304 lines
41 KiB
XML
304 lines
41 KiB
XML
<document id="9F2F251BF520E8AF72925744F15806C9" ID-DOI="10.5281/zenodo.185751" ID-GBIF-Dataset="b6e978e1-3073-4192-88d1-e631ed77a26d" ID-ISSN="1175-5326" ID-Zenodo-Dep="185751" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460013875103" checkinUser="plazi" docAuthor="Schmidt, Joachim & Hartmann, Matthias" docDate="2009" docId="D5727D136368FFE5FF20F927FCEDD82D" docLanguage="en" docName="zt02009p026.pdf" docOrigin="Zootaxa 2009" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pristosia atrema Andrewes 1926" docType="treatment" docVersion="9" lastPageNumber="12" masterDocId="294B056B6361FFEEFFB7FFEEFF94DE0F" masterDocTitle="Pristosia Motschulsky, 1865 from the Nepal Himalaya: Taxonomy and Biogeography (Coleoptera: Carabidae: Sphodrini)" masterLastPageNumber="26" masterPageNumber="1" pageNumber="10" updateTime="1698233570847" updateUser="plazi">
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<mods:title id="A8B627116E664D05BDAD926A78F90176">Pristosia Motschulsky, 1865 from the Nepal Himalaya: Taxonomy and Biogeography (Coleoptera: Carabidae: Sphodrini)</mods:title>
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<mods:namePart id="99C94F507D11DD922B82B02595898931">Schmidt, Joachim</mods:namePart>
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<mods:namePart id="198FC3B22A07BF9C5D454BCE5A817B9D">Hartmann, Matthias</mods:namePart>
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<mods:date id="1998E7C6F628956153B59850C64D6F41">2009</mods:date>
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<treatment id="D5727D136368FFE5FF20F927FCEDD82D" ID-DOI="http://doi.org/10.5281/zenodo.6222560" ID-GBIF-Taxon="119381560" ID-Zenodo-Dep="6222560" LSID="urn:lsid:plazi:treatment:D5727D136368FFE5FF20F927FCEDD82D" httpUri="http://treatment.plazi.org/id/D5727D136368FFE5FF20F927FCEDD82D" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10">
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<subSubSection id="15C19F8E6368FFE7FF20F927FEC1D909" pageId="9" pageNumber="10" type="nomenclature">
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<paragraph id="5D64CC056368FFE7FF20F927FDA9D8EC" blockId="9.[151,573,1737,1798]" box="[151,573,1737,1763]" pageId="9" pageNumber="10">
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<heading id="062C7B696368FFE7FF20F927FDA9D8EC" bold="true" box="[151,573,1737,1763]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
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<taxonomicName id="9ADBB7866368FFE7FF20F927FDA9D8EC" ID-CoL="8M87R" authority="Andrewes, 1926" authorityName="Andrewes" authorityYear="1926" box="[151,573,1737,1763]" class="Insecta" family="Carabidae" genus="Pristosia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="atrema">
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<emphasis id="6FAF10176368FFE7FF20F927FDA9D8EC" bold="true" box="[151,573,1737,1763]" pageId="9" pageNumber="10">
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<emphasis id="6FAF10176368FFE7FF20F927FEC9D8EC" bold="true" box="[151,349,1737,1763]" italics="true" pageId="9" pageNumber="10">Pristosia atrema</emphasis>
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(
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<bibRefCitation id="394AB1F46368FFE7FEDBF927FDA1D8EC" author="Andrewes" box="[364,565,1737,1763]" pageId="9" pageNumber="25" refString="Andrewes, H. E. (1926) On a collection of Carabidae from the Kumaon-Tibetan frontier. The Entomologist's Monthly Magazine, 62, 65 - 80." type="journal article" year="1926">Andrewes, 1926</bibRefCitation>
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)
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</emphasis>
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||
</taxonomicName>
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||
</heading>
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</paragraph>
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<paragraph id="5D64CC056368FFE7FF20F902FEC1D909" blockId="9.[151,573,1737,1798]" box="[151,341,1772,1798]" pageId="9" pageNumber="10">
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<figureCitation id="C5E0D0806368FFE7FF20F902FF63D909" box="[151,247,1772,1798]" captionStart="FIGURES 5 – 15" captionStartId="6.[151,272,1702,1726]" captionTargetBox="[177,1434,196,1674]" captionTargetId="figure@6.[151,1436,194,1678]" captionTargetPageId="6" captionText="FIGURES 5 – 15. Pristosia spp., pronotum. Fig. 5, P. dahud polita ssp. n., Paratype, male. Fig. 6, P. dahud Morvan, 1994, non-type, male, Rara Lake. Fig. 7, P. dahud Morvan, 1994, non-type, male, Maharigaon. Fig. 8, P. dahud Morvan, 1994, non-type, male, Khari Lagna (transitional form). Fig. 9, P. dahud Morvan, 1994, non-type, male, Khari Lagna (typical form). Fig. 10, P. similata sp. n., Paratype, male. Fig. 11, P. glabella sp. n., Paratype, male. Fig. 12, P. amaroides (Putzeys, 1877), non-type, female, India, Darjeeling. Fig. 13, P. a t re m a (Andrewes, 1926), Holotype. Fig. 14, P. nepalensis sp. n., Paratype, male. Fig. 15, P. championi (Andrewes, 1934), Holotype. Scale bar = 2 mm." httpUri="https://zenodo.org/record/185753/files/figure.png" pageId="9" pageNumber="10">Figs. 13</figureCitation>
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,
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<figureCitation id="C5E0D0806368FFE7FEB3F902FEC5D909" box="[260,337,1772,1798]" captionStart="FIGURES 24 – 29" captionStartId="10.[151,269,1393,1417]" captionTargetBox="[179,1406,500,1358]" captionTargetId="figure@10.[151,1436,475,1427]" captionTargetPageId="10" captionText="FIGURES 24 – 29. Pristosia spp., aedeagal median lobe, dorsal view (Figs. 24, 26, 28) and left lateral view (Figs. 25, 27, 29, the internal sac is figured only in dorsal view). Figs. 24, 25, P. s i m i l a t a sp. n., Paratype. Figs. 26, 27, P. a trema (Andrewes, 1926), Holotype. Figs. 28, 29, P. a t re m a (Andrewes, 1926), non-type, India, Pindar Valley. Scale bar = 1 mm." httpUri="https://zenodo.org/record/185755/files/figure.png" pageId="9" pageNumber="10">26–29</figureCitation>
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.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="15C19F8E6368FFE7FF20F8D7FD32D9DF" pageId="9" pageNumber="10" type="description">
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<paragraph id="5D64CC056368FFE7FF20F8D7FD8CD97F" blockId="9.[151,1436,1849,2000]" pageId="9" pageNumber="10">
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Catalogue:
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<taxonomicName id="9ADBB7866368FFE7FEA5F8D7FD3FD95F" authority="Andrewes, 1926: 77" authorityName="Andrewes" authorityPageNumber="77" authorityYear="1926" box="[274,683,1849,1872]" class="Insecta" family="Carabidae" genus="Calathus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="atrema">
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<emphasis id="6FAF10176368FFE7FEA5F8D7FE52D95F" box="[274,454,1849,1872]" italics="true" pageId="9" pageNumber="10">Calathus atrema</emphasis>
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<bibRefCitation id="394AB1F46368FFE7FE7AF8D7FD15D95F" author="Andrewes" box="[461,641,1849,1872]" pageId="9" pageNumber="25" refString="Andrewes, H. E. (1926) On a collection of Carabidae from the Kumaon-Tibetan frontier. The Entomologist's Monthly Magazine, 62, 65 - 80." type="journal article" year="1926">Andrewes, 1926</bibRefCitation>
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: 77
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||
</taxonomicName>
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||
.
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<typeStatus id="826072A76368FFE7FD01F8D7FD7FD95F" box="[694,747,1849,1872]" pageId="9" pageNumber="10">Type</typeStatus>
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locality: N
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<collectingCountry id="25CC8C956368FFE7FCD9F8D7FC33D95F" box="[878,935,1849,1872]" name="India" pageId="9" pageNumber="10">India</collectingCountry>
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, Kumaon Himalaya, Uttaranchal district, Gori Valley, Burphu, ca. 3500 mNN.
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</paragraph>
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<paragraph id="5D64CC056368FFE7FF20F897FD2CD99F" blockId="9.[151,1436,1849,2000]" box="[151,696,1913,1936]" pageId="9" pageNumber="10">
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<treatmentCitationGroup id="7DCBEB2B6368FFE7FF20F897FD2CD99F" box="[151,696,1913,1936]" pageId="9" pageNumber="10">
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<taxonomicName id="9ADBB7866368FFE7FF20F897FE44D99F" ID-CoL="8M87R" authority="Andrewes" authorityName="Andrewes" box="[151,464,1913,1936]" class="Insecta" family="Carabidae" genus="Pristosia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="atrema">
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<emphasis id="6FAF10176368FFE7FF20F897FEDCD99F" box="[151,328,1913,1936]" italics="true" pageId="9" pageNumber="10">Pristosia atrema</emphasis>
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(Andrewes)
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</taxonomicName>
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:
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<treatmentCitation id="DC7AEA146368FFE7FE6CF897FD20D99F" author="Lindroth" box="[475,692,1913,1936]" page="551" pageId="9" pageNumber="10" year="1956">
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||
<bibRefCitation id="394AB1F46368FFE7FE6CF897FDEBD99F" author="Lindroth" box="[475,639,1913,1936]" pageId="9" pageNumber="26" refString="Lindroth, C. H. (1956) A revision of the genus Synuchus Gyllenhal (Coleoptera: Carabidae) in the widest sense, with notes on Pristosia Motschulsky (Eucalathus Bates) and Calathus Bonelli. The Transactions of the Royal Entomological Society of London 108, 485 - 576." type="journal article" year="1956">Lindroth, 1956</bibRefCitation>
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: 551
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
|
||
</paragraph>
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||
<paragraph id="5D64CC056368FFE7FF20F877FCB5D9BF" blockId="9.[151,1436,1849,2000]" box="[151,801,1945,1968]" pageId="9" pageNumber="10">
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<treatmentCitationGroup id="7DCBEB2B6368FFE7FF20F877FCB5D9BF" box="[151,801,1945,1968]" pageId="9" pageNumber="10">
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<taxonomicName id="9ADBB7866368FFE7FF20F877FE45D9BF" ID-CoL="8M87R" authority="Andrewes" authorityName="Andrewes" box="[151,465,1945,1968]" class="Insecta" family="Carabidae" genus="Pristosia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="atrema">
|
||
<emphasis id="6FAF10176368FFE7FF20F877FEDCD9BF" box="[151,328,1945,1968]" italics="true" pageId="9" pageNumber="10">Pristosia atrema</emphasis>
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||
(Andrewes)
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||
</taxonomicName>
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||
:
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||
<treatmentCitation id="DC7AEA146368FFE7FE6CF877FC89D9BF" author="Hovorka" box="[475,797,1945,1968]" page="530" pageId="9" pageNumber="10" year="2003">
|
||
<bibRefCitation id="394AB1F46368FFE7FE6CF877FD73D9BF" author="Hovorka" box="[475,743,1945,1968]" pageId="9" pageNumber="26" refString="Hovorka, O. & Sciaky, R. (2003) Subtribe Pristosiina Lindroth, 1956. In: Lobl, I. & Smetana, A. (Eds), Catalogue of Palaearctic Coleoptera, Volume 1. Archostemata-Myxophaga-Adephaga. Apollo Books, Stenstrup, pp. 530 - 531." type="book chapter" year="2003">Hovorka & Sciaky, 2003</bibRefCitation>
|
||
: 530
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="5D64CC056368FFE7FF20F857FD32D9DF" blockId="9.[151,1436,1849,2000]" box="[151,678,1977,2000]" pageId="9" pageNumber="10">
|
||
<treatmentCitationGroup id="7DCBEB2B6368FFE7FF20F857FD32D9DF" box="[151,678,1977,2000]" pageId="9" pageNumber="10">
|
||
<taxonomicName id="9ADBB7866368FFE7FF20F857FE44D9DF" ID-CoL="8M87R" authority="Andrewes" authorityName="Andrewes" box="[151,464,1977,2000]" class="Insecta" family="Carabidae" genus="Pristosia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="atrema">
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<emphasis id="6FAF10176368FFE7FF20F857FEDCD9DF" box="[151,328,1977,2000]" italics="true" pageId="9" pageNumber="10">Pristosia atrema</emphasis>
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(Andrewes)
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</taxonomicName>
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:
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<treatmentCitation id="DC7AEA146368FFE7FE6CF857FD36D9DF" author="Lorenz" box="[475,674,1977,2000]" page="400" pageId="9" pageNumber="10" year="2005">
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||
<bibRefCitation id="394AB1F46368FFE7FE6CF857FDF9D9DF" author="Lorenz" box="[475,621,1977,2000]" pageId="9" pageNumber="26" refString="Lorenz, W. (2005) A systematic list of extant ground beetles of the world (Coleoptera " Geadephaga ": Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), Second edition. Published by the author, Tutzing, 530 pp." type="book" year="2005">Lorenz, 2005</bibRefCitation>
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||
: 400
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="15C19F8E636BFFE5FF20FF79FC6CDA5D" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="materials_examined">
|
||
<paragraph id="5D64CC05636BFFE4FF20FF79FC2FDEF0" blockId="10.[151,1437,151,448]" pageId="10" pageNumber="11">
|
||
<emphasis id="6FAF1017636BFFE4FF20FF79FED9DEBE" bold="true" box="[151,333,151,177]" pageId="10" pageNumber="11">
|
||
<typeStatus id="826072A7636BFFE4FF20FF79FF46DEBE" box="[151,210,151,177]" pageId="10" pageNumber="11">Type</typeStatus>
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||
material:
|
||
</emphasis>
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||
<typeStatus id="826072A7636BFFE4FEE1FF76FE57DEBD" box="[342,451,152,178]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
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||
male, with label data ”
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||
<typeStatus id="826072A7636BFFE4FD67FF76FC84DEBD" box="[720,784,152,178]" pageId="10" pageNumber="11">Type</typeStatus>
|
||
” (round and red bordered printed label), “Burphu Gori- V.,
|
||
<quantity id="9A2361E0636BFFE4FF74FF50FEA5DED7" box="[195,305,190,216]" metricMagnitude="3" metricUnit="m" metricValue="3.5052" pageId="10" pageNumber="11" unit="ft" value="11500.0">11500 ft</quantity>
|
||
.
|
||
<collectingCountry id="25CC8C95636BFFE4FE89FF50FE17DED7" box="[318,387,190,216]" name="India" pageId="10" pageNumber="11">India</collectingCountry>
|
||
. H.G.C.”, “G.C. Champion Brit. Mus.
|
||
<date id="2965EAC5636BFFE4FCDDFF50FC40DED7" box="[874,980,190,216]" pageId="10" pageNumber="11" value="1925" valueMax="1942">1925-42</date>
|
||
”,“
|
||
<taxonomicName id="9ADBB786636BFFE4FC58FF50FB57DED7" box="[1007,1219,190,216]" class="Insecta" family="Carabidae" genus="Calathus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="atrema">Calathus atrema</taxonomicName>
|
||
<typeStatus id="826072A7636BFFE4FB7AFF50FA9FDED7" box="[1229,1291,190,216]" pageId="10" pageNumber="11">Type</typeStatus>
|
||
<collectingCountry id="25CC8C95636BFFE4FAA4FF50FAC2DED7" box="[1299,1366,190,216]" name="Andorra" pageId="10" pageNumber="11">Andr</collectingCountry>
|
||
. H.E. Andrewes det.” (the latter handwritten by H.E. Andrewes) (
|
||
<collectionCode id="3BCA54C0636BFFE4FCE5FF0BFC3BDEF0" box="[850,943,229,255]" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="10" pageNumber="11">NHML</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="5D64CC05636BFFE4FF72FEE2FE04DF7C" blockId="10.[151,1437,151,448]" pageId="10" pageNumber="11">
|
||
<typeStatus id="826072A7636BFFE4FF72FEE2FEAFDF29" box="[197,315,268,294]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="4BDD078C636BFFE4FEF1FEE2FE23DF29" box="[326,439,268,294]" pageId="10" pageNumber="11" type="female">2 females</specimenCount>
|
||
with same label data as
|
||
<typeStatus id="826072A7636BFFE4FD64FEE2FCA9DF29" box="[723,829,268,294]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
|
||
, but “
|
||
<typeStatus id="826072A7636BFFE4FCC8FEE2FC7FDF29" box="[895,1003,268,294]" pageId="10" pageNumber="11" type="paratype">Paratype</typeStatus>
|
||
” (round and yellow bordered printed label), and one specimen “
|
||
<taxonomicName id="9ADBB786636BFFE4FE7BFEDCFC7ADF43" authority="Cotype Andr. H.E. Andrewes" authorityName="Cotype Andr. H.E. Andrewes" box="[460,1006,306,332]" class="Insecta" family="Carabidae" genus="Calathus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="atrema">
|
||
Calathus atrema Cotype
|
||
<collectingCountry id="25CC8C95636BFFE4FD47FEDCFCBBDF43" box="[752,815,306,332]" name="Andorra" pageId="10" pageNumber="11">Andr</collectingCountry>
|
||
. H.E. Andrewes
|
||
</taxonomicName>
|
||
det.” (the latter handwritten by H.E. Andrewes) (
|
||
<collectionCode id="3BCA54C0636BFFE4FE90FEB7FE10DF7C" box="[295,388,345,371]" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="10" pageNumber="11">NHML</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="5D64CC05636BFFE4FF71FE91FBD3DFCF" blockId="10.[151,1437,151,448]" pageId="10" pageNumber="11">
|
||
<emphasis id="6FAF1017636BFFE4FF71FE91FE58DF96" bold="true" box="[198,460,383,409]" pageId="10" pageNumber="11">Additional material:</emphasis>
|
||
<collectingCountry id="25CC8C95636BFFE4FE6FFE6EFDA4DF95" box="[472,560,384,410]" name="India" pageId="10" pageNumber="11">INDIA</collectingCountry>
|
||
: UTTARANCHAL DISTRICT:
|
||
<specimenCount id="4BDD078C636BFFE4FC0BFE6EFB87DF95" box="[956,1043,384,410]" pageId="10" pageNumber="11" type="male">1 male</specimenCount>
|
||
, with label data “Pindar Valley,
|
||
<quantity id="9A2361E0636BFFE4FF20FE48FEB5DFCF" box="[151,289,422,448]" metricMagnitude="3" metricUnit="m" metricValue="1.6776191999999999" metricValueMax="3.3528" metricValueMin="0.0024384" pageId="10" pageNumber="11" unit="ft" value="5504.0" valueMax="11000.0" valueMin="8.0">8–11,000 ft</quantity>
|
||
. Kumaon. H.G.C.”, “H.E. Andrewes Coll. B.M.
|
||
<date id="2965EAC5636BFFE4FCEFFE48FC2BDFCF" box="[856,959,422,448]" pageId="10" pageNumber="11" value="1945" valueMax="1997">1945-97</date>
|
||
.” (
|
||
<collectionCode id="3BCA54C0636BFFE4FC6AFE48FBAEDFCF" box="[989,1082,422,448]" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="10" pageNumber="11">NHML</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<caption id="09A49C8D636BFFE4FF20FA9FFF5CDBFF" httpUri="https://zenodo.org/record/185755/files/figure.png" pageId="10" pageNumber="11" targetBox="[179,1406,500,1358]" targetPageId="10">
|
||
<paragraph id="5D64CC05636BFFE4FF20FA9FFF5CDBFF" blockId="10.[151,1435,1393,1520]" pageId="10" pageNumber="11">
|
||
<emphasis id="6FAF1017636BFFE4FF20FA9FFEC9DB86" bold="true" box="[151,349,1393,1417]" pageId="10" pageNumber="11">FIGURES 24–29.</emphasis>
|
||
<taxonomicName id="9ADBB786636BFFE4FED4FA9CFE57DB86" box="[355,451,1394,1417]" class="Insecta" family="Carabidae" genus="Pristosia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6FAF1017636BFFE4FED4FA9CFE57DB86" box="[355,451,1394,1417]" italics="true" pageId="10" pageNumber="11">Pristosia</emphasis>
|
||
</taxonomicName>
|
||
spp., aedeagal median lobe, dorsal view (Figs. 24, 26, 28) and left lateral view (Figs. 25, 27, 29, the internal sac is figured only in dorsal view). Figs. 24, 25,
|
||
<emphasis id="6FAF1017636BFFE4FCD1FA7AFC43DBA4" box="[870,983,1428,1451]" italics="true" pageId="10" pageNumber="11">P. s i m i l a t a</emphasis>
|
||
<emphasis id="6FAF1017636BFFE4FC54FA7DFBB5DBA4" bold="true" box="[995,1057,1427,1451]" pageId="10" pageNumber="11">sp. n.</emphasis>
|
||
, Paratype. Figs. 26, 27,
|
||
<emphasis id="6FAF1017636BFFE4FA84FA7AFA0DDBA4" box="[1331,1433,1428,1451]" italics="true" pageId="10" pageNumber="11">P. a trema</emphasis>
|
||
(Andrewes, 1926), Holotype. Figs. 28, 29,
|
||
<emphasis id="6FAF1017636BFFE4FDC7FA59FD40DBC1" box="[624,724,1463,1486]" italics="true" pageId="10" pageNumber="11">P. a t re m a</emphasis>
|
||
(Andrewes, 1926), non-type, India, Pindar Valley. Scale bar = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5D64CC05636BFFE4FF71F9CCFD36D832" blockId="10.[151,1436,1570,2022]" box="[198,674,1570,1597]" pageId="10" pageNumber="11">
|
||
<emphasis id="6FAF1017636BFFE4FF71F9CCFEEAD833" bold="true" box="[198,382,1570,1596]" pageId="10" pageNumber="11">Redescription:</emphasis>
|
||
Four specimens studied.
|
||
</paragraph>
|
||
<paragraph id="5D64CC05636BFFE4FF72F9A7FD82D86C" blockId="10.[151,1436,1570,2022]" box="[197,534,1609,1635]" pageId="10" pageNumber="11">
|
||
Body length:
|
||
<quantity id="9A2361E0636BFFE4FED3F9A7FD86D86C" box="[356,530,1609,1635]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" metricValueMax="1.05" metricValueMin="1.01" pageId="10" pageNumber="11" unit="mm" value="10.3" valueMax="10.5" valueMin="10.1">10.1–10.5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5D64CC05636BFFE4FF72F99EFD18D8D8" blockId="10.[151,1436,1570,2022]" pageId="10" pageNumber="11">Colour: Dorsal and ventral surface of body and most parts of femora dark brown, upper anterior side of femora, tibiae, tarsi, antennae and palpi yellowish brown. Male dorsal surface shiny throughout; female shiny on head and pronotum but with elytra dull.</paragraph>
|
||
<paragraph id="5D64CC05636BFFE4FF72F90AFE79D97D" blockId="10.[151,1436,1570,2022]" pageId="10" pageNumber="11">Microsculpture: Head with mesh pattern isodiametric, moderately engraved, and pronotum with very weakly engraved transverse meshes, visible under high magnification (100x). Meshes of microsculpture on elytra in male weakly engraved, slightly transverse; in female isodiametric and much more deeply engraved, and scale-like in anterior 2/3.</paragraph>
|
||
<paragraph id="5D64CC05636BFFE4FF72F891FED3D9B0" blockId="10.[151,1436,1570,2022]" pageId="10" pageNumber="11">Head: Temporae about half of eye diameter. Antennomeres I–III completely smooth except for primary apical setation.</paragraph>
|
||
<paragraph id="5D64CC05636BFFE5FF72F822FE4ADF29" blockId="10.[151,1436,1570,2022]" lastBlockId="11.[151,1437,152,1570]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
|
||
Pronotum: Slightly transverse (ratio PW/PL 1.08–1.12, PW/HW 1.51–1.57). Base distinctly wider than anterior margin. Sides evenly rounded in anterior 3/4 but concave in posterior quarter, hind angles almost rectangular (95–100°) (
|
||
<figureCitation id="C5E0D080636AFFE5FE19FF50FD92DED7" box="[430,518,190,216]" captionStart="FIGURES 5 – 15" captionStartId="6.[151,272,1702,1726]" captionTargetBox="[177,1434,196,1674]" captionTargetId="figure@6.[151,1436,194,1678]" captionTargetPageId="6" captionText="FIGURES 5 – 15. Pristosia spp., pronotum. Fig. 5, P. dahud polita ssp. n., Paratype, male. Fig. 6, P. dahud Morvan, 1994, non-type, male, Rara Lake. Fig. 7, P. dahud Morvan, 1994, non-type, male, Maharigaon. Fig. 8, P. dahud Morvan, 1994, non-type, male, Khari Lagna (transitional form). Fig. 9, P. dahud Morvan, 1994, non-type, male, Khari Lagna (typical form). Fig. 10, P. similata sp. n., Paratype, male. Fig. 11, P. glabella sp. n., Paratype, male. Fig. 12, P. amaroides (Putzeys, 1877), non-type, female, India, Darjeeling. Fig. 13, P. a t re m a (Andrewes, 1926), Holotype. Fig. 14, P. nepalensis sp. n., Paratype, male. Fig. 15, P. championi (Andrewes, 1934), Holotype. Scale bar = 2 mm." httpUri="https://zenodo.org/record/185753/files/figure.png" pageId="11" pageNumber="12">Fig. 13</figureCitation>
|
||
). Base almost straight in middle, slightly bent anteriorly toward hind angles. Lateral gutter slightly expanded toward base. Basolateral seta located somewhat distant (1–2 times pore diameter) from lateral edge.
|
||
</paragraph>
|
||
<paragraph id="5D64CC05636AFFE5FF72FEDCFF64DF95" blockId="11.[151,1437,152,1570]" pageId="11" pageNumber="12">Elytra: Oval, with relatively wide shoulders, ratio EL/EW 1.47–1.53, EW/PW 1.41–1.47. Basal groove strongly concave, forming an almost right angle with scutellar stria and a slightly obtuse angle with lateral groove.</paragraph>
|
||
<paragraph id="5D64CC05636AFFE5FF72FE48FE5ADFCF" blockId="11.[151,1437,152,1570]" box="[197,462,422,448]" pageId="11" pageNumber="12">Legs: Relatively stout.</paragraph>
|
||
<paragraph id="5D64CC05636AFFE5FF72FE23FBA5DC3B" blockId="11.[151,1437,152,1570]" pageId="11" pageNumber="12">
|
||
Male genitalia: Aedeagal median lobe relatively small, in lateral view almost straight toward apex, but with apical lamella slightly bent down, and with terminal bead not angular (
|
||
<figureCitation id="C5E0D080636AFFE5FB8DFE1AFB44DC01" box="[1082,1232,500,526]" captionStart="FIGURES 24 – 29" captionStartId="10.[151,269,1393,1417]" captionTargetBox="[179,1406,500,1358]" captionTargetId="figure@10.[151,1436,475,1427]" captionTargetPageId="10" captionText="FIGURES 24 – 29. Pristosia spp., aedeagal median lobe, dorsal view (Figs. 24, 26, 28) and left lateral view (Figs. 25, 27, 29, the internal sac is figured only in dorsal view). Figs. 24, 25, P. s i m i l a t a sp. n., Paratype. Figs. 26, 27, P. a trema (Andrewes, 1926), Holotype. Figs. 28, 29, P. a t re m a (Andrewes, 1926), non-type, India, Pindar Valley. Scale bar = 1 mm." httpUri="https://zenodo.org/record/185755/files/figure.png" pageId="11" pageNumber="12">Figs. 27, 29</figureCitation>
|
||
). In dorsal view, apical lamella thin and moderately long. Internal sac folding as in
|
||
<figureCitation id="C5E0D080636AFFE5FC29FDF4FBB9DC3B" box="[926,1069,538,564]" captionStart="FIGURES 24 – 29" captionStartId="10.[151,269,1393,1417]" captionTargetBox="[179,1406,500,1358]" captionTargetId="figure@10.[151,1436,475,1427]" captionTargetPageId="10" captionText="FIGURES 24 – 29. Pristosia spp., aedeagal median lobe, dorsal view (Figs. 24, 26, 28) and left lateral view (Figs. 25, 27, 29, the internal sac is figured only in dorsal view). Figs. 24, 25, P. s i m i l a t a sp. n., Paratype. Figs. 26, 27, P. a trema (Andrewes, 1926), Holotype. Figs. 28, 29, P. a t re m a (Andrewes, 1926), non-type, India, Pindar Valley. Scale bar = 1 mm." httpUri="https://zenodo.org/record/185755/files/figure.png" pageId="11" pageNumber="12">Figs. 26, 28</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5D64CC05636AFFE5FF71FDAEFC6CDA5D" blockId="11.[151,1437,152,1570]" pageId="11" pageNumber="12">
|
||
<emphasis id="6FAF1017636AFFE5FF71FDAEFEEDDC55" bold="true" box="[198,377,576,602]" pageId="11" pageNumber="12">Identification:</emphasis>
|
||
Within the fauna of Kumaon and
|
||
<collectingCountry id="25CC8C95636AFFE5FCA4FDAFFCCDDC54" box="[787,857,577,603]" name="Nepal" pageId="11" pageNumber="12">Nepal</collectingCountry>
|
||
Himalaya
|
||
<emphasis id="6FAF1017636AFFE5FC6AFDAFFBDEDC55" box="[989,1098,577,602]" italics="true" pageId="11" pageNumber="12">P. a t re m a</emphasis>
|
||
differs from other
|
||
<taxonomicName id="9ADBB786636AFFE5FA85FDAFFA08DC55" box="[1330,1436,577,602]" class="Insecta" family="Carabidae" genus="Pristosia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6FAF1017636AFFE5FA85FDAFFA08DC55" box="[1330,1436,577,602]" italics="true" pageId="11" pageNumber="12">Pristosia</emphasis>
|
||
</taxonomicName>
|
||
species by the combination of characters of pronotal and elytral shape, elytral chaetotaxy and form of aedeagal median lobe.
|
||
<taxonomicName id="9ADBB786636AFFE5FE1EFD61FDD0DCA7" box="[425,580,655,680]" class="Insecta" family="Carabidae" genus="Calathus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="championi">
|
||
<emphasis id="6FAF1017636AFFE5FE1EFD61FDD0DCA7" box="[425,580,655,680]" italics="true" pageId="11" pageNumber="12">P. championi</emphasis>
|
||
</taxonomicName>
|
||
is a species of sympatric distribution, and has a similar pronotal shape, but has a smaller pronotal base with hind angles more obtuse, has elytra narrower at shoulders and has usually two dorsal setiferous pores in interval III. In addition, the aedeagal median lobe is much more elongated.
|
||
<taxonomicName id="9ADBB786636AFFE5FA31FD32FE04DD13" authority="Morvan, 1994" authorityName="Morvan" authorityYear="1994" class="Insecta" family="Carabidae" genus="Pristosia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="dahud">
|
||
<emphasis id="6FAF1017636AFFE5FA31FD32FF74DD14" italics="true" pageId="11" pageNumber="12">P. dahud</emphasis>
|
||
<bibRefCitation id="394AB1F4636AFFE5FF50FCECFE04DD13" author="Morvan" box="[231,400,770,796]" pageId="11" pageNumber="26" refString="Morvan, D. m. (1994) Carabidae nouveaux du Nepal et de Malaisie (Coleoptera, Carabidae). Bulletin de la Societe entomologique de France 99 (3), 323 - 334." type="journal article" year="1994">Morvan, 1994</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="6FAF1017636AFFE5FE2BFCECFD81DD14" box="[412,533,770,795]" italics="true" pageId="11" pageNumber="12">P. g l a b e l l a</emphasis>
|
||
<emphasis id="6FAF1017636AFFE5FDA8FCECFDF5DD13" bold="true" box="[543,609,770,796]" pageId="11" pageNumber="12">sp. n.</emphasis>
|
||
, and
|
||
<taxonomicName id="9ADBB786636AFFE5FD16FCECFC8FDD14" box="[673,795,770,795]" class="Insecta" family="Carabidae" genus="Pristosia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="similata" status="sp. nov.">
|
||
<emphasis id="6FAF1017636AFFE5FD16FCECFC8FDD14" box="[673,795,770,795]" italics="true" pageId="11" pageNumber="12">P. similata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="749CAD6C636AFFE5FC94FCECFCF1DD13" box="[803,869,770,796]" pageId="11" pageNumber="12" rank="species">
|
||
<emphasis id="6FAF1017636AFFE5FC94FCECFCF1DD13" bold="true" box="[803,869,770,796]" pageId="11" pageNumber="12">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, all from western
|
||
<collectingCountry id="25CC8C95636AFFE5FB8CFCECFB15DD13" box="[1083,1153,770,796]" name="Nepal" pageId="11" pageNumber="12">Nepal</collectingCountry>
|
||
Himalaya, are darker in colour of body, have legs more slender, pronotal posterior angles rounded, aedeagal median lobe larger with terminal bead of apical lamella more protruded dorsally, and with different form of internal sac folding (see
|
||
<figureCitation id="C5E0D080636AFFE5FF20FC98FE96DD9F" box="[151,258,886,912]" captionStart="FIGURES 24 – 29" captionStartId="10.[151,269,1393,1417]" captionTargetBox="[179,1406,500,1358]" captionTargetId="figure@10.[151,1436,475,1427]" captionTargetPageId="10" captionText="FIGURES 24 – 29. Pristosia spp., aedeagal median lobe, dorsal view (Figs. 24, 26, 28) and left lateral view (Figs. 25, 27, 29, the internal sac is figured only in dorsal view). Figs. 24, 25, P. s i m i l a t a sp. n., Paratype. Figs. 26, 27, P. a trema (Andrewes, 1926), Holotype. Figs. 28, 29, P. a t re m a (Andrewes, 1926), non-type, India, Pindar Valley. Scale bar = 1 mm." httpUri="https://zenodo.org/record/185755/files/figure.png" pageId="11" pageNumber="12">Figs. 24</figureCitation>
|
||
,
|
||
<figureCitation id="C5E0D080636AFFE5FEA7FC98FEF0DD9F" box="[272,356,886,912]" captionStart-0="FIGURES 30 – 35" captionStart-1="FIGURES 36 – 41" captionStartId-0="14.[151,269,1259,1283]" captionStartId-1="15.[151,269,1591,1615]" captionTargetBox-0="[157,1430,486,1209]" captionTargetBox-1="[158,1427,921,1532]" captionTargetId-0="figure@14.[151,1436,465,1236]" captionTargetId-1="figure@15.[151,1436,887,1567]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURES 30 – 35. Pristosia spp., aedeagal median lobe, dorsal view (Figs. 30, 32, 34) and left lateral view (Figs. 31, 33, 35, the internal sac is figured only in dorsal view). Figs. 30, 31, P. dahud polita ssp. n., Paratype. Figs. 32, 33, P. dahud Morvan, 1994, non-type, Khari Lagna (transitional form). Figs. 34, 35, P. dahud Morvan, 1994, non-type, Khari Lagna (typical form). Scale bar = 1 mm." captionText-1="FIGURES 36 – 41. Pristosia spp., aedeagal median lobe, dorsal view (Figs. 36, 38, 40) and left lateral view (Figs. 37, 39, 41, the internal sac is figured only in dorsal view). Figs. 36, 37, P. dahud Morvan, 1994, non-type, Maharigaon. Figs. 38, 39, P. dahud Morvan, 1994, non-type, Rara Lake. Figs. 40, 41, P. glabella sp. n., Holotype. Scale bar = 1 mm." httpUri-0="https://zenodo.org/record/185757/files/figure.png" httpUri-1="https://zenodo.org/record/185758/files/figure.png" pageId="11" pageNumber="12">30–40</figureCitation>
|
||
). Both the species
|
||
<taxonomicName id="9ADBB786636AFFE5FDE2FC99FC2FDD9F" authority="Andrewes, 1934" authorityName="Andrewes" authorityYear="1934" box="[597,955,886,912]" class="Insecta" family="Carabidae" genus="Pristosia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="leptodes">
|
||
<emphasis id="6FAF1017636AFFE5FDE2FC99FD74DD9F" box="[597,736,887,912]" italics="true" pageId="11" pageNumber="12">P. leptodes</emphasis>
|
||
<bibRefCitation id="394AB1F4636AFFE5FD5FFC98FC2FDD9F" author="Andrewes" box="[744,955,886,912]" pageId="11" pageNumber="25" refString="Andrewes, H. E. (1934) Keys to some Indian genera of Carabidae (Col.). IV. The genus Calathus. Stylops 3 (9), 209 - 222." type="journal article" year="1934">Andrewes, 1934</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="9ADBB786636AFFE5FBB7FC99FEC7DDB8" authority="Deuve, Lassalle & Queinnec, 1985" authorityName="Deuve, Lassalle & Queinnec" authorityYear="1985" class="Insecta" family="Carabidae" genus="Pristosia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="brancuccii">
|
||
<emphasis id="6FAF1017636AFFE5FBB7FC99FB3CDD9F" box="[1024,1192,887,912]" italics="true" pageId="11" pageNumber="12">P. brancuccii</emphasis>
|
||
<bibRefCitation id="394AB1F4636AFFE5FB06FC98FEC7DDB8" author="Deuve" pageId="11" pageNumber="25" refString="Deuve, T., Lassalle, B. & Queinnec, E. (1985) Nouveaux Pristosia Motschulsky et Calathus Bonelli de la region himalayenne (Coleoptera, Carabidae, Pterostichinae). Entomologica Basiliensia 10, 75 - 84." type="journal article" year="1985">Deuve, Lassalle & Quéinnec, 1985</bibRefCitation>
|
||
</taxonomicName>
|
||
, which have a more western distribution in the Kumaon Himalaya, are easily distinguished in pronotal shape, too, because sides are convexly rounded throughout and posterior angles are strongly obtuse (for the latter see
|
||
<bibRefCitation id="394AB1F4636AFFE5FEDCFC04FDD1DA0B" author="Deuve" box="[363,581,1002,1028]" pageId="11" pageNumber="25" refString="Deuve, T., Lassalle, B. & Queinnec, E. (1985) Nouveaux Pristosia Motschulsky et Calathus Bonelli de la region himalayenne (Coleoptera, Carabidae, Pterostichinae). Entomologica Basiliensia 10, 75 - 84." type="journal article" year="1985">
|
||
Deuve
|
||
<emphasis id="6FAF1017636AFFE5FE77FC04FE69DA0C" box="[448,509,1002,1027]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
|
||
1985
|
||
</bibRefCitation>
|
||
: p. 78,
|
||
<figureCitation id="C5E0D080636AFFE5FD2BFC04FD70DA0B" box="[668,740,1002,1028]" captionStart="FIGURES 1 – 4" captionStartId="5.[151,272,1939,1963]" captionTargetBox="[366,1218,198,1938]" captionTargetId="figure@5.[357,1233,188,1938]" captionTargetPageId="5" captionText="FIGURES 1 – 4. Pristosia spp., habitus. Fig. 1, P. crenata (Putzeys, 1873), non-type, male, Nepal, Dailekh. Fig. 2, P. amaroides (Putzeys, 1877), non-type, female, India, Darjeeling. Fig. 3, P. glabella sp. n., Paratype, female. Fig. 4, P. nepalensis sp. n., Paratype, male. Scale bar = 5 mm." httpUri="https://zenodo.org/record/185752/files/figure.png" pageId="11" pageNumber="12">Fig. 3</figureCitation>
|
||
). In addition,
|
||
<taxonomicName id="9ADBB786636AFFE5FC3AFC04FB9ADA0C" box="[909,1038,1002,1027]" class="Insecta" family="Carabidae" genus="Pristosia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="leptodes">
|
||
<emphasis id="6FAF1017636AFFE5FC3AFC04FB9ADA0C" box="[909,1038,1002,1027]" italics="true" pageId="11" pageNumber="12">P. leptodes</emphasis>
|
||
</taxonomicName>
|
||
is a smaller species (body length
|
||
<quantity id="9A2361E0636AFFE5FF20FBFFFE82DA24" box="[151,278,1041,1067]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="11" pageNumber="12" unit="mm" value="6.75" valueMax="7.0" valueMin="6.5">6.5–7 mm</quantity>
|
||
), has usually 2 setiferous pore punctures in elytral interval III, and terminal lamella of aedeagal median lobe more strongly angular (see
|
||
<bibRefCitation id="394AB1F4636AFFE5FDD8FBD6FC8FDA5D" author="Lindroth" box="[623,795,1080,1106]" pageId="11" pageNumber="26" refString="Lindroth, C. H. (1956) A revision of the genus Synuchus Gyllenhal (Coleoptera: Carabidae) in the widest sense, with notes on Pristosia Motschulsky (Eucalathus Bates) and Calathus Bonelli. The Transactions of the Royal Entomological Society of London 108, 485 - 576." type="journal article" year="1956">Lindroth 1956</bibRefCitation>
|
||
: p. 550,
|
||
<figureCitation id="C5E0D080636AFFE5FCC9FBD6FC46DA5D" box="[894,978,1080,1106]" captionStart="FIGURES 30 – 35" captionStartId="14.[151,269,1259,1283]" captionTargetBox="[157,1430,486,1209]" captionTargetId="figure@14.[151,1436,465,1236]" captionTargetPageId="14" captionText="FIGURES 30 – 35. Pristosia spp., aedeagal median lobe, dorsal view (Figs. 30, 32, 34) and left lateral view (Figs. 31, 33, 35, the internal sac is figured only in dorsal view). Figs. 30, 31, P. dahud polita ssp. n., Paratype. Figs. 32, 33, P. dahud Morvan, 1994, non-type, Khari Lagna (transitional form). Figs. 34, 35, P. dahud Morvan, 1994, non-type, Khari Lagna (typical form). Scale bar = 1 mm." httpUri="https://zenodo.org/record/185757/files/figure.png" pageId="11" pageNumber="12">Fig. 31</figureCitation>
|
||
H).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="15C19F8E636AFFE5FF71FBB0FCEDD82D" pageId="11" pageNumber="12" type="distribution">
|
||
<paragraph id="5D64CC05636AFFE5FF71FBB0FE59DB88" blockId="11.[151,1437,152,1570]" pageId="11" pageNumber="12">
|
||
<emphasis id="6FAF1017636AFFE5FF71FBB0FD42DA77" bold="true" box="[198,726,1118,1144]" pageId="11" pageNumber="12">Distribution and geographical variation:</emphasis>
|
||
<emphasis id="6FAF1017636AFFE5FD53FBB1FCC2DA77" box="[740,854,1119,1144]" italics="true" pageId="11" pageNumber="12">P. a t re m a</emphasis>
|
||
seems endemic to the Nanda Devi massif of Kumaon Himalaya. The species is currently only known from the Gori Valley on SE slope of Nanda Devi which is located close to the Indian-Nepalese border, and from the Pindar Valley on SW slope of Nanda Devi massif. Based on the material studied, the two populations slightly differ in aedeagal characters: The single male specimen from Pindar valley has the median lobe distinctly slenderer and the internal sac more strongly sclerotized before median lobe apex (see
|
||
<figureCitation id="C5E0D080636AFFE5FDC8FACEFC9ADB35" box="[639,782,1312,1338]" captionStart="FIGURES 24 – 29" captionStartId="10.[151,269,1393,1417]" captionTargetBox="[179,1406,500,1358]" captionTargetId="figure@10.[151,1436,475,1427]" captionTargetPageId="10" captionText="FIGURES 24 – 29. Pristosia spp., aedeagal median lobe, dorsal view (Figs. 24, 26, 28) and left lateral view (Figs. 25, 27, 29, the internal sac is figured only in dorsal view). Figs. 24, 25, P. s i m i l a t a sp. n., Paratype. Figs. 26, 27, P. a trema (Andrewes, 1926), Holotype. Figs. 28, 29, P. a t re m a (Andrewes, 1926), non-type, India, Pindar Valley. Scale bar = 1 mm." httpUri="https://zenodo.org/record/185755/files/figure.png" pageId="11" pageNumber="12">Figs. 28, 29</figureCitation>
|
||
). According to these differences,
|
||
<emphasis id="6FAF1017636AFFE5FB2CFACEFA93DB36" box="[1179,1287,1312,1337]" italics="true" pageId="11" pageNumber="12">P. a t re m a</emphasis>
|
||
seems to be a polytypic species, but more material from additional localities needs to be studied before taxonomical conclusions can be drawn.
|
||
</paragraph>
|
||
<paragraph id="5D64CC05636AFFE5FF71FA7DFCEDD82D" blockId="11.[151,1437,152,1570]" pageId="11" pageNumber="12">
|
||
<emphasis id="6FAF1017636AFFE5FF71FA7DFEA4DBA2" bold="true" box="[198,304,1427,1453]" pageId="11" pageNumber="12">Habitat:</emphasis>
|
||
During original description and subsequent species group revisions (
|
||
<bibRefCitation id="394AB1F4636AFFE5FB31FA7AFADCDBA1" author="Andrewes" box="[1158,1352,1428,1454]" pageId="11" pageNumber="25" refString="Andrewes, H. E. (1926) On a collection of Carabidae from the Kumaon-Tibetan frontier. The Entomologist's Monthly Magazine, 62, 65 - 80." type="journal article" year="1926">Andrewes 1926</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="394AB1F4636AFFE5FAEDFA7AFA03DBA1" author="Andrewes" box="[1370,1431,1428,1454]" pageId="11" pageNumber="25" refString="Andrewes, H. E. (1934) Keys to some Indian genera of Carabidae (Col.). IV. The genus Calathus. Stylops 3 (9), 209 - 222." type="journal article" year="1934">1934</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="394AB1F4636AFFE5FF20FA55FED3DBDA" author="Lindroth" box="[151,327,1467,1493]" pageId="11" pageNumber="26" refString="Lindroth, C. H. (1956) A revision of the genus Synuchus Gyllenhal (Coleoptera: Carabidae) in the widest sense, with notes on Pristosia Motschulsky (Eucalathus Bates) and Calathus Bonelli. The Transactions of the Royal Entomological Society of London 108, 485 - 576." type="journal article" year="1956">Lindroth 1956</bibRefCitation>
|
||
) no notes about species habitat where given. However, during the last decade three closely related species were frequently found in the adjacent
|
||
<collectingCountry id="25CC8C95636AFFE5FCBDFA0FFCC4DBF4" box="[778,848,1505,1531]" name="Nepal" pageId="11" pageNumber="12">Nepal</collectingCountry>
|
||
Himalaya. These species probably have similar if not identical habitat preferences (see below for further details).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |