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<document id="D64DC9DF3A01F18BC4ED4C1749279523" ID-DOI="10.5281/zenodo.5393362" ID-ISSN="1638-9387" ID-Zenodo-Dep="5393362" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630318630459" checkinUser="marcus" docAuthor="Ohler, Annemarie &amp; Dubois, Alain" docDate="2006" docId="D4590212FFFEFFFCFC3A87ADFB0D3A48" docLanguage="en" docName="Zoosystema.2006.28.3.769-784.pdf" docOrigin="Zoosystema 28 (3)" docStyle="DocumentStyle:64A3174DE33E974D4EB6573A0A793B12.4:Zoosystema.2000-2008.journal_article.type2" docStyleId="64A3174DE33E974D4EB6573A0A793B12" docStyleName="Zoosystema.2000-2008.journal_article.type2" docStyleVersion="4" docTitle="Allopaa Ohler &amp; Dubois 2006, n. gen." docType="treatment" docVersion="4" lastPageNumber="780" masterDocId="28607A6AFFF5FFF7FFBF8656FF943F7E" masterDocTitle="Phylogenetic relationships and generic taxonomy of the tribe Paini (Amphibia, Anura, Ranidae, Dicroglossinae), with diagnoses of two new genera" masterLastPageNumber="784" masterPageNumber="769" pageNumber="780" updateTime="1699225126584" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="135C60CF34E3E593B5122DF43296A1B1">Phylogenetic relationships and generic taxonomy of the tribe Paini (Amphibia, Anura, Ranidae, Dicroglossinae), with diagnoses of two new genera</mods:title>
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<mods:namePart id="BE72B7CA523853EEC4D6A5369C7D6881">Ohler, Annemarie</mods:namePart>
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<mods:namePart id="CC83F6BB476C8DC223F6C280DB7DB493">Dubois, Alain</mods:namePart>
<mods:affiliation id="EB9FD8151306F0BDC98FAD379D6B8842">Muséum national dHistoire naturelle, Département Systématique et Évolution, Reptiles et Amphibiens, USM 0602, case postale 30, 25 rue Cuvier, F- 75231 Paris cedex 05 (France) ohler @ mnhn. fr ..</mods:affiliation>
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<mods:title id="2A12A85BBB9B502D28619CF9519C5B88">Zoosystema</mods:title>
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<mods:date id="6C1F3E296472B8531D5067D99B256CEA">2006</mods:date>
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<taxonomicName id="9BF0C887FFFEFFFCFC3A87ADFC403D6B" ID-CoL="SG8" authority="Ohler &amp; Dubois, 2006" authorityName="Ohler &amp; Dubois" authorityYear="2006" box="[901,980,507,533]" class="Amphibia" family="Dicroglossidae" genus="Allopaa" kingdom="Animalia" order="Anura" pageId="11" pageNumber="780" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="6E846F16FFFEFFFCFC3A87ADFC403D6B" box="[901,980,507,533]" italics="true" pageId="11" pageNumber="780">Allopaa</emphasis>
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<taxonomicNameLabel id="75B7D26DFFFEFFFCFC6487ADFBB13D6B" box="[987,1061,507,533]" pageId="11" pageNumber="780" rank="genus">n. gen.</taxonomicNameLabel>
</heading>
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SPECIES
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. —
<taxonomicName id="9BF0C887FFFEFFFCFCD18462FCA53D19" authority="Dubois &amp; Khan, 1980: 403" authorityName="Dubois &amp; Khan" authorityPageNumber="403" authorityYear="1980" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="11" pageNumber="780" phylum="Chordata" rank="species" species="hazarensis" subGenus="Paa">
<emphasis id="6E846F16FFFEFFFCFCD18462FC343D32" box="[878,928,564,588]" italics="true" pageId="11" pageNumber="780">Rana</emphasis>
(
<emphasis id="6E846F16FFFEFFFCFC138462FC5B3D32" box="[940,975,564,588]" italics="true" pageId="11" pageNumber="780">Paa</emphasis>
)
<emphasis id="6E846F16FFFEFFFCFC638462FBAE3D32" box="[988,1082,564,588]" italics="true" pageId="11" pageNumber="780">hazarensis</emphasis>
<bibRefCitation id="3861CEF5FFFEFFFCFB818462FCA53D19" author="DUBOIS A. &amp; KHAN M. S." pageId="11" pageNumber="780" pagination="403 - 410" refId="ref10154" refString="DUBOIS A. &amp; KHAN M. S. 1980. - A new species of frog (genus Rana, subgenus Paa) from Northern Pakistan (Amphibia, Anura). Journal of Herpetology 13 (4): 403 - 410 (dated 1979, published 1980)." type="journal article" year="1980">Dubois &amp; Khan, 1980: 403</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="14EAE08FFFFEFFFCFD758429FC3B3B13" pageId="11" pageNumber="780" type="diagnosis">
<paragraph id="5C4FB304FFFEFFFCFD758429FCE13B77" blockId="11.[712,1248,638,1033]" pageId="11" pageNumber="780">
DIAGNOSIS. — Genus distinguished from all other genera of
<taxonomicName id="9BF0C887FFFEFFFCFD5A84CFFC8C3DCF" authorityName="Dubois" authorityYear="1992" box="[741,792,665,689]" class="Amphibia" family="Ranidae" kingdom="Animalia" order="Anura" pageId="11" pageNumber="783" phylum="Chordata" rank="tribe" tribe="Paini">Paini</taxonomicName>
by the following combination of characters: 1) first finger slightly longer than second; 2) extremities of fingers and toes blunt; 3) hind legs short, tibia slightly shorter than half of snout-vent length, without significant sex dimorphism in length; 4) a feeble tarsal fold present; 5) webbing complete; 6) flap of skin along external side of toe V extending proximally beyond first subarticular tubercle of toe but not reaching basis of metatarsus; 7) middorsal chevron present in some specimens; 8) breeding males with slightly enlarged forearm, with black nuptial spines scattered on dorsal part of metacarpal tubercle and finger I alone, with internal vocal sacs; 9) intersexuality not known to exist; and 10) tadpoles with a keratodont formula of 6-8/3.
</paragraph>
<paragraph id="5C4FB304FFFEFFFCFD758277FC3B3B13" blockId="11.[712,1248,1056,1133]" pageId="11" pageNumber="780">
INCLUDED SPECIES. —
<taxonomicName id="9BF0C887FFFEFFFCFC268276FC903B2D" authority="(Dubois &amp; Khan, 1980)" authorityName="Ohler &amp; Dubois" authorityYear="2006" baseAuthorityName="Dubois &amp; Khan" baseAuthorityYear="1980" class="Amphibia" family="Dicroglossidae" genus="Allopaa" kingdom="Animalia" order="Anura" pageId="11" pageNumber="780" phylum="Chordata" rank="species" species="hazarensis" status="comb. nov.">
<emphasis id="6E846F16FFFEFFFCFC268276FBAD3B46" box="[921,1081,1056,1080]" italics="true" pageId="11" pageNumber="780">Allopaa hazarensis</emphasis>
(
<bibRefCitation id="3861CEF5FFFEFFFCFBFB8276FD693B2D" author="DUBOIS A. &amp; KHAN M. S." pageId="11" pageNumber="780" pagination="403 - 410" refId="ref10154" refString="DUBOIS A. &amp; KHAN M. S. 1980. - A new species of frog (genus Rana, subgenus Paa) from Northern Pakistan (Amphibia, Anura). Journal of Herpetology 13 (4): 403 - 410 (dated 1979, published 1980)." type="journal article" year="1980">Dubois &amp; Khan, 1980</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="75B7D26DFFFEFFFCFCB3826DFCFC3B2D" box="[780,872,1083,1107]" pageId="11" pageNumber="780" rank="species">n. comb.</taxonomicNameLabel>
Synonym:
<taxonomicName id="9BF0C887FFFEFFFCFC60826DFC3F3B13" authority="Khan &amp; Tasnim, 1989" authorityName="Khan &amp; Tasnim" authorityYear="1989" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="11" pageNumber="780" phylum="Chordata" rank="species" species="barmoachensis" subGenus="Paa">
<emphasis id="6E846F16FFFEFFFCFC60826DFB863B2D" box="[991,1042,1083,1107]" italics="true" pageId="11" pageNumber="780">Rana</emphasis>
(
<emphasis id="6E846F16FFFEFFFCFB9D826DFBD23B2D" box="[1058,1094,1083,1107]" italics="true" pageId="11" pageNumber="780">Paa</emphasis>
)
<emphasis id="6E846F16FFFEFFFCFBE9826DFB4B3B2D" box="[1110,1247,1083,1107]" italics="true" pageId="11" pageNumber="780">barmoachensis</emphasis>
<bibRefCitation id="3861CEF5FFFEFFFCFD758203FC3F3B13" author="KHAN M. S. &amp; TASNIM R." box="[714,939,1109,1133]" pageId="11" pageNumber="780" pagination="419 - 423" refId="ref10685" refString="KHAN M. S. &amp; TASNIM R. 1989. - A new frog of the genus Rana, subgenus Paa, from Southwestern Azad Kashmir. Journal of Herpetology 23: 419 - 423." type="journal article" year="1989">Khan &amp; Tasnim, 1989</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="14EAE08FFFFEFFFCFD7582D3FBD43BE3" box="[714,1088,1157,1181]" pageId="11" pageNumber="780" type="distribution">
<paragraph id="5C4FB304FFFEFFFCFD7582D3FBD43BE3" blockId="11.[714,1088,1157,1181]" box="[714,1088,1157,1181]" pageId="11" pageNumber="780">
DISTRIBUTION. — Northern
<collectingCountry id="24E7F394FFFEFFFCFC5882D3FBA83BE3" box="[999,1084,1157,1181]" name="Pakistan" pageId="11" pageNumber="780">Pakistan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="14EAE08FFFFEFFFCFD7582E3FB0D3A48" pageId="11" pageNumber="780" type="etymology">
<paragraph id="5C4FB304FFFEFFFCFD7582E3FB0D3A48" blockId="11.[712,1248,1204,1334]" pageId="11" pageNumber="780">
ETYMOLOGY OF GENERIC NAME. — From the Greek
<emphasis id="6E846F16FFFEFFFCFB0E82E2FB4D3BB2" box="[1201,1241,1204,1228]" italics="true" pageId="11" pageNumber="780">allos</emphasis>
, “other, different”, and from the generic name
<taxonomicName id="9BF0C887FFFEFFFCFBD28299FD6F3A7F" authority="Dubois, 1975" authorityName="Dubois" authorityYear="1975" class="Amphibia" family="Dicroglossidae" genus="Paa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="780" phylum="Chordata" rank="genus">
<emphasis id="6E846F16FFFEFFFCFBD28299FB1B3B99" box="[1133,1167,1231,1255]" italics="true" pageId="11" pageNumber="780">Paa</emphasis>
<bibRefCitation id="3861CEF5FFFEFFFCFB2C8299FD6F3A7F" author="DUBOIS A." pageId="11" pageNumber="780" pagination="1093 - 1115" refId="ref9742" refString="DUBOIS A. 1975. - Un nouveau sous-genre (Paa) et trois nouvelles especes du genre Rana. Remarques sur la phylogenie des Ranides (Amphibiens, Anoures). Bulletin du Museum national d'Histoire naturelle, 3 e ser., 324, Zoologie 231: 1093 - 1115." type="journal article" year="1975">Dubois, 1975</bibRefCitation>
</taxonomicName>
(from the Tamang name
<emphasis id="6E846F16FFFEFFFCFC4582BFFB883A7F" box="[1018,1052,1257,1281]" italics="true" pageId="11" pageNumber="780">paa</emphasis>
, “frog”). This name refers to the unusual combination of characters of this taxon as compared to the other genera of
<taxonomicName id="9BF0C887FFFEFFFCFBDF8348FB013A48" authorityName="Dubois" authorityYear="1992" box="[1120,1173,1310,1334]" class="Amphibia" family="Ranidae" kingdom="Animalia" order="Anura" pageId="11" pageNumber="783" phylum="Chordata" rank="tribe" tribe="Paini">Paini</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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