxonomicName>
differs from all of these by the width and length of the MD, the length of the lateral branches, and the shape and relative size of the atrium (Figs
<figureCitation id="833711675D376366AA2E6A6915E89899" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Anapistula panensis A male palp, prolateral B male palp, retrolateral C epigyne, ventral D vulva, ventral E vulva, dorsal. Abbreviations: A = epigynal atrium; Co = conductor; C 1 = anterior projection of conductor; C 2 = posterior projection of conductor; Cy = cymbium; E = embolus; Fd = fertilization duct; Lb = lateral branch of the MD; Llb = distal lobe of lateral branch; MD = median duct of vulva; Pa = palpal patella; S = spermatheca; Sd = sperm duct; Te = palpal tibia. Scale bars: 0.10 (A-E)." figureDoi="10.3897/zookeys.1130.91467.figure6" httpUri="https://binary.pensoft.net/fig/772119" pageId="0" pageNumber="167">6C-E</figureCitation>
,
<figureCitation id="86956BF40B0D4B576F5ED03EC781A208" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Anapistula panensis A male palp, prolateral B male palp, retrolateral C vulva, ventral D vulva, dorsal. Abbreviations: A = epigynal atrium; Co = conductor; C 1 = anterior projection of conductor; C 2 = posterior projection of conductor; Cy = cymbium; E = embolus; Fd = fertilization duct; Lb = lateral branch of the MD; Llb = distal lobe of lateral branch; MD = median duct of vulva; Pa = palpal patella; S = spermatheca; Sd = sperm duct; Te = palpal tibia. Scale bars: 0.10 (A-D)." figureDoi="10.3897/zookeys.1130.91467.figure9" httpUri="https://binary.pensoft.net/fig/772122" pageId="0" pageNumber="167">9C, D</figureCitation>
and
<bibRefCitation id="0BE7151F8A77AAAB7821D6B01474058F" author="Lin, Y" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="0" pageNumber="167" pagination="53 - 58" refId="B28" refString="Lin, Y, Tao, Y, Li, S, 2013. Two new species of the genus Anapistula (Araneae, Symphytognathidae) from southern China. Acta Zootaxonomica Sinica 38: 53 - 58" title="Two new species of the genus Anapistula (Araneae, Symphytognathidae) from southern China." volume="38" year="2013">Lin et al. 2013</bibRefCitation>
: figs 3, 4 vs.
<bibRefCitation id="7A3EECD472578A69A1526406C2ED037F" author="Forster, RR" journalOrPublisher="American Museum Novitates" pageId="0" pageNumber="167" pagination="1 - 29" refId="B13" refString="Forster, RR, Platnick, NI, 1977. A review of the spider family Symphytognathidae (Arachnida, Araneae). American Museum Novitates 2619: 1 - 29" title="A review of the spider family Symphytognathidae (Arachnida, Araneae)." volume="2619" year="1977">Forster and Platnick 1977</bibRefCitation>
: fig. 19;
<bibRefCitation id="2FCCBA7D280C498A098C69827F7C07E6" author="Harvey, MS" journalOrPublisher="Records of the Western Australian Museum" pageId="0" pageNumber="167" pagination="111 - 120" refId="B18" refString="Harvey, MS, 1998. A review of the Australasian species of Anapistula Gertsch (Araneae: Symphytognathidae). Records of the Western Australian Museum 19: 111 - 120" title="A review of the Australasian species of Anapistula Gertsch (Araneae: Symphytognathidae)." volume="19" year="1998">Harvey 1998</bibRefCitation>
: figs 9, 19;
<bibRefCitation id="B6F5FCEF85E66B3B89B5B1810EECD7F6" DOI="https://doi.org/10.3897/zookeys.1012.57047" author="Rivera-Quiroz, FA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="167" pagination="21 - 53" refId="B39" refString="Rivera-Quiroz, FA, Petcharad, B, Miller, JA, 2021. First records and three new species of the family Symphytognathidae (Arachnida, Araneae) from Thailand, and the circumscription of the genus Crassignatha Wunderlich, 1995. ZooKeys 1012: 21 - 53, DOI: https://doi.org/10.3897/zookeys.1012.57047" title="First records and three new species of the family Symphytognathidae (Arachnida, Araneae) from Thailand, and the circumscription of the genus Crassignatha Wunderlich, 1995." url="https://doi.org/10.3897/zookeys.1012.57047" volume="1012" year="2021">Rivera-Quiroz et al. 2021</bibRefCitation>
: figs 8d, 9c;
<bibRefCitation id="509B7B32B5D75906F2D138510208CDBD" DOI="https://doi.org/10.11646/zootaxa.4341.3.3" author="Duperre, N" journalOrPublisher="Zootaxa" pageId="0" pageNumber="167" pagination="375 - 399" refId="B7" refString="Duperre, N, Tapia, E, 2017. On some minuscule spiders (Araneae: Theridiosomatidae, Symphytognathidae) from the Choco region of Ecuador with the description of ten new species. Zootaxa 4341 (3): 375 - 399, DOI: https://doi.org/10.11646/zootaxa.4341.3.3" title="On some minuscule spiders (Araneae: Theridiosomatidae, Symphytognathidae) from the Choco region of Ecuador with the description of ten new species." url="https://doi.org/10.11646/zootaxa.4341.3.3" volume="4341" year="2017">
<normalizedToken id="FD11054453865A46ECEBE9C9FA7C4B74" originalValue="Dupérré">Duperre</normalizedToken>
and Tapia 2017
</bibRefCitation>
: fig. 33;
<bibRefCitation id="0B45D22965CD767CE0D4D154B5DF8E50" author="Lin, Y" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="0" pageNumber="167" pagination="53 - 58" refId="B28" refString="Lin, Y, Tao, Y, Li, S, 2013. Two new species of the genus Anapistula (Araneae, Symphytognathidae) from southern China. Acta Zootaxonomica Sinica 38: 53 - 58" title="Two new species of the genus Anapistula (Araneae, Symphytognathidae) from southern China." volume="38" year="2013">Lin et al. 2013</bibRefCitation>
: figs 8, 9).
</paragraph>
<caption id="702C1E12114496CDE497483409281872" doi="10.3897/zookeys.1130.91467.figure9" httpUri="https://binary.pensoft.net/fig/772122" pageId="0" pageNumber="167" start="Figure 9" startId="F9">
<paragraph id="31010052F7754B276884B35B6B5841E6" pageId="0" pageNumber="167">
<emphasis id="C576064C0311DB48ACA944EAAB83D0C4" bold="true" pageId="0" pageNumber="167">Figure 9.</emphasis>
<taxonomicName id="28FE60E7F0C0EE957A0F7DEA74DB6CA7" authorityName="Lin, Tao &amp; S. Li" authorityYear="2013" class="Arachnida" family="Symphytognathidae" genus="Anapistula" kingdom="Animalia" lsidName="Anapistula panensis" order="Araneae" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="panensis">
<emphasis id="D7CAB040C49C468C8EB1A1317D090B40" italics="true" pageId="0" pageNumber="167">Anapistula panensis</emphasis>
</taxonomicName>
<emphasis id="B4F0FD1C9DAD071774063B11171B6816" bold="true" pageId="0" pageNumber="167">A</emphasis>
male palp, prolateral
<emphasis id="899B789FE3E8C10736E5FBBA89237082" bold="true" pageId="0" pageNumber="167">B</emphasis>
male palp, retrolateral
<emphasis id="03BB7C6C2782DB2EC7F8820486CD142A" bold="true" pageId="0" pageNumber="167">C</emphasis>
vulva, ventral
<emphasis id="E2707B6A08740716B815F33AE7061E15" bold="true" pageId="0" pageNumber="167">D</emphasis>
vulva, dorsal. Abbreviations: A = epigynal atrium; Co = conductor; C1 = anterior projection of conductor; C2 = posterior projection of conductor; Cy = cymbium; E = embolus; Fd = fertilization duct; Lb = lateral branch of the MD; Llb = distal lobe of lateral branch; MD = median duct of vulva; Pa = palpal patella; S = spermatheca; Sd = sperm duct; Te = palpal tibia. Scale bars: 0.10 (
<emphasis id="74CF6B0BC978F0C7EF806B9413B74FAA" bold="true" pageId="0" pageNumber="167">A-D</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="368796C51F8700993FB67DFDC81EF8C0" pageId="0" pageNumber="167" type="redescription">
<paragraph id="387A774E557A7D461F1309D9C1EEB9B1" pageId="0" pageNumber="167">Redescription.</paragraph>
<paragraph id="66B2005CB90D609D13D80B1E3B6E5C5E" pageId="0" pageNumber="167">
<emphasis id="C753B4E0FAAE7040C2607A61BD41A44E" bold="true" pageId="0" pageNumber="167">Male</emphasis>
: habitus as in Fig.
<figureCitation id="E06516FC25C91359886FABAD56991149" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Anapistula sanjiao sp. nov. (A, D, G, J), Anapistula walayaku sp. nov. (B, E, H, K), and Anapistula panensis (C, F, I, L) A, C male habitus, dorsal D, F male habitus, ventral B male prosoma, dorsal E male prosoma, ventral G-I female habitus, dorsal J-L female habitus, ventral. Scale bars: 0.20 (A-L)." figureDoi="10.3897/zookeys.1130.91467.figure3" httpUri="https://binary.pensoft.net/fig/772116" pageId="0" pageNumber="167">3C, F</figureCitation>
. Body yellow. Legs pale yellow. Carapace nearly round, cephalic area moderately elevated. Four vestigial eyes in diads. Chelicerae distinctly sclerotized and fused basally, concave at inner margins, with two promarginal teeth. Endites as long as wide. Labium rectangular, length ca equal to ⅓ of width, fused to sternum. Sternum flat, with sparse setae, truncated posteriorly. Femur I and II swollen retrolatero-basally, tiny serrations and granulations on surface. Patellae I-IV each with a distal-dorsal seta. Abdomen ovoid dorsally, higher than long, covered with sparse, grey, long setae, posterior expanded beyond spinnerets. Colulus absent. Somatic measurement: body length 0.56, carapace 0.28 long, 0.28 wide, 0.24 high; sternum 0.20 long, 0.24 wide; abdomen 0.32 long, 0.28 wide, 0.36 high; length of legs: I 1.18 (0.34, 0.14, 0.26, 0.14, 0.30), II 0.96 (0.28, 0.12, 0.18, 0.12, 0.26), III 0.80 (0.20, 0.10, 0.16, 0.10, 0.24), IV 0.98 (0.30, 0.12, 0.18, 0.16, 0.22); leg formula I-IV-II-III.
</paragraph>
<paragraph id="8CA2B6231B8CED320DF3DC4192F3D32D" pageId="0" pageNumber="167">
<emphasis id="A9A7332806487C1539ACC88C949E98D4" bold="true" italics="true" pageId="0" pageNumber="167">Palp</emphasis>
: small and weakly sclerotized. Femur slightly swollen distally, with a long seta at retrolateral base. Patella short, semilunar shaped. Tibia contracted proximally, broad distally. Cymbium transparent, with 7 retrolateral short and 2 dorsal long setae. Conductor sheet shaped, with two projections (C1 and C2), C1 sharp, C2 lamellar, nearly invisible. Embolus short, needle shaped, posterior to conductor. Sd coiled ca 2 times inside bulb (Figs
<figureCitation id="98F2FFBEC25E4B51CC7C4DA2A158E14C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Anapistula panensis A male palp, prolateral B male palp, retrolateral C epigyne, ventral D vulva, ventral E vulva, dorsal. Abbreviations: A = epigynal atrium; Co = conductor; C 1 = anterior projection of conductor; C 2 = posterior projection of conductor; Cy = cymbium; E = embolus; Fd = fertilization duct; Lb = lateral branch of the MD; Llb = distal lobe of lateral branch; MD = median duct of vulva; Pa = palpal patella; S = spermatheca; Sd = sperm duct; Te = palpal tibia. Scale bars: 0.10 (A-E)." figureDoi="10.3897/zookeys.1130.91467.figure6" httpUri="https://binary.pensoft.net/fig/772119" pageId="0" pageNumber="167">6A, B</figureCitation>
,
<figureCitation id="3C3ACF63DC435D922ECEA155F7038C22" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Anapistula panensis A male palp, prolateral B male palp, retrolateral C vulva, ventral D vulva, dorsal. Abbreviations: A = epigynal atrium; Co = conductor; C 1 = anterior projection of conductor; C 2 = posterior projection of conductor; Cy = cymbium; E = embolus; Fd = fertilization duct; Lb = lateral branch of the MD; Llb = distal lobe of lateral branch; MD = median duct of vulva; Pa = palpal patella; S = spermatheca; Sd = sperm duct; Te = palpal tibia. Scale bars: 0.10 (A-D)." figureDoi="10.3897/zookeys.1130.91467.figure9" httpUri="https://binary.pensoft.net/fig/772122" pageId="0" pageNumber="167">9A, B</figureCitation>
).
</paragraph>
<paragraph id="79F387B11220621A062547634AC9265A" pageId="0" pageNumber="167">
<emphasis id="AD7FBE85268450A74D4F6EB49884FCF0" bold="true" pageId="0" pageNumber="167">Female</emphasis>
: habitus see Fig.
<figureCitation id="202765D5016877924C0B2FAC912202E9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Anapistula sanjiao sp. nov. (A, D, G, J), Anapistula walayaku sp. nov. (B, E, H, K), and Anapistula panensis (C, F, I, L) A, C male habitus, dorsal D, F male habitus, ventral B male prosoma, dorsal E male prosoma, ventral G-I female habitus, dorsal J-L female habitus, ventral. Scale bars: 0.20 (A-L)." figureDoi="10.3897/zookeys.1130.91467.figure3" httpUri="https://binary.pensoft.net/fig/772116" pageId="0" pageNumber="167">3I, L</figureCitation>
. Carapace darker yellow than abdomen. Palps absent, others as in male. Somatic measurements: body length 0.68, carapace 0.28 long, 0.28 wide, 0.24 high; sternum 0.20 long, 0.18 wide; abdomen 0.50 long, 0.52 wide, 0.52 high; length of legs: I 1.18 (0.36, 0.14, 0.24, 0.20, 0.24), II 1.08 (0.30, 0.14, 0.24, 0.18, 0.22), III 0.82 (0.18, 0.12, 0.16, 0.16, 0.20), IV 1.12 (0.30, 0.14, 0.26, 0.18, 0.24); leg formula I-IV-II-III.
</paragraph>
<paragraph id="18B46A7CF2841557FA51FAB768FA3473" pageId="0" pageNumber="167">
<emphasis id="A869C3C516DD75B2DC9966EC0E90F977" bold="true" italics="true" pageId="0" pageNumber="167">Epigyne</emphasis>
: flat, without scape. Atrium ovoid, narrower than space between spermathecae. Spermathecae spherical, separated by ca 1.3
<normalizedToken id="5C6BD5D9A81CAA70289E6AD31C8EDCD6" originalValue="×">x</normalizedToken>
their diameter, obviously sclerotized (Figs
<figureCitation id="6A5814D48775C23A524B96B02FD5F7DB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Anapistula sanjiao sp. nov. (A, D, G, J), Anapistula walayaku sp. nov. (B, E, H, K), and Anapistula panensis (C, F, I, L) A, C male habitus, dorsal D, F male habitus, ventral B male prosoma, dorsal E male prosoma, ventral G-I female habitus, dorsal J-L female habitus, ventral. Scale bars: 0.20 (A-L)." figureDoi="10.3897/zookeys.1130.91467.figure3" httpUri="https://binary.pensoft.net/fig/772116" pageId="0" pageNumber="167">3L</figureCitation>
,
<figureCitation id="F14FBD8F55C1450C1425E2E956B5E053" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Anapistula panensis A male palp, prolateral B male palp, retrolateral C epigyne, ventral D vulva, ventral E vulva, dorsal. Abbreviations: A = epigynal atrium; Co = conductor; C 1 = anterior projection of conductor; C 2 = posterior projection of conductor; Cy = cymbium; E = embolus; Fd = fertilization duct; Lb = lateral branch of the MD; Llb = distal lobe of lateral branch; MD = median duct of vulva; Pa = palpal patella; S = spermatheca; Sd = sperm duct; Te = palpal tibia. Scale bars: 0.10 (A-E)." figureDoi="10.3897/zookeys.1130.91467.figure6" httpUri="https://binary.pensoft.net/fig/772119" pageId="0" pageNumber="167">6C</figureCitation>
,
<figureCitation id="5AA9BA1A9F8AE8D941E2FA6FF6969B84" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Anapistula panensis A male palp, prolateral B male palp, retrolateral C vulva, ventral D vulva, dorsal. Abbreviations: A = epigynal atrium; Co = conductor; C 1 = anterior projection of conductor; C 2 = posterior projection of conductor; Cy = cymbium; E = embolus; Fd = fertilization duct; Lb = lateral branch of the MD; Llb = distal lobe of lateral branch; MD = median duct of vulva; Pa = palpal patella; S = spermatheca; Sd = sperm duct; Te = palpal tibia. Scale bars: 0.10 (A-D)." figureDoi="10.3897/zookeys.1130.91467.figure9" httpUri="https://binary.pensoft.net/fig/772122" pageId="0" pageNumber="167">9C</figureCitation>
). Lateral branches diverging from MD, forming Y-shape (Figs
<figureCitation id="442606ADBCF157B157C7FB2337A948E4" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Anapistula panensis A male palp, prolateral B male palp, retrolateral C epigyne, ventral D vulva, ventral E vulva, dorsal. Abbreviations: A = epigynal atrium; Co = conductor; C 1 = anterior projection of conductor; C 2 = posterior projection of conductor; Cy = cymbium; E = embolus; Fd = fertilization duct; Lb = lateral branch of the MD; Llb = distal lobe of lateral branch; MD = median duct of vulva; Pa = palpal patella; S = spermatheca; Sd = sperm duct; Te = palpal tibia. Scale bars: 0.10 (A-E)." figureDoi="10.3897/zookeys.1130.91467.figure6" httpUri="https://binary.pensoft.net/fig/772119" pageId="0" pageNumber="167">6D, E</figureCitation>
,
<figureCitation id="DF57FC92CA685B8404B2CBBE3E7B3B24" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Anapistula panensis A male palp, prolateral B male palp, retrolateral C vulva, ventral D vulva, dorsal. Abbreviations: A = epigynal atrium; Co = conductor; C 1 = anterior projection of conductor; C 2 = posterior projection of conductor; Cy = cymbium; E = embolus; Fd = fertilization duct; Lb = lateral branch of the MD; Llb = distal lobe of lateral branch; MD = median duct of vulva; Pa = palpal patella; S = spermatheca; Sd = sperm duct; Te = palpal tibia. Scale bars: 0.10 (A-D)." figureDoi="10.3897/zookeys.1130.91467.figure9" httpUri="https://binary.pensoft.net/fig/772122" pageId="0" pageNumber="167">9C, D</figureCitation>
), as wide as ⅓ of MD, as long as MD (Fig.
<figureCitation id="D47870243F2849220071C1ABA768452E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Anapistula panensis A male palp, prolateral B male palp, retrolateral C epigyne, ventral D vulva, ventral E vulva, dorsal. Abbreviations: A = epigynal atrium; Co = conductor; C 1 = anterior projection of conductor; C 2 = posterior projection of conductor; Cy = cymbium; E = embolus; Fd = fertilization duct; Lb = lateral branch of the MD; Llb = distal lobe of lateral branch; MD = median duct of vulva; Pa = palpal patella; S = spermatheca; Sd = sperm duct; Te = palpal tibia. Scale bars: 0.10 (A-E)." figureDoi="10.3897/zookeys.1130.91467.figure6" httpUri="https://binary.pensoft.net/fig/772119" pageId="0" pageNumber="167">6E</figureCitation>
;
<bibRefCitation id="56C00A49E1643E0A00204ECBBE0DFAD5" author="Lin, Y" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="0" pageNumber="167" pagination="53 - 58" refId="B28" refString="Lin, Y, Tao, Y, Li, S, 2013. Two new species of the genus Anapistula (Araneae, Symphytognathidae) from southern China. Acta Zootaxonomica Sinica 38: 53 - 58" title="Two new species of the genus Anapistula (Araneae, Symphytognathidae) from southern China." volume="38" year="2013">Lin et al. 2013</bibRefCitation>
: figs 3, 4). Fertilization ducts very short, translucent, nearly invisible, Llb appear as tiny bumps distally on lateral branches (Figs
<figureCitation id="E2E6C682E6B583ABE95CF5B2743F966F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Anapistula panensis A male palp, prolateral B male palp, retrolateral C epigyne, ventral D vulva, ventral E vulva, dorsal. Abbreviations: A = epigynal atrium; Co = conductor; C 1 = anterior projection of conductor; C 2 = posterior projection of conductor; Cy = cymbium; E = embolus; Fd = fertilization duct; Lb = lateral branch of the MD; Llb = distal lobe of lateral branch; MD = median duct of vulva; Pa = palpal patella; S = spermatheca; Sd = sperm duct; Te = palpal tibia. Scale bars: 0.10 (A-E)." figureDoi="10.3897/zookeys.1130.91467.figure6" httpUri="https://binary.pensoft.net/fig/772119" pageId="0" pageNumber="167">6D, E</figureCitation>
,
<figureCitation id="D299E3A6AB7271CEFB3B3C303C586FB2" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Anapistula panensis A male palp, prolateral B male palp, retrolateral C vulva, ventral D vulva, dorsal. Abbreviations: A = epigynal atrium; Co = conductor; C 1 = anterior projection of conductor; C 2 = posterior projection of conductor; Cy = cymbium; E = embolus; Fd = fertilization duct; Lb = lateral branch of the MD; Llb = distal lobe of lateral branch; MD = median duct of vulva; Pa = palpal patella; S = spermatheca; Sd = sperm duct; Te = palpal tibia. Scale bars: 0.10 (A-D)." figureDoi="10.3897/zookeys.1130.91467.figure9" httpUri="https://binary.pensoft.net/fig/772122" pageId="0" pageNumber="167">9C, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="AA2E62E94FD8D936C259B550D2EE538E" pageId="0" pageNumber="167" type="natural history">
<paragraph id="C7D065C3B64956C7537C8518B44F97E1" pageId="0" pageNumber="167">Natural history.</paragraph>
<paragraph id="E5DE58D3D201EEBA7680F1B07246793A" pageId="0" pageNumber="167">This species spins a small, flat circular web in the crevices of stalagmites or stalactites in caves.</paragraph>
</subSubSection>
<subSubSection id="B452DEDCF200F6A421ABAEBE5EFD2C0F" pageId="0" pageNumber="167" type="distribution">
<paragraph id="A764DC6AB812A460E387911B0A1D0E0B" pageId="0" pageNumber="167">Distribution.</paragraph>
<paragraph id="FBAED9CEB8328F299FE74239DD6C8F6C" pageId="0" pageNumber="167">
China (Guizhou) (Fig.
<figureCitation id="DA8E2E7250A90F5C52BC5BCBBE7EBF8A" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Distribution records of three Chinese cave-dwelling Anapistula species." figureDoi="10.3897/zookeys.1130.91467.figure10" httpUri="https://binary.pensoft.net/fig/772123" pageId="0" pageNumber="167">10</figureCitation>
).
</paragraph>
<caption id="1A619ACBDF6ADA7964A3109BF0181D34" doi="10.3897/zookeys.1130.91467.figure10" httpUri="https://binary.pensoft.net/fig/772123" pageId="0" pageNumber="167" start="Figure 10" startId="F10">
<paragraph id="7B9F0DB227AEB0C861D10D9DFB5CE307" pageId="0" pageNumber="167">
<emphasis id="99829E80FD9D66612FDE3379C3BA01B4" bold="true" pageId="0" pageNumber="167">Figure 10.</emphasis>
Distribution records of three Chinese cave-dwelling
<taxonomicName id="C9A11BD4553A1B4196CA4597DFD76953" authorityName="Gertsch" authorityYear="1941" class="Arachnida" family="Symphytognathidae" genus="Anapistula" kingdom="Animalia" lsidName="Anapistula" order="Araneae" pageId="0" pageNumber="167" phylum="Arthropoda" rank="genus">
<emphasis id="B75E5684B84E2258603B3E8752FFD670" italics="true" pageId="0" pageNumber="167">Anapistula</emphasis>
</taxonomicName>
species.
</paragraph>
</caption>
</subSubSection>
</treatment>
</subSection>
</document>
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