210 lines
19 KiB
XML
210 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.646.10787" ID-GBIF-Dataset="118044a5-2659-44d1-846a-9e0a4861b42d" ID-PMC="PMC5299439" ID-Pensoft-Pub="1313-2970-646-45" ID-PubMed="28228675" ID-ZBK="498CDCA3D6344414B3BF87C8F649154C" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-646-45" ModsDocOrigin="ZooKeys 646" ModsDocTitle="New species of Habronattus and Pellenes jumping spiders (Araneae, Salticidae, Harmochirina)" checkinTime="1484692248706" checkinUser="pensoft" docAuthor="Maddison, Wayne P." docDate="2017" docId="D449898E65ECFED1E4701F0E8CCFAF51" docLanguage="en" docName="ZooKeys 646: 45-72" docOrigin="ZooKeys 646" docSource="http://dx.doi.org/10.3897/zookeys.646.10787" docTitle="Habronattus empyrus Maddison, sp. n." docType="treatment" docUuid="1FCF2BA2-B4E8-4C5C-B384-7198F62E5D17" docUuidSource="ZooBank" docVersion="5" lastPageNumber="49" masterDocId="FF97CF79FFE01865B910956AFFDFFFF3" masterDocTitle="New species of Habronattus and Pellenes jumping spiders (Araneae, Salticidae, Harmochirina)" masterLastPageNumber="72" masterPageNumber="45" pageNumber="49" updateTime="1668163903918" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New species of Habronattus and Pellenes jumping spiders (Araneae, Salticidae, Harmochirina)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Maddison, Wayne P.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>646</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>45</mods:start>
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<mods:end>72</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.646.10787</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.646.10787</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-646-45</mods:identifier>
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<mods:identifier type="ZBK">498CDCA3D6344414B3BF87C8F649154C</mods:identifier>
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<mods:identifier type="ZooBank">498CDCA3D6344414B3BF87C8F649154C</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="127875133" LSID="urn:lsid:zoobank.org:act:1FCF2BA2-B4E8-4C5C-B384-7198F62E5D17" httpUri="http://treatment.plazi.org/id/D449898E65ECFED1E4701F0E8CCFAF51" lastPageNumber="49" pageId="4" pageNumber="49">
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<subSubSection pageId="4" pageNumber="49" type="nomenclature">
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<paragraph pageId="4" pageNumber="49">
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<taxonomicName LSID="http://zoobank.org/1FCF2BA2-B4E8-4C5C-B384-7198F62E5D17" authority="Maddison" class="Arachnida" family="Salticidae" genus="Habronattus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Habronattus empyrus" order="Araneae" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="empyrus">Habronattus empyrus Maddison</taxonomicName>
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<taxonomicNameLabel pageId="4" pageNumber="49">sp. n.</taxonomicNameLabel>
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Figs 26-37
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="49" type="holotype">
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<paragraph pageId="4" pageNumber="49">Holotype.</paragraph>
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<paragraph pageId="4" pageNumber="49">
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Male in CNAN- IBUNAM, with data:
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<normalizedToken originalValue="México">Mexico</normalizedToken>
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: Sonora: Puerto
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<normalizedToken originalValue="Peñasco">Penasco</normalizedToken>
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, Estero
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<normalizedToken originalValue="Morúa">Morua</normalizedToken>
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, 31.293 - 31.295°N 113.456 - 113.459°W, 17 August 2013, Maddison/Proctor/Evans/Leduc-Robert/Meza, WPM#13-084.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="49" type="paratypes">
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<paragraph pageId="4" pageNumber="49">Paratypes</paragraph>
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<paragraph pageId="4" pageNumber="49">
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(5♂♂ 4♀♀). Same data as holotype (1♀ specimen AZS13-7828 in CNAN- IBUNAM, 1♂ 1♀ in UBC-SEM, 1♂ 1♀ in MCZ, 1♂ 1♀ in AMNH).
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<normalizedToken originalValue="México">Mexico</normalizedToken>
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: Sonora: Puerto
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<normalizedToken originalValue="Peñasco">Penasco</normalizedToken>
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, Estero
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<normalizedToken originalValue="Morúa">Morua</normalizedToken>
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, 31.293 - 31.294°N 113.456 - 113.458°W, 1 m elev., 16 August 2013, Maddison/Proctor/Evans/Leduc-Robert, WPM#13-079 (2♂♂ in UBC-SEM specimens AZS13-7562 [Fig. 30] and AZS13-7582 [Fig. 31]).
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</paragraph>
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<caption pageId="4" pageNumber="49">
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<paragraph pageId="4" pageNumber="49">
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Figures 26-37.
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<taxonomicName class="Arachnida" family="Salticidae" genus="Habronattus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Habronattus empyrus" order="Araneae" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="empyrus">Habronattus empyrus</taxonomicName>
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sp. n. 26 Left male palp, ventral view (holotype male) 27 Same, retrolateral view of palp tibia 28 Epigynum, ventral view (paratype female AZS13-7828) 29 Femur, patella and tibia of third leg, prolateral view (holotype male) 30 Male AZS13-7562 (paratype) 31 Male AZS13-7582 (paratype) 32-33 Male AZS13-7575 34 Male AZS13-7834 35 Female AZS13-7617 36-37 Female AZS13-7828 (paratype). All specimens are from Estero
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<normalizedToken originalValue="Morúa">Morua</normalizedToken>
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(WPM#13-084) except Figs 30-33, 35 from Estero Cerro Prieto (WPM#13-079). Figs 30-37 are copyright © 2015 W. P. Maddison, released under a Creative Commons Attribution (CC-BY) 3.0 license.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="49" type="etymology">
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<paragraph pageId="4" pageNumber="49">Etymology.</paragraph>
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<paragraph pageId="4" pageNumber="49">
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From the Greek empyros, burning, referring to the
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<normalizedToken originalValue="male’s">male's</normalizedToken>
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flaming colors: a brilliant red face against a pale yellow-orange body and legs. Also, to the
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<normalizedToken originalValue="author’s">author's</normalizedToken>
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synesthesia, the dominant letters of the name match the colours of males perfectly:
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<normalizedToken originalValue="“e”">"e"</normalizedToken>
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for the green first legs,
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<normalizedToken originalValue="“r”">"r"</normalizedToken>
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for the red face, and
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<normalizedToken originalValue="“y”">"y"</normalizedToken>
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and
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<normalizedToken originalValue="“s”">"s"</normalizedToken>
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for the pale yellow-orange of the body and legs.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="49" type="diagnosis">
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<paragraph pageId="4" pageNumber="49">Diagnosis.</paragraph>
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<paragraph pageId="4" pageNumber="49">
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A member of the
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<taxonomicName class="Arachnida" family="Salticidae" genus="Harmochirina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Harmochirina coecatus" order="Araneae" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="coecatus">coecatus</taxonomicName>
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group distinctive for its pale colours. The
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<normalizedToken originalValue="male’s">male's</normalizedToken>
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red face and form of the third legs (Figs 29-31, 33, 34) distinguish it from all other members of the
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<taxonomicName class="Arachnida" family="Salticidae" genus="Harmochirina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Harmochirina coecatus" order="Araneae" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="coecatus">coecatus</taxonomicName>
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group except
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<taxonomicName class="Arachnida" family="Salticidae" genus="Habronattus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Habronattus pyrrithrix" order="Araneae" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="pyrrithrix">Habronattus pyrrithrix</taxonomicName>
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, from which it differs in having a much paler body, the green first legs paler in life, and the red facial band narrower. The third leg is much like that of
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<taxonomicName class="Arachnida" family="Salticidae" genus="Habronattus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Habronattus pyrrithrix" order="Araneae" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="pyrrithrix">Habronattus pyrrithrix</taxonomicName>
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,
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<taxonomicName class="Arachnida" family="Salticidae" genus="Habronattus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Habronattus carpus" order="Araneae" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="carpus">Habronattus carpus</taxonomicName>
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and
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<taxonomicName class="Arachnida" family="Salticidae" genus="Habronattus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Habronattus mexicanus" order="Araneae" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="mexicanus">Habronattus mexicanus</taxonomicName>
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, with an orange tuft on the dorsal distal side of the femur and a dark patella with pale speckles, a bright white dorsal-basal tuft, and a moderate but thin thumb-like dorsal-distal apophysis (Figs 29-31; compare to
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<bibRefCitation author="Griswold, CE" journalOrPublisher="The University of California Publications in Entomology" pageId="20" pageNumber="65" pagination="1 - 344" title="A revision of the jumping spider genus Habronattus F. O. P. - Cambridge (Araneae; Salticidae), with phenetic and cladistic analyses." volume="107" year="1987">Griswold 1987</bibRefCitation>
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figures 84-85). However, the femur of
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<taxonomicName class="Arachnida" family="Salticidae" genus="Habronattus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Habronattus empyrus" order="Araneae" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="empyrus">Habronattus empyrus</taxonomicName>
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has an additional black streak just ventral to the prominent macroseta on the prolateral distal face of the femur (Fig. 29). Typical
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<taxonomicName class="Arachnida" family="Salticidae" genus="Habronattus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Habronattus pyrrithrix" order="Araneae" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="pyrrithrix">Habronattus pyrrithrix</taxonomicName>
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were found only 6 km away from the type locality, lessening concerns that
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<taxonomicName class="Arachnida" family="Salticidae" genus="Habronattus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Habronattus empyrus" order="Araneae" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="empyrus">Habronattus empyrus</taxonomicName>
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might be a only a geographical variant.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="49" type="description">
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<paragraph pageId="4" pageNumber="49">Description.</paragraph>
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<paragraph pageId="4" pageNumber="49">
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Male (focal specimen: holotype). Carapace length 2.2; abdomen length 2.1. Palp typical for
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<taxonomicName class="Arachnida" family="Salticidae" genus="Harmochirina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Harmochirina coecatus" order="Araneae" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="coecatus">coecatus</taxonomicName>
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group, with sickle-shaped TmA. Embolus arises at 180° (Fig. 26). Colour and ornaments in alcohol: Chelicerae dark at base, paler at tips. Palp femur and tibia pale except dark patch prolaterally and ventrally. Cymbium pale yellow-brown with long white hairs. Legs pale yellowish except for dorsal black stripe on first femur and markings of third leg. First leg with fringes and modified spatulate setae typical of
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<taxonomicName class="Arachnida" family="Salticidae" genus="Harmochirina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Harmochirina coecatus" order="Araneae" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="coecatus">coecatus</taxonomicName>
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group. Third femur with longitudinal black lines on dorsal and ventral edges of prolateral face, up to the expanded distal area which bears two black spots and a dorsal tuft of orange setae (Figs 29, 30). Third patella with a typical expanded triangular ridge above and a thumb-like apophysis distally. Clypeus red, transitioning abruptly to black between the AME and ALE. The black is a fairly narrow region beneath the ALE. Otherwise, the carapace is covered fairly uniformly with cream to light yellowish-brown scales, with the usual
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<taxonomicName class="Arachnida" family="Salticidae" genus="Harmochirina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Harmochirina coecatus" order="Araneae" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="coecatus">coecatus</taxonomicName>
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-group markings indistinct. Abdomen with standard
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<taxonomicName class="Arachnida" family="Salticidae" genus="Harmochirina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Harmochirina coecatus" order="Araneae" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="coecatus">coecatus</taxonomicName>
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-group markings of a central pale chevroned longitudinal band with a transverse band cutting across it, but less distinct than usual, because the background is light brown rather than black. In life (Figs 30-34), the palp femur is light brown, not red as in
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<taxonomicName class="Arachnida" family="Salticidae" genus="Habronattus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Habronattus pyrrithrix" order="Araneae" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="pyrrithrix">Habronattus pyrrithrix</taxonomicName>
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. The integument of the first leg is light green. The third tibia is also green.
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</paragraph>
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<paragraph pageId="4" pageNumber="49">
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Female (focal specimen: paratype, specimen AZS13-7828; Figs 28, 36-37). Carapace length 2.6; abdomen length 2.7. Structure (including epigynum, Fig. 28) typical for
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<taxonomicName class="Arachnida" family="Salticidae" genus="Harmochirina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Harmochirina coecatus" order="Araneae" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="coecatus">coecatus</taxonomicName>
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group. Colour (Figs 35-37) typical for
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<taxonomicName class="Arachnida" family="Salticidae" genus="Harmochirina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Harmochirina coecatus" order="Araneae" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="coecatus">coecatus</taxonomicName>
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group, pale beige to light brown. Clypeus white (Fig. 37). Abdomen shows only a trace of the markings of the male.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="49" type="materials_examined">
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<paragraph pageId="4" pageNumber="49">Additional material examined.</paragraph>
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<paragraph pageId="4" pageNumber="49">
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10♂♂ 7♀♀ in UBC-SEM:
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<normalizedToken originalValue="México">Mexico</normalizedToken>
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: Sonora: Puerto
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<normalizedToken originalValue="Peñasco">Penasco</normalizedToken>
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, Estero
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<normalizedToken originalValue="Morúa">Morua</normalizedToken>
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,
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<geoCoordinate direction="north" orientation="latitude" precision="55" value="31.293">31.293°N</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="55" value="-113.452">113.452°W</geoCoordinate>
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, 1 m elev., 16 August 2013, S.C. Evans, WPM#13-078 (1♂).
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<normalizedToken originalValue="México">Mexico</normalizedToken>
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: Sonora: Puerto
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<normalizedToken originalValue="Peñasco">Penasco</normalizedToken>
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, Estero
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<normalizedToken originalValue="Morúa">Morua</normalizedToken>
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, 31.293 - 31.294°N 113.456 - 113.458°W, 1 m elev., 16 August 2013, Maddison/Proctor/Evans/Leduc-Robert, WPM#13-079 (3♂♂).
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<normalizedToken originalValue="México">Mexico</normalizedToken>
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: Sonora: Puerto
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<normalizedToken originalValue="Peñasco">Penasco</normalizedToken>
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, Estero
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<normalizedToken originalValue="Morúa">Morua</normalizedToken>
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, 31.293 - 31.295°N 113.456 - 113.459°W, 17 August 2013, Maddison/ Proctor/Evans/Leduc-Robert/Meza, WPM#13-084 (3♂♂ 7♀♀).
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<normalizedToken originalValue="México">Mexico</normalizedToken>
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: Sonora: Puerto
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<normalizedToken originalValue="Peñasco">Penasco</normalizedToken>
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, Estero
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<normalizedToken originalValue="Morúa">Morua</normalizedToken>
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, 31.296 - 31.297°N 113.487 - 113.493°W, 17 August 2013, Maddison/ Proctor/Evans/Leduc-Robert, WPM#13-085 (2♂♂).
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<normalizedToken originalValue="México">Mexico</normalizedToken>
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: Sonora: Puerto
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<normalizedToken originalValue="Peñasco">Penasco</normalizedToken>
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, Estero Cerro Prieto,
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<geoCoordinate direction="north" orientation="latitude" precision="55" value="31.418">31.418°N</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="55" value="-113.626">113.626°W</geoCoordinate>
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, 1 m elev., 18 August 2013, W. Maddison & A. Meza
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<normalizedToken originalValue="López">Lopez</normalizedToken>
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, WPM#13-086 (1♂).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="49" type="natural history">
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<paragraph pageId="4" pageNumber="49">Natural history.</paragraph>
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<paragraph pageId="4" pageNumber="49">
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Found with
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<taxonomicName class="Arachnida" family="Salticidae" genus="Habronattus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Habronattus aestus" order="Araneae" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="aestus">Habronattus aestus</taxonomicName>
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in the negative tidal estuaries of Sonora,
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<normalizedToken originalValue="México">Mexico</normalizedToken>
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.
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<taxonomicName class="Arachnida" family="Salticidae" genus="Habronattus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Habronattus empyrus" order="Araneae" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="empyrus">Habronattus empyrus</taxonomicName>
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, however, was found in the flatter areas with short salt-tolerant plants including
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<taxonomicName class="Magnoliopsida" family="Amaranthaceae" genus="Salicornia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Salicornia" order="Caryophyllales" pageId="4" pageNumber="49" phylum="Tracheophyta" rank="genus">Salicornia</taxonomicName>
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(Fig. 97), unlike
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<taxonomicName class="Arachnida" family="Salticidae" genus="Habronattus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Habronattus aestus" order="Araneae" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="aestus">Habronattus aestus</taxonomicName>
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which was associated with large salt-tolerant plants along the tidal channels. Individuals of
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<taxonomicName class="Arachnida" family="Salticidae" genus="Habronattus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Habronattus empyrus" order="Araneae" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="empyrus">Habronattus empyrus</taxonomicName>
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were found on the wet sand/mud, or hopping from one short plant to another like little monkeys. A portion of courtship was observed, and appears typical for the
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<taxonomicName class="Arachnida" family="Salticidae" genus="Habronattus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Habronattus coecatus" order="Araneae" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="coecatus">Habronattus coecatus</taxonomicName>
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species group (https://youtu.be/Lwa678NVC3U).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |