202 lines
35 KiB
XML
202 lines
35 KiB
XML
<document id="85AA82446E2CD9051EF16DF44F9F8C24" ID-DOI="10.5281/zenodo.4665318" ID-ISSN="1638-9395" ID-Zenodo-Dep="4665318" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617712144027" checkinUser="felipe" docAuthor="Duy-Jacquemin, Monique Nguyen & Azar, Dany" docDate="2004" docId="D57D87F7FFCEBF3105D0EC3AD071FDA1" docLanguage="en" docName="Geodiversitas.26.4.631-641.pdf" docOrigin="Geodiversitas 26 (4)" docStyle="DocumentStyle:94524207C6714CF92F879FE43E306202.3:Geodiversitas.2000-2008.journal_article.type2" docStyleId="94524207C6714CF92F879FE43E306202" docStyleName="Geodiversitas.2000-2008.journal_article.type2" docStyleVersion="3" docTitle="Libanoxenus hammanaensis Duy-Jacquemin & Azar 2004, n. sp." docType="treatment" docVersion="3" lastPageNumber="636" masterDocId="2944FF8FFFCABF340726EE57D34AFFC6" masterDocTitle="The oldest records of Polyxenida (Myriapoda, Diplopoda): new discoveries from the Cretaceous ambers of Lebanon and France" masterLastPageNumber="641" masterPageNumber="631" pageNumber="635" updateTime="1698900732333" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:mods id="3D0FA6E92F83D3B771020F5D3820BFF6" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="97E03422791C28D0E9A2E39911534F67">
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<mods:title id="F5AC27ED52835F4522DF2657E0B8A7EC">The oldest records of Polyxenida (Myriapoda, Diplopoda): new discoveries from the Cretaceous ambers of Lebanon and France</mods:title>
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</mods:titleInfo>
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<mods:name id="A0824FEE429A8D29FF338896F1D1E7CF" type="personal">
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<mods:role id="4E74E5DB8758E9C0EF70BF17DBCA46E8">
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<mods:roleTerm id="07B280E78979F8BE899EB9AE2999BFED">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="845C32027973E7A8EC124C98B49F5C33">Duy-Jacquemin, Monique Nguyen</mods:namePart>
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<mods:affiliation id="55D4F3FA578C031D731AC311F75D52DB">Histoire naturelle, Département Systématique et Évolution, Arthropodes, CP 53, 61 rue Buffon, F- 75231 Paris cedex 05 (France) monguyen @ mnhn. fr</mods:affiliation>
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<mods:nameIdentifier id="1E96F6BC4A2E5216EAF8D48512E598A4" type="email">monguyen@mnhn.fr</mods:nameIdentifier>
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</mods:name>
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<mods:name id="B2E23339AEDB557733027FFA1F0531BB" type="personal">
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<mods:roleTerm id="5D6372541D778ED458244058BD51A70B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F01AEEB36B2C6D934A4F7CA8A60A8DE4">Azar, Dany</mods:namePart>
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<mods:affiliation id="149C202B1993EC188586A1D995191600">Lebanese University, Faculty of Science II, Biology Department, BP 26110217 Fanar-Matn (Lebanon) and Department, Saint-Joseph University, Campus of Sciences and Technology, Mar Roukos (Mkalles), BP 11 - 1514 Beirut (Lebanon) azar @ mnhn. fr</mods:affiliation>
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<mods:nameIdentifier id="48B35A5D983FF6B322C4040D2259F110" type="email">azar@mnhn.fr</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="AAF14E8EE2FA2CBF669AFD51631B1EE1">text</mods:typeOfResource>
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<mods:titleInfo id="357CC11DE6D16D6B5FED49930560D161">
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<mods:title id="7EE5F4607D76FE85F47C035E32E6D852">Geodiversitas</mods:title>
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</mods:titleInfo>
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<mods:part id="8DD6B44AE2B9F1260145949D8333760C">
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<mods:date id="E91D456391C1E1AED95B08EEB0E85FB2">2004</mods:date>
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<mods:detail id="1B17AD84443812777E33743B42AAB7A4" type="volume">
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<mods:number id="701B00F351A24C2BB0226185497E6B50">26</mods:number>
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</mods:detail>
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<mods:detail id="1709E57F817ED534B21C0EFBF8CD3595" type="issue">
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<mods:number id="1B01F6D6873F52395E5554D966CDE6DB">4</mods:number>
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</mods:detail>
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<mods:extent id="C48B1DA8687CD21574B39E942142C861" unit="page">
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<mods:start id="AFBED940FEF97FC07F3DF04F07DC6C62">631</mods:start>
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<mods:end id="4C2DF1B6A150751243E98CA80A9B0114">641</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="8AA406D7BE65F16A7E79714D000848A7">journal article</mods:classification>
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<mods:identifier id="3401F666E0FEC78088518AEFC42CACBB" type="DOI">10.5281/zenodo.4665318</mods:identifier>
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<mods:identifier id="C3B0225F220C2E0AA772337C36887FFD" type="ISSN">1638-9395</mods:identifier>
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<mods:identifier id="A7983652898B3EF3F8F647C37A8164E1" type="Zenodo-Dep">4665318</mods:identifier>
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</mods:mods>
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<treatment id="D57D87F7FFCEBF3105D0EC3AD071FDA1" LSID="urn:lsid:plazi:treatment:D57D87F7FFCEBF3105D0EC3AD071FDA1" httpUri="http://treatment.plazi.org/id/D57D87F7FFCEBF3105D0EC3AD071FDA1" lastPageId="5" lastPageNumber="636" pageId="4" pageNumber="635">
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<subSubSection id="15CE656AFFCEBF3005D0EC3AD0ACFD61" pageId="4" pageNumber="635" type="nomenclature">
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<paragraph id="5D6B36E1FFCEBF3005D0EC3AD0ACFD61" blockId="4.[676,1209,620,1018]" pageId="4" pageNumber="635">
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<heading id="0623818DFFCEBF3005D0EC3AD0ACFD61" centered="true" fontSize="11" level="2" pageId="4" pageNumber="635" reason="2">
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<taxonomicName id="9AD44D62FFCEBF3005D0EC3AD76CFD41" ID-CoL="6Q8P8" authority="Duy-Jacquemin & Azar, 2004" authorityName="Duy-Jacquemin & Azar" authorityYear="2004" box="[758,1062,621,647]" class="Diplopoda" family="Polyxenidae" genus="Libanoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="635" phylum="Arthropoda" rank="species" species="hammanaensis" status="sp. nov.">
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<emphasis id="6FA0EAF3FFCEBF3005D0EC3AD76CFD41" bold="true" box="[758,1062,621,647]" italics="true" pageId="4" pageNumber="635">Libanoxenus hammanaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="74935788FFCEBF30030BEC3BD722FD40" box="[1069,1128,620,646]" pageId="4" pageNumber="635" rank="species">n. sp.</taxonomicNameLabel>
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(
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<figureCitation id="C5EF2A64FFCEBF300459ECDAD088FD61" box="[895,962,653,679]" captionStart="FIG" captionStartId="5.[113,124,1265,1282]" captionTargetBox="[115,644,204,1227]" captionTargetId="figure-405@5.[115,644,204,1227]" captionTargetPageId="5" captionText="FIG. 3. — Libanoxenus hammanaensis n. gen., n. sp., holotype adult body length: 2.90 mm (633), Azar collection, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4665324" httpUri="https://zenodo.org/record/4665324/files/figure.png" pageId="4" pageNumber="635">Figs 3</figureCitation>
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;
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<figureCitation id="C5EF2A64FFCEBF3004E9ECDAD094FD61" box="[975,990,653,679]" captionStart="FIG" captionStartId="6.[113,124,1580,1597]" captionTargetBox="[123,1201,215,1543]" captionTargetId="figure-49@6.[112,1208,131,1567]" captionTargetPageId="6" captionText="FIG. 4. — A-J, Libanoxenus hammanaensis n. gen., n. sp., holotype adult (633), Azar collection; A, head, dorsal face, dotted line indicates the missing part; B, detail of left antenna; C, labrum, only the visible cuticular spines are represented in two areas; D, right palpus and left middle palpus of gnathochilarium; E, left lateral posterior part of tergite IV; F, left lateral part of tergite III with detail of a trichome; G, right leg IX; H, trochanter, prefemora, femora and tibia of right leg XIII; I, part of tarsus of left leg VIII showing the seta; J, hooked caudal trichome; K, Polyxenus lagurus, distal part of crooked caudal trichomes. Abbreviations: b, sensillum basiconicum; s, setiform sensillum; t, trichobothria. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4665327" httpUri="https://zenodo.org/record/4665327/files/figure.png" pageId="4" pageNumber="635">4</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="15CE656AFFCEBF300582ECE1D1B5FCC6" pageId="4" pageNumber="635" type="materials_examined">
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<paragraph id="5D6B36E1FFCEBF300582ECE1D1B5FCC6" blockId="4.[676,1209,620,1018]" pageId="4" pageNumber="635">
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<materialsCitation id="EDBC3CBCFFCEBF300582ECE1D1B5FCC6" collectionCode="TYPE, MATERIAL, MNHN" pageId="4" pageNumber="635" specimenCount="1" typeStatus="holotype">
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<collectionCode id="3BC5AE24FFCEBF300582ECE1D070FD0A" box="[676,826,694,717]" pageId="4" pageNumber="635">TYPE MATERIAL</collectionCode>
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. —
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<typeStatus id="826F8843FFCEBF300441ECE0D08DFD09" box="[871,967,695,719]" pageId="4" pageNumber="635" type="holotype">Holotype</typeStatus>
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: adult specimen number 633, Azar collection, provisionally deposited in the
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<collectionCode id="3BC5AE24FFCEBF300582ECBED1B0FCC6" box="[676,762,745,768]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="635">MNHN</collectionCode>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="15CE656AFFCEBF300582ED46D7D0FCEC" box="[676,1178,785,810]" pageId="4" pageNumber="635" type="etymology">
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<paragraph id="5D6B36E1FFCEBF300582ED46D7D0FCEC" blockId="4.[676,1209,620,1018]" box="[676,1178,785,810]" pageId="4" pageNumber="635">
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ETYMOLOGY. — After Hammana, the
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<typeStatus id="826F8843FFCEBF30033AED46D70CFCEF" box="[1052,1094,785,809]" pageId="4" pageNumber="635">type</typeStatus>
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locality.
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</paragraph>
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</subSubSection>
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<subSubSection id="15CE656AFFCEBF300582ED6DD710FC17" pageId="4" pageNumber="635" type="materials_examined">
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<paragraph id="5D6B36E1FFCEBF300582ED6DD710FC17" blockId="4.[676,1209,620,1018]" pageId="4" pageNumber="635">
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<collectionCode id="3BC5AE24FFCEBF300582ED6DD035FC96" box="[676,895,826,849]" pageId="4" pageNumber="635">
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<typeStatus id="826F8843FFCEBF300582ED6DD193FC89" box="[676,729,826,849]" pageId="4" pageNumber="635">TYPE</typeStatus>
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LOCALITY AND
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</collectionCode>
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<materialsCitation id="EDBC3CBCFFCEBF3004AFED68D0AAFC43" collectionCode="HORIZON" country="Lebanon" county="Central" location="Mdeirij" municipality="Mouhafazit Jabal Loubnan" pageId="4" pageNumber="635" specimenCount="1" typeStatus="holotype">
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<typeStatus id="826F8843FFCEBF3004AFED68D0F0FC89" box="[905,954,831,847]" pageId="4" pageNumber="635">TYPE</typeStatus>
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<collectionCode id="3BC5AE24FFCEBF3004E2ED68D762FC96" box="[964,1064,831,848]" pageId="4" pageNumber="635">HORIZON</collectionCode>
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. —
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<location id="580B603AFFCEBF30037BED6CD7FBFC95" LSID="urn:lsid:plazi:treatment:D57D87F7FFCEBF3105D0EC3AD071FDA1:580B603AFFCEBF30037BED6CD7FBFC95" box="[1117,1201,827,851]" country="Lebanon" county="Central" municipality="Mouhafazit Jabal Loubnan" name="Mdeirij" pageId="4" pageNumber="635">Mdeirij</location>
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/
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<location id="580B603AFFCEBF300582ED04D046FCAC" LSID="urn:lsid:plazi:treatment:D57D87F7FFCEBF3105D0EC3AD071FDA1:580B603AFFCEBF300582ED04D046FCAC" box="[676,780,851,874]" country="Lebanon" county="Central" municipality="Mouhafazit Jabal Loubnan" name="Hammana" pageId="4" pageNumber="635">Hammana</location>
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locality,
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<location id="580B603AFFCEBF30044FED03D74FFCAA" LSID="urn:lsid:plazi:treatment:D57D87F7FFCEBF3105D0EC3AD071FDA1:580B603AFFCEBF30044FED03D74FFCAA" box="[873,1029,852,876]" country="Lebanon" county="Central" municipality="Mouhafazit Jabal Loubnan" name="Caza of Baabda" pageId="4" pageNumber="635">Caza of Baabda</location>
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,
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<collectingMunicipality id="BD0FAC9BFFCEBF300336ED03D046FC43" pageId="4" pageNumber="635">Mouhafazit Jabal Loubnan</collectingMunicipality>
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,
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<collectingCounty id="B40A4E6DFFCEBF30043DED3AD03AFC43" box="[795,880,877,901]" pageId="4" pageNumber="635">Central</collectingCounty>
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<collectingCountry id="25C37671FFCEBF30045DED3AD0AAFC43" box="[891,992,877,901]" name="Lebanon" pageId="4" pageNumber="635">Lebanon</collectingCountry>
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</materialsCitation>
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;
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<materialsCitation id="EDBC3CBCFFCEBF3004C9ED3AD71CFC17" country="Canada" location="lower Aptian" pageId="4" pageNumber="635" specimenCount="1" typeStatus="holotype">
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Early Cretaceous, Valanginian-Hauterivian to
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<location id="580B603AFFCEBF3004C5EDD0D739FC59" LSID="urn:lsid:plazi:treatment:D57D87F7FFCEBF3105D0EC3AD071FDA1:580B603AFFCEBF3004C5EDD0D739FC59" box="[995,1139,903,927]" country="Canada" name="lower Aptian" pageId="4" pageNumber="635">lower Aptian</location>
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(135- 125 million years). The amber piece is mounted on a microscope slide in
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<collectingCountry id="25C37671FFCEBF300440EDEED0FBFC17" box="[870,945,953,977]" name="Canada" pageId="4" pageNumber="635">Canada</collectingCountry>
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balsam medium
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="15CE656AFFCEBF300582EDB6D0F6FC3F" box="[676,956,993,1018]" pageId="4" pageNumber="635" type="diagnosis">
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<paragraph id="5D6B36E1FFCEBF300582EDB6D0F6FC3F" blockId="4.[676,1209,620,1018]" box="[676,956,993,1018]" pageId="4" pageNumber="635">DIAGNOSIS. — As for genus.</paragraph>
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</subSubSection>
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<subSubSection id="15CE656AFFCEBF310582EA7BD071FDA1" lastPageId="5" lastPageNumber="636" pageId="4" pageNumber="635" type="description">
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<paragraph id="5D6B36E1FFCEBF300582EA7BD07FFB82" blockId="4.[676,1210,1068,1703]" box="[676,821,1068,1094]" pageId="4" pageNumber="635">DESCRIPTION</paragraph>
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<paragraph id="5D6B36E1FFCEBF300582EA1AD03FFBA0" blockId="4.[676,1210,1068,1703]" box="[676,885,1100,1127]" pageId="4" pageNumber="635">
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<emphasis id="6FA0EAF3FFCEBF300582EA1AD03FFBA0" box="[676,885,1100,1127]" italics="true" pageId="4" pageNumber="635">
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Adult (
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<figureCitation id="C5EF2A64FFCEBF3005CBEA1BD067FBA0" box="[749,813,1100,1126]" captionStart="FIG" captionStartId="5.[113,124,1265,1282]" captionTargetBox="[115,644,204,1227]" captionTargetId="figure-405@5.[115,644,204,1227]" captionTargetPageId="5" captionText="FIG. 3. — Libanoxenus hammanaensis n. gen., n. sp., holotype adult body length: 2.90 mm (633), Azar collection, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4665324" httpUri="https://zenodo.org/record/4665324/files/figure.png" pageId="4" pageNumber="635">Figs 3</figureCitation>
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;
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<figureCitation id="C5EF2A64FFCEBF30041DEA1BD00DFBA0" box="[827,839,1100,1126]" captionStart="FIG" captionStartId="6.[113,124,1580,1597]" captionTargetBox="[123,1201,215,1543]" captionTargetId="figure-49@6.[112,1208,131,1567]" captionTargetPageId="6" captionText="FIG. 4. — A-J, Libanoxenus hammanaensis n. gen., n. sp., holotype adult (633), Azar collection; A, head, dorsal face, dotted line indicates the missing part; B, detail of left antenna; C, labrum, only the visible cuticular spines are represented in two areas; D, right palpus and left middle palpus of gnathochilarium; E, left lateral posterior part of tergite IV; F, left lateral part of tergite III with detail of a trichome; G, right leg IX; H, trochanter, prefemora, femora and tibia of right leg XIII; I, part of tarsus of left leg VIII showing the seta; J, hooked caudal trichome; K, Polyxenus lagurus, distal part of crooked caudal trichomes. Abbreviations: b, sensillum basiconicum; s, setiform sensillum; t, trichobothria. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4665327" httpUri="https://zenodo.org/record/4665327/files/figure.png" pageId="4" pageNumber="635">4</figureCitation>
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A-J)
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</emphasis>
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</paragraph>
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<paragraph id="5D6B36E1FFCEBF300582EA3AD093FB01" blockId="4.[676,1210,1068,1703]" pageId="4" pageNumber="635">
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The trunk of the
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<typeStatus id="826F8843FFCEBF300452EA3AD093FB41" box="[884,985,1133,1159]" pageId="4" pageNumber="635" type="holotype">holotype</typeStatus>
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is open over all its length along the axis of symmetry, hence the tergites are not seen entirely.
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</paragraph>
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<paragraph id="5D6B36E1FFCEBF300582EA9BD750FAE0" blockId="4.[676,1210,1068,1703]" pageId="4" pageNumber="635">
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<emphasis id="6FA0EAF3FFCEBF300582EA9BD007FB20" bold="true" box="[676,845,1228,1254]" pageId="4" pageNumber="635">Measurements.</emphasis>
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Body length =
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<quantity id="9A2C9B04FFCEBF3004DDEA9AD72EFB20" box="[1019,1124,1228,1255]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="635" unit="mm" value="2.5">2.50 mm</quantity>
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; penicil length =
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<quantity id="9A2C9B04FFCEBF300435EABAD0C9FAC0" box="[787,899,1260,1287]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="635" unit="mm" value="0.3">0.30 mm</quantity>
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; head trichomes length =
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<quantity id="9A2C9B04FFCEBF300582EB5AD041FAE0" box="[676,779,1292,1319]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="635" unit="mm" value="0.1">0.10 mm</quantity>
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; total length =
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<quantity id="9A2C9B04FFCEBF300488EB5AD75CFAE0" box="[942,1046,1292,1319]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="4" pageNumber="635" unit="mm" value="2.9">2.90 mm</quantity>
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.
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</paragraph>
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<paragraph id="5D6B36E1FFCEBF300582EB7BD079F961" blockId="4.[676,1210,1068,1703]" pageId="4" pageNumber="635">
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<emphasis id="6FA0EAF3FFCEBF300582EB7BD1A0FA80" bold="true" box="[676,746,1324,1350]" pageId="4" pageNumber="635">Head.</emphasis>
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Eight ocelli (
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<figureCitation id="C5EF2A64FFCEBF3004A5EB7AD096FA81" box="[899,988,1325,1351]" captionStart="FIG" captionStartId="6.[113,124,1580,1597]" captionTargetBox="[123,1201,215,1543]" captionTargetId="figure-49@6.[112,1208,131,1567]" captionTargetPageId="6" captionText="FIG. 4. — A-J, Libanoxenus hammanaensis n. gen., n. sp., holotype adult (633), Azar collection; A, head, dorsal face, dotted line indicates the missing part; B, detail of left antenna; C, labrum, only the visible cuticular spines are represented in two areas; D, right palpus and left middle palpus of gnathochilarium; E, left lateral posterior part of tergite IV; F, left lateral part of tergite III with detail of a trichome; G, right leg IX; H, trochanter, prefemora, femora and tibia of right leg XIII; I, part of tarsus of left leg VIII showing the seta; J, hooked caudal trichome; K, Polyxenus lagurus, distal part of crooked caudal trichomes. Abbreviations: b, sensillum basiconicum; s, setiform sensillum; t, trichobothria. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4665327" httpUri="https://zenodo.org/record/4665327/files/figure.png" pageId="4" pageNumber="635">Fig. 4A</figureCitation>
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). Left antenna with eight articles (
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<figureCitation id="C5EF2A64FFCEBF300472EB1AD0FDFAA1" box="[852,951,1357,1383]" captionStart="FIG" captionStartId="6.[113,124,1580,1597]" captionTargetBox="[123,1201,215,1543]" captionTargetId="figure-49@6.[112,1208,131,1567]" captionTargetPageId="6" captionText="FIG. 4. — A-J, Libanoxenus hammanaensis n. gen., n. sp., holotype adult (633), Azar collection; A, head, dorsal face, dotted line indicates the missing part; B, detail of left antenna; C, labrum, only the visible cuticular spines are represented in two areas; D, right palpus and left middle palpus of gnathochilarium; E, left lateral posterior part of tergite IV; F, left lateral part of tergite III with detail of a trichome; G, right leg IX; H, trochanter, prefemora, femora and tibia of right leg XIII; I, part of tarsus of left leg VIII showing the seta; J, hooked caudal trichome; K, Polyxenus lagurus, distal part of crooked caudal trichomes. Abbreviations: b, sensillum basiconicum; s, setiform sensillum; t, trichobothria. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4665327" httpUri="https://zenodo.org/record/4665327/files/figure.png" pageId="4" pageNumber="635">Fig. 4A</figureCitation>
|
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). Length to diameter ratio of the sixth antennal article = 1.80. Only the end of one sensillum basiconicum is visible on articles VI and VII (
|
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<figureCitation id="C5EF2A64FFCEBF30048BEBFAD74CFA01" box="[941,1030,1453,1479]" captionStart="FIG" captionStartId="6.[113,124,1580,1597]" captionTargetBox="[123,1201,215,1543]" captionTargetId="figure-49@6.[112,1208,131,1567]" captionTargetPageId="6" captionText="FIG. 4. — A-J, Libanoxenus hammanaensis n. gen., n. sp., holotype adult (633), Azar collection; A, head, dorsal face, dotted line indicates the missing part; B, detail of left antenna; C, labrum, only the visible cuticular spines are represented in two areas; D, right palpus and left middle palpus of gnathochilarium; E, left lateral posterior part of tergite IV; F, left lateral part of tergite III with detail of a trichome; G, right leg IX; H, trochanter, prefemora, femora and tibia of right leg XIII; I, part of tarsus of left leg VIII showing the seta; J, hooked caudal trichome; K, Polyxenus lagurus, distal part of crooked caudal trichomes. Abbreviations: b, sensillum basiconicum; s, setiform sensillum; t, trichobothria. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4665327" httpUri="https://zenodo.org/record/4665327/files/figure.png" pageId="4" pageNumber="635">Fig. 4B</figureCitation>
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); some setiform sensilla are seen on article III, V, VI and VII; right antenna with four articles, the article V is broken. Labrum with numerous small setae (
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<figureCitation id="C5EF2A64FFCEBF30058AE87AD042F981" box="[684,776,1581,1607]" captionStart="FIG" captionStartId="6.[113,124,1580,1597]" captionTargetBox="[123,1201,215,1543]" captionTargetId="figure-49@6.[112,1208,131,1567]" captionTargetPageId="6" captionText="FIG. 4. — A-J, Libanoxenus hammanaensis n. gen., n. sp., holotype adult (633), Azar collection; A, head, dorsal face, dotted line indicates the missing part; B, detail of left antenna; C, labrum, only the visible cuticular spines are represented in two areas; D, right palpus and left middle palpus of gnathochilarium; E, left lateral posterior part of tergite IV; F, left lateral part of tergite III with detail of a trichome; G, right leg IX; H, trochanter, prefemora, femora and tibia of right leg XIII; I, part of tarsus of left leg VIII showing the seta; J, hooked caudal trichome; K, Polyxenus lagurus, distal part of crooked caudal trichomes. Abbreviations: b, sensillum basiconicum; s, setiform sensillum; t, trichobothria. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4665327" httpUri="https://zenodo.org/record/4665327/files/figure.png" pageId="4" pageNumber="635">Fig. 4C</figureCitation>
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). Outer right palp of gnathochilarium with 12 or 13 sensilla (
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<figureCitation id="C5EF2A64FFCEBF3004E2E81AD760F9A1" box="[964,1066,1613,1639]" captionStart="FIG" captionStartId="6.[113,124,1580,1597]" captionTargetBox="[123,1201,215,1543]" captionTargetId="figure-49@6.[112,1208,131,1567]" captionTargetPageId="6" captionText="FIG. 4. — A-J, Libanoxenus hammanaensis n. gen., n. sp., holotype adult (633), Azar collection; A, head, dorsal face, dotted line indicates the missing part; B, detail of left antenna; C, labrum, only the visible cuticular spines are represented in two areas; D, right palpus and left middle palpus of gnathochilarium; E, left lateral posterior part of tergite IV; F, left lateral part of tergite III with detail of a trichome; G, right leg IX; H, trochanter, prefemora, femora and tibia of right leg XIII; I, part of tarsus of left leg VIII showing the seta; J, hooked caudal trichome; K, Polyxenus lagurus, distal part of crooked caudal trichomes. Abbreviations: b, sensillum basiconicum; s, setiform sensillum; t, trichobothria. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4665327" httpUri="https://zenodo.org/record/4665327/files/figure.png" pageId="4" pageNumber="635">Fig. 4D</figureCitation>
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); it is more than twice as long as the diameter of the middle palp.
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</paragraph>
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<caption id="09AB6669FFCFBF310757EAA6D133FAD1" ID-DOI="http://doi.org/10.5281/zenodo.4665324" ID-Zenodo-Dep="4665324" httpUri="https://zenodo.org/record/4665324/files/figure.png" pageId="5" pageNumber="636" startId="5.[113,124,1265,1282]" targetBox="[115,644,204,1227]" targetPageId="5">
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<paragraph id="5D6B36E1FFCFBF310757EAA6D133FAD1" blockId="5.[113,646,1265,1303]" pageId="5" pageNumber="636">
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FIG. 3. —
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<taxonomicName id="9AD44D62FFCFBF3107E0EAA6D2F9FAC4" authority="Duy-Jacquemin & Azar, 2004" authorityName="Duy-Jacquemin & Azar" authorityYear="2004" box="[198,435,1265,1282]" class="Diplopoda" family="Polyxenidae" genus="Libanoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polyxenida" pageId="5" pageNumber="636" phylum="Arthropoda" rank="species" species="hammanaensis" status="n. gen., n. sp.">
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<emphasis id="6FA0EAF3FFCFBF3107E0EAA6D2F9FAC4" box="[198,435,1265,1282]" italics="true" pageId="5" pageNumber="636">Libanoxenus hammanaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="74935788FFCFBF31069CEAA6D17FFAC4" box="[442,565,1265,1282]" pageId="5" pageNumber="636" rank="species">n. gen., n. sp.</taxonomicNameLabel>
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, holotype adult body length: 2.90 mm (633), Azar collection, dorsal view.
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</paragraph>
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</caption>
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<paragraph id="5D6B36E1FFCFBF310757EB3BD043FDC1" blockId="5.[113,646,1388,1703]" lastBlockId="5.[676,1209,205,519]" pageId="5" pageNumber="636">
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<emphasis id="6FA0EAF3FFCFBF310757EB3BD388FA40" bold="true" box="[113,194,1388,1414]" pageId="5" pageNumber="636">Trunk.</emphasis>
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The tergal trichomes (seen only on the lateral part of tergites), are short (
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<figureCitation id="C5EF2A64FFCFBF3106DBEBDAD132FA60" box="[509,632,1420,1447]" captionStart="FIG" captionStartId="6.[113,124,1580,1597]" captionTargetBox="[123,1201,215,1543]" captionTargetId="figure-49@6.[112,1208,131,1567]" captionTargetPageId="6" captionText="FIG. 4. — A-J, Libanoxenus hammanaensis n. gen., n. sp., holotype adult (633), Azar collection; A, head, dorsal face, dotted line indicates the missing part; B, detail of left antenna; C, labrum, only the visible cuticular spines are represented in two areas; D, right palpus and left middle palpus of gnathochilarium; E, left lateral posterior part of tergite IV; F, left lateral part of tergite III with detail of a trichome; G, right leg IX; H, trochanter, prefemora, femora and tibia of right leg XIII; I, part of tarsus of left leg VIII showing the seta; J, hooked caudal trichome; K, Polyxenus lagurus, distal part of crooked caudal trichomes. Abbreviations: b, sensillum basiconicum; s, setiform sensillum; t, trichobothria. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4665327" httpUri="https://zenodo.org/record/4665327/files/figure.png" pageId="5" pageNumber="636">Fig. 4E, F</figureCitation>
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); the longer lateral trichomes of tergite reach
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|
<quantity id="9A2C9B04FFCFBF31057DEBFAD3A4FA20" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="5" pageNumber="636" unit="mm" value="125.0" valueMax="130.0" valueMin="120.0">120 to 130 µm</quantity>
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while in the middle of the tergite the trichomes measure
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<quantity id="9A2C9B04FFCFBF310652EBBAD149F9C0" box="[372,515,1516,1543]" metricMagnitude="-2" metricUnit="m" metricValue="5.15" metricValueMax="5.3" metricValueMin="5.0" pageId="5" pageNumber="636" unit="mm" value="51.5" valueMax="53.0" valueMin="50.0">50 to 53 µm</quantity>
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. They seem to be arranged in two rows: one anterior row with trichomes directed anteriad and one posterior row with trichomes directed caudal (
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<figureCitation id="C5EF2A64FFCFBF31075FE83AD399F941" box="[121,211,1645,1671]" captionStart="FIG" captionStartId="6.[113,124,1580,1597]" captionTargetBox="[123,1201,215,1543]" captionTargetId="figure-49@6.[112,1208,131,1567]" captionTargetPageId="6" captionText="FIG. 4. — A-J, Libanoxenus hammanaensis n. gen., n. sp., holotype adult (633), Azar collection; A, head, dorsal face, dotted line indicates the missing part; B, detail of left antenna; C, labrum, only the visible cuticular spines are represented in two areas; D, right palpus and left middle palpus of gnathochilarium; E, left lateral posterior part of tergite IV; F, left lateral part of tergite III with detail of a trichome; G, right leg IX; H, trochanter, prefemora, femora and tibia of right leg XIII; I, part of tarsus of left leg VIII showing the seta; J, hooked caudal trichome; K, Polyxenus lagurus, distal part of crooked caudal trichomes. Abbreviations: b, sensillum basiconicum; s, setiform sensillum; t, trichobothria. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4665327" httpUri="https://zenodo.org/record/4665327/files/figure.png" pageId="5" pageNumber="636">Fig. 4F</figureCitation>
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). One clearly visible seta on tarsus II (
|
|
<figureCitation id="C5EF2A64FFCFBF31075FE8DAD3A5F960" box="[121,239,1676,1703]" captionStart="FIG" captionStartId="6.[113,124,1580,1597]" captionTargetBox="[123,1201,215,1543]" captionTargetId="figure-49@6.[112,1208,131,1567]" captionTargetPageId="6" captionText="FIG. 4. — A-J, Libanoxenus hammanaensis n. gen., n. sp., holotype adult (633), Azar collection; A, head, dorsal face, dotted line indicates the missing part; B, detail of left antenna; C, labrum, only the visible cuticular spines are represented in two areas; D, right palpus and left middle palpus of gnathochilarium; E, left lateral posterior part of tergite IV; F, left lateral part of tergite III with detail of a trichome; G, right leg IX; H, trochanter, prefemora, femora and tibia of right leg XIII; I, part of tarsus of left leg VIII showing the seta; J, hooked caudal trichome; K, Polyxenus lagurus, distal part of crooked caudal trichomes. Abbreviations: b, sensillum basiconicum; s, setiform sensillum; t, trichobothria. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4665327" httpUri="https://zenodo.org/record/4665327/files/figure.png" pageId="5" pageNumber="636">Fig. 4G, I</figureCitation>
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), nude setae (
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|
<figureCitation id="C5EF2A64FFCFBF3106B6E8DAD2A6F961" box="[400,492,1677,1703]" captionStart="FIG" captionStartId="6.[113,124,1580,1597]" captionTargetBox="[123,1201,215,1543]" captionTargetId="figure-49@6.[112,1208,131,1567]" captionTargetPageId="6" captionText="FIG. 4. — A-J, Libanoxenus hammanaensis n. gen., n. sp., holotype adult (633), Azar collection; A, head, dorsal face, dotted line indicates the missing part; B, detail of left antenna; C, labrum, only the visible cuticular spines are represented in two areas; D, right palpus and left middle palpus of gnathochilarium; E, left lateral posterior part of tergite IV; F, left lateral part of tergite III with detail of a trichome; G, right leg IX; H, trochanter, prefemora, femora and tibia of right leg XIII; I, part of tarsus of left leg VIII showing the seta; J, hooked caudal trichome; K, Polyxenus lagurus, distal part of crooked caudal trichomes. Abbreviations: b, sensillum basiconicum; s, setiform sensillum; t, trichobothria. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4665327" httpUri="https://zenodo.org/record/4665327/files/figure.png" pageId="5" pageNumber="636">Fig. 4H</figureCitation>
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) observed on other article (coxa, trochanter, prefemur, femur and tibia). Telotarsus with clearly visible poste- rior lamellate process longer than claw (
|
|
<figureCitation id="C5EF2A64FFCFBF31058AEF7AD04DFE81" box="[684,775,301,327]" captionStart="FIG" captionStartId="6.[113,124,1580,1597]" captionTargetBox="[123,1201,215,1543]" captionTargetId="figure-49@6.[112,1208,131,1567]" captionTargetPageId="6" captionText="FIG. 4. — A-J, Libanoxenus hammanaensis n. gen., n. sp., holotype adult (633), Azar collection; A, head, dorsal face, dotted line indicates the missing part; B, detail of left antenna; C, labrum, only the visible cuticular spines are represented in two areas; D, right palpus and left middle palpus of gnathochilarium; E, left lateral posterior part of tergite IV; F, left lateral part of tergite III with detail of a trichome; G, right leg IX; H, trochanter, prefemora, femora and tibia of right leg XIII; I, part of tarsus of left leg VIII showing the seta; J, hooked caudal trichome; K, Polyxenus lagurus, distal part of crooked caudal trichomes. Abbreviations: b, sensillum basiconicum; s, setiform sensillum; t, trichobothria. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4665327" httpUri="https://zenodo.org/record/4665327/files/figure.png" pageId="5" pageNumber="636">Fig. 4G</figureCitation>
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). Dorsal face of telson with an anterior row including three to four visible barbate trichomes. Telson is not of
|
|
<taxonomicName id="9AD44D62FFCFBF31032DEF3AD7CBFE41" authorityName="Latreille" authorityYear="1803" box="[1035,1153,365,391]" class="Diplopoda" family="Polyxenidae" genus="Polyxenus" kingdom="Animalia" order="Polyxenida" pageId="5" pageNumber="636" phylum="Arthropoda" rank="genus">
|
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<emphasis id="6FA0EAF3FFCFBF31032DEF3AD7CBFE41" box="[1035,1153,365,391]" italics="true" pageId="5" pageNumber="636">Polyxenus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="9AD44D62FFCFBF310582EFDAD07AFE61" authorityName="Silvestri" authorityYear="1948" box="[676,816,397,423]" class="Diplopoda" family="Polyxenidae" genus="Propolyxenus" kingdom="Animalia" order="Polyxenida" pageId="5" pageNumber="636" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="6FA0EAF3FFCFBF310582EFDAD07AFE61" box="[676,816,397,423]" italics="true" pageId="5" pageNumber="636">Propolyxenus</emphasis>
|
|
</taxonomicName>
|
|
<typeStatus id="826F8843FFCFBF31041CEFDBD020FE60" box="[826,874,396,422]" pageId="5" pageNumber="636">type</typeStatus>
|
|
(latter with trichomes ended by multiple hooks:
|
|
<figureCitation id="C5EF2A64FFCFBF3104A5EFFAD096FE01" box="[899,988,429,455]" captionStart="FIG" captionStartId="6.[113,124,1580,1597]" captionTargetBox="[123,1201,215,1543]" captionTargetId="figure-49@6.[112,1208,131,1567]" captionTargetPageId="6" captionText="FIG. 4. — A-J, Libanoxenus hammanaensis n. gen., n. sp., holotype adult (633), Azar collection; A, head, dorsal face, dotted line indicates the missing part; B, detail of left antenna; C, labrum, only the visible cuticular spines are represented in two areas; D, right palpus and left middle palpus of gnathochilarium; E, left lateral posterior part of tergite IV; F, left lateral part of tergite III with detail of a trichome; G, right leg IX; H, trochanter, prefemora, femora and tibia of right leg XIII; I, part of tarsus of left leg VIII showing the seta; J, hooked caudal trichome; K, Polyxenus lagurus, distal part of crooked caudal trichomes. Abbreviations: b, sensillum basiconicum; s, setiform sensillum; t, trichobothria. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4665327" httpUri="https://zenodo.org/record/4665327/files/figure.png" pageId="5" pageNumber="636">Fig. 4K</figureCitation>
|
|
); trichomes of caudal penicil with three or four aligned hooks (
|
|
<figureCitation id="C5EF2A64FFCFBF31058AEFBAD1B6FDC1" box="[684,764,493,519]" captionStart="FIG" captionStartId="6.[113,124,1580,1597]" captionTargetBox="[123,1201,215,1543]" captionTargetId="figure-49@6.[112,1208,131,1567]" captionTargetPageId="6" captionText="FIG. 4. — A-J, Libanoxenus hammanaensis n. gen., n. sp., holotype adult (633), Azar collection; A, head, dorsal face, dotted line indicates the missing part; B, detail of left antenna; C, labrum, only the visible cuticular spines are represented in two areas; D, right palpus and left middle palpus of gnathochilarium; E, left lateral posterior part of tergite IV; F, left lateral part of tergite III with detail of a trichome; G, right leg IX; H, trochanter, prefemora, femora and tibia of right leg XIII; I, part of tarsus of left leg VIII showing the seta; J, hooked caudal trichome; K, Polyxenus lagurus, distal part of crooked caudal trichomes. Abbreviations: b, sensillum basiconicum; s, setiform sensillum; t, trichobothria. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4665327" httpUri="https://zenodo.org/record/4665327/files/figure.png" pageId="5" pageNumber="636">Fig. 4J</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="5D6B36E1FFCFBF310582EC1AD071FDA1" blockId="5.[676,827,589,615]" box="[676,827,589,615]" pageId="5" pageNumber="636">French amber</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |